228:
246:
263:
131:
102:
1078:
J. F. Lawrence and A. F. Newton, Jr. 1995. Families and subfamilies of
Coleoptera (with selected genera, notes, references and data on family-group names). pp. 779-1006 In: J. Pakaluk & S. A. Slipinski (Eds.): Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th
985:
313:
Adults can be recognised by the hind wings having no accessory posterior ridge (locking device), no medial loop, no wedge cell and no apical hinge. The 8th segment of the
1069:
Beutel, R. G. and
Leschen, R.A.B. 2005. Phylogenetic analysis of Staphyliniformia (Coleoptera) based on characters of larvae and adults. Systematic Entomology
320:
Larval staphylinoids have 3-segmented (rarely 4-segmented) maxillary palps with distinct (often fused) galia and lacinia. The body usually has well-developed
250:
787:
529:
643:
688:
433:
424:
373:
520:
463:
364:
607:
538:
3232:
886:
832:
697:
406:
454:
481:
3027:
3180:
3206:
984:
Polilov, Alexey A.; Ribera, Ignacio; Yavorskaya, Margarita I.; Cardoso, Anabela; Grebennikov, Vasily V.; Beutel, Rolf G. (2019).
317:
is not entirely invaginated within the 7th. The head usually lacks a coronal suture (rarely with a short, rudimentary suture).
3211:
1140:
306:
can reach 45 mm. The superfamily includes the smallest beetles (and the smallest of all non-parasitic insects) in family
986:"The phylogeny of Ptiliidae (Coleoptera: Staphylinoidea) – the smallest beetles and their evolutionary transformations"
388:
957:
3159:
3237:
3172:
3288:
3219:
109:
130:
3102:
944:
325:
1015:
Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011).
3247:
310:. Most Ptiliidae do not exceed 1 mm long as adults, while the smallest species is just 325 μm long.
217:
3080:
1133:
3283:
267:
3260:
3042:
3255:
329:
298:
Adult staphylinoids are generally small beetles no more than a few millimetres long, though
3141:
3089:
2279:
227:
8:
2498:
1126:
1112:
Grimaldi, D. and Engel, M.S. (2005). Evolution of the
Insects. Cambridge University Press
245:
3094:
1047:
125:
3107:
262:
3128:
2767:
2184:
1052:
1034:
963:
953:
3133:
2864:
2643:
2638:
2561:
2319:
1335:
1042:
1024:
1016:
997:
202:
2553:
2506:
2351:
2335:
2011:
1771:
1740:
1655:
1573:
1464:
1432:
670:
3224:
3167:
3154:
2907:
2762:
2569:
2237:
2221:
2160:
2055:
2026:
1954:
1918:
1902:
1818:
1763:
1668:
1565:
1414:
1376:
1356:
1252:
946:
Phylogeny and classification of the staphyliniform beetle families (Coleoptera)
760:
661:
499:
283:
3277:
3065:
2938:
2742:
2733:
2474:
2445:
2437:
2429:
2403:
2378:
2274:
2261:
2200:
2118:
2021:
2016:
1897:
1831:
1802:
1786:
1750:
1720:
1673:
1627:
1594:
1510:
1502:
1473:
1311:
1118:
1038:
868:
841:
706:
679:
634:
568:
384:
117:
59:
1089:
1001:
967:
2915:
2839:
2585:
2577:
2532:
2516:
2466:
2387:
2343:
2295:
2213:
2126:
1995:
1982:
1962:
1923:
1823:
1810:
1781:
1715:
1703:
1599:
1556:
1539:
1531:
1481:
1319:
1289:
1220:
1209:
1193:
1056:
1029:
918:
877:
859:
823:
814:
805:
652:
598:
415:
3193:
3074:
3001:
2972:
2946:
2933:
2873:
2831:
2815:
2807:
2799:
2783:
2775:
2711:
2669:
2661:
2630:
2545:
2524:
2490:
2482:
2395:
2327:
2311:
2303:
2266:
2253:
2229:
2147:
2139:
2089:
2081:
2039:
2003:
1974:
1946:
1941:
1857:
1852:
1844:
1794:
1730:
1725:
1639:
1604:
1589:
1581:
1497:
1489:
1306:
796:
778:
616:
588:
396:
299:
238:
34:
3120:
2928:
2889:
2881:
2823:
2686:
2648:
2601:
2593:
2413:
2361:
2245:
2176:
2047:
2031:
1987:
1928:
1892:
1872:
1836:
1745:
1735:
1609:
1343:
1327:
751:
742:
733:
472:
79:
44:
3198:
2993:
2985:
2977:
2964:
2920:
2719:
2694:
2622:
2450:
2421:
2105:
1936:
1910:
1862:
1776:
1678:
1647:
1619:
1452:
1427:
1422:
1403:
1368:
1257:
1240:
1232:
1181:
1169:
1014:
914:
769:
724:
715:
625:
577:
558:
547:
510:
490:
444:
355:
345:
307:
303:
272:
255:
192:
162:
142:
84:
3185:
3036:
1079:
Birthday of Roy A. Crowson. Museum i
Instytut Zoologii PAN, Warszawa
290:. It is a very large and diverse group with worldwide distribution.
101:
3115:
3059:
2897:
2847:
2791:
2750:
2699:
2356:
2287:
2168:
1880:
1686:
1660:
1518:
1351:
1298:
1277:
1187:
902:
898:
850:
233:
74:
69:
54:
49:
39:
2614:
2606:
2458:
2192:
2113:
2097:
314:
89:
64:
2205:
2065:
1698:
1175:
1163:
1150:
1090:"Superfamily Staphylinoidea - Rove, Carrion and Fungus Beetles"
332:
are annular or annular-biforous. There are no epistomal lobes.
