847:
53:
719:
888:
74:
3267:
1129:
1088:
957:
762:
1022:
667:
1191:
998:
1063:
618:
Boobies hunt fish by diving from a height into the sea and pursuing their prey underwater. Facial air sacs under their skin cushion the impact with the water. Boobies are colonial breeders on islands and coasts. They normally lay one or more chalky-blue eggs on the ground or sometimes in a tree nest.
500:, meaning "stupid", as these tame birds had a habit of landing on board sailing ships, where they were easily captured and eaten. Owing to this, boobies are often mentioned as having been caught and eaten by shipwrecked sailors, notably
619:
Selective pressures, likely through competition for resource, have shaped the ecomorphology and foraging behaviours of the six species of boobies in the
Pacific.
1251:
3224:
3558:
3437:
2916:
1467:
Friesen, V. L.; Anderson, D. J.; Steeves, T.E.; Jones, H.; Schreiber, E.A. (2002). "Molecular support for species status of the Nazca Booby".
3476:
3244:
3140:
2767:
2757:
3150:
2892:
2827:
596:(when gannet diversity was at its highest), or because the booby fossil species record is as yet incomplete due to most localities being
3207:
3180:
2984:
2950:
2838:
2659:
1018:
eastern
Pacific from the islands in Baja California to the Galapagos islands and the Isla de la Plata in Ecuador and Malpelo in Colombia
3190:
3160:
3411:
1860:
3450:
2637:
1177:
1011:
1351:
3455:
1892:
1272:
Patterson, S.A.; Morris-Pocock, J.A.; Friesen, V.L (2011). "A multilocus phylogeny of the
Sulidae (Aves: Pelecaniformes)".
1526:
1451:
1320:
Ornithologie, ou, Méthode contenant la division des oiseaux en ordres, sections, genres, especes & leurs variétés
3525:
3390:
613:
3481:
1243:
3403:
2652:
1547:
van Oordt, F.; Torres-Mura, J. C.; Hertel, F. (2018). "Ecomorphology and foraging behaviour of
Pacific boobies".
563:. The distinct lineages of gannets and boobies are known to have existed in such form, since at least the Middle
1480:
3463:
3346:
3333:
814:
17:
3351:
2645:
1828:
1782:
1736:
1690:
1644:
1598:
901:
3266:
597:
3530:
3491:
2335:
1413:
1379:
846:
1350:. Vol. 1 (2nd ed.). Cambridge, Massachusetts: Museum of Comparative Zoology. p. 181.
52:
2470:
2365:
1885:
1315:
1077:
636:
455:
152:
2667:
1978:
1418:
1384:
831:
73:
3517:
3286:
2345:
680:
31:
3442:
3512:
2551:
2432:
2406:
2340:
1988:
1983:
1426:
1392:
1214:
1111:
1045:
980:
870:
742:
718:
3504:
3372:
2520:
2480:
2450:
1870:
1281:
715:
Gulf of
California down along the western coasts of Central and South America down to Peru
8:
2475:
2442:
2241:
2145:
1878:
1805:
1759:
1713:
1667:
1621:
1575:
601:
1853:
1285:
2618:
2370:
1998:
1484:
511:
475:
68:
887:
552:
3499:
3359:
2880:
2593:
2437:
2391:
2315:
2115:
2072:
1993:
1447:
1297:
532:
318:
60:
1488:
3553:
2546:
2396:
2350:
2268:
2172:
2092:
2025:
1923:
1823:
1777:
1731:
1685:
1639:
1593:
1556:
1518:
1476:
1289:
769:
705:
490:
424:
201:
3364:
2967:
2401:
2360:
2231:
2135:
1901:
1864:
1502:
1441:
895:
776:
173:
3468:
1345:
1327:
1323:
1293:
3398:
3385:
3324:
3234:
2613:
2608:
2603:
2411:
2355:
2320:
2236:
2077:
1070:
338:
1506:
843:
islands and coasts in the pantropical areas of the
Atlantic and Pacific oceans
3547:
3309:
3015:
2957:
2587:
2577:
2572:
2500:
2306:
1814:
1768:
1722:
1676:
1630:
1584:
1143:
674:
501:
3338:
1128:
3197:
3170:
3089:
3082:
3024:
3001:
2974:
2869:
2567:
2557:
2511:
2330:
2082:
1301:
818:
459:
262:
161:
3424:
3318:
3251:
3096:
3038:
2940:
2817:
2669:
2598:
2582:
2461:
2140:
2097:
2087:
1969:
1919:
1546:
1005:
756:
463:
282:
228:
1373:
761:
3416:
3075:
3066:
2923:
2906:
2850:
2797:
2778:
2733:
2718:
2712:
2562:
2531:
2382:
1905:
1407:
1341:
1168:
956:
578:
510:
and his adherents during their famous voyage after being set adrift by
505:
125:
1918:
1560:
1522:
551:. Some authorities consider that all ten species should be considered
3130:
3052:
2933:
2807:
2677:
2540:
2325:
2264:
1136:
1087:
560:
540:
471:
85:
3429:
3280:
666:
3303:
3214:
3031:
2700:
836:– Red Sea through the Indian Ocean to the west and central Pacific
699:
Milne-Edwards, 1882 – Pacific coast of
Southern and Middle America
593:
105:
1021:
584:
The fossil record of boobies is not as well documented as that of
3377:
3118:
3103:
2899:
2859:
2787:
2681:
2491:
2292:
2226:
2203:
2049:
2021:
1955:
1322:(in French and Latin). Vol. 1. Paris: Jean-Baptiste Bauche.
