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Partula (gastropod)

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Individuals are being reintroduced to Tahiti from captive breeding programmes since 2014. In April 2023, over 5,000 individual snails from zoos in the United States and the United Kingdom were released on Tahiti and
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that were endemic to Tahiti and the nearby islands, causing all but 12 species to go extinct within a decade. Scientists were able to save 12 of these species prior to their becoming
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ceremonial wear and jewelry, these small snails (averaging about one-half to three-quarters of an inch in length) gained the attention of science when Dr.
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act as an example of the possible deleterious effects of attempted biological control. After an infestation of the introduced giant African land snails (
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Myers, P.; Espinosa, R.; Parr, C. S.; Jones, T.; Hammond, G. S. & Dewey, T. A. (2006). The Animal Diversity Web (online). Accessed at
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runs the Partula Programme Consortium which maintains a captive-breeding programme in the United Kingdom, France, and the United States.
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Studies on the variation, distribution and evolution of the genus Partula. The species of the Mariana Islands, Guam and Saipan.
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Gerlach, J. (2016) Icons of Evolution - Pacific island tree snails, family Partulidae. Phelsuma Press, Cambridge
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Studies on the variation, distribution and evolution of the genus Partula. The species inhabiting Moorea.
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Studies on the variation, distribution and evolution of the genus Partula. The species inhabiting Tahiti.
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The list may be currently missing some species. If you notice a missing species please add it.
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Proc. Joint Conference - American Malacological Society and Western Society of Malacologists
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Type population is almost certainly extinct. Remaining populations have not been surveyed.
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Currently only known from a dead subadult in one location and a live one in another.
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Lee, T.; Burch, J. B.; Jung, Y.; Coote, T.; Pearce-Kelly, P.; ó Foighil, D. (2007).
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Survives in populations above 950m altitude and in one population on Tahiti Iti.
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The species pages are currently outdated because of the IUCN 2024-1 update.
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Only known from a single location. Not seen since its discovery in 2005.
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species on Tahiti were usually found on the undersides of the leaves of
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This article incorporates public domain text from the reference.
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Could not be located after 1991. Old shells were found in 1994.