321:
287:
182:
172:
152:
983:
3146:
1521:(furniture beetles, death watch beetles, spider beetles)
952:. Copenhagen: Kongelige Danske Videnskabernes Selskab.
913:A 2019 molecular phylogenetic study confirmed the
340:Staphylinoidea contains the following subgroups:
3275:
2398:(jewel beetles, or metallic wood-boring beetles)
2121:(true weevils, bark beetles, ambrosia beetles)
1288:
1148:
1134:
2364:(ironclad beetles, cylindrical bark beetles)
1017:"Family-group names in Coleoptera (Insecta)"
897:The unambiguous fossil record dates back to
335:
1008:
1141:
1127:
100:
3028:List of subgroups of the order Coleoptera
2108:(straight snout weevils, New York weevil)
1207:
1046:
1028:
2535:(cicada beetle, cicada parasite beetles)
261:
244:
226:
3276:
2625:(false fireflies, long-lipped beetles)
1450:
1401:
1082:
942:
3041:
3040:
1449:
1400:
1275:
1274:
1206:
1122:
990:Arthropod Systematics & Phylogeny
3248:162156fd-0063-4f43-95f8-2d90af85adaf
3173:73e14325-76d2-4fab-b216-1cee1548691f
2651:(false metallic wood-boring beetles)
979:
977:
938:
936:
934:
13:
14:
3300:
974:
931:
917:of Ptilidae and found that it is
905:origin of the group is probable.
2262:Perimylopidae, or Promecheilidae
1847:(minute brown scavenger beetles)
129:
32:
2477:(variegated mud-loving beetles)
1106:
1072:
1063:
293:
1:
2306:(narrow-waisted bark beetles)
1998:(parasitic flat bark beetles)
1949:(short-winged flower beetles)
1805:(lady beetles, or God's cows)
1542:(tooth-necked fungus beetles)
924:
2818:(sand-loving scarab beetles)
2208:(minute tree-fungus beetles)
2034:(silvanid flat bark beetles)
908:
351:(primitive carrion beetles)
7:
2967:(primitive carrion beetles)
2923:(minute mud-loving beetles)
2786:(bumble bee scarab beetles)
2556:(soft-bodied plant beetles)
2527:(soft bodied plant beetles)
1330:(predaceous diving beetles)
10:
3305:
2802:(scavenger scarab beetles)
2794:(enigmatic scarab beetles)
1967:Lamingtonium binnaberrense
1681:(soft-wing flower beetles)
110:Devil's coach horse beetle
3049:
3025:
2955:
2941:(water scavenger beetles)
2906:
2872:
2863:
2741:
2732:
2685:
2674:Rhinorhipus tamborinensis
2660:
2544:
2515:
2453:(long-toed water beetles)
2412:
2386:
2377:
2346:(polypore fungus beetles)
2248:(palm and flower beetles)
2232:(tumbling flower beetles)
2179:(ant-like flower beetles)
2159:
2138:
2064:
1957:(lined flat bark beetles)
1931:(pleasing fungus beetles)
1871:
1826:(handsome fungus beetles)
1762:
1618:
1564:
1555:
1530:
1472:
1463:
1459:
1445:
1410:
1396:
1371:(burrowing water beetles)
1284:
1270:
1216:
1202:
1158:
336:Systematics and evolution
302:can reach 50 mm long and
126:Scientific classification
124:
108:
99:
23:
2988:(feather-winged beetles)
2322:(false longhorn beetles)
2269:(jugular-horned beetles)
2216:(false darkling beetles)
2187:(cryptic fungus beetles)
2006:(shining flower beetles)
1346:(crawling water beetles)
1245:Sikhotealinia zhiltzovae
943:Hansen, Michael (1997).