589:
564:
525:, while the three gannet species are usually placed in the genus
479:
427:
411:
135:
2167:
1190:
3045:
2694:
2217:
2168:
2126:
585:
527:
483:
431:
95:
997:
2991:
2706:
2423:
2281:
2192:
2176:
2038:
1944:
1927:
1271:
1062:
493:
415:
1900:
1162:(Linnaeus, 1766) – Caribbean and southwest Atlantic islands
1517:. Vol. 8. New York: Academic Press. pp. 79–238 .
1481:
10.1642/0004-8038(2002)119[0820:MSFSSO]2.0.CO;2
1466:
115:
2263:
588:, either because booby speciation was lower from the late
2020:
547:, which represents an ancient lineage perhaps closer to
30:
This article is about the seabird. For other uses, see
1803:
1757:
1711:
1665:
1619:
1573:
1443:
The Bounty: The True Story of the Mutiny on the Bounty
559:. However, they are readily distinguished by means of
3240:
3230:
3220:
3203:
3186:
3176:
3166:
3156:
3146:
3136:
3126:
3063:
3012:
2997:
2980:
2963:
2946:
2929:
2912:
2888:
2865:
2833:
2823:
2813:
2803:
2793:
2763:
2753:
2447:
1513:. In Farner, D.S.; King, D.S.; Parkes, K.C. (eds.).
1405:
1187:
Sri Lanka, Christmas Island, eastern central
Pacific
1371:
489:The English name "booby" was possibly based on the
1829:10.2305/IUCN.UK.2018-2.RLTS.T22696694A132589278.en
1783:10.2305/IUCN.UK.2018-2.RLTS.T22696686A132589026.en
1737:10.2305/IUCN.UK.2018-2.RLTS.T22728990A132659882.en
1691:10.2305/IUCN.UK.2018-2.RLTS.T22736173A132666363.en
1645:10.2305/IUCN.UK.2018-2.RLTS.T22696698A132590197.en
1599:10.2305/IUCN.UK.2018-2.RLTS.T22696683A132588719.en
1406:
1372:
937:van Tets, Meredith, Fullagar & Davidson, 1988
823:– Pacific coasts of Central America and Colombia
3545:
614:Seabird breeding behavior § Mating dances
2653:
1886:
1507:"The Fossil Record of Birds (Section X.G.5.a
517:Six of the ten extant Sulidae species called
1340:
1265:
1171:, 1838 – tropical Pacific and Indian Oceans
445:
396:Cladogram showing the species in the genus
45:Temporal range: Miocene (Langhian) – recent
3062:
2660:
2646:
1893:
1879:
51:
1827:
1781:
1735:
1689:
1643:
1597:
1439:
478:word for the other member of the family
1314:
454:was introduced by the French zoologist
14:
3559:Taxa named by Mathurin Jacques Brisson
3546:
1460:
3285:
3284:
2731:
2641:
2262:
2166:
2019:
1917:
1874:
1501:
1344:; Cottrell, G. William, eds. (1979).
1274:Molecular Phylogenetics and Evolution
658:IUCN status and estimated population
430:. Boobies are closely related to the
3492:9b87fb12-8102-4109-a626-1d0910fdcd6f
3404:a5daf67e-1c11-4566-94ab-23f41cf5fc8d
2590:(imperial cormorant, blue-eyed shag)
1427:participating institution membership
1393:participating institution membership
1854:Videos of several Booby species on
1815:IUCN Red List of Threatened Species
1769:IUCN Red List of Threatened Species
1723:IUCN Red List of Threatened Species
1677:IUCN Red List of Threatened Species
1631:IUCN Red List of Threatened Species
1585:IUCN Red List of Threatened Species
1440:Alexander, Caroline (25 May 2004).
807:– Pacific coasts of USA and Mexico
438:), which were formerly included in
24:
622:
25:
3570:
1847:
1180:, 1898 – eastern central Pacific
798:– Caribbean and Atlantic Islands
3265:
1347:Check-list of Birds of the World
1189:
1127:
1086:
1061:
1020:
996:
955:
886:
845:
760:
717:
665:
72:
1804:BirdLife International (2018).
1797:
1758:BirdLife International (2018).
1751:
1712:BirdLife International (2018).
1705:
1666:BirdLife International (2018).
1659:
1620:BirdLife International (2018).
1613:
1574:BirdLife International (2018).
1567:
1540:
1529:from the original on 2021-04-10
1354:from the original on 2021-08-08
1254:from the original on 2021-08-08
649:Scientific name and subspecies
2732:
1495:
1433:
1399:
1365:
1334:
1308:
1236:
13:
1:
1229:
805:Nathaniel Stickney Goss, 1888
63:displaying by raising a foot
1989:Christmas Island frigatebird
1856:The Internet Bird Collection
607:
7:
2290:
2201:
2047:
1953:
1294:10.1016/j.ympev.2010.11.021
1244:"Sula Brisson 1760 (booby)"
708:, 1948 – Galápagos Islands
466:. The name is derived from
10:
3575:
953:islands in tropical oceans
611:
29:
3293:
3263:
3117:
3011:
2879:
2849:
2777:
2744:
2740:
2727:
2689:
2529:
2509:
2489:
2459:
2421:
2380:
2304:
2289:
2280:
2276:
2258:
2215:
2200:
2191:
2187:
2162:
2124:
2106:
2061:
2046:
2037:
2033:
2015:
1967:
1952:
1943:
1939:
1913:
1822:: e.T22696694A132589278.
