730:
3479:
5594:
3472:
1976:
2801:
2787:
5046:
4267:
5032:
643:
2780:
5039:
1982:
3486:
140:
2794:
631:
439:
5607:
4285:
4273:
4292:
5601:
5026:
118:
4280:
818:
662:
gave a possible length of 11 metres (36.3 feet). In 2016, its length was estimated to be 7.4 metres (24 ft 3 in) in a comprehensive analysis of abelisaur size. Others authors gave a similar size at 7.2 metres (23.6 feet) and 1.65 tonnes (1.82 short tons).
1282:
Sampson, S.D.; Witmer, L.M.; Forster, C.A.; Krause, D.A.; O'Connor, P.M.; Dodson, P.; Ravoavy, F. (1998). "Predatory
Dinosaur Remains from Madagascar: Implications for the Cretaceous Biogeography of Gondwana".
534:
consists of a skull, lacking the lower jaws, that is incomplete, especially on the right side. Most of the connections between the snout and the back of the skull are absent. It is also missing most of the
805:, a group unrelated to other species assigned to the Abelisauridae, and, since it is known only from a skull, it has been suggested that future discoveries may show that
1408:
1026:
Leanza, H.A.; ApesteguĂa, S.; Novas, F.E.; Fuente, M.S. (2004). "Cretaceous terrestrial beds from the NeuquĂ©n Basin (Argentina) and their tetrapod assemblages".
5750:
680: in) in 1985. The skull is relatively deep. Although there are no bony crests or horns, like those found in some other abelisaurids, such as
526:, was in 1983 uncovered at the "Cantera de la Pala MĂ©canica"-site in the Lago Pellegrini quarries exploited by Abel since 1975. This single known
1329:
Lamanna, M.C.; MartĂnez, R.D.; Smith, J.B. (2002). "A definitive abelisaurid theropod dinosaur from the early Late
Cretaceous of Patagonia".
969:
Gianechini, Federico A.; ApesteguĂa, SebastiĂĄn; Landini, Walter; Finotti, Franco; JuĂĄrez
Valieri, Rubén; Zandonai, Fabiana (2015-05-01).
426:
that probably reached about 7.4 metres (24 ft 3 in) in length, although this is uncertain as it is known from only one partial
832:
was relatively broad at the back, Bonaparte and Novas drew a comparison between the newly described species and the also wide-skulled
5686:
694:(window-like openings) in the skull, which are found in many dinosaurs and reduce skull weight. One of these is a large triangular
5775:
5770:
698:
in the side of the snout. The eye socket behind it is rather high. It is constricted in the middle by bony projections of the
5780:
658:
estimated its body length at ten metres (32 ft 10 in), its weight at 3 tonnes (3 long tons; 3 short tons). In 2012
3478:
3471:
1459:
5593:
1438:
1381:
1217:
1192:
1128:
1103:
1073:
4266:
2800:
895:
539:(roof of the mouth). Despite the missing pieces, it could be estimated at over 85 centimetres (33 in) long.
2786:
1342:
5031:
5760:
5755:
5472:
2779:
1975:
4284:
1981:
5045:
3485:
5765:
2793:
139:
5038:
4291:
1208:
Tykoski, R.S.; Rowe, T. (2004). "Ceratosauria". In Dodson, P.; OsmĂłlska, H.; Weishamepl, D.B. (eds.).
5606:
970:
729:
1526:
950:[Abelisaurus comahuensis, n.g., n.sp., Carnosauria from the Late Cretaceous of Patagonia].
686:, rough ridges on the snout and above the eyes might have supported some kind of crest made out of
706:
at the rear. The eye was located above the constriction. Behind the eye socket a large triangular
5745:
5600:
4272:
1431:
915:
690:, which would not have become fossilized. The skull roof is thickened. There are also very large
505:
5717:
5626:
707:
971:"New abelisaurid remains from the Anacleto Formation (Upper Cretaceous), Patagonia, Argentina"
765:
Many other abelisaurids have since been discovered, including much more complete specimens of
5740:
5712:
5704:
5664:
3997:
3909:
802:
790:
781:. They showed that abelisaurids were not carnosaurs in the modern sense, but belonged to the
472:
5673:
5558:
3686:
3654:
1292:
1157:
1035:
982:
650:
As only the skull is known, it has proven difficult to provide a reliable size estimate of
5025:
8:
4010:
3797:
1144:
Grillo, O.N.; Delcourt, R. (2016). "Allometry and body length of abelisauroid theropods:
695:
659:
4279:
1296:
1161:
1039:
986:
5465:
3986:
3846:
3702:
3642:
3077:
2987:
2870:
1734:
1424:
1354:
1259:
1234:
577:
562:
554:
462:
443:
308:
134:
5699:
5347:
5264:
5238:
5077:
5064:
4942:
4500:
4452:
3284:
2634:
1713:
1387:
1377:
1346:
1308:
1264:
1213:
1188:
1124:
1121:
Dinosaurs: The Most
Complete, Up-to-date Encyclopedia for Dinosaur Lovers of All Ages
1099:
1069:
1008:
948:"Abelisaurus comahuensis, n.g., n.sp., Carnosauria del Crétacico Tardio de Patagonia"
947:
710:
is present. Its form reflects a strong forward inclination of the back of the skull.
589:
1358:
468:
5451:
5208:
5118:
4907:
4839:
4702:
4621:
4593:
4553:
4434:
4404:
3922:
3675:
3539:
3396:
3296:
3145:
3052:
3010:
2577:
1887:
1338:
1300:
1254:
1246:
1165:
1043:
998:
990:
870:
748:
703:
408:
346:
1304:
1169:
1047:
994:
642:
5514:
5252:
5215:
5147:
5002:
4995:
4823:
4765:
4738:
4728:
4688:
4523:
4382:
4366:
4184:
4156:
4134:
4021:
3866:
3573:
3347:
3230:
3208:
3129:
2958:
2935:
2928:
2906:
2892:
2885:
2846:
2697:
2544:
2471:
2460:
2396:
2284:
2171:
1748:
1657:
1511:
1486:
833:
782:
655:
611:
585:
573:
550:
465:
5479:
5442:
5423:
5385:
5374:
5171:
4884:
4850:
4832:
4812:
4709:
4423:
4168:
4030:
3857:
3839:
3667:
3634:
3620:
3581:
3355:
3216:
3173:
3166:
3088:
3064:
3017:
2976:
2913:
2899:
2627:
2612:
2439:
2273:
2137:
2105:
2094:
2071:
2033:
1894:
1864:
1788:
1727:
1626:
864:
603:
299:
295:
5734:
5649:
5521:
5367:
5328:
5307:
5186:
5179:
5106:
4900:
4873:
4747:
4664:
4642:
4635:
4628:
4571:
4488:
4351:
4323:
4310:
4234:
4111:
3951:
3785:
3716:
3602:
3589:
3334:
3312:
3305:
3269:
3242:
3194:
3118:
2854:
2645:
2619:
2535:
2516:
2509:
2446:
2425:
2404:
2389:
2375:
2334:
2300:
2293:
2212:
2064:
1951:
1944:
1871:
1841:
1830:
1720:
1691:
1684:
1610:
1391:
1350:
1012:
882:
876:
841:
794:
777:
751:
721:
behind it had at least seven, but perhaps as many as thirteen, larger teeth.
699:
630:
558:
488:
476:
438:
416:
239:
223:
68:
654:. It has transpired that abelisaurids have relatively short heads. In 2010,
553:
but subsequent research proved the remains were actually found in the older
5493:
5431:
5401:
5321:
5314:
5292:
5284:
5164:
5156:
5136:
4859:
4778:
4716:
4607:
4578:
4530:
4359:
4344:
4119:
4051:
3979:
3958:
3937:
3753:
3709:
3517:
3504:
3434:
3387:
3262:
3152:
3104:
3097:
3034:
3003:
2996:
2877:
2704:
2556:
2453:
2355:
2348:
2243:
2227:
2163:
2114:
2046:
1823:
1706:
1699:
1670:
1633:
1602:
1372:
Novas, F.E. (1997). "Abelisauridae". In Currie, P.J.; Padian, K.P. (eds.).
1268:
1250:
1185:
RĂ©cords y curiosidades de los dinosaurios TerĂłpodos y otros dinosauromorfos
856:
475:
in 1985. The generic name recognizes
Roberto Abel as the discoverer of the
455:
278:
252:
1312:
5658:
5547:
5528:
5458:
5408:
5394:
5336:
5277:
5222:
5099:
4893:
4785:
4754:
4671:
4657:
4389:
4337:
4205:
4127:
4090:
3972:
3944:
3897:
3817:
3695:
3552:
3403:
3376:
3276:
3223:
3159:
3138:
3111:
2862:
2675:
2668:
2480:
2327:
2318:
2220:
2191:
2121:
2086:
2079:
1879:
1857:
1850:
1809:
1801:
1777:
1762:
1741:
1649:
1641:
755:
714:
682:
395:
43:
5691:
5486:
5415:
5360:
4981:
4796:
4614:
4600:
4585:
4537:
4471:
4397:
4219:
4212:
4198:
4104:
4037:
3965:
3929:
3882:
3824:
3810:
3777:
3769:
3762:
3743:
3566:
3531:
3319:
2920:
2711:
2661:
2598:
2569:
2524:
2502:
2495:
2487:
2411:
2307:
2266:
2259:
2251:
2235:
2155:
2026:
1902:
1755:
1618:
1477:
1003:
767:
597:
480:
405:
318:
197:
88:
53:
809:
was in fact a carcharodontosaurid. However, this is thought unlikely.