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were misidentified as the species in parentheses next to them.
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Many populations are unstable and inviable in the long-term.
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2005, 2006 and 2008 surveys found only 4 old, eroded shells.
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Little is known about this species. May already be extinct.
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Instead, the rosy wolfsnail hunted the nearly 76 species of
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Single valley endemic. A single shell was found in 1992.
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Tolerant of current levels of disturbance and predators.
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Has become much rarer but several populations persist.
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Only one population remains. Viable but vulnerable.
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Only dead shells could be found in the 1991 survey.
56:. Unsourced material may be challenged and removed. 1538:All populations appear to be tolerant of threats. 5748: 964:, though currently relatively free of predators. 800:As of the 2024-1 update released on June 27, the 5986: 5736:Carnegie Institution of Washington, 228a: 1-116. 3052:Phylogenetic analyses revealed that many of the 2437:Tolerant of habitat loss and free of predators. 2096:. Population declined severely in both islands. 5745:Carnegie Institution of Washington, 410: 1-335. 1036:The last captive individual died in the 1980s. 773:spp.), the carnivorous Florida rosy wolfsnail ( 5790:Partula evolution, diversity and conservation 5470: 2810:The list of surviving species are as follows: 1777:Survival confirmed from a single individual. 5696:Memoirs of the Museum of Comparative Zoölogy 1995:No major threats to this species are known. 642:snails on the underside of a single leaf of 634:Black-and-white photo taken in July 1920 in 916:but reasonable numbers exist in captivity. 5620:Elizabeth Claire Alberts (28 April 2023). 5369: 3060:. The resulting cladogram is shown below. 1677:The last captive individual died in 2002. 1431:The last captive individual died in 2016. 939:The last captive individual died in 1994. 712:are known under the general common names " 140: 5766: 2138:Endemic to nine islands. Eliminated from 1236:Could not be located in the 1992 survey. 751:, and behavioral aspects that illustrate 116:Learn how and when to remove this message 5683: 5681: 2783: 1911:. Could not be located in 2006 or 2017. 662: 650: 629: 5533: 5531: 5987: 2539:Last captive individual died in 1992. 1886:. Little is known about this species. 1698:One population may survive in central 1120:Last captive individual died in 1996. 408:; Leptopartula is a replacement name) 5801: 5800: 5678: 5574:"IUCN Red List of Threatened Species" 5490:ITIS Standard Report Page: Partulidae 1141:One population photographed in 2004. 1007:Extirpated from its type locality in 5694:from Tahiti. A study in variation". 5594: 5568: 5566: 5528: 2805: 2314:Little is known about this species. 808:species. Of these, 32 are listed as 54:adding citations to reliable sources 25: 5483: 5314: 5085: 1932:Only known from a single specimen. 