1776:: e.T22696686A132589026.
1730:: e.T22728990A132659882.
1684:: e.T22736173A132666363.
1638:: e.T22696698A132590197.
1592:: e.T22696683A132588719.
1414:Oxford English Dictionary
1380:Oxford English Dictionary
1316:Brisson, Mathurin Jacques
1124:
1058:
993:
883:
755:
662:
657:
654:
651:
648:
645:
535:was formerly included in
446:Systematics and evolution
336:
316:
309:
280:
260:
253:
246:
226:
219:
199:
192:
167:
160:
69:Scientific classification
67:
59:
50:
41:
2684:and their extinct allies
2471:Double-crested cormorant
2366:White-breasted cormorant
456:Mathurin Jacques Brisson
2414:(long-tailed cormorant)
1979:Magnificent frigatebird
1419:Oxford University Press
1385:Oxford University Press
539:but is now placed in a
2543:(Magellanic cormorant)
2373:(Temminck's cormorant)
2346:Little black cormorant
1908:(Phalacrocoraciformes)
32:Booby (disambiguation)
3513:Paleobiology Database
2552:New Zealand king shag
2407:Little pied cormorant
2341:Black-faced cormorant
1984:Ascension frigatebird
1863:12 April 2016 at the
169:Pelecanus leucogaster
3399:Fauna Europaea (new)
2521:Red-legged cormorant
2481:Flightless cormorant
2451:Spectacled cormorant
2271:: Phalacrocoracidae)
2175:: Anhingidae ·
1926:: Fregatidae ·
2476:Neotropic cormorant
2443:Red-faced cormorant
2242:Australasian darter
2146:Australasian gannet
2022:Gannets and boobies
1417:(Online ed.).
1383:(Online ed.).
1286:2011MolPE..58..181P
642:
602:Southern Hemisphere
514:and his followers.
2619:South Georgia shag
2554:(rough-faced shag)
2433:Brandt's cormorant
2371:Japanese cormorant
1999:Lesser frigatebird
663:Blue-footed booby
627:
512:Fletcher Christian
3541:
3540:
3500:Open Tree of Life
3287:Taxon identifiers
3278:
3277:
3274:
3273:
3261:
3260:
3113:
3112:
2881:Phalacrocoracidae
2635:
2634:
2631:
2630:
2627:
2626:
2594:Heard Island shag
2438:Pelagic cormorant
2392:Crowned cormorant
2316:Socotra cormorant
2300:
2299:
2254:
2253:
2250:
2249:
2211:
2210:
2158:
2157:
2154:
2153:
2073:Blue-footed booby
2057:
2056:
2011:
2010:
2007:
2006:
1994:Great frigatebird
1963:
1962:
1561:10.1111/ibi.12545
1446:. Penguin Books.
1425:(Subscription or
1391:(Subscription or
1227:
1226:
1147:
1125:Red-footed booby
1081:
1015:
946:
938:
930:
922:
905:
835:
822:
806:
797:
793:S. l. leucogaster
780:
684:
655:Size and ecology
640:
521:are in the genus
404:
403:
390:
389:
381:
380:
372:
371:
363:
362:
354:
353:
319:Blue-footed booby
298:
297:
181:
180:
156:
61:Blue-footed booby
16:(Redirected from
3566:
3534:
3533:
3521:
3520:
3508:
3507:
3495:
3494:
3485:
3484:
3472:
3471:
3469:NHMSYS0000533779
3459:
3458:
3446:
3445:
3433:
3432:
3420:
3419:
3407:
3406:
3394:
3393:
3381:
3380:
3368:
3367:
3355:
3354:
3342:
3341:
3329:
3328:
3327:
3314:
3313:
3312:
3282:
3281:
3269:
3242:
3232:
3225:Prophalacrocorax
3222:
3205:
3188:
3178:
3168:
3158:
3148:
3138:
3128:
3065:
3060:
3059:
3014:
2999:
2982:
2965:
2948:
2931:
2914:
2890:
2867:
2835:
2825:
2815:
2805:
2795:
2765:
2755:
2742:
2741:
2729:
2728:
2662:
2655:
2648:
2639:
2638:
2547:Guanay cormorant
2449:
2397:Little cormorant
2351:Indian cormorant
2287:
2286:
2278:
2277:
2260:
2259:
2198:
2197:
2189:
2188:
2164:
2163:
2093:Red-footed booby
2044:
2043:
2035:
2034:
2017:
2016:
1950:
1949:
1941:
1940:
1915:
1914:
1895:
1888:
1881:
1872:
1871:
1841:
1840:
1838:
1836:
1831:
1801:
1795:
1794:
1792:
1790:
1785:
1755:
1749:
1748:
1746:
1744:
1739:
1709:
1703:
1702:
1700:
1698:
1693:
1663:
1657:
1656:
1654:
1652:
1647:
1624:Sula leucogaster
1617:
1611:
1610:
1608:
1606:
1601:
1571:
1565:
1564:
1544:
1538:
1537:
1535:
1534:
1503:Olson, Storrs L.