5507:
5500:
5200:
5193:
4866:
4805:
4459:
4177:
3723:
3627:
3362:
2947:
2832:
2819:
2746:
2682:
2605:
2584:
2341:
2204:
2012:
1999:
1917:
1769:
1588:
1575:
1538:
1447:
859:
845:
581:
570:
546:
484:
423:
412:
210:
151:
93:
37:
30:
5620:
801:
shares some skull features, such as a relative elongation, with the
576:, which makes its stratigraphic position unclear. The Anacleto is a
569:
is known to have come from the Sr. Fernandez field, which is in the
117:
5643:
5091:
4516:
4509:
4481:
4374:
3874:
3255:
2591:
2418:
2179:
1816:
1495:
1471:
519:
401:
398:
184:
171:
83:
78:
63:
58:
48:
968:
4649:
4411:
4141:
4044:
3369:
2690:
817:
718:
687:
512:
98:
73:
5678:
1212:(2nd ed.). University of California Press. pp. 47â70.
5300:
4988:
2432:
1465:
536:
527:
161:
1235:"New dinosaurs link southern landmasses in the Mid-Cretaceous"
479:, and also as the former director of the provincial Museum of
4148:
3889:
3831:
498:
491:
427:
420:
391:
373:
1416:
5564:
4191:
1343:
10.1671/0272-4634(2002)022[0058:ADATDF]2.0.CO;2
1025:
379:
358:
349:
1281:
545:
is one of the many dinosaurs that have been discovered in
735:
487:, where the specimen is housed. It also incorporates the
364:
129:
skull with original bones of the holotype. Scale = 10 cm
1239:
Proceedings of the Royal
Society B: Biological Sciences
461:, both genus and species were named and described by
382:
370:
352:
1328:
376:
361:
355:
285:
4973:
367:
1232:
1233:Sereno, P.C.; Wilson, J.A.; Conrad, J.L. (2004).
549:. It was originally described as coming from the
515:region of Argentina, where the fossil was found.
5732:
5075:
4941:
4432:
2956:
1510:
1182:
1143:
945:
666:Skull length was estimated at 85 centimetres (
1432:
836:. They assumed that, just as tyrannosaurids,
754:in 1985. They thought it was a member of the
4321:
3515:
3433:
2555:
1376:. San Diego: Academic Press. pp. 1â2.
1207:
1187:. Barcelona, Spain: Larousse. p. 257.
797:. Others are less certain of its position.
5751:Late Cretaceous dinosaurs of South America
1439:
1425:
116:
1258:
1068:. Indiana University Press. p. 281.
1002:
2830:
2745:
2010:
1586:
816:
728:
717:, bore four relatively small teeth. The
641:
629:
580:in South America, dating from the early
437:
433:
5733:
1096:The Princeton Field Guide to Dinosaurs
5625:
5624:
5063:
5062:
4308:
3502:
2817:
1997:
1573:
1458:
1420:
1371:
1324:
1322:
1118:
1066:The Age of Dinosaurs in South America
1063:
1093:
1089:
1087:
1085:
1059:
1057:
941:
939:
937:
935:
933:
931:
958:: 259â265 – via ResearchGate.
946:Bonaparte, J.; Novas, E.E. (1985).
793:abelisaurid, outside the subfamily
13:
1331:Journal of Vertebrate Paleontology
1319:
14:
5792:
4309:
1402:
1183:M-P., R.; Larramendi, A. (2016).
1082:
1054:
928:
762:was the first abelisaurid named.
724:
5605:
5599:
5592:
5044:
5037:
5030:
5024:
4290:
4283:
4278:
4271:
4265:
3503:
3484:
3477:
3470:
2799:
2792:
2785:
2778:
1980:
1974:
622:. This has found no acceptance.
345:
138:
41:
1365:
1275:
1226:
1201:
896:Timeline of ceratosaur research
812:
785:instead. Some scientists place
1176:
1137:
1112:
1098:. Princeton University Press.
1019:
962:
908:
625:
595:In 2009, Novas suggested that
565:, Argentina. However, locally
1:
5771:Fossil taxa described in 1985
2818:
1998:
1574:
1446:
1305:10.1126/science.280.5366.1048
1170:10.1016/j.cretres.2016.09.001
1048:10.1016/j.cretres.2003.10.005
995:10.1016/j.cretres.2014.11.009
901:
638:(red) with other carnotaurins
5781:Taxa named by Fernando Novas
5776:Taxa named by José Bonaparte
1123:. Random House. p. 83.
851:In 2010, Paul proposed that
499:
7:
889:
821:Restoration of the head of
743:Bonaparte and Novas placed
272:Bonaparte & Novas, 1985
10:
5797:
713:The front snout bone, the
588:Period, between 83 and 80
492:
16:Extinct genus of dinosaurs
5633:
5589:
5546:
5441:
5384:
5355:
5346:
5272:
5263:
5247:
5236:
5146:
5131:
5127:
5116:
5086:
5071:
5058:
5022:
4969:
4937:
4883:
4849:
4822:
4795:
4773:
4764:
4737:
4726:
4697:
4686:
4566:
4562:
4551:
4499:
4470:
4447:
4443:
4421:
4332:
4317:
4304:
4263:
4233:
4167:
4099:
4089:
4085:
4020:
4005:
3996:
3917:
3908:
3856:
3805:
3796:
3752:
3741:
3685:
3662:
3653:
3615:
3611:
3600:
3561:
3550:
3526:
3511:
3498:
3467:
3429:
3386:
3342:
3333:
3295:
3250:
3241:
3203:
3192:
3128:
3087:
3072:
3063:
3047:
3043:
3032:
2986:
2971:
2967:
2945:
2841:
2826:
2813:
2775:
2741:
2644:
2564:
2534:
2470:
2384:
2373:
2317:
2283:
2199:
2190:
2150:
2146:
2135:
2104:
2059:
2055:
2044:
2021:
2006:
1993:
1971:
1936:
1916:
1840:
1796:
1787:
1679:
1668:
1597:
1582:
1569:
1537:
1519:
1506:
1454:
1374:Encyclopedia of Dinosaurs
504:), meaning 'lizard'. The
314:
307:
284:
277:
135:Scientific classification
133:
124:
115:
23:
390:; "Abel's lizard") is a
608:Abelisaurus comahuensis
459:Abelisaurus comahuensis
450:remains have been found
290:Abelisaurus comahuensis
1251:10.1098/rspb.2004.2692
1146:Pycnonemosaurus nevesi
825:
740:
708:infratemporal fenestra
647:
639:
451:
404:alive during the Late
125:Reconstruction of the
5713:Paleobiology Database
3998:Carcharodontosaurinae
3910:Carcharodontosauridae
1412:in the Dino Directory
820:
747:in the newly created
732:
702:in the front and the
645:
633:
473:Fernando Emilio Novas
441:
5761:Fossils of Argentina
5756:Cretaceous Argentina
4974:Dubious coelurosaurs
3687:Metriacanthosaurinae
3655:Metriacanthosauridae
1937:Dubious neotheropods
1119:Holtz, T.R. (2012).
1064:Novas, F.E. (2009).
803:carcharodontosaurids
646:Artist's restoration
620:Abelisaurus garridoi
434:Discovery and naming
4011:Carcharodontosaurus
3798:Carcharodontosauria
1297:1998Sci...280.1048S
1291:(5366): 1048â1051.
1245:(1546): 1325â1330.
1162:2017CrRes..69...71G
1150:Cretaceous Research
1094:Paul, G.S. (2010).
1040:2004CrRes..25...61L
1028:Cretaceous Research
987:2015CrRes..54....1G
975:Cretaceous Research
696:antorbital fenestra
634:Size comparison of
616:Aucasaurus garridoi
5766:Anacleto Formation
5466:Archaeornithomimus
3987:Veterupristisaurus
3847:Veterupristisaurus
3703:Metriacanthosaurus
3643:Piatnitzkysauridae
3078:Eustreptospondylus
2988:Piatnitzkysauridae
2871:Chuandongocoelurus
1735:Notatesseraeraptor
1148:is the new king".