2469:(forest stream beetles)
2330:(synchroa bark beetles)
2290:(dead log bark beetles)
2282:(fire-coloured beetles)
2171:(ant-like leaf beetles)
1905:(silken fungus beetles)
1839:(well polished beetles)
1813:(minute fungus beetles)
1632:Acanthocnemus nigricans
1002:10.26049/ASP77-3-2019-4
836:Newton and Thayer, 1992
791:Newton and Thayer, 1995
629:Newton and Thayer, 1992
450:(round fungus beetles)
268:Nicrophorus interruptus
2980:(round fungus beetles)
2314:(false flower beetles)
2298:(wedge-shaped beetles)
2256:(false blister beetle)
2240:(hairy fungus beetles)
2195:(conifer bark beetles)
2092:(leaf-rolling weevils)
1753:(bark-gnawing beetles)
1513:(wounded-tree beetles)
1379:(false ground beetles)
1301:(trout-stream beetles)
1030:10.3897/zookeys.88.807
516:(featherwing beetles)
402:(minute moss beetles)
276:
259:
242:
16:Superfamily of beetles
3256:Paleobiology Database
2892:(false clown beetles)
2714:(plate-thigh beetles)
2596:(false click beetles)
2501:(water-penny beetles)
2150:(ship-timber beetles)
2129:(pine flower weevils)
1977:(root-eating beetles)
1797:(minute bark beetles)
1708:Phloiophilus edwardsi
1235:(reticulated beetles)
656:Mulsant and Rey, 1876
265:
248:
230:
3289:Beetle superfamilies
3168:Fauna Europaea (new)
2770:(false stag beetles)
2755:Belohina inexpectata
2617:(net-winged beetles)
2580:(rare click beetles)
2493:(travertine beetles)
2485:(minute mud beetles)
2050:(dry-fungus beetles)
1885:Agapytho foveicollis
1642:(false skin beetles)
1505:(Jacobson's beetles)
1225:Crowsoniella relicta
2768:Diphyllostomatidae
2100:(primitive weevils)
1913:(flat bark beetles)
1663:(checkered beetles)
1650:(fruitworm beetles)
1338:(whirligig beetles)
419:C. G. Thomson, 1860
251:Ptenidium punctatum
2633:(glowworm beetles)
2338:(darkling beetles)
2185:Archeocrypticidae
2042:(palmetto beetles)
1789:(dry bark beetles)
1576:(longhorn beetles)
564:(carrion beetles)
277:
260:
243:
3271:
3270:
3043:Taxon identifiers
3034:
3033:
3021:
3020:
3017:
3016:
3013:
3012:
3005:
2997:
2996:(carrion beetles)
2989:
2981:
2968:
2942:
2924:
2893:
2885:
2859:
2858:
2851:
2843:
2835:
2827:
2819:
2811:
2803:
2795:
2787:
2779:
2771:
2758:
2728:
2727:
2715:
2707:
2677:
2652:
2634:
2626:
2618:
2610:
2597:
2589:
2581:
2573:
2572:(soldier beetles)
2565:
2562:Brachypsectridae
2557:
2536:
2528:
2502:
2494:
2486:
2478:
2470:
2462:
2454:
2441:
2433:
2425:
2399:
2373:
2372:
2365:
2347:
2339:
2331:
2323:
2320:Stenotrachelidae
2315:
2307:
2299:
2291:
2283:
2270:
2257:
2249:
2241:
2233:
2225:
2224:(blister beetles)
2217:
2209:
2196:
2188:
2180:
2172:
2151:
2130:
2122:
2109:
2101:
2093:
2085:
2075:
2051:
2043:
2035:
2007:
1999:
1991:
1978:
1970:
1958:
1950:
1932:
1914:
1906:
1888:
1848:
1840:
1827:
1814:
1806:
1798:
1790:
1772:Akalyptoischiidae
1754:
1711:
1694:
1682:
1664:
1651:
1643:
1635:
1585:
1577:
1551:
1550:
1543:
1522:
1514:
1506:
1493:
1485:
1441:
1440:
1418:
1392:
1391:
1388:
1387:
1380:
1372:
1364:
1347:
1339:
1331:
1323:
1315:
1302:
1266:
1265:
1248:
1236:
1228:
891:
882:
873:
864:
855:
846:
837:
828:
819:
810:
801:
792:
783:
774:
765:
756:
747:
738:
729:
720:
711:
702:
693:
684:
675:
666:
657:
648:
639:
630:
621:
612:
603:
593:
582:
573:
563:
552:
543:
534:
525:
515:
504:
495:
486:
477:
468:
459:
449:
438:
429:
420:
411:
401:
392:
378:
369:
360:
350:
349:C.G. Thomson 1859
225:
224:
221:
3296:
3264:
3263:
3251:
3250:
3241:
3240:
3228:
3227:
3225:NHMSYS0001720142
3215:
3214:
3202:
3201:
3189:
3188:
3176:
3175:
3163:
3162:
3150:
3149:
3137:
3136:
3124:
3123:
3111:
3110:
3098:
3097:
3085:
3084:
3083:
3070:
3069:
3068:
3038:
3037:
3003:
2995:
2987:
2979:
2966:
2940:
2922:
2891:
2883:
2870:
2869:
2865:Staphyliniformia
2849:
2841:
2833:
2825:
2817:
2809:
2801:
2793:
2785:
2777:
2769:
2752:
2739:
2738:
2713:
2704:Declinia relicta
2701:
2671:
2650:
2644:Sinopyrophoridae
2639:Rhagophthalmidae
2632:
2624:
2616:
2608:
2595:
2587:
2579:
2571:
2563:
2555:
2554:Artematopodidae
2534:
2526:
2500:
2492:
2484:
2476:
2468:
2461:(riffle beetles)
2460:
2452:
2440:(turtle beetles)
2439:
2431:
2423:
2397:
2384:
2383:
2363:
2345:
2337:
2329:
2321:
2313:
2305:
2297:
2289:
2281:
2268:
2255:
2247:
2239:
2231:
2223:
2215:
2207:
2194:
2186:
2178:
2170:
2149:
2128:
2120:
2107:
2099:
2091:
2084:(fungus weevils)
2083:
2074:
2073:
2069:
2056:Tasmosalpingidae
2049:
2041:
2033:
2005:
1997:
1989:
1976:
1964:
1956:
1948:
1930:
1912:
1904:
1882:
1846:
1838:
1825:
1812:
1804:
1796:
1788:
1752:
1705:
1688:
1680:
1662:
1649:
1641:
1629:
1628:Acanthocnemidae
1583:
1575:
1562:
1561:
1541:
1520:
1512:
1504:
1491:
1483:
1470:
1469:
1461:
1460:
1447:
1446:
1416:
1398:
1397:
1378:
1370:
1358:
1345:
1337:
1329:
1321:
1314:(ground beetles)
1313:
1300:
1286:
1285:
1272:
1271:
1242:
1234:
1222:
1221:Crowsoniellidae
1204:
1203:
1143:
1136:
1129:
1120:
1119:
1113:
1110:
1104:
1103:
1101:
1100:
1086:
1080:
1076:
1070:
1067:
1061:
1060:
1050:
1032:
1012:
1006:
1005:
981:
972:
971:
951:
940:
921:to Hydraenidae.