1861:Could not be located in the 1990s. 802:IUCN Red List of Threatened Species 524:accepted, alternate representation 13: 5725:Carnegie Institution of Washington 5711: 5550:"Partula snail conservation | ZSL" 5275: 5188: 2610:Collected for ex situ conservation 14: 6021: 5784: 5690:(January 1902). "Some species of 5563: 2560:Could not be located after 1980. 2518:Could not be located after 1990. 2395:Could not be located after 1990. 2335:Could not be located after 1980. 2230:Could not be located after 1980. 2209:Could not be located after 1980. 1819:Could not be located after 1980. 1798:Could not be located after 1980. 1625:All individuals are hybrids with 1410:Could not be located after 1980. 1368:Could not be located after 1992. 1347:Could not be located after 1980. 1326:Could not be located after 1980. 1078:Could not be located after 1993. 735:Once used as decorative items in 5424: 5408: 5392: 5376: 5353: 5337: 5321: 5298: 5282: 5259: 5243: 5227: 5211: 5195: 5172: 5156: 5140: 5124: 5108: 5092: 2624:breeding include the following: 2416:Dead shells were found in 1989. 912:A single population persists on 162: 30: 5660: 5639: 5613: 763:The partulids of the island of 41:needs additional citations for 5588: 5542: 5507: 5494: 5451: 2188:Could not be located in 1994. 1: 6010:Molluscs of the Pacific Ocean 5513:Jung, Younghun, Taehwan Lee, 5441: 2791:snail with its shell removed. 718:Moorean viviparous tree snail 3047: 1168:. Old shells found in 2005. 855: 795:Zoological Society of London 572:(invalid: junior homonym of 533:(invalid: junior homonym of 508:(invalid: junior homonym of 492:(invalid: junior homonym of 436:(invalid: junior homonym of 417:(invalid: junior homonym of 401:(invalid: junior homonym of 385:(invalid: junior homonym of 361:(invalid: junior homonym of 7: 10: 6026: 5595:Kuta, Sarah (2 May 2023). 5478:http://animaldiversity.org 5080: 5049: 843: 758: 684:of air-breathing tropical 18: 5809: 5768:10.1016/j.cub.2007.05.006 5004: 4997: 4980: 4973: 4915: 4898: 4891: 4869: 4862: 4845: 4828: 4793: 4776: 4769: 4752: 4745: 4738: 4707: 4690: 4673: 4647: 4602: 4584: 4577: 4541: 4534: 4517: 4510: 4503: 4449: 4442: 4425: 4418: 4401: 4394: 4377: 4360: 4353: 4346: 4339: 4313: 4296: 4279: 4262: 4236: 4219: 4212: 4168: 4161: 4144: 4137: 4093: 4086: 4069: 4052: 4045: 4028: 4021: 4014: 3988: 3970: 3953: 3935: 3917: 3873: 3855: 3848: 3821: 3803: 3785: 3778: 3761: 3754: 3737: 3730: 3704: 3687: 3661: 3644: 3637: 3611: 3594: 3577: 3560: 3543: 3526: 3509: 3502: 3495: 3451: 3444: 3427: 3392: 3385: 3368: 3361: 3354: 3328: 3311: 3294: 3277: 3270: 3235: 3228: 3211: 3185: 3168: 3151: 3134: 3127: 3120: 3113: 3106: 3099: 3092: 3080: 3073: 848:Species within the genus 277: 270: 159:Scientific classification 157: 148: 139: 132: 3924:(Austral Islands, etc.) 3765:P. otaheitana otaheitana 2618:that were collected for 2479:3/4 subspecies survive. 2458:2/2 subspecies survive. 2374:Not located after 1936. 65:"Partula" gastropod 21:Partula (disambiguation) 5739:Crampton H. E. (1932). 5730:Crampton H. E. (1925). 5370:Mariana Islands species 3155:P. suturalis dendronica 1257:Isolated population on 828:and only 4 species are 497:H. & A. Adams, 1855 427:is a replacement name) 5458:Férussac A. É. d'A. de 2792: 1907:Only species found in 672: 660: 648: 447:Partula (Leptopartula) 5954:Paleobiology Database 4381:P. otaheitana affinis 3315:P. suturalis vexillum 3239:P. suturalis vexillum 3215:P. suturalis strigosa 3189:P. suturalis vexillum 3172:P. suturalis strigosa 2787: 1756:Not located in 1992. 