826:
741:
648:
640:
578:geologic formation
563:Rio Negro Province
555:Anacleto Formation
452:
444:Anacleto Formation
5728:
5727:
5700:Open Tree of Life
5627:Taxon identifiers
5618:
5617:
5614:
5613:
5587:
5586:
5583:
5582:
5579:
5578:
5575:
5574:
5542:
5541:
5538:
5537:
5348:Macrocheiriformes
5265:Ornithomimosauria
5239:Maniraptoriformes
5232:
5231:
5078:Maniraptoromorpha
5065:Maniraptoromorpha
5054:
5053:
5020:
5019:
5016:
5015:
5012:
5011:
4965:
4964:
4961:
4960:
4943:Maniraptoromorpha
4933:
4932:
4929:
4928:
4925:
4924:
4921:
4920:
4917:
4916:
4682:
4681:
4547:
4546:
4501:Proceratosauridae
4453:Chingkankousaurus
4300:
4299:
4261:
4260:
4257:
4256:
4253:
4252:
4229:
4228:
4081:
4080:
4077:
4076:
4073:
4072:
4069:
4068:
4065:
4064:
4061:
4060:
3737:
3736:
3733:
3732:
3494:
3493:
3465:
3464:
3461:
3460:
3457:
3456:
3453:
3452:
3425:
3424:
3421:
3420:
3417:
3416:
3413:
3412:
3329:
3328:
3285:Vallibonavenatrix
3188:
3187:
3184:
3183:
3028:
3027:
2809:
2808:
2773:
2772:
2769:
2768:
2765:
2764:
2737:
2736:
2733:
2732:
2729:
2728:
2725:
2724:
2721:
2720:
2369:
2368:
2365:
2364:
2131:
2130:
1989:
1988:
1969:
1968:
1965:
1964:
1961:
1960:
1912:
1911:
1565:
1564:
1561:
1560:
1557:
1556:
590:million years ago
415:) of what is now
336:
335:
330:
273:
5788:
5721:
5720:
5708:
5707:
5695:
5694:
5682:
5681:
5669:
5668:
5667:
5654:
5653:
5652:
5622:
5621:
5609:
5603:
5596:
5452:Aepyornithomimus
5353:
5352:
5270:
5269:
5245:
5244:
5209:Sinocalliopteryx
5129:
5128:
5125:
5124:
5119:Neocoelurosauria
5084:
5083:
5073:
5072:
5060:
5059:
5048:
5041:
5034:
5028:
4971:
4970:
4939:
4938:
4908:Zhuchengtyrannus
4840:Thanatotheristes
4771:
4770:
4735:
4734:
4703:Appalachiosaurus
4695:
4694:
4564:
4563:
4560:
4559:
4554:Pantyrannosauria
4445:
4444:
4441:
4440:
4435:Tyrannosauroidea
4430:
4429:
4405:Xinjiangovenator
4330:
4329:
4319:
4318:
4306:
4305:
4294:
4287:
4282:
4275:
4269:
4097:
4096:
4087:
4086:
4003:
4002:
3923:Acrocanthosaurus
3915:
3914:
3803:
3802:
3750:
3749:
3676:Yangchuanosaurus
3660:
3659:
3613:
3612:
3609:
3608:
3559:
3558:
3540:Lourinhanosaurus
3524:
3523:
3513:
3512:
3500:
3499:
3488:
3481:
3474:
3431:
3430:
3397:Sigilmassasaurus
3340:
3339:
3297:Ceratosuchopsini
3248:
3247:
3201:
3200:
3146:Dubreuillosaurus
3070:
3069:
3053:Streptospondylus
3045:
3044:
3041:
3040:
3011:Piatnitzkysaurus
2969:
2968:
2965:
2964:
2954:
2953:
2839:
2838:
2828:
2827:
2815:
2814:
2803:
2796:
2789:
2782:
2743:
2742:
2578:Ekrixinatosaurus
2562:
2561:
2553:
2552:
2382:
2381:
2197:
2196:
2148:
2147:
2144:
2143:
2057:
2056:
2053:
2052:
2019:
2018:
2008:
2007:
1995:
1994:
1984:
1978:
1888:Procompsognathus
1794:
1793:
1677:
1676:
1595:
1594:
1584:
1583:
1571:
1570:
1517:
1516:
1508:
1507:
1501:
1500:
1456:
1455:
1441:
1434:
1427:
1418:
1417:
1396:
1395:
1369:
1363:
1362:
1326:
1317:
1316:
1279:
1273:
1272:
1262:
1230:
1224:
1223:
1205:
1199:
1198:
1180:
1174:
1173:
1141:
1135:
1134:
1116:
1110:
1109:
1091:
1080:
1079:
1061:
1052:
1051:
1023:
1017:
1016:
1006:
966:
960:
959:
943:
926:
925:
923:
922:
912:
871:Pellegrinisaurus
828:As the skull of
704:postorbital bone
679:
678:
674:
671:
502:
495:
494:
389:
388:
385:
384:
381:
378:
375:
372:
369:
366:
363:
360:
357:
354:
351:
324:
287:
271:
264:
251:
238:
222:
143:
142:
120:
110:
40:
29:Temporal range:
21:
20:
5796:
5795:
5791:
5790:
5789:
5787:
5786:
5785:
5731:
5730:
5729:
5724:
5716:
5711:
5703:
5698:
5690:
5685:
5677:
5672:
5663:
5662:
5657:
5648:
5647:
5642:
5629:
5619:
5610:
5597:
5571:
5534:
5515:Sinornithomimus
5437:
5380:
5342:
5259:
5253:Compsognathidae
5241:
5228:
5216:Sinosauropteryx
5148:Compsognathidae
5142:
5121:
5112:
5080:
5067:
5050:
5049:
5035:
5008:
5003:Shanyangosaurus
4996:Phaedrolosaurus
4957:
4913:
4879:
4845:
4824:Daspletosaurini
4818:
4791:
4766:Tyrannosaurinae
4760:
4739:Albertosaurinae
4731:
4729:Tyrannosauridae
4722:
4691:
4689:Eutyrannosauria
4678:
4556:
4543:
4524:Proceratosaurus
4495:
4466:
4437:
4426:
4417:
4383:Richardoestesia
4367:Chilantaisaurus
4326:
4313:
4296:
4295:
4276:
4249:
4225:
4185:Australovenator
4163:
4157:Chilantaisaurus
4135:Phuwiangvenator
4057:
4022:Giganotosaurini
4016:
3992:
3904:
3867:Chilantaisaurus
3852:
3792:
3746:
3729:
3681:
3649:
3605:
3596:
3574:Monolophosaurus
3555:
3546:
3520:
3507:
3490:
3489:
3475:
3449:
3409:
3382:
3348:Camarillasaurus
3325:
3291:
3237:
3231:Ostafrikasaurus
3209:Camarillasaurus
3197:
3180:
3130:Afrovenatorinae
3124:
3083:
3059:
3037:
3024:
2982:
2961:
2959:Megalosauroidea
2950:
2941:
2936:Vectaerovenator
2929:Szechuanosaurus
2907:Monolophosaurus
2893:Kaijiangosaurus
2886:Cryolophosaurus
2847:Calamospondylus
2835:
2822:
2805:
2804:
2797:
2790:
2783:
2761:
2717:
2698:Pycnonemosaurus
2640:
2551:
2545:Majungasaurinae
2530:
2472:Majungasaurinae
2466:
2461:Xenotarsosaurus
2397:Dryptosauroides
2378:
2361:
2313:
2285:Elaphrosaurinae
2279:
2186:
2172:Ornithomimoides
2140:
2127:
2100:
2049:
2040:
2015:
2002:
1985:
1979:
1957:
1932:
1908:
1836:
1783:
1749:Shuangbaisaurus
1673:
1664:
1591:
1578:
1553:
1533:
1527:Avemetatarsalia
1512:Avemetatarsalia
1502:
1487:Avemetatarsalia
1461:
1460:
1450:
1445:
1405:
1400:
1399:
1384:
1370:
1366:
1327:
1320:
1280:
1276:
1231:
1227:
1220:
1206:
1202:
1195:
1181:
1177:
1142:
1138:
1131:
1117:
1113:
1106:
1092:
1083:
1076:
1062:
1055:
1024:
1020:
967:
963:
944:
929:
920:
918:
914:
913:
909:
904:
892:
834:Tyrannosauridae
815:
783:Neoceratosauria
727:
676:
672:
669:
667:
656:Gregory S. Paul
628:
612:Gregory S. Paul
586:Late Cretaceous
574:Allen Formation
551:Allen Formation
466:paleontologists
436:
348:
344:
303:
293:
270:
262:
249:
236:
220:
137:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
5794:
5784:
5783:
5778:
5773:
5768:
5763:
5758:
5753:
5748:
5746:Campanian life
5743:
5726:
5725:
5723:
5722:
5709:
5696:
5683:
5670:
5655:
5639:
5637:
5631:
5630:
5616:
5615:
5612:
5611:
5591:
5590:
5588:
5585:
5584:
5581:
5580:
5577:
5576:
5573:
5572:
5570:
5569:
5568:
5567:
5561:
5552:
5550:
5544:
5543:
5540:
5539:
5536:
5535:
5533:
5532:
5525:
5518:
5511:
5504:
5497:
5490:
5483:
5480:Dromiceiomimus
5476:
5469:
5462:
5455:
5447:
5445:
5443:Ornithomimidae
5439:
5438:
5436:
5435:
5428:
5424:Paraxenisaurus
5420:
5412:
5405:
5398:
5390:
5388:
5386:Deinocheiridae
5382:
5381:
5379:
5378:
5375:Shenzhousaurus
5371:
5364:
5356:
5350:
5344:
5343:
5341:
5340:
5333:
5325:
5318:
5311:
5304:
5297:
5289:
5281:
5273:
5267:
5261:
5260:
5258:
5257:
5248:
5242:
5237:
5234:
5233:
5230:
5229:
5227:
5226:
5219:
5212:
5205:
5197:
5190:
5183:
5176:
5172:Beipiaognathus
5168:
5161:
5152:
5150:
5144:
5143:
5141:
5140:
5132:
5122:
5117:
5114:
5113:
5111:
5110:
5103:
5096:
5087:
5081:
5076:
5069:
5068:
5056:
5055:
5052:
5051:
5043:
5029:
5023:
5021:
5018:
5017:
5014:
5013:
5010:
5009:
5007:
5006:
4999:
4992:
4985:
4977:
4975:
4967:
4966:
4963:
4962:
4959:
4958:
4956:
4955:
4954:
4953:
4947:
4945:
4935:
4934:
4931:
4930:
4927:
4926:
4923:
4922:
4919:
4918:
4915:
4914:
4912:
4911:
4904:
4897:
4889:
4887:
4885:Tyrannosaurini
4881:
4880:
4878:
4877:
4870:
4863:
4855:
4853:
4851:Teratophoneini
4847:
4846:
4844:
4843:
4836:
4833:Daspletosaurus
4828:
4826:
4820:
4819:
4817:
4816:
4813:Qianzhousaurus
4809:
4801:
4799:
4793:
4792:
4790:
4789:
4782:
4774:
4768:
4762:
4761:
4759:
4758:
4751:
4743:
4741:
4732:
4727:
4724:
4723:
4721:
4720:
4713:
4710:Bistahieversor
4706:
4698:
4692:
4687:
4684:
4683:
4680:
4679:
4677:
4676:
4668:
4661:
4654:
4646:
4639:
4632:
4625:
4618:
4611:
4604:
4597:
4590:
4582:
4575:
4567:
4557:
4552:
4549:
4548:
4545:
4544:
4542:
4541:
4534:
4527:
4520:
4513:
4505:
4503:
4497:
4496:
4494:
4493:
4485:
4477:
4475:
4468:
4467:
4465:
4464:
4456:
4448:
4438:
4433:
4427:
4424:Tyrannoraptora
4422:
4419:
4418:
4416:
4415:
4408:
4401:
4394:
4386:
4379:
4371:
4363:
4356:
4348:
4341:
4333:
4327:
4322:
4315:
4314:
4302:
4301:
4298:
4297:
4289:
4270:
4264:
4262:
4259:
4258:
4255:
4254:
4251:
4250:
4248:
4247:
4246:
4245:
4239:
4237:
4231:
4230:
4227:
4226:
4224:
4223:
4216:
4209:
4202:
4195:
4188:
4181:
4173:
4171:
4169:Megaraptoridae
4165:
4164:
4162:
4161:
4153:
4145:
4138:
4131:
4124:
4116:
4108:
4100:
4094:
4083:
4082:
4079:
4078:
4075:
4074:
4071:
4070:
4067:
4066:
4063:
4062:
4059:
4058:
4056:
4055:
4048:
4041:
4034:
4031:Giganotosaurus
4026:
4024:
4018:
4017:
4015:
4014:
4006:
4000:
3994:
3993:
3991:
3990:
3983:
3976:
3969:
3962:
3955:
3948:
3941:
3934:
3926:
3918:
3912:
3906:
3905:
3903:
3902:
3894:
3886:
3879:
3871:
3862:
3860:
3858:Neovenatoridae
3854:
3853:
3851:
3850:
3843:
3840:Ulughbegsaurus
3836:
3828:
3821:
3814:
3806:
3800:
3794:
3793:
3791:
3790:
3782:
3774:
3766:
3758:
3756:
3747:
3742:
3739:
3738:
3735:
3734:
3731:
3730:
3728:
3727:
3720:
3713:
3706:
3699:
3691:
3689:
3683:
3682:
3680:
3679:
3672:
3668:Xuanhanosaurus
3663:
3657:
3651:
3650:
3648:
3647:
3639:
3635:Xuanhanosaurus
3631:
3624:
3621:Asfaltovenator
3616:
3606:
3601:
3598:
3597:
3595:
3594:
3586:
3582:Megalosauridae
3578:
3570:
3562:
3556:
3551:
3548:
3547:
3545:
3544:
3536:
3527:
3521:
3516:
3509:
3508:
3496:
3495:
3492:
3491:
3483:
3469:
3468:
3466:
3463:
3462:
3459:
3458:
3455:
3454:
3451:
3450:
3448:
3447:
3446:
3445:
3439:
3437:
3427:
3426:
3423:
3422:
3419:
3418:
3415:
3414:
3411:
3410:
3408:
3407:
3400:
3392:
3390:
3384:
3383:
3381:
3380:
3373:
3366:
3359:
3356:Ichthyovenator
3352:
3343:
3337:
3331:
3330:
3327:
3326:
3324:
3323:
3316:
3309:
3301:
3299:
3293:
3292:
3290:
3289:
3281:
3273:
3266:
3259:
3251:
3245:
3239:
3238:
3236:
3235:
3227:
3220:
3217:Cristatusaurus
3213:
3204:
3198:
3193:
3190:
3189:
3186:
3185:
3182:
3181:
3179:
3178:
3174:Poekilopleuron
3170:
3167:Piveteausaurus
3163:
3156:
3149:
3142:
3134:
3132:
3126:
3125:
3123:
3122:
3115:
3108:
3101:
3093:
3091:
3089:Megalosaurinae
3085:
3084:
3082:
3081:
3073:
3067:
3065:Megalosauridae
3061:
3060:
3058:
3057:
3048:
3038:
3033:
3030:
3029:
3026:
3025:
3023:
3022:
3018:Xuanhanosaurus
3014:
3007:
3000:
2992:
2990:
2984:
2983:
2981:
2980:
2977:Yunyangosaurus
2972:
2962:
2957:
2951:
2946:
2943:
2942:
2940:
2939:
2932:
2925:
2917:
2914:Pandoravenator
2910:
2903:
2900:Kayentavenator
2896:
2889:
2882:
2874:
2867:
2859:
2851:
2842:
2836:
2831:
2824:
2823:
2811:
2810:
2807:
2806:
2798:
2791:
2784:
2777:
2776:
2774:
2771:
2770:
2767:
2766:
2763:
2762:
2760:
2759:
2758:
2757:
2751:
2749:
2739:
2738:
2735:
2734:
2731:
2730:
2727:
2726:
2723:
2722:
2719:
2718:
2716:
2715:
2708:
2701:
2694:
2687:
2679:
2672:
2665:
2658:
2650:
2648:
2642:
2641:
2639:
2638:
2631:
2628:Skorpiovenator
2624:
2616:
2609:
2602:
2595:
2588:
2581:
2574:
2565:
2559:
2550:
2549:
2540:
2538:
2532:
2531:
2529:
2528:
2521:
2513:
2506:
2499:
2492:
2484:
2476:
2474:
2468:
2467:
2465:
2464:
2457:
2450:
2443:
2440:Spectrovenator
2436:
2429:
2422:
2415:
2408:
2401:
2393:
2385:
2379:
2374:
2371:
2370:
2367:
2366:
2363:
2362:
2360:
2359:
2352:
2345:
2338:
2331:
2323:
2321:
2315:
2314:
2312:
2311:
2304:
2297:
2289:
2287:
2281:
2280:
2278:
2277:
2274:Spinostropheus
2270:
2263:
2256:
2248:
2240:
2232:
2224:
2217:
2209:
2200:
2194:
2188:
2187:
2185:
2184:
2176:
2168:
2160:
2151:
2141:
2138:Abelisauroidea
2136:
2133:
2132:
2129:
2128:
2126:
2125:
2118:
2110:
2108:
2106:Ceratosauridae
2102:
2101:
2099:
2098:
2095:Saltriovenator
2091:
2083:
2076:
2072:Fosterovenator
2068:
2060:
2050:
2045:
2042:
2041:
2039:
2038:
2034:Lophostropheus
2030:
2022:
2016:
2011:
2004:
2003:
1991:
1990:
1987:
1986:
1973:
1972:
1970:
1967:
1966:
1963:
1962:
1959:
1958:
1956:
1955:
1948:
1940:
1938:
1934:
1933:
1931:
1930:
1929:
1928:
1922:
1920:
1914:
1913:
1910:
1909:
1907:
1906:
1899:
1895:Pterospondylus
1891:
1884:
1876:
1868:
1865:Lucianovenator
1861:
1854:
1846:
1844:
1838:
1837:
1835:
1834:
1827:
1820:
1813:
1806:
1797:
1791:
1789:Coelophysoidea
1785:
1784:
1782:
1781:
1774:
1766:
1759:
1752:
1745:
1738:
1731:
1728:Lophostropheus
1724:
1717:
1710:
1703:
1696:
1688:
1680:
1674:
1669:
1666:
1665:
1663:
1662:
1654:
1646:
1638:
1630:
1627:Erythrovenator
1623:
1615:
1607:
1598:
1592:
1587:
1580:
1579:
1567:
1566:
1563:
1562:
1559:
1558:
1555:
1554:
1552:
1551:
1550:
1549:
1543:
1541:
1535:
1534:
1532:
1531:
1530:
1529:
1520:
1514:
1504:
1503:
1499:
1498:
1489:
1480:
1474:
1468:
1452:
1451:
1444:
1443:
1436:
1429:
1421:
1415:
1414:
1404:
1403:External links
1401:
1398:
1397:
1382:
1364:
1318:
1274:
1225:
1218:
1210:The Dinosauria
1200:
1193:
1175:
1136:
1129:
1111:
1104:
1081:
1074:
1053:
1018:
961:
927:
906:
905:
903:
900:
899:
898:
891:
888:
865:Antarctosaurus
842:apex predators
814:
811:
726:
725:Classification
723:
627:
624:
604:junior synonym
469:José Bonaparte
435:
432:
334:
333:
332:
331:
312:
311:
305:
304:
294:
282:
281:
275:
274:
260:
256:
255:
247:
243:
242:
234:
227:
226:
218:
214:
213:
208:
201:
200:
195:
188:
187:
182:
175:
174:
169:
165:
164:
159:
155:
154:
149:
145:
144:
131:
130:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
28:
15:
9:
6:
4:
3:
2:
5793:
5782:
5779:
5777:
5774:
5772:
5769:
5767:
5764:
5762:
5759:
5757:
5754:
5752:
5749:
5747:
5744:
5742:
5739:
5738:
5736:
5719:
5714:
5710:
5706:
5701:
5697:
5693:
5688:
5684:
5680:
5675:
5671:
5666:
5660:
5656:
5651:
5645:
5641:
5640:
5638:
5636:
5632:
5628:
5623:
5608:
5604:
5602:
5595:
5566:
5562:
5560:
5556:
5555:
5554:
5553:
5551:
5549:
5545:
5531:
5530:
5526:
5524:
5523:
5522:Struthiomimus
5519:
5517:
5516:
5512:
5510:
5509:
5505:
5503:
5502:
5498:
5496:
5495:
5491:
5489:
5488:
5484:
5482:
5481:
5477:
5475:
5474:
5473:"Coelosaurus"
5470:
5468:
5467:
5463:
5461:
5460:
5456:
5454:
5453:
5449:
5448:
5446:
5444:
5440:
5434:
5433:
5429:
5426:
5425:
5421:
5418:
5417:
5413:
5411:
5410:
5406:
5404:
5403:
5399:
5397:
5396:
5392:
5391:
5389:
5387:
5383:
5377:
5376:
5372:
5370:
5369:
5368:Pelecanimimus
5365:
5363:
5362:
5358:
5357:
5354:
5351:
5349:
5345:
5339:
5338:
5334:
5331:
5330:
5329:Thecocoelurus
5326:
5324:
5323:
5319:
5317:
5316:
5312:
5310:
5309:
5308:Kinnareemimus
5305:
5303:
5302:
5298:
5295:
5294:
5290:
5287:
5286:
5282:
5280:
5279:
5275:
5274:
5271:
5268:
5266:
5262:
5255:
5254:
5250:
5249:
5246:
5243:
5240:
5235:
5225:
5224:
5220:
5218:
5217:
5213:
5211:
5210:
5206:
5203:
5202:
5198:
5196:
5195:
5191:
5189:
5188:
5187:Huaxiagnathus
5184:
5182:
5181:
5180:Compsognathus
5177:
5174:
5173:
5169:
5167:
5166:
5162:
5159:
5158:
5154:
5153:
5151:
5149:
5145:
5139:
5138:
5134:
5133:
5130:
5126:
5123:
5120:
5115:
5109:
5108:
5107:Ornitholestes
5104:
5102:
5101:
5097:
5094:
5093:
5089:
5088:
5085:
5082:
5079:
5074:
5070:
5066:
5061:
5057:
5047:
5042:
5040:
5033:
5027:
5005:
5004:
5000:
4998:
4997:
4993:
4991:
4990:
4986:
4984:
4983:
4979:
4978:
4976:
4972:
4968:
4951:
4950:
4949:
4948:
4946:
4944:
4940:
4936:
4910:
4909:
4905:
4903:
4902:
4901:Tyrannosaurus
4898:
4896:
4895:
4891:
4890:
4888:
4886:
4882:
4876:
4875:
4874:Teratophoneus
4871:
4869:
4868:
4864:
4862:
4861:
4857:
4856:
4854:
4852:
4848:
4842:
4841:
4837:
4835:
4834:
4830:
4829:
4827:
4825:
4821:
4815:
4814:
4810:
4808:
4807:
4803:
4802:
4800:
4798:
4794:
4788:
4787:
4783:
4781:
4780:
4776:
4775:
4772:
4769:
4767:
4763:
4757:
4756:
4752:
4750:
4749:
4748:Albertosaurus
4745:
4744:
4742:
4740:
4736:
4733:
4730:
4725:
4719:
4718:
4714:
4712:
4711:
4707:
4705:
4704:
4700:
4699:
4696:
4693:
4690:
4685:
4674:
4673:
4669:
4667:
4666:
4665:Xiongguanlong
4662:
4660:
4659:
4655:
4652:
4651:
4647:
4645:
4644:
4643:Suskityrannus
4640:
4638:
4637:
4636:Stokesosaurus
4633:
4631:
4630:
4629:Santanaraptor
4626:
4624:
4623:
4619:
4617:
4616:
4612:
4610:
4609:
4605:
4603:
4602:
4598:
4596:
4595:
4591:
4588:
4587:
4583:
4581:
4580:
4576:
4574:
4573:
4572:Alectrosaurus
4569:
4568:
4565:
4561:
4558:
4555:
4550:
4540:
4539:
4535:
4533:
4532:
4528:
4526:
4525:
4521:
4519:
4518:
4514:
4512:
4511:
4507:
4506:
4504:
4502:
4498:
4491:
4490:
4489:Tanycolagreus
4486:
4484:
4483:
4479:
4478:
4476:
4473:
4469:
4462:
4461:
4457:
4455:
4454:
4450:
4449:
4446:
4442:
4439:
4436:
4431:
4428:
4425:
4420:
4414:
4413:
4409:
4407:
4406:
4402:
4400:
4399:
4395:
4392:
4391:
4387:
4385:
4384:
4380:
4377:
4376:
4372:
4369:
4368:
4364:
4362:
4361:
4357:
4354:
4353:
4352:Bahariasaurus
4349:
4347:
4346:
4342:
4340:
4339:
4335:
4334:
4331:
4328:
4325:
4324:Coelurosauria
4320:
4316:
4312:
4311:Coelurosauria
4307:
4303:
4293:
4288:
4286:
4281:
4274:
4268:
4243:
4242:
4241:
4240:
4238:
4236:
4235:Coelurosauria
4232:
4222:
4221:
4217:
4215:
4214:
4210:
4208:
4207:
4203:
4201:
4200:
4196:
4194:
4193:
4189:
4187:
4186:
4182:
4180:
4179:
4175:
4174:
4172:
4170:
4166:
4160:
4158:
4154:
4152:
4150:
4146:
4144:
4143:
4139:
4137:
4136:
4132:
4130:
4129:
4125:
4122:
4121:
4117:
4114:
4113:
4112:Bahariasaurus
4109:
4107:
4106:
4102:
4101:
4098:
4095:
4092:
4088:
4084:
4054:
4053:
4049:
4047:
4046:
4042:
4040:
4039:
4035:
4033:
4032:
4028:
4027:
4025:
4023:
4019:
4013:
4012:
4008:
4007:
4004:
4001:
3999:
3995:
3989:
3988:
3984:
3982:
3981:
3977:
3975:
3974:
3970:
3968:
3967:
3963:
3961:
3960:
3956:
3954:
3953:
3952:Kelmayisaurus
3949:
3947:
3946:
3942:
3940:
3939:
3935:
3932:
3931:
3927:
3925:
3924:
3920:
3919:
3916:
3913:
3911:
3907:
3900:
3899:
3895:
3892:
3891:
3887:
3885:
3884:
3880:
3877:
3876:
3872:
3869:
3868:
3864:
3863:
3861:
3859:
3855:
3849:
3848:
3844:
3842:
3841:
3837:
3834:
3833:
3829:
3827:
3826:
3822:
3820:
3819:
3815:
3813:
3812:
3808:
3807:
3804:
3801:
3799:
3795:
3788:
3787:
3786:Saurophaganax
3783:
3780:
3779:
3775:
3772:
3771:
3767:
3765:
3764:
3760:
3759:
3757:
3755:
3751:
3748:
3745:
3740:
3726:
3725:
3721:
3719:
3718:
3717:Siamotyrannus
3714:
3712:
3711:
3707:
3705:
3704:
3700:
3698:
3697:
3693:
3692:
3690:
3688:
3684:
3678:
3677:
3673:
3670:
3669:
3665:
3664:
3661:
3658:
3656:
3652:
3645:
3644:
3640:
3637:
3636:
3632:
3630:
3629:
3625:
3623:
3622:
3618:
3617:
3614:
3610:
3607:
3604:
3603:Allosauroidea
3599:
3592:
3591:
3590:Spinosauridae
3587:
3584:
3583:
3579:
3576:
3575:
3571:
3569:
3568:
3564:
3563:
3560:
3557:
3554:
3549:
3542:
3541:
3537:
3534:
3533:
3529:
3528:
3525:
3522:
3519:
3514:
3510:
3506:
3501:
3497:
3487:
3482:
3480:
3473:
3443:
3442:
3441:
3440:
3438:
3436:
3432:
3428:
3406:
3405:
3401:
3399:
3398:
3394:
3393:
3391:
3389:
3385:
3379:
3378:
3374:
3372:
3371:
3367:
3365:
3364:
3360:
3358:
3357:
3353:
3350:
3349:
3345:
3344:
3341:
3338:
3336:
3335:Spinosaurinae
3332:
3322:
3321:
3317:
3315:
3314:
3313:Riparovenator
3310:
3308:
3307:
3306:Ceratosuchops
3303:
3302:
3300:
3298:
3294:
3287:
3286:
3282:
3279:
3278:
3274:
3272:
3271:
3270:Riojavenatrix
3267:
3265:
3264:
3260:
3258:
3257:
3253:
3252:
3249:
3246:
3244:
3243:Baryonychinae
3240:
3233:
3232:
3228:
3226:
3225:
3221:
3219:
3218:
3214:
3211:
3210:
3206:
3205:
3202:
3199:
3196:
3195:Spinosauridae
3191:
3176:
3175:
3171:
3169:
3168:
3164:
3162:
3161:
3157:
3155:
3154:
3150:
3148:
3147:
3143:
3141:
3140:
3136:
3135:
3133:
3131:
3127:
3121:
3120:
3119:Wiehenvenator
3116:
3114:
3113:
3109:
3107:
3106:
3102:
3100:
3099:
3095:
3094:
3092:
3090:
3086:
3080:
3079:
3075:
3074:
3071:
3068:
3066:
3062:
3055:
3054:
3050:
3049:
3046:
3042:
3039:
3036:
3031:
3020:
3019:
3015:
3013:
3012:
3008:
3006:
3005:
3001:
2999:
2998:
2994:
2993:
2991:
2989:
2985:
2979:
2978:
2974:
2973:
2970:
2966:
2963:
2960:
2955:
2952:
2949:
2944:
2938:
2937:
2933:
2931:
2930:
2926:
2923:
2922:
2918:
2916:
2915:
2911:
2909:
2908:
2904:
2902:
2901:
2897:
2895:
2894:
2890:
2888:
2887:
2883:
2880:
2879:
2875:
2873:
2872:
2868:
2865:
2864:
2860:
2857:
2856:
2855:Chienkosaurus
2852:
2849:
2848:
2844:
2843:
2840:
2837:
2834:
2829:
2825:
2821:
2816:
2812:
2802:
2795:
2788:
2781:
2755:
2754:
2753:
2752:
2750:
2748:
2744:
2740:
2714:
2713:
2709:
2707:
2706:
2702:
2700:
2699:
2695:
2693:
2692:
2688:
2685:
2684:
2680:
2678:
2677:
2673:
2671:
2670:
2666:
2664:
2663:
2659:
2657:
2656:
2652:
2651:
2649:
2647:
2646:Furileusauria
2643:
2637:
2636:
2632:
2630:
2629:
2625:
2622:
2621:
2620:Rahiolisaurus
2617:
2615:
2614:
2610:
2608:
2607:
2603:
2601:
2600:
2596:
2594:
2593:
2589:
2587:
2586:
2582:
2580:
2579:
2575:
2572:
2571:
2567:
2566:
2563:
2560:
2558:
2554:
2547:
2546:
2542:
2541:
2539:
2537:
2536:Carnotaurinae
2533:
2527:
2526:
2522:
2519:
2518:
2517:Rahiolisaurus
2514:
2512:
2511:
2510:Majungasaurus
2507:
2505:
2504:
2500:
2498:
2497:
2493:
2490:
2489:
2485:
2483:
2482:
2478:
2477:
2475:
2473:
2469:
2463:
2462:
2458:
2456:
2455:
2451:
2449:
2448:
2447:Tarascosaurus
2444:
2442:
2441:
2437:
2435:
2434:
2430:
2428:
2427:
2423:
2421:
2420:
2416:
2414:
2413:
2409:
2407:
2406:
2405:Eoabelisaurus
2402:
2399:
2398:
2394:
2392:
2391:
2390:Chenanisaurus
2387:
2386:
2383:
2380:
2377:
2376:Abelisauridae
2372:
2358:
2357:
2353:
2351:
2350:
2346:
2344:
2343:
2339:
2337:
2336:
2335:Masiakasaurus
2332:
2330:
2329:
2325:
2324:
2322:
2320:
2316:
2310:
2309:
2305:
2303:
2302:
2301:Huinculsaurus
2298:
2296:
2295:
2294:Elaphrosaurus
2291:
2290:
2288:
2286:
2282:
2276:
2275:
2271:
2269:
2268:
2264:
2262:
2261:
2257:
2254:
2253:
2249:
2246:
2245:
2241:
2238:
2237:
2233:
2230:
2229:
2225:
2223:
2222:
2218:
2215:
2214:
2213:Austrocheirus
2210:
2207:
2206:
2202:
2201:
2198:
2195:
2193:
2189:
2182:
2181:
2177:
2174:
2173:
2169:
2166:
2165:
2161:
2158:
2157:
2153:
2152:
2149:
2145:
2142:
2139:
2134:
2124:
2123:
2119:
2117:
2116:
2112:
2111:
2109:
2107:
2103:
2097:
2096:
2092:
2089:
2088:
2084:
2082:
2081:
2077:
2074:
2073:
2069:
2067:
2066:
2065:Berberosaurus
2062:
2061:
2058:
2054:
2051:
2048:
2043:
2036:
2035:
2031:
2029:
2028:
2024:
2023:
2020:
2017:
2014:
2009:
2005:
2001:
1996:
1992:
1983:
1977:
1954:
1953:
1952:Halticosaurus
1949:
1947:
1946:
1945:Teinurosaurus
1942:
1941:
1939:
1935:
1926:
1925:
1924:
1923:
1921:
1919:
1915:
1905:
1904:
1900:
1897:
1896:
1892:
1890:
1889:
1885:
1882:
1881:
1877:
1874:
1873:
1872:Megapnosaurus
1869:
1867:
1866:
1862:
1860:
1859:
1855:
1853:
1852:
1848:
1847:
1845:
1843:
1842:Coelophysidae
1839:
1833:
1832:
1831:Powellvenator
1828:
1826:
1825:
1821:
1819:
1818:
1814:
1812:
1811:
1807:
1804:
1803:
1799:
1798:
1795:
1792:
1790:
1786:
1780:
1779:
1775:
1772:
1771:
1767:
1765:
1764:
1760:
1758:
1757:
1753:
1751:
1750:
1746:
1744:
1743:
1739:
1737:
1736:
1732:
1730:
1729:
1725:
1723:
1722:
1721:Liliensternus
1718:
1716:
1715:
1711:
1709:
1708:
1704:
1702:
1701:
1697:
1694:
1693:
1692:Dolichosuchus
1689:
1687:
1686:
1685:Dilophosaurus
1682:
1681:
1678:
1675:
1672:
1667:
1660:
1659:
1658:Herrerasauria
1655:
1652:
1651:
1647:
1644:
1643:
1639:
1636:
1635:
1631:
1629:
1628:
1624:
1621:
1620:
1616:
1613:
1612:
1611:Daemonosaurus
1608:
1605:
1604:
1600:
1599:
1596:
1593:
1590:
1585:
1581:
1577:
1572:
1568:
1547:
1546:
1545:
1544:
1542:
1540:
1536:
1528:
1524:
1523:
1522:
1521:
1518:
1515:
1513:
1509:
1505:
1497:
1493:
1490:
1488:
1484:
1481:
1479:
1475:
1473:
1469:
1467:
1463:
1462:
1457:
1453:
1449:
1442:
1437:
1435:
1430:
1428:
1423:
1422:
1419:
1413:
1411:
1407:
1406:
1393:
1389:
1385:
1383:9780080494746
1379:
1375:
1368:
1360:
1356:
1352:
1348:
1344:
1340:
1336:
1332:
1325:
1323:
1314:
1310:
1306:
1302:
1298:
1294:
1290:
1286:
1278:
1270:
1266:
1261:
1256:
1252:
1248:
1244:
1240:
1236:
1229:
1221:
1219:9780520941434
1215:
1211:
1204:
1196:
1194:9788416641154
1190:
1186:
1179:
1171:
1167:
1163:
1159:
1155:
1151:
1147:
1140:
1132:
1130:9780375824197
1126:
1122:
1115:
1107:
1105:9781400836154
1101:
1097:
1090:
1088:
1086:
1077:
1075:9780253352897
1071:
1067:
1060:
1058:
1049:
1045:
1041:
1037:
1033:
1029:
1022:
1014:
1010:
1005:
1000:
996:
992:
988:
984:
980:
976:
972:
965:
957:
953:
949:
942:
940:
938:
936:
934:
932:
917:
916:"Abelisaurus"
911:
907:
897:
894:
893:
887:
885:
884:
883:Neuquensaurus
879:
878:
877:Barrosasaurus
873:
872:
867:
866:
861:
858:
857:titanosaurian
854:
849:
847:
843:
839:
835:
831:
824:
819:
810:
808:
804:
800:
796:
795:Carnotaurinae
792:
788:
784:
780:
779:
778:Majungasaurus
774:
770:
769:
763:
761:
757:
753:
752:Abelisauridae
750:
746:
739:
737:
731:
722:
720:
716:
711:
709:
705:
701:
700:lacrimal bone
697:
693:
689:
685:
684:
664:
661:
657:
653:
644:
637:
632:
623:
621:
617:
613:
609:
605:
601:
599:
593:
591:
587:
584:stage of the
583:
579:
575:
572:
568:
564:
560:
559:Neuquén Group
557:(part of the
556:
552:
548:
544:
540:
538:
533:
529:
525:
521:
516:
514:
510:
507:
506:specific name
503:
501:
490:
486:
482:
478:
477:type specimen
474:
470:
467:
464:
460:
457:
449:
445:
440:
431:
429:
425:
422:
418:
417:South America
414:
410:
407:
403:
400:
397:
394:of predatory
393:
387:
342:
341:
328:
322:
320:
316:
315:
313:
310:
306:
301:
297:
292:
291:
283:
280:
276:
269:
268:
261:
258:
257:
254:
248:
245:
244:
241:
240:Furileusauria
235:
232:
229:
228:
225:
224:Abelisauridae
219:
216:
215:
212:
209:
206:
203:
202:
199:
196:
193:
190:
189:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
153:
150:
147:
146:
141:
136:
132:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
5741:Abelisaurids
5634:
5598:
5527:
5520:
5513:
5506:
5499:
5494:Ornithomimus
5492:
5485:
5478:
5471:
5464:
5457:
5450:
5432:Tyrannomimus
5430:
5422:
5414:
5407:
5402:Deinocheirus
5400:
5393:
5373:
5366:
5359:
5335:
5327:
5322:Nqwebasaurus
5320:
5315:Nedcolbertia
5313:
5306:
5299:
5293:Calamosaurus
5291:
5285:Aviatyrannis
5283:
5276:
5251:
5221:
5214:
5207:
5199:
5192:
5185:
5178:
5170:
5165:Aristosuchus
5163:
5157:Aniksosaurus
5155:
5137:Aniksosaurus
5135:
5105:
5098:
5090:
5036:
5001:
4994:
4987:
4980:
4906:
4899:
4892:
4872:
4865:
4860:Dynamoterror
4858:
4838:
4831:
4811:
4804:
4784:
4779:Asiatyrannus
4777:
4753:
4746:
4717:Dryptosaurus
4715:
4708:
4701:
4670:
4663:
4656:
4648:
4641:
4634:
4627:
4620:
4613:
4608:Jinbeisaurus
4606:
4599:
4592:
4584:
4579:Aviatyrannis
4577:
4570:
4536:
4531:Sinotyrannus
4529:
4522:
4515:
4508:
4487:
4480:
4458:
4451:
4410:
4403:
4396:
4388:
4381:
4373:
4365:
4360:Bicentenaria
4358:
4350:
4345:Asiamericana
4343:
4336:
4277:
4218:
4211:
4204:
4197:
4190:
4183:
4176:
4155:
4147:
4140:
4133:
4126:
4120:Deltadromeus
4118:
4110:
4103:
4052:Tyrannotitan
4050:
4043:
4036:
4029:
4009:
3985:
3980:Taurovenator
3978:
3971:
3964:
3959:Lajasvenator
3957:
3950:
3943:
3938:Concavenator
3936:
3928:
3921:
3896:
3888:
3881:
3873:
3865:
3845:
3838:
3830:
3823:
3816:
3809:
3784:
3776:
3768:
3761:
3754:Allosauridae
3722:
3715:
3710:Shidaisaurus
3708:
3701:
3694:
3674:
3666:
3641:
3633:
3626:
3619:
3588:
3580:
3572:
3565:
3538:
3530:
3518:Avetheropoda
3505:Avetheropoda
3476:
3435:Avetheropoda
3402:
3395:
3388:Spinosaurini
3375:
3368:
3361:
3354:
3346:
3318:
3311:
3304:
3283:
3275:
3268:
3263:Protathlitis
3261:
3254:
3229:
3222:
3215:
3207:
3172:
3165:
3158:
3153:Leshansaurus
3151:
3144:
3137:
3117:
3110:
3105:Megalosaurus
3103:
3098:Duriavenator
3096:
3076:
3051:
3035:Megalosauria
3016:
3009:
3004:Marshosaurus
3002:
2997:Condorraptor
2995:
2975:
2934:
2927:
2919:
2912:
2905:
2898:
2891:
2884:
2878:Cruxicheiros
2876:
2869:
2861:
2853:
2845:
2710:
2705:Quilmesaurus
2703:
2696:
2689:
2681:
2674:
2667:
2660:
2654:
2653:
2633:
2626:
2618:
2611:
2604:
2597:
2590:
2583:
2576:
2568:
2557:Brachyrostra
2543:
2523:
2515:
2508:
2501:
2494:
2486:
2479:
2459:
2454:Tralkasaurus
2452:
2445:
2438:
2431:
2424:
2417:
2410:
2403:
2395:
2388:
2356:Vespersaurus
2354:
2349:Velocisaurus
2347:
2340:
2333:
2326:
2306:
2299:
2292:
2272:
2265:
2258:
2250:
2244:Deltadromeus
2242:
2234:
2228:Compsosuchus
2226:
2219:
2211:
2203:
2178:
2170:
2164:Lametasaurus
2162:
2154:
2120:
2115:Ceratosaurus
2113:
2093:
2085:
2078:
2070:
2063:
2047:Ceratosauria
2032:
2025:
1950:
1943:
1901:
1893:
1886:
1878:
1870:
1863:
1856:
1849:
1829:
1824:Podokesaurus
1822:
1815:
1808:
1800:
1776:
1768:
1761:
1754:
1747:
1740:
1733:
1726:
1719:
1712:
1707:Gojirasaurus
1705:
1700:Dracovenator
1698:
1690:
1683:
1671:Neotheropoda
1656:
1648:
1640:
1634:Guaibasaurus
1632:
1625:
1617:
1609:
1603:Chindesaurus
1601:
1491:
1482:
1409:
1373:
1367:
1337:(1): 58â69.
1334:
1330:
1288:
1284:
1277:
1242:
1238:
1228:
1209:
1203:
1184:
1178:
1153:
1149:
1145:
1139:
1120:
1114:
1095:
1065:
1034:(1): 61â87.