1499:
1493:
1492:
1464:
1458:
1457:
1437:
1431:
1430:
1422:
1410:
1403:
1397:
1396:
1388:
1376:
1369:
1363:
1362:
1360:
1359:
1338:
1332:
1331:
1312:
1306:
1305:
1269:
1263:
1262:
1260:
1259:
1240:
1218:
1193:
1154:Three subspecies
1141:
1131:
1115:
1090:
1075:
1065:
1049:
1024:
1010:
1000:
984:
959:
944:
936:
928:
920:
918:S. d. dactylatra
900:
890:
874:
849:
829:
813:
804:
796:(Boddaert, 1783)
795:
774:
770:Sula leucogaster
764:
746:
721:
679:
669:
643:
635:
626:
577:
573:
312:
311:
256:
255:
249:
248:
233:Sula leucogaster
222:
221:
202:Red-footed booby
195:
194:
188:
187:
183:
182:
151:
77:
76:
55:
39:
38:
21:
3574:
3573:
3569:
3568:
3567:
3565:
3564:
3563:
3544:
3543:
3542:
3537:
3529:
3524:
3516:
3511:
3503:
3498:
3490:
3488:
3480:
3475:
3467:
3462:
3454:
3449:
3441:
3436:
3428:
3423:
3415:
3410:
3402:
3397:
3389:
3384:
3376:
3371:
3363:
3358:
3350:
3345:
3337:
3332:
3323:
3322:
3317:
3308:
3307:
3302:
3289:
3279:
3270:
3257:
3109:
3058:
3007:
2875:
2845:
2773:
2736:
2723:
2685:
2666:
2636:
2623:
2525:
2505:
2485:
2455:
2417:
2402:Pygmy cormorant
2376:
2361:Great cormorant
2296:
2272:
2246:
2232:Oriental darter
2207:
2183:
2150:
2136:Northern gannet
2120:
2102:
2053:
2029:
2003:
1959:
1935:
1909:
1899:
1865:Wayback Machine
1850:
1845:
1844:
1834:
1832:
1802:
1798:
1788:
1786:
1756:
1752:
1742:
1740:
1710:
1706:
1696:
1694:
1670:Sula dactylatra
1664:
1660:
1650:
1648:
1618:
1614:
1604:
1602:
1572:
1568:
1545:
1541:
1532:
1530:
1500:
1496:
1465:
1461:
1454:
1438:
1434:
1424:
1404:
1400:
1390:
1370:
1366:
1357:
1355:
1339:
1335:
1313:
1309:
1270:
1266:
1257:
1255:
1242:
1241:
1237:
1232:
1222:
1216:
1207:
1206:
1201:
1200:
1188:
1184:
1183:
1156:
1155:
1149:
1148:
1140:
1119:
1113:
1104:
1103:
1098:
1097:
1085:
1074:
1059:Peruvian booby
1053:
1047:
1038:
1037:
1032:
1031:
1019:
1009:
988:
982:
973:
972:
967:
966:
954:
950:
949:
942:S. d. personata
914:
913:
912:Four subspecies
907:
906:
899:
896:Sula dactylatra
878:
872:
863:
862:
857:
856:
844:
840:
839:
802:S. l. brewsteri
789:
788:
787:Four subspecies
782:
781:
773:
750:
744:
735:
734:
729:
728:
716:
712:
711:
693:
692:
686:
685:
678:
625:
623:List of species
616:
610:
575:
571:
448:
391:
382:
373:
364:
355:
299:
267:Sula dactylatra
177:
171:
150:
71:
46:
43:
35:
28:
23:
22:
15:
12:
11:
5:
3572:
3562:
3561:
3556:
3539:
3538:
3536:
3535:
3522:
3509:
3496:
3486:
3473:
3460:
3447:
3434:
3421:
3408:
3395:
3386:Fauna Europaea
3382:
3369:
3356:
3343:
3330:
3315:
3299:
3297:
3291:
3290:
3276:
3275:
3272:
3271:
3264:
3262:
3259:
3258:
3256:
3255:
3248:
3238:
3235:Rhamphastosula
3228:
3218:
3211:
3201:
3194:
3184:
3174:
3164:
3154:
3144:
3134:
3123:
3121:
3115:
3114:
3111:
3110:
3108:
3107:
3100:
3093:
3086:
3079:
3071:
3069:
3057:
3056:
3049:
3042:
3035:
3028:
3020:
3018:
3009:
3008:
3006:
3005:
2995:
2988:
2978:
2971:
2961:
2954:
2944:
2937:
2927:
2920:
2910:
2903:
2896:
2885:
2883:
2877:
2876:
2874:
2873:
2863:
2855:
2853:
2847:
2846:
2844:
2843:
2831:
2821:
2811:
2801:
2791:
2783:
2781:
2775:
2774:
2772:
2771:
2761:
2750:
2748:
2746:incertae sedis
2738:
2737:
2725:
2724:
2722:
2721:
2715:
2709:
2703:
2697:
2690:
2687:
2686:
2665:
2664:
2657:
2650:
2642:
2633:
2632:
2629:
2628:
2625:
2624:
2622:
2621:
2616:
2614:Macquarie shag
2611:
2609:Kerguelen shag
2606:
2604:Antarctic shag
2601:
2596:
2591:
2585:
2580:
2575:
2570:
2565:
2560:
2555:
2549:
2544:
2537:
2535:
2527:
2526:
2524:
2523:
2517:
2515:
2507:
2506:
2504:
2503:
2497:
2495:
2487:
2486:
2484:
2483:
2478:
2473:
2467:
2465:
2457:
2456:
2454:
2453:
2445:
2440:
2435:
2429:
2427:
2419:
2418:
2416:
2415:
2412:Reed cormorant
2409:
2404:
2399:
2394:
2388:
2386:
2378:
2377:
2375:
2374:
2368:
2363:
2358:
2356:Cape cormorant
2353:
2348:
2343:
2338:
2336:Pied cormorant
2333:
2328:
2323:
2321:Bank cormorant
2318:
2312:
2310:
2302:
2301:
2298:
2297:
2284:
2274:
2273:
2256:
2255:
2252:
2251:
2248:
2247:
2245:
2244:
2239:
2237:African darter
2234:
2229:
2223:
2221:
2213:
2212:
2209:
2208:
2195:
2185:
2184:
2160:
2159:
2156:
2155:
2152:
2151:
2149:
2148:
2143:
2138:
2132:
2130:
2122:
2121:
2119:
2118:
2116:Abbott's booby
2112:
2110:
2104:
2103:
2101:
2100:
2095:
2090:
2085:
2080:
2078:Peruvian booby
2075:
2069:
2067:
2059:
2058:
2055:
2054:
2041:
2031:
2030:
2013:
2012:
2009:
2008:
2005:
2004:
2002:
2001:
1996:
1991:
1986:
1981:
1975:
1973:
1965:
1964:
1961:
1960:
1947:
1937:
1936:
1911:
1910:
1898:
1897:
1890:
1883:
1875:
1869:
1868:
1849:
1848:External links
1846:
1843:
1842:
1796:
1762:Sula variegata
1750:
1704:
1658:
1612:
1566:
1555:(2): 313–326.
1539:
1494:
1475:(3): 820–826.
1459:
1453:978-0142004692
1452:
1432:
1398:
1364:
1333:
1307:
1280:(2): 181–191.
1264:
1234:
1233:
1231:
1228:
1225:
1224:
1217: LC
1212:
1194:
1185:
1182:
1181:
1175:S. s. websteri
1172:
1166:S. s. rubripes
1163:
1153:
1152:
1151:
1150:
1133:
1122:
1121:
1114: LC
1109:
1091:
1082:
1071:Sula variegata
1067:
1056:
1055:
1048: LC
1043:
1025:
1016:
1002:
991:
990:
983: LC
978:
960:
951:
948:
947:
939:
931:
929:Hartlaub, 1859
926:S. d. melanops
923:
911:
910:
909:
908:
892:
881:
880:
873: LC
868:
850:
841:
838:
837:
824:
811:S. l. etesiaca
808:
799:
786:
785:
784:
783:
766:
753:
752:
745: LC
740:
722:
713:
710:
709:
700:
697:S. n. nebouxii
691:Two subspecies
690:
689:
688:
687:
671:
660:
659:
656:
653:
650:
647:
641:– six species
624:
621:
609:
606:
533:Abbott's booby
458:in 1760. The
447:
444:
423:, part of the
402:
401:
393:
392:
388:
387:
384:
383:
379:
378:
375:
374:
370:
369:
366:
365:
361:
360:
357:
356:
352:
351:
348:
347:
343:Sula variegata
339:Peruvian booby
335:
332:
331:
328:
327:
315:
310:
308:
305:
304:
301:
300:
296:
295:
292:
291:
279:
276:
275:
272:
271:
259:
254:
252:
247:
245:
242:
241:
238:
237:
225:
220:
218:
215:
214:
211:
210:
198:
193:
191:
186:
179:
178:
172:
165:
164:
158:
157:
143:
139:
138:
133:
129:
128:
123:
119:
118:
113:
109:
108:
103:
99:
98:
93:
89:
88:
83:
79:
78:
65:
64:
57:
56:
48:
47:
44:
27:Genus of birds
26:
9:
6:
4:
3:
2:
3571:
3560:
3557:
3555:
3552:
3551:
3549:
3532:
3527:
3523:
3519:
3514:
3510:
3506:
3501:
3497:
3493:
3487:
3483:
3478:
3474:
3470:
3465:
3461:
3457:
3452:
3448:
3444:
3439:
3435:
3431:
3426:
3422:
3418:
3413:
3409:
3405:
3400:
3396:
3392:
3387:
3383:
3379:
3374:
3370:
3366:
3361:
3357:
3353:
3348:
3344:
3340:
3335:
3331:
3326:
3320:
3316:
3311:
3305:
3301:
3300:
3298:
3296:
3292:
3288:
3283:
3268:
3254:
3253:
3249:
3247:
3246:
3239:
3237:
3236:
3229:
3227:
3226:
3219:
3217:
3216:
3212:
3210:
3209:
3202:
3200:
3199:
3195:
3193:
3192:
3185:
3183:
3182:
3175:
3173:
3172:
3165:
3163:
3162:
3155:
3153:
3152:
3145:
3143:
3142:
3135:
3133:
3132:
3125:
3124:
3122:
3120:
3116:
3106:
3105:
3101:
3099:
3098:
3094:
3092:
3091:
3087:
3085:
3084:
3080:
3078:
3077:
3073:
3072:
3070:
3068:
3061:
3055:
3054:
3050:
3048:
3047:
3043:
3041:
3040:
3036:
3034:
3033:
3029:
3027:
3026:
3022:
3021:
3019:
3017:
3016:Plotopteridae
3010:
3004:
3003:
2996:
2994:
2993:
2989:
2987:
2986:
2979:
2977:
2976:
2972:
2970:
2969:
2962:
2960:
2959:
2958:Phalacrocorax
2955:
2953:
2952:
2945:
2943:
2942:
2938:
2936:
2935:
2928:
2926:
2925:
2921:
2919:
2918:
2911:
2909:
2908:
2904:
2902:
2901:
2897:
2895:
2894:
2887:
2886:
2884:
2882:
2878:
2872:
2871:
2864:
2862:
2861:
2857:
2856:
2854:
2852:
2848:
2841:
2840:
2832:
2830:
2829:
2822:
2820:
2819:
2812:
2810:
2809:
2802:
2800:
2799:
2792:
2790:
2789:
2785:
2784:
2782:
2780:
2776:
2770:
2769:
2762:
2760:
2759:
2752:
2751:
2749:
2747:
2743:
2739:
2735:
2730:
2726:
2720:
2716:
2714:
2710:
2708:
2704:
2702:
2698:
2696:
2692:
2691:
2688:
2683:
2679:
2675:
2671:
2663:
2658:
2656:
2651:
2649:
2644:
2643:
2640:
2620:
2617:
2615:
2612:
2610:
2607:
2605:
2602:
2600:
2597:
2595:
2592:
2589:
2588:Imperial shag
2586:
2584:
2581:
2579:
2578:Campbell shag
2576:
2574:
2573:Auckland shag
2571:
2569:
2566:
2564:
2561:
2559:
2556:
2553:
2550:
2548:
2545:
2542:
2539:
2538:
2536:
2534:
2533:
2528:
2522:
2519:
2518:
2516:
2514:
2513:
2508:
2502:
2501:European shag
2499:
2498:
2496:
2494:
2493:
2488:
2482:
2479:
2477:
2474:
2472:
2469:
2468:
2466:
2464:
2463:
2458:
2452:
2446:
2444:
2441:
2439:
2436:
2434:
2431:
2430:
2428:
2426:
2425:
2420:
2413:
2410:
2408:
2405:
2403:
2400:
2398:
2395:
2393:
2390:
2389:
2387:
2385:
2384:
2379:
2372:
2369:
2367:
2364:
2362:
2359:
2357:
2354:
2352:
2349:
2347:
2344:
2342:
2339:
2337:
2334:
2332:
2329:
2327:
2324:
2322:
2319:
2317:
2314:
2313:
2311:
2309:
2308:
2307:Phalacrocorax
2303:
2295:
2294:
2288:
2285:
2283:
2279:
2275:
2270:
2266:
2261:
2257:
2243:
2240:
2238:
2235:
2233:
2230:
2228:
2225:
2224:
2222:
2220:
2219:
2214:
2206:
2205:
2199:
2196:
2194:
2190:
2186:
2181:
2178:
2174:
2170:
2165:
2161:
2147:
2144:
2142:
2139:
2137:
2134:
2133:
2131:
2129:
2128:
2123:
2117:
2114:
2113:
2111:
2109:
2105:
2099:
2096:
2094:
2091:
2089:
2086:
2084:
2081:
2079:
2076:
2074:
2071:
2070:
2068:
2066:
2065:
2060:
2052:
2051:
2045:
2042:
2040:
2036:
2032:
2027:
2023:
2018:
2014:
2000:
1997:
1995:
1992:
1990:
1987:
1985:
1982:
1980:
1977:
1976:
1974:
1972:
1971:
1966:
1958:
1957:
1951:
1948:
1946:
1942:
1938:
1933:
1929:
1925:
1921:
1916:
1912:
1907:
1903:
1896:
1891:
1889:
1884:
1882:
1877:
1876:
1873:
1866:
1862:
1858:
1857:
1852:
1851:
1830:
1825:
1821:
1817:
1816:
1811:
1809:
1800:
1784:
1779:
1775:
1771:
1770:
1765:
1763:
1754:
1738:
1733:
1729:
1725:
1724:
1719:
1717:
1708:
1692:
1687:
1683:
1679:
1678:
1673:
1671:
1662:
1646:
1641:
1637:
1633:
1632:
1627:
1625:
1616:
1600:
1595:
1591:
1587:
1586:
1581:
1579:
1578:Sula nebouxii
1570:
1562:
1558:
1554:
1550:
1543:
1528:
1524:
1520:
1516:
1515:Avian Biology
1512:
1510:
1504:
1498:
1490:
1486:
1482:
1478:
1474:
1470:
1463:
1455:
1449:
1445:
1444:
1436:
1428:
1420:
1416:
1415:
1409:
1402:
1394:
1386:
1382:
1381:
1375:
1368:
1353:
1349:
1348:
1343:
1337:
1329:
1325:
1321:
1317:
1311:
1303:
1299:
1295:
1291:
1287:
1283:
1279:
1275:
1268:
1253:
1249:
1245:
1239:
1235:
1223:
1220:
1219:
1213:
1210:
1204:
1198:
1195:
1192:
1186:
1179:
1176:
1173:
1170:
1167:
1164:
1161:
1158:
1157:
1145:
1139:
1138:
1134:
1132:
1130:
1123:
1120:
1117:
1116:
1110:
1107:
1101:
1095:
1092:
1089:
1083:
1079:
1073:
1072:
1068:
1066:
1064:
1057:
1054:
1051:
1050:
1044:
1041:
1035:
1029:
1026:
1023:
1017:
1013:
1008:
1007:
1003:
1001:
999:
992:
989:
986:
985:
979:
976:
970:
964:
961:
958:
952:
943:
940:
935:
934:S. d. tasmani
932:
927:
924:
919:
916:
915:
903:
898:
897:
893:
891:
889:
884:Masked booby
882:
879:
876:
875:
869:
866:
860:
854:
851:
848:
842:
833:
828:
825:
820:
816:
812:
809:
803:
800:
794:
791:
790:
778:
772:
771:
767:
765:
763:
758:
754:
751:
748:
747:
741:
738:
732:
726:
723:
720:
714:
707:
704:
701:
698:
695:
694:
682:
681:Milne-Edwards
677:
676:
675:Sula nebouxii
672:
670:
668:
661:
644:
638:
633:
632:
620:
615:
605:
603:
599:
595:
591:
587:
582:
580:
570:
566:
562:
558:
554:
550:
546:
542:
538:
534:
530:
529:
524:
520:
515:
513:
509:
508:
503:
502:William Bligh
499:
495:
492:
487:
485:
481:
477:
473:
469:
465:
461:
457:
453:
443:
441:
437:
433:
429:
426:
422:
421:
417:
413:
409:
399:
395:
394:
386:
385:
377:
376:
368:
367:
359:
358:
350:
349:
346:
344:
340:
334:
333:
330:
329:
326:
324:
323:Sula nebouxii
320:
314:
313:
307:
306:
303:
302:
294:
293:
290:
288:
284:
278:
277:
274:
273:
270:
268:
264:
258:
257:
251:
250:
244:
243:
240:
239:
236:
234:
230:
224:
223:
217:
216:
213:
212:
209:
207:
203:
197:
196:
190:
189:
185:
184:
175:
170:
166:
163:
159:
154:
149:
148:
144:
141:
140:
137:
134:
131:
130:
127:
124:
121:
120:
117:
114:
111:
110:
107:
104:
101:
100:
97:
94:
91:
90:
87:
84:
81:
80:
75:
70:
66:
62:
58:
54:
49:
40:
37:
33:
19:
3294:
3250:
3243:
3233:
3223:
3213:
3206:
3196:
3189:
3179:
3171:Masillastega
3169:
3159:
3149:
3139:
3129:
3102:
3095:
3090:Klallamornis
3088:
3083:Hokkaidornis
3081:
3074:
3051:
3044:
3037:
3030:
3025:Empeirodytes
3023:
3002:Valenticarbo
3000:
2990:
2983:
2975:Poikilocarbo
2973:
2966:
2956:
2949:
2939:
2932:
2922:
2917:Limicorallus
2915:
2905:
2898:
2891:
2870:Limnofregata
2868:
2858:
2837:
2826:
2816:
2806:
2796:
2786:
2766:
2756:
2745:
2711:Infraclass:
2673:
2670:frigatebirds
2568:Chatham shag
2558:Foveaux shag
2530:
2512:Poikilocarbo
2510:
2490:
2460:
2422:
2381:
2331:Spotted shag
2305:
2291:
2216:
2202:
2179:
2125:
2107:
2083:Masked booby
2063:
2062:
2048:
1968:
1954:
1931:
1920:Frigatebirds
1855:
1833:. Retrieved
1819:
1813:
1807:
1799:
1787:. Retrieved
1773:
1767:
1761:
1753:
1741:. Retrieved
1727:
1721:
1715:
1707:
1695:. Retrieved
1681:
1675:
1669:
1661:
1649:. Retrieved
1635:
1629:
1623:
1615:
1603:. Retrieved
1589:
1583:
1577:
1569:
1552:
1548:
1542:
1531:. Retrieved
1514:
1508:
1497:
1472:
1468:
1462:
1442:
1435:
1412:
1401:
1378:
1367:
1356:. Retrieved
1346:
1336:
1328:Vol. 6 p.494
1324:Vol. 1 p. 60
1319:
1310:
1277:
1273:
1267:
1256:. Retrieved
1247:
1238:
1221:
1215:
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1196:
1174:
1165:
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1135:
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1099:
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1069:
1060:
1052:
1046:
1039:
1033:
1027:
1004:
995:
994:Nazca booby
987:
981:
974:
968:
962:
941:
933:
925:
921:Lesson, 1831
917:
894:
885:
877:
871:
864:
858:
852:
827:S. l. plotus
826:
810:
801:
792:
768:
759:
749:
743:
736:
730:
724:
703:S. n. excisa
702:
696:
673:
664:
646:Common name
630:
629:
617:
583:
568:
556:
548:
544:
536:
526:
522:
518:
516:
506:
497:
488:
467:
460:type species
451:
449:
439:
435:
419:
418:
407:
405:
397:
342:
337:
322:
317:
286:
281:
266:
263:Masked booby
261:
232:
227:
205:
200:
168:
162:Type species
146:
145:
36:
3425:iNaturalist
3319:Wikispecies
3245:Sarmatosula
3141:Empheresula
3097:Olympidytes
3039:Plotopterum
2941:Nannopterum
2818:Macranhinga
2768:Protoplotus
2758:Mangystania
2599:Crozet shag
2583:Bounty shag
2462:Nannopterum
2141:Cape gannet
2098:Brown booby
2088:Nazca booby
1835:13 November
1789:13 November
1743:13 November
1716:Sula granti
1697:13 November
1651:13 November
1605:13 November
1408:"booby, n."