1193:, though survives in 818:critically endangered 714:Polynesian tree snail 666: 654: 633: 372:Partula (Carolinella) 5654:University of Hawaii 5601:Smithsonian Magazine 4919:Palaopartula calypso 3665:P. taeniata nucleola 3598:P. taeniata simulans 3581:P. taeniata nucleola 3564:P. taeniata elongata 2567:Partula vanikorensis 2153:Partula pearcekellyi 1638:Partula jackieburchi 1333:Partula dolichostoma 1204:Partula crassilabris 1011:but persists in the 583:Partula (Thakombaua) 479:Partula (Melanesica) 455:Partula (Marianella) 50:improve this article 19:For other uses, see 6005:Molluscs of Oceania 5995:Partula (gastropod) 5704:, Cambridge, U.S.A. 5362:Partula navigatoria 4902:Palaopartula thetis 4756:Samoana decussatula 4521:undescribed species 3741:undescribed species 2174:Partula planilabrum 2038:Partula navigatoria 1805:Partula levistriata 1261:confirmed in 2005. 814:extinct in the wild 638:, showing numerous 558:Partula (Rennellia) 16:Genus of gastropods 5462:Journ. de Physique 5181:Partula otaheitana 2793: 2546:Partula umbilicata 2300:Partula recluziana 2103:Partula otaheitana 1784:Partula leptochila 1709:Partula lanceolata 1043:Partula auriculata 753:adaptive radiation 724:islands, from the 673: 661: 649: 528:Partula (Pasithea) 522:A. Férussac, 1821· 463:Partula (Marianna) 412:Partula (Harmonia) 292:Bulimus (Partula) 5982: 5981: 5941:Open Tree of Life 5803:Taxon identifiers 5761:(13): R502–R503. 5433:Partula langfordi 5417:Partula radiolata 5401:Partula lutaensis 5330:Partula garrettii 5268:Partula tohiveana 5236:Partula suturalis 5204:Partula mirabilis 5046: 5045: 5037: 5036: 5028: 5027: 5019: 5018: 4957: 4956: 4948: 4947: 4939: 4938: 4930: 4929: 4849:Samoana thurstoni 4817: 4816: 4808: 4807: 4780:Samoana attenuata 4722: 4721: 4662: 4661: 4636: 4635: 4627: 4626: 4618: 4617: 4566: 4565: 4557: 4556: 4492: 4491: 4483: 4482: 4474: 4473: 4465: 4464: 4328: 4327: 4251: 4250: 4201: 4200: 4192: 4191: 4183: 4182: 4126: 4125: 4117: 4116: 4108: 4107: 4003: 4002: 3906: 3905: 3897: 3896: 3888: 3887: 3837: 3836: 3719: 3718: 3676: 3675: 3626: 3625: 3484: 3483: 3475: 3474: 3466: 3465: 3416: 3415: 3407: 3406: 3343: 3342: 3259: 3258: 3250: 3249: 3200: 3199: 2806:Surviving species 2607: 2606: 2486:Partula tohiveana 2444:Partula suturalis 2423:Partula similaris 2258:Partula radiolata 2216:Partula protracta 2002:Partula mirabilis 1868:Partula lutaensis 1742:Partula langfordi 1684:Partula laevigata 1600:Wild populations 1545:Partula guamensis 1463:Partula garrettii 1268:Partula dentifera 1175:Partula cramptoni 1127:Partula compressa 1064:Partula bilineata 991:Partula auraniana 946:Partula assimilis 749:ecological niches 669:Partula langfordi 628: 627: 621: 613: 605:(junior synonym) 604: 596:(junior synonym) 595: 587: 578: 571: 567:Partula (Sterope) 563:(junior synonym) 562: 553: 546: 539: 532: 523: 519:Partula (Partula) 514: 507: 498: 491: 483: 475: 467: 459: 451: 442: 435: 423: 416: 407: 400: 391: 384: 376: 367: 360: 356:Partula (Astraea) 352: 344: 336: 328: 320: 312: 304: 298: 289: 281:list of synonyms: 266: 151:Partula radiolata 126: 125: 118: 100: 6017: 6000:Gastropod genera 5975: 5974: 5962: 5961: 5949: 5948: 5936: 5935: 5923: 5922: 5910: 5909: 5897: 5896: 5884: 5883: 5871: 5870: 5858: 5857: 5845: 5844: 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3129: 3123: 3122: 3116: 3115: 3109: 3108: 3102: 3101: 3095: 3094: 3083: 3082: 3076: 3075: 3064: 3063: 3056:species are not 3020:Papua New Guinea 2465:Partula taeniata 2402:Partula salifana 2195:Partula producta 2124:Partula pacifica 2020:Partula mooreana 1953:Fossil species. 