889:
880:
871:
862:
853:
844:
835:
826:
817:
809:Ganglbauer, 1895
808:
799:
790:
788:Protopselaphinae
781:
772:
763:
754:
745:
736:
727:
718:
709:
700:
691:
682:
673:
664:
655:
646:
637:
628:
619:
610:
601:
591:
580:
571:
561:
550:
541:
532:
530:Cephaloplectinae
523:
513:
502:
493:
484:
475:
466:
457:
447:
436:
427:
418:
409:
399:
387:
376:
367:
358:
348:
231:Some species of
216:
203:Staphyliniformia
134:
133:
104:
94:
31:
27:Temporal range:
21:
20:
3304:
3303:
3299:
3298:
3297:
3295:
3294:
3293:
3274:
3273:
3272:
3267:
3259:
3254:
3246:
3244:
3236:
3231:
3223:
3218:
3210:
3205:
3197:
3192:
3184:
3179:
3171:
3166:
3158:
3153:
3145:
3140:
3132:
3127:
3119:
3114:
3106:
3101:
3093:
3088:
3079:
3078:
3073:
3064:
3063:
3058:
3045:
3035:
3030:
3009:
2951:
2902:
2884:(clown beetles)
2855:
2826:(betsy beetles)
2734:Scarabaeiformia
2724:
2681:
2656:
2588:(click beetles)
2564:(Texas beetles)
2540:
2511:
2507:Ptilodactylidae
2432:(cedar beetles)
2408:
2369:
2352:Trictenotomidae
2238:Mycetophagidae
2155:
2134:
2071:
2070:
2068:
2060:
2012:Phloeostichidae
1963:Lamingtoniidae
1955:Laemophloeidae
1903:Cryptophagidae
1867:
1758:
1741:Thanerocleridae
1704:Phloiophilidae
1691:Metaxina ornata
1656:Chaetosomatidae
1614:
1547:
1526:
1484:(auger beetles)
1465:Bostrichiformia
1455:
1437:
1433:Torridincolidae
1417:(skiff beetles)
1415:Hydroscaphidae
1406:
1384:
1377:Trachypachidae
1322:(tiger beetles)
1280:
1262:
1212:
1198:
1154:
1147:
1117:
1116:
1111:
1107:
1098:
1096:
1088:
1087:
1083:
1077:
1073:
1068:
1064:
1013:
1009:
982:
975:
960:
949:
941:
932:
927:
911:
901:, and an early
845:Latreille, 1802
818:Latreille, 1806
800:Latreille, 1802
671:Megalopsidiinae
644:Glypholomatinae
594:(rove beetles)
581:Latreille, 1806
338:
296:
215:
128:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
30:Rhaetian–Recent
29:
28:
25:
17:
12:
11:
5:
3302:
3292:
3291:
3286:
3284:Staphylinoidea
3269:
3268:
3266:
3265:
3252:
3242:
3229:
3216:
3203:
3190:
3177:
3164:
3155:Fauna Europaea
3151:
3138:
3125:
3112:
3108:Staphylinoidea
3099:
3095:Staphylinoidea
3086:
3081:Staphylinoidea
3071:
3055:
3053:
3051:Staphylinoidea
3047:
3046:
3032:
3031:
3026:
3023:
3022:
3019:
3018:
3015:
3014:
3011:
3010:
3008:
3007:
3004:(rove beetles)
3002:Staphylinidae
2999:
2991:
2983:
2975:
2970:
2961:
2959:
2957:Staphylinoidea
2953:
2952:
2950:
2949:
2944:
2939:Hydrophilidae
2936:
2931:
2926:
2918:
2912:
2910:
2908:Hydrophiloidea
2904:
2903:
2901:
2900:
2895:
2887:
2878:
2876:
2867:
2861:
2860:
2857:
2856:
2854:
2853:
2850:(hide beetles)
2845:
2837:
2834:(rain beetles)
2829:
2821:
2813:
2810:(stag beetles)
2805:
2797:
2789:
2781:
2773:
2765:
2763:Bolboceratidae
2760:
2747:
2745:
2736:
2730:
2729:
2726:
2725:
2723:
2722:
2717:
2709:
2697:
2691:
2689:
2683:
2682:
2680:
2679:
2670:Rhinorhipidae
2666:
2664:
2662:Rhinorhipoidea
2658:
2657:
2655:
2654:
2646:
2641:
2636:
2628:
2620:
2612:
2604:
2599:
2591:
2583:
2575:
2567:
2559:
2550:
2548:
2542:
2541:
2539:
2538:
2530:
2521:
2519:
2513:
2512:
2510:
2509:
2504:
2496:
2488:
2480:
2475:Heteroceridae
2472:
2464:
2456:
2448:
2443:
2438:Chelonariidae
2435:
2430:Callirhipidae
2427:
2424:(pill beetles)
2418:
2416:
2410:
2409:
2407:
2406:
2401:
2392:
2390:
2381:
2375:
2374:
2371:
2370:
2368:
2367:
2359:
2354:
2349:
2341:
2336:Tenebrionidae
2333:
2325:
2317:
2309:
2301:
2293:
2285:
2277:
2272:
2264:
2259:
2251:
2243:
2235:
2227:
2219:
2211:
2203:
2198:
2190:
2182:
2174:
2165:
2163:
2161:Tenebrionoidea