1342:Mayr, Ernst
1006:Sula granti
945:Gould, 1846
832:Forster, JR
757:Brown booby
464:brown booby
287:Sula granti
283:Nazca booby
229:Brown booby
18:Sula (bird)
3548:Categories
3151:Enkurosula
3076:Copepteryx
3067:Tonsalinae
2924:Microcarbo
2907:Leucocarbo
2893:Borvocarbo
2851:Fregatidae
2828:Meganhinga
2798:Giganhinga
2779:Anhingidae
2734:Suliformes
2719:Suliformes
2713:Neognathae
2678:cormorants
2668:Genera of
2563:Otago shag
2532:Leucocarbo
2383:Microcarbo
2265:Cormorants
2028:: Sulidae)
1906:Suliformes
1533:2017-11-22
1523:10088/6553
1429:required.)
1395:required.)
1374:"Sula, n."
1358:2017-11-21
1258:2019-07-08
1230:References
1178:Rothschild
1160:S. s. sula
1012:Rothschild
612:See also:
600:or in the
598:equatorial
553:congeneric
450:The genus
126:Suliformes
3208:Paleosula
3181:Microsula
3131:Bimbisula
3053:Stenornis
2985:Praecarbo
2951:Nectornis
2934:Nambashag
2839:Paranavis
2808:Liptornis
2693:Kingdom:
2541:Rock shag
2326:Pitt shag
1808:Sula sula
1137:Sula sula
608:Behaviour
561:osteology
541:monotypic
476:Icelandic
472:Old Norse
206:Sula sula
92:Kingdom:
86:Eukaryota
3378:45509066
3304:Wikidata
3215:Papasula
3032:Phocavis
2968:Piscator
2701:Chordata
2699:Phylum:
2695:Animalia
2682:anhingas
2108:Papasula
1861:Archived
1527:Archived
1505:(1985).
1489:82903466
1352:Archived
1318:(1760).
1302:21144905
1252:Archived
1144:Linnaeus
777:Boddaert
594:Pliocene
545:Papasula
174:Boddaert
132:Family:
106:Chordata
102:Phylum:
96:Animalia
82:Domain:
3554:Boobies
3443:1294449
3417:2480966
3310:Q612817
3191:Miosula
3161:Eostega
3119:Sulidae
3104:Tonsala
2900:Gulosus
2860:Fregata
2788:Anhinga
2717:Clade:
2705:Class:
2674:boobies
2492:Gulosus
2293:Species
2227:Anhinga
2218:Anhinga
2204:Species
2180:Anhinga
2169:Darters
2050:Species
1970:Fregata
1956:Species
1932:Fregata
1509:Sulidae
1469:The Auk
1282:Bibcode
1203:Habitat
1146:, 1766)
1100:Habitat
1080:, 1843)
1078:Tschudi
1034:Habitat
969:Habitat
859:Habitat
834:, 1844)
779:, 1783)
731:Habitat
637:Brisson
592:to the
590:Miocene
586:gannets
572:
565:Miocene
519:boobies
504:of the
491:Spanish
480:Sulidae
462:is the
432:gannets
428:Sulidae
414:in the
412:seabird
153:Brisson
142:Genus:
136:Sulidae
122:Order:
112:Class:
3531:137055
3505:160486
3489:NZOR:
3456:174697
3046:Stemec
2269:family
2173:family
2026:family
1924:family
1487:
1450:
1300:
1014:, 1902
904:, 1831
902:Lesson
821:, 1905
817:&
815:Thayer
683:, 1882
652:Range
639:, 1760
628:Genus
576:
543:genus
507:Bounty
484:gannet
482:, the
470:, the
425:family
176:, 1783
155:, 1760
3526:WoRMS
3518:36639
3482:32641
3438:IRMNG
3391:96609
3198:Morus
2992:Urile
2424:Urile
2282:Genus
2193:Genus
2177:genus
2127:Morus
2039:Genus
1945:Genus
1928:genus
1902:Order
1485:S2CID
1423:
1389:
1169:Gould
819:Bangs
549:Morus
528:Morus
496:term
494:slang
436:Morus
416:genus
410:is a
408:booby
42:Booby
3477:NCBI
3451:ITIS
3430:3785
3412:GBIF
3365:7Q76
3352:4598
3347:BOLD
3339:Sula
3325:Sula
3295:Sula
3252:Sula
2707:Aves
2064:Sula
1837:2021
1820:2018
1791:2021
1774:2018
1745:2021
1728:2018
1699:2021
1682:2018
1653:2021
1636:2018
1607:2021
1590:2018
1549:Ibis
1448:ISBN
1298:PMID
1248:PBDB
1209:Diet
1197:Size
1106:Diet
1094:Size
1084:Peru
1040:Diet
1028:Size
975:Diet
963:Size
865:Diet
853:Size
737:Diet
725:Size
706:Todd
631:Sula
557:Sula
537:Sula
523:Sula
498:bobo
474:and
468:súla
452:Sula
440:Sula
420:Sula
398:Sula
147:Sula
116:Aves
3464:NBN
3373:EoL
3360:CoL
3334:AFD
1824:doi
1778:doi
1732:doi
1686:doi
1640:doi
1594:doi
1557:doi
1553:160
1519:hdl
1477:doi
1473:119
1290:doi
579:mya
555:in
3550::
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634:–
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574:15
569:c.
567:,
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406:A
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