1918:Partula magistri 1847:Partula lugubris 1663:Partula labrusca 1611:Partula incrassa 1438:Partula flexuosa 1375:Partula emersoni 1354:Partula dolorosa 1312:Partula diminuta 1305:Fossil species. 1289:Partula desolata 1243:Partula cytherea 1189:Extirpated from 1022:Partula aurantia 868: 867: 776:Euglandina rosea 708:Many species of 619: 611: 602: 593: 585: 576: 569: 560: 551: 544: 537: 530: 521: 512: 505: 503:Partula (Oenone) 496: 489: 481: 473: 471:Partula (Matata) 465: 457: 449: 440: 433: 431:Partula (Helena) 421: 414: 405: 398: 389: 382: 380:Partula (Clytia) 374: 365: 358: 350: 342: 334: 326: 318: 310: 302: 296: 287: 261: 167: 166: 144: 130: 129: 121: 114: 110: 107: 101: 99: 58: 34: 26: 6025: 6024: 6020: 6019: 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4637: 4628: 4619: 4567: 4558: 4493: 4484: 4475: 4466: 4329: 4300:P. carteriensis 4252: 4202: 4193: 4184: 4127: 4118: 4109: 4004: 3907: 3898: 3889: 3838: 3720: 3677: 3627: 3485: 3476: 3467: 3417: 3408: 3344: 3260: 3251: 3201: 3050: 3003:Solomon Islands 2808: 2612: 2525:Partula turgida 2504:Partula tristis 2381:Partula sagitta 2279:Partula radiosa 2237:Partula pyramis 1939:Partula makatea 1652:May survive on 1604:be recovering. 1584:Partula hyalina 1396:Partula eremita 1222:Partula cuneata 1165:Partula hyalina 1162:Outcompeted by 1106:Partula clarkei 896:Partula affinis 858: 846: 761: 726:Society Islands 624: 487:Partula (Nenia) 390:Lamouroux, 1812 283: 282: 260: 216:Stylommatophora 161: 122: 111: 105: 102: 59: 57: 47: 35: 24: 17: 12: 11: 5: 6023: 6013: 6012: 6007: 6002: 5997: 5980: 5979: 5977: 5976: 5963: 5950: 5937: 5924: 5911: 5898: 5885: 5872: 5859: 5846: 5831: 5815: 5813: 5807: 5806: 5795: 5794: 5786: 5785:External links 5783: 5782: 5781: 5746: 5737: 5728: 5718:Crampton H. E. 5713: 5710: 5707: 5706: 5677: 5659: 5638: 5612: 5587: 5576:. 27 June 2024 5562: 5541: 5527: 5506: 5493: 5482: 5469: 5449: 5448: 5443: 5440: 5439: 5438: 5430: 5423: 5421: 5414: 5407: 5405: 5398: 5391: 5389: 5382: 5375: 5371: 5368: 5367: 5366: 5359: 5352: 5350: 5343: 5336: 5334: 5327: 5320: 5316: 5313: 5312: 5311: 5304: 5297: 5295: 5288: 5281: 5277: 5274: 5273: 5272: 5265: 5258: 5256: 5249: 5242: 5240: 5233: 5226: 5224: 5217: 5210: 5208: 5201: 5194: 5190: 5187: 5186: 5185: 5178: 5171: 5169: 5165:Partula nodosa 5162: 5155: 5153: 5146: 5139: 5137: 5130: 5123: 5121: 5114: 5107: 5105: 5098: 5091: 5087: 5084: 5082: 5079: 5051: 5048: 5044: 5043: 5040: 5039: 5035: 5034: 5031: 5030: 5026: 5025: 5022: 5021: 5017: 5016: 5013: 5012: 5003: 4998: 4996: 4993: 4992: 4989: 4988: 4979: 4974: 4972: 4964: 4963: 4960: 4959: 4955: 4954: 4951: 4950: 4946: 4945: 4942: 4941: 4937: 4936: 4933: 4932: 4928: 4927: 4924: 4923: 4914: 4911: 4910: 4907: 4906: 4897: 4892: 4890: 4882: 4881: 4878: 4877: 4868: 4863: 4861: 4858: 4857: 4854: 4853: 4844: 4841: 4840: 4837: 4836: 4832:Samoana conica 4827: 4824: 4823: 4820: 4819: 4815: 4814: 4811: 4810: 4806: 4805: 4802: 4801: 4797:Samoana burchi 4792: 4789: 4788: 4785: 4784: 4775: 4770: 4768: 4765: 4764: 4761: 4760: 4751: 4746: 4744: 4739: 4737: 4729: 4728: 4725: 4724: 4720: 4719: 4716: 4715: 4706: 4703: 4702: 4699: 4698: 4689: 4686: 4685: 4682: 4681: 4672: 4669: 4668: 4665: 4664: 4660: 4659: 4656: 4655: 4646: 4643: 4642: 4639: 4638: 4634: 4633: 4630: 4629: 4625: 4624: 4621: 4620: 4616: 4615: 4612: 4611: 4601: 4598: 4597: 