2157:
2156:
2154:
2153:
2144:
2142:
2136:
2135:
2133:
2132:
2124:
2119:Curculionidae
2116:
2111:
2103:
2095:
2087:
2078:
2076:
2066:Curculionoidea
2062:
2061:
2059:
2058:
2053:
2045:
2037:
2029:
2027:Protocucujidae
2024:
2019:
2014:
2009:
2001:
1993:
1985:
1980:
1972:
1960:
1952:
1944:
1939:
1934:
1926:
1921:
1919:Cybocephalidae
1916:
1908:
1900:
1895:
1890:
1877:
1875:
1869:
1868:
1866:
1865:
1860:
1855:
1850:
1842:
1834:
1829:
1821:
1819:Discolomatidae
1816:
1808:
1803:Coccinellidae
1800:
1792:
1787:Bothrideridae
1784:
1779:
1774:
1768:
1766:
1764:Coccinelloidea
1760:
1759:
1757:
1756:
1751:Trogossitidae
1748:
1743:
1738:
1733:
1728:
1723:
1718:
1713:
1701:
1696:
1684:
1676:
1671:
1669:Lophocateridae
1666:
1658:
1653:
1645:
1637:
1624:
1622:
1616:
1615:
1613:
1612:
1607:
1602:
1597:
1592:
1587:
1584:(leaf beetles)
1582:Chrysomelidae
1579:
1570:
1568:
1566:Chrysomeloidea
1559:
1553:
1552:
1549:
1548:
1546:
1545:
1536:
1534:
1528:
1527:
1525:
1524:
1516:
1511:Nosodendridae
1508:
1503:Jacobsoniidae
1500:
1495:
1492:(skin beetles)
1487:
1478:
1476:
1467:
1457:
1456:
1443:
1442:
1439:
1438:
1436:
1435:
1430:
1425:
1420:
1411:
1408:
1407:
1394:
1393:
1390:
1389:
1386:
1385:
1383:
1382:
1374:
1366:
1361:Meru phyllisae
1354:
1349:
1341:
1333:
1325:
1317:
1309:
1304:
1295:
1293:
1282:
1281:
1268:
1267:
1264:
1263:
1261:
1260:
1255:
1253:Micromalthidae
1250:
1238:
1230:
1217:
1214:
1213:
1200:
1199:
1197:
1196:
1190:
1184:
1178:
1172:
1166:
1159:
1156:
1155:
1146:
1145:
1138:
1131:
1123:
1115:
1114:
1105:
1081:
1071:
1062:
1007:
996:(3): 433–455.
973:
958:
929:
928:
926:
923:
910:
907:
895:
894:
893:
892:
883:
874:
865:
856:
847:
838:
829:
820:
811:
802:
793:
784:
782:Erichson, 1839
775:
773:Erichson, 1839
766:
764:Erichson, 1839
761:Phloeocharinae
757:
748:
739:
730:
728:Erichson, 1839
721:
712:
703:
694:
689:Microsilphinae
685:
676:
667:
662:Leptotyphlinae
658:
649:
640:
631:
622:
613:
604:
592:Latreille 1802
585:
584:
583:
574:
562:Latreille 1807
555:
554:
553:
551:Erichson, 1845
544:
535:
526:
507:
506:
505:
500:Platypsyllinae
496:
487:
478:
469:
460:
441:
440:
439:
434:Prosthetopinae
430:
425:Orchymontiinae
421:
412:
393:
381:
380:
379:
374:Pterolomatinae
370:
361:
337:
334:
295:
292:
280:Staphylinoidea
223:
222:
213:Staphylinoidea
210:
206:
205:
200:
196:
195:
190:
186:
185:
180:
176:
175:
170:
166:
165:
160:
156:
155:
150:
146:
145:
140:
136:
135:
122:
121:
106:
105:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
26:
24:Staphylinoidea
15:
9:
6:
4:
3:
2:
3301:
3290:
3287:
3285:
3282:
3281:
3279:
3262:
3257:
3253:
3249:
3243:
3239:
3234:
3230:
3226:
3221:
3217:
3213:
3208:
3204:
3200:
3195:
3191:
3187:
3182:
3178:
3174:
3169:
3165:
3161:
3156:
3152:
3148:
3143:
3139:
3135:
3130:
3126:
3122:
3117:
3113:
3109:
3104:
3100:
3096:
3091:
3087:
3082:
3076:
3072:
3067:
3061:
3057:
3056:
3054:
3052:
3048:
3044:
3039:
3029:
3024:
3006:
3000:
2998:
2992:
2990:
2984:
2982:
2976:
2974:
2971:
2969:
2963:
2962:
2960:
2958:
2954:
2948:
2945:
2943:
2937:
2935:
2932:
2930:
2927:
2925:
2919:
2917:
2914:
2913:
2911:
2909:
2905:
2899:
2896:
2894:
2890:Sphaeritidae
2888:
2886:
2880:
2879:
2877:
2875:
2871:
2868:
2866:
2862:
2852:
2846:
2844:
2840:Scarabaeidae
2838:
2836:
2830:
2828:
2822:
2820:
2814:
2812:
2806:
2804:
2798:
2796:
2790:
2788:
2782:
2780:
2778:(dor beetles)
2774:
2772:
2766:
2764:
2761:
2759:
2756:
2749:
2748:
2746:
2744:
2743:Scarabaeoidea
2740:
2737:
2735:
2731:
2721:
2718:
2716:
2710:
2708:
2705:
2698:
2696:
2693:
2692:
2690:
2688:
2684:
2678:
2675:
2668:
2667:
2665:
2663:
2659:
2653:
2647:
2645:
2642:
2640:
2637:
2635:
2629:
2627:
2621:
2619:
2613:
2611:
2605:
2603:
2600:
2598:
2592:
2590:
2584:
2582:
2578:Cerophytidae
2576:
2574:
2568:
2566:
2560:
2558:
2552:
2551:
2549:
2547:
2543:
2537:
2533:Rhipiceridae
2531:
2529:
2523:
2522:
2520:
2518:
2514:
2508:
2505:
2503:
2497:
2495:
2489:
2487:
2481:
2479:
2473:
2471:
2467:Eulichadidae
2465:
2463:
2457:
2455:
2449:
2447:
2446:Cneoglossidae
2444:
2442:
2436:
2434:
2428:
2426:
2420:
2419:
2417:
2415:
2411:
2405:
2404:Schizopodidae
2402:
2400:
2394:
2393:
2391:
2389:
2385:
2382:
2380:
2379:Elateriformia
2376:
2366:
2360:
2358:
2355:
2353:
2350:
2348:
2344:Tetratomidae
2342:
2340:
2334:
2332:
2326:
2324:
2318:
2316:
2310:
2308:
2302:
2300:
2296:Ripiphoridae
2294:
2292:
2286:
2284:
2280:Pyrochroidae
2278:
2276:
2275:Pterogeniidae
2273:
2271:
2265:
2263:
2260:
2258:
2252:
2250:
2244:
2242:
2236:
2234:
2228:
2226:
2220:
2218:
2214:Melandryidae
2212:
2210:
2204:
2202:
2201:Chalcodryidae
2199:
2197:
2191:
2189:
2183:
2181:
2175:
2173:
2167:
2166:
2164:
2162:
2158:
2152:
2146:
2145:
2143:
2141:
2137:
2131:
2127:Nemonychidae
2125:
2123:
2117:
2115:
2112:
2110:
2104:
2102:
2096:
2094:
2088:
2086:
2080:
2079:
2077:
2067:
2063:
2057:
2054:
2052:
2046:
2044:
2038:
2036:
2030:
2028:
2025:
2023:
2022:Propalticidae
2020:
2018:
2017:Priasilphidae
2015:
2013:
2010:
2008:
2002:
2000:
1996:Passandridae
1994:
1992:
1990:(sap beetles)
1986:
1984:
1981:
1979:
1973:
1971:
1968:
1961:
1959:
1953:
1951:
1945:
1943:
1940:
1938:
1935:
1933:
1927:
1925:
1922:
1920:
1917:
1915:
1909:
1907:
1901:
1899:
1898:Cavognathidae
1896:
1894:
1891:
1889:
1886:
1879:
1878:
1876:
1874:
1870:
1864:
1861:
1859:
1856:
1854:
1851:
1849:
1843:
1841:
1835:
1833:
1832:Eupsilobiidae
1830:
1828:
1824:Endomychidae
1822:
1820:
1817:
1815:
1811:Corylophidae
1809:
1807:
1801:
1799:
1793:
1791:
1785:
1783:
1780:
1778:
1775:
1773:
1770:
1769:
1767:
1765:
1761:
1755:
1749:
1747:
1744:
1742:
1739:
1737:
1734:
1732:
1729:
1727:
1726:Protopeltidae
1724:
1722:
1721:Prionoceridae
1719:
1717:
1714:
1712:
1709:
1702:
1700:
1697:
1695:
1692:
1685:
1683:
1677:
1675:
1674:Mauroniscidae
1672:
1670:
1667:
1665:
1659:
1657:
1654:
1652:
1646:
1644:
1638:
1636:
1633:
1626:
1625:
1623:
1621:
1617:
1611:
1608:
1606:
1603:
1601:
1598:
1596:
1595:Megalopodidae
1593:
1591:
1588:
1586:
1580:
1578:
1574:Cerambycidae
1572:
1571:
1569:
1567:
1563:
1560:
1558:
1554:
1544:
1540:Derodontidae
1538:
1537:
1535:
1533:
1532:Derodontoidea
1529:
1523:
1517:
1515:
1509:
1507:
1501:
1499:
1498:Endecatomidae
1496:
1494:
1488:
1486:
1482:Bostrichidae
1480:
1479:
1477:
1475:
1474:Bostrichoidea
1471:
1468:
1466:
1462:
1458:
1454:
1448:
1444:
1434:
1431:
1429:
1428:Sphaeriusidae
1426:
1424:
1421:
1419:
1413:
1412:
1409:
1405:
1399:
1395:
1381:
1375:
1373:
1367:
1365:
1362:
1355:
1353:
1350:
1348:
1342:
1340:
1334:
1332:
1326:
1324:
1320:Cicindelidae
1318:
1316:
1310:
1308:
1305:
1303:
1297:
1296:
1294:
1291:
1287:
1283:
1279:
1273:
1269:
1259:
1256:
1254:
1251:
1249:
1246:
1239:
1237:
1231:
1229:
1226:
1219:
1218:
1215:
1211:
1205:
1201:
1195:
1191:
1189:
1185:
1183:
1179:
1177:
1173:
1171:
1167:
1165:
1161:
1160:
1157:
1152:
1144:
1139:
1137:
1132:
1130:
1125:
1124:
1121:
1109:
1095:
1091:
1085:
1075:
1066:
1058:
1054:
1049:
1044:
1040:
1036:
1031:
1026:
1023:(88): 1–972.