1031:
1027:
1021:
978:
974:
964:
955:
951:
919:. Retrieved
910:
881:
875:
869:
863:
852:
850:
837:
829:
827:
822:
813:Paleobiology
806:
798:
786:
776:
772:
766:
764:
759:
744:
742:
734:
712:
691:
681:
665:
660:Thomas Holtz
651:
649:
635:
619:
615:
607:
596:
594:
566:
542:
541:
531:
523:
517:
508:
497:
458:
456:type species
454:Coining the
453:
447:
339:
338:
337:
326:
317:
289:
288:
279:Type species
266:
265:
253:Carnotaurini
230:
204:
191:
178:
126:
24:
18:
5665:Abelisaurus
5659:Wikispecies
5635:Abelisaurus
5559:Maniraptora
5548:Maniraptora
5529:Tototlmimus
5459:Anserimimus
5409:Garudimimus
5395:Beishanlong
5337:Valdoraptor
5278:Arkansaurus
5223:Xunmenglong
5100:Juravenator
4894:Tarbosaurus
4786:Nanuqsaurus
4755:Gorgosaurus
4672:Megaraptora
4658:Timurlengia
4390:Sciurumimus
4338:Aratasaurus
4206:Murusraptor
4128:Fukuiraptor
4091:Megaraptora
3973:Shaochilong
3945:Eocarcharia
3898:Megaraptora
3818:Lusovenator
3696:Alpkarakush
3553:Carnosauria
3404:Spinosaurus
3377:Siamosaurus
3277:Suchosaurus
3224:Iberospinus
3160:Magnosaurus
3139:Afrovenator
3112:Torvosaurus
2863:Chilesaurus
2676:Carnotaurus
2669:Caletodraco
2655:Abelisaurus
2481:Arcovenator
2328:Laevisuchus
2319:Noasaurinae
2221:Berthasaura
2192:Noasauridae
2122:Genyodectes
2087:Lukousaurus
2080:Jubbulpuria
1880:Panguraptor
1858:Coelophysis
1851:Camposaurus
1810:Panguraptor
1802:Dracoraptor
1778:Zupaysaurus
1763:Tachiraptor
1742:Sarcosaurus
1642:Nhandumirim
1410:Abelisaurus
1004:11336/37727
952:Ameghiniana
853:Abelisaurus
838:Abelisaurus
830:Abelisaurus
823:Abelisaurus
807:Abelisaurus
799:Abelisaurus
787:Abelisaurus
773:Carnotaurus
760:Abelisaurus
756:Carnosauria
745:Abelisaurus
738:Abelisaurus
683:Carnotaurus
652:Abelisaurus
636:Abelisaurus
626:Description
610:. In 2010,
602:might be a
567:Abelisaurus
543:Abelisaurus
532:Abelisaurus
511:honors the
509:comahuensis
448:Abelisaurus
419:. It was a
396:abelisaurid
340:Abelisaurus
267:Abelisaurus
127:Abelisaurus
36:74â70
25:Abelisaurus
5735:Categories
5487:Gallimimus
5416:Harpymimus
5361:Harpymimus
4982:Iliosuchus
4952:see belowâ
4797:Alioramini
4615:Juratyrant
4601:Eotyrannus
4586:Bagaraatan
4538:Yutyrannus
4472:Coeluridae
4398:Vayuraptor
4244:see belowâ
4220:Tratayenia
4213:Orkoraptor
4199:Megaraptor
4105:Aoniraptor
4038:Mapusaurus
3966:Sauroniops
3930:Altispinax
3883:Neovenator
3825:Siamraptor
3811:Datanglong
3778:Epanterias
3770:Antrodemus
3763:Allosaurus
3744:Allosauria
3567:Altispinax
3532:Gasosaurus
3444:see belowâ
3320:Suchomimus
2921:Sinosaurus
2756:see belowâ
2712:Viavenator
2662:Aucasaurus
2599:Ilokelesia
2570:Dahalokely
2525:Rajasaurus
2503:Indosaurus
2496:Genusaurus
2488:Dahalokely
2412:Indosuchus
2308:Limusaurus
2267:Ligabueino
2260:Kiyacursor
2252:Genusaurus
2236:Dahalokely
2156:Betasuchus
2027:Dornraptor
1927:see belowâ
1903:Segisaurus
1756:Sinosaurus
1619:Eodromaeus
1548:see belowâ
1496:Dinosauria
1478:Sauropsida
921:2024-08-08
902:References
862:, such as
768:Aucasaurus
715:premaxilla
598:Aucasaurus
481:Cipolletti
406:Cretaceous
319:Aucasaurus
198:Saurischia
185:Dinosauria
5563:includes
5508:Rativates
5501:Qiupalong
5201:Scipionyx
5194:Mirischia
4867:Lythronax
4806:Alioramus
4460:Labocania
4178:Aerosteon
3724:Sinraptor
3628:Erectopus
3363:Irritator
2948:Orionides
2833:Tetanurae
2820:Tetanurae
2747:Tetanurae
2683:Elemgasem
2606:Llukalkan
2585:Elemgasem
2342:Noasaurus
2205:Afromimus
2013:Averostra
2000:Averostra
1918:Averostra
1770:Velocipes
1589:Theropoda
1576:Theropoda
1539:Theropoda
1464:Kingdom:
1448:Theropoda
1392:436848919
1351:0272-4634
1156:: 71â89.
1013:0195-6671
860:sauropods
846:ecosystem
844:of their
840:were the
733:Tooth of
692:fenestrae
582:Campanian
571:Campanian
547:Patagonia
485:Argentina
463:Argentine
424:carnivore
413:Campanian
296:Bonaparte
211:Theropoda
158:Kingdom:
152:Eukaryota
31:Campanian
5644:Wikidata
5092:Coelurus
4517:Kileskus
4510:Guanlong
4482:Coelurus
4375:Gualicho
3875:Gualicho
3256:Baryonyx
2592:Guemesia
2419:Kryptops
2180:Ozraptor
1817:Pendraig
1472:Chordata
1470:Phylum:
1466:Animalia
1359:86153297
1269:15306329
981:: 1â16.
890:See also
614:renamed
600:garridoi
524:MC 11098
520:holotype
402:dinosaur
399:theropod
321:garridoi
309:Synonyms
217:Family:
172:Chordata
168:Phylum:
162:Animalia
148:Domain:
5705:4129823
5692:4822836
5679:4532776
5650:Q130921
4650:Timimus
4412:Zuolong
4142:Rapator
4045:Meraxes
3370:Oxalaia
2691:Koleken
1714:Lepidus
1476:Class:
1313:9582112
1293:Bibcode
1285:Science
1260:1691741
1158:Bibcode
1036:Bibcode
983:Bibcode
855:hunted
719:maxilla
688:keratin
675:⁄
513:Comahue
421:bipedal
259:Genus:
246:Tribe:
105:↓
5301:Hexing
4989:Kakuru
4594:Dilong
2635:Thanos
2613:Niebla
2433:Rugops
2426:Kurupi
1390:
1380:
1357:
1349:
1311:
1267:
1257:
1216:
1191:
1127:
1102:
1072:
1011:
749:family
537:palate
528:fossil
500:sauros
493:ÏαÏ
ÏÎżÏ
446:where
409:Period
329:, 2002
327:et al.
325:Coria
302:, 1985
298:&
5718:53946
5565:birds
4622:Moros
4149:Siats
3890:Siats
3832:Siats
1492:Clade
1483:Clade
1355:S2CID
791:basal
789:as a
618:into
561:) of
489:Greek
428:skull
392:genus
300:Novas
231:Clade
205:Clade
192:Clade
179:Clade
5687:GBIF
5557:see
4192:Maip
1650:Tawa
1525:see
1388:OCLC
1378:ISBN
1347:ISSN
1309:PMID
1265:PMID
1214:ISBN
1189:ISBN
1125:ISBN
1100:ISBN
1070:ISBN
1009:ISSN
880:and
775:and
518:The
471:and
442:The
44:Preê
5674:EoL
1339:doi
1301:doi
1289:280
1255:PMC
1247:doi
1243:271
1166:doi
1044:doi
999:hdl
991:doi
736:cf.
606:of
530:of
483:in
5737::
5715::
5702::
5689::
5676::
5661::
5646::
1494::
1485::
1386:.
1353:.
1345:.
1335:22
1333:.
1321:^
1307:.
1299:.
1287:.
1263:.
1253:.
1241:.
1237:.
1164:.
1154:69
1152:.
1084:^
1056:^
1042:.
1032:25
1030:.
1007:.
997:.
989:.
979:54
977:.
973:.
956:21
954:.
930:^
886:.
874:,
868:,
848:.
771:,
758:.
668:33
592:.
522:,
430:.
374:ÉË
323:?
233::
207::
194::
181::
94:Pg
38:Ma
33:,
5427:?
5419:?
5332:?
5296:?
5288:?
5256:?
5204:?
5175:?
5160:?
5095:?
4675:?
4653:?
4589:?
4492:?
4474:?
4463:?
4393:?
4378:?
4370:?
4355:?
4159:?
4151:?
4123:?
4115:?
4093:?
3933:?
3901:?
3893:?
3878:?
3870:?
3835:?
3789:?
3781:?
3773:?
3671:?
3646:?
3638:?
3593:?
3585:?
3577:?
3543:?
3535:?
3351:?
3288:?
3280:?
3234:?
3212:?
3177:?
3056:?
3021:?
2924:?
2881:?
2866:?
2858:?
2850:?
2686:?
2623:?
2573:?
2548:?
2520:?
2491:?
2400:?
2255:?
2247:?
2239:?
2231:?
2216:?
2208:?
2183:?
2175:?
2167:?
2159:?
2090:?
2075:?
2037:?
1898:?
1883:?
1875:?
1805:?
1773:?
1695:?
1661:?
1653:?
1645:?
1637:?
1622:?
1614:?
1606:?
1440:e
1433:t
1426:v
1394:.
1361:.
1341::
1315:.
1303::
1295::
1271:.
1249::
1222:.
1197:.
1172:.
1168::
1160::
1133:.
1108:.
1078:.
1050:.
1046::
1038::
1015:.
1001::
993::
985::
924:.
677:2
673:1
670:+
496:(
411:(
386:/
383:s
380:É
377:r
371:s
368:Ë
365:ÉȘ
362:l
359:É
356:b
353:Ë
350:É
347:/
343:(
286:â
263:â
250:â
237:â
221:â
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:ê
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