1022:
1018:
1011:
1003:
999:
995:
991:
987:
980:
978:
969:
965:
961:
959:87-7304-286-2
955:
948:
947:
939:
937:
935:
930:
922:
920:
916:
906:
904:
900:
888:
884:
879:
875:
872:Thomson, 1858
870:
869:Trichophyinae
866:
863:MacLeay, 1825
861:
857:
854:MacLeay, 1825
852:
848:
843:
842:Staphylininae
839:
834:
830:
825:
821:
816:
812:
807:
803:
798:
794:
789:
785:
780:
776:
771:
767:
762:
758:
755:Fleming, 1821
753:
749:
746:Fleming, 1821
744:
740:
737:Fleming, 1821
735:
731:
726:
722:
719:MacLeay, 1825
717:
713:
710:Thomson, 1858
708:
707:Olisthaerinae
704:
699:
695:
692:Crowson, 1950
690:
686:
681:
680:Micropeplinae
677:
672:
668:
663:
659:
654:
650:
647:Jeannel, 1962
645:
641:
638:Thomson, 1859
636:
635:Euaesthetinae
632:
627:
623:
620:Reitter, 1887
618:
614:
609:
605:
602:Fleming, 1821
600:
596:
595:
590:
589:Staphylinidae
586:
579:
575:
570:
569:Nicrophorinae
566:
565:
560:
556:
549:
545:
540:
536:
531:
527:
524:Reitter, 1909
522:
521:Acrotrichinae
518:
517:
514:Erichson 1845
512:
508:
503:Ritsema, 1869
501:
497:
494:Fleming, 1821
492:
488:
483:
479:
474:
470:
465:
464:Catopocerinae
461:
458:Jeannel, 1911
456:
452:
451:
446:
442:
437:Perkins, 1994
435:
431:
428:Perkins, 1997
426:
422:
417:
413:
410:Mulsant, 1844
408:
404:
403:
398:
394:
390:
386:
385:Jacobsoniidae
382:
377:Thomson, 1862
375:
371:
366:
365:Necrophilinae
362:
359:Thomson, 1859
357:
353:
352:
347:
343:
342:
341:
333:
331:
327:
323:
318:
316:
311:
309:
305:
301:
300:Staphylinidae
291:
289:
285:
281:
274:
270:
269:
264:
257:
253:
252:
247:
240:
236:
235:
229:
219:
214:
211:
209:Superfamily:
208:
207:
204:
201:
198:
197:
194:
191:
188:
187:
184:
181:
178:
177:
174:
171:
168:
167:
164:
161:
158:
157:
154:
151:
148:
147:
144:
141:
138:
137:
132:
127:
123:
119:
118:Staphylinidae
115:
111:
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
22:
19:
3050:
2956:
2929:Helophoridae
2921:Georissidae
2916:Epimetopidae
2832:Pleocomidae
2816:Ochodaeidae
2800:Hybosoridae
2784:Glaphyridae
2776:Geotrupidae
2754:
2751:Belohinidae
2712:Eucinetidae
2703:
2700:Decliniidae
2673:
2631:Phengodidae
2570:Cantharidae
2525:Dascillidae
2517:Dascilloidea
2499:Psephenidae
2491:Lutrochidae
2483:Limnichidae
2396:Buprestidae
2388:Buprestoidea
2328:Synchroidae
2312:Scraptiidae
2304:Salpingidae
2267:Prostomidae
2254:Oedemeridae
2230:Mordellidae
2148:Lymexylidae
2140:Lymexyloidea
2090:Attelabidae
2082:Anthribidae
2040:Smicripidae
2004:Phalacridae
1988:Nitidulidae
1983:Myraboliidae
1975:Monotomidae
1966:
1947:Kateretidae
1924:Cyclaxyridae
1884:
1881:Agapythidae
1845:Latridiidae
1795:Cerylonidae
1782:Anamorphidae
1716:Phycosecidae
1707:
1690:
1687:Metaxinidae
1640:Biphyllidae
1631:
1600:Orsodacnidae
1557:Cucujiformia
1490:Dermestidae
1360:
1299:Amphizoidae
1244:
1224:
1210:Archostemata
1194:Holometabola
1192:Superorder:
1186:Infraclass:
1108:
1097:. Retrieved
1094:BugGuide.net
1093:
1084:
1074:
1065:
1020:
1010:
993:
989:
945:
919:sister group
912:
896:
885:† Subfamily
881:Reiche, 1866
878:Trigonurinae
860:Tachyporinae
824:Scydmaeninae
815:Scaphidiinae
806:Pseudopsinae
701:Fauvel, 1905
665:Fauvel, 1874
653:Habrocerinae
611:Fauvel, 1895
608:Apateticinae
599:Aleocharinae
542:Barber, 1924
539:Nanosellinae
448:Fleming 1821
416:Ochthebiinae
400:Mulsant 1844
368:Newton, 1997
339:
319:
312:
297:
279:
278:
266:
249:
232:
212:
199:Infraorder:
114:Ocypus olens
113:
18:
3194:iNaturalist
3075:Wikispecies
2973:Hydraenidae
2947:Spercheidae
2934:Hydrochidae
2898:Synteliidae
2882:Histeridae
2874:Histeroidea
2824:Passalidae
2792:Glaresidae
2649:Throscidae
2609:(fireflies)
2607:Lampyridae
2594:Eucnemidae
2586:Elateridae
2546:Elateroidea
2362:Zopheridae
2246:Mycteridae
2177:Anthicidae
2048:Sphindidae
2032:Silvanidae
1942:Hobartiidae
1929:Erotylidae
1858:Mycetaeidae
1853:Murmidiidae
1837:Euxestidae
1731:Rentoniidae
1605:Oxypeltidae
1590:Disteniidae
1423:Lepiceridae
1352:Hygrobiidae
1344:Haliplidae
1328:Dytiscidae
1307:Aspidytidae
887:Protactinae
833:Solieriinae
827:Leach, 1815
797:Pselaphinae
779:Proteininae
698:Neophoninae
683:Leach, 1815
617:Dasycerinae
572:Kirby, 1837
533:Sharp, 1883
476:Kirby, 1837
467:Hatch, 1927
407:Hydraeninae
397:Hydraenidae
294:Description
284:superfamily
239:Hydraenidae
3278:Categories
2994:Silphidae
2986:Ptiliidae
2978:Leiodidae
2965:Agyrtidae
2808:Lucanidae
2687:Scirtoidea
2623:Omethidae
2602:Jurasaidae
2451:Dryopidae
2422:Byrrhidae
2414:Byrrhoidea
2106:Brentidae
1911:Cucujidae
1893:Boganiidae
1873:Cucujoidea
1746:Thymalidae
1736:Rhadalidae
1679:Melyridae
1648:Byturidae
1610:Vesperidae
1519:Ptiniidae
1369:Noteridae
1336:Gyrinidae
1312:Carabidae
1241:Jurodidae
1233:Cupedidae
1180:Subclass:
1170:Arthropoda
1151:Coleoptera
1099:2019-06-16
925:References
890:Heer, 1847
876:Subfamily
867:Subfamily
858:Subfamily
849:Subfamily
840:Subfamily
831:Subfamily
822:Subfamily
813:Subfamily
804:Subfamily
795:Subfamily
786:Subfamily
777:Subfamily
768:Subfamily
759:Subfamily
752:Paederinae
750:Subfamily
743:Oxytelinae
741:Subfamily
734:Oxyporinae
732:Subfamily
723:Subfamily
714:Subfamily
705:Subfamily
696:Subfamily
687:Subfamily
678:Subfamily
674:Leng, 1920
669:Subfamily
660:Subfamily
651:Subfamily
642:Subfamily
633:Subfamily
624:Subfamily
615:Subfamily
606:Subfamily
597:Subfamily
576:Subfamily
567:Subfamily
546:Subfamily
537:Subfamily
528:Subfamily
519:Subfamily
498:Subfamily
489:Subfamily
485:Horn, 1880
480:Subfamily
473:Cholevinae
471:Subfamily
462:Subfamily
455:Camiarinae
453:Subfamily
432:Subfamily
423:Subfamily
414:Subfamily
405:Subfamily
372:Subfamily
363:Subfamily
354:Subfamily
189:Suborder:
183:Coleoptera
163:Arthropoda
2848:Trogidae
2842:(scarabs)
2720:Scirtidae
2695:Clambidae
2288:Pythidae
2222:Meloidae
2169:Aderidae
2072:(weevils)
1937:Helotidae
1863:Teredidae
1777:Alexiidae
1661:Cleridae
1620:Cleroidea
1453:Polyphaga
1451:Suborder
1404:Myxophaga
1402:Suborder
1357:Meruidae
1276:Suborder
1258:Ommatidae
1208:Suborder
1182:Pterygota
1162:Kingdom:
1039:1313-2989
915:monophyly
909:Phylogeny
770:Piestinae
725:Osoriinae
716:Omaliinae
626:Empelinae
578:Silphinae
559:Silphidae
548:Ptiliinae
511:Ptiliidae
491:Leiodinae
482:Coloninae
445:Leiodidae
356:Agyrtinae
346:Agyrtidae
330:spiracles
326:sternites
308:Ptiliidae
304:Silphidae
273:Silphidae
256:Ptiliidae
218:Latreille
193:Polyphaga
149:Kingdom:
143:Eukaryota
3116:BugGuide
3060:Wikidata
2615:Lycidae
2459:Elmidae
2357:Ulodidae
2193:Boridae
2098:Belidae
1699:Peltidae
1292:families
1278:Adephaga
1188:Neoptera
1168:Phylum:
1164:Animalia
1153:families
1057:21594053
968:38156728
903:Mesozoic
899:Triassic
851:Steninae
322:tergites
234:Hydraena
159:Phylum:
153:Animalia
139:Domain:
3066:Q132752
2206:Ciidae
2114:Caridae
1176:Insecta
1174:Class:
1149:Extant
1048:3088472
1021:ZooKeys
587:Family
557:Family
509:Family
443:Family
395:Family
383:Family
344:Family
315:abdomen
288:beetles
179:Order:
173:Insecta
169:Class:
3245:NZOR:
3212:113264
1290:Extant
1055:
1045:
1037:
966:
956:
391:, 1926
389:Heller
328:. The
220:, 1802
3261:69274
3238:75543
3199:48311
3160:11103
3134:B6LT7
3121:40608
950:(PDF)
282:is a
3233:NCBI
3207:ITIS
3186:1583
3181:GBIF
1053:PMID
1035:ISSN
964:OCLC
954:ISBN
324:and
35:PreꞒ
3220:NBN
3147:365
3142:EoL
3129:CoL
3103:AFD
3090:ADW
1043:PMC
1025:doi
998:doi
286:of
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994:77
992:.
988:.
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933:^
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112:,
85:Pg
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55:D
50:S
45:O
40:Ꞓ
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
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