4594: 4593: 4583: 4578: 4576: 4573: 4572: 4569: 4568: 4564: 4563: 4560: 4559: 4555: 4554: 4551: 4550: 4540: 4535: 4533: 4530: 4529: 4526: 4525: 4516: 4511: 4509: 4504: 4502: 4499: 4498: 4495: 4494: 4490: 4489: 4486: 4485: 4481: 4480: 4477: 4476: 4472: 4471: 4468: 4467: 4463: 4462: 4459: 4458: 4448: 4443: 4441: 4438: 4437: 4434: 4433: 4424: 4419: 4417: 4414: 4413: 4410: 4409: 4400: 4395: 4393: 4390: 4389: 4386: 4385: 4376: 4373: 4372: 4369: 4368: 4359: 4354: 4352: 4347: 4345: 4340: 4338: 4335: 4334: 4331: 4330: 4326: 4325: 4322: 4321: 4312: 4309: 4308: 4305: 4304: 4295: 4292: 4291: 4288: 4287: 4278: 4275: 4274: 4271: 4270: 4261: 4258: 4257: 4254: 4253: 4249: 4248: 4245: 4244: 4235: 4232: 4231: 4228: 4227: 4218: 4213: 4211: 4208: 4207: 4204: 4203: 4199: 4198: 4195: 4194: 4190: 4189: 4186: 4185: 4181: 4180: 4177: 4176: 4167: 4162: 4160: 4157: 4156: 4153: 4152: 4143: 4138: 4136: 4133: 4132: 4129: 4128: 4124: 4123: 4120: 4119: 4115: 4114: 4111: 4110: 4106: 4105: 4102: 4101: 4092: 4087: 4085: 4082: 4081: 4078: 4077: 4068: 4065: 4064: 4061: 4060: 4051: 4046: 4044: 4041: 4040: 4037: 4036: 4027: 4022: 4020: 4015: 4013: 4010: 4009: 4006: 4005: 4001: 4000: 3997: 3996: 3987: 3984: 3983: 3980: 3979: 3969: 3966: 3965: 3962: 3961: 3952: 3949: 3948: 3945: 3944: 3934: 3931: 3930: 3927: 3926: 3916: 3913: 3912: 3909: 3908: 3904: 3903: 3900: 3899: 3895: 3894: 3891: 3890: 3886: 3885: 3882: 3881: 3872: 3869: 3868: 3865: 3864: 3854: 3849: 3847: 3844: 3843: 3840: 3839: 3835: 3834: 3831: 3830: 3820: 3817: 3816: 3813: 3812: 3802: 3799: 3798: 3795: 3794: 3784: 3779: 3777: 3774: 3773: 3770: 3769: 3760: 3755: 3753: 3750: 3749: 3746: 3745: 3736: 3731: 3729: 3726: 3725: 3722: 3721: 3717: 3716: 3713: 3712: 3703: 3700: 3699: 3696: 3695: 3686: 3683: 3682: 3679: 3678: 3674: 3673: 3670: 3669: 3660: 3657: 3656: 3653: 3652: 3643: 3638: 3636: 3633: 3632: 3629: 3628: 3624: 3623: 3620: 3619: 3610: 3607: 3606: 3603: 3602: 3593: 3590: 3589: 3586: 3585: 3576: 3573: 3572: 3569: 3568: 3559: 3556: 3555: 3552: 3551: 3542: 3539: 3538: 3535: 3534: 3525: 3522: 3521: 3518: 3517: 3508: 3503: 3501: 3496: 3494: 3491: 3490: 3487: 3486: 3482: 3481: 3478: 3477: 3473: 3472: 3469: 3468: 3464: 3463: 3460: 3459: 3450: 3445: 3443: 3440: 3439: 3436: 3435: 3426: 3423: 3422: 3419: 3418: 3414: 3413: 3410: 3409: 3405: 3404: 3401: 3400: 3391: 3386: 3384: 3381: 3380: 3377: 3376: 3367: 3362: 3360: 3355: 3353: 3350: 3349: 3346: 3345: 3341: 3340: 3337: 3336: 3327: 3324: 3323: 3320: 3319: 3310: 3307: 3306: 3303: 3302: 3293: 3290: 3289: 3286: 3285: 3276: 3271: 3269: 3266: 3265: 3262: 3261: 3257: 3256: 3253: 3252: 3248: 3247: 3244: 3243: 3234: 3229: 3227: 3224: 3223: 3220: 3219: 3210: 3207: 3206: 3203: 3202: 3198: 3197: 3194: 3193: 3184: 3181: 3180: 3177: 3176: 3167: 3164: 3163: 3160: 3159: 3150: 3147: 3146: 3143: 3142: 3133: 3128: 3126: 3121: 3119: 3114: 3112: 3107: 3105: 3100: 3098: 3093: 3091: 3081: 3079: 3074: 3072: 3062: 3049: 3046: 2929:P. navigatoria 2807: 2804: 2800:P. navigatoria 2752:P. navigatoria 2611: 2608: 2605: 2604: 2601: 2598: 2595: 2587: 2586: 2579: 2576: 2573: 2570: 2562: 2561: 2558: 2555: 2552: 2549: 2541: 2540: 2537: 2534: 2531: 2528: 2520: 2519: 2516: 2515:Misidentified 2513: 2510: 2507: 2499: 2498: 2495: 2492: 2489: 2481: 2480: 2477: 2474: 2471: 2468: 2460: 2459: 2456: 2453: 2450: 2447: 2439: 2438: 2435: 2432: 2429: 2426: 2418: 2417: 2414: 2411: 2408: 2405: 2397: 2396: 2393: 2390: 2387: 2384: 2376: 2375: 2372: 2369: 2366: 2363: 2355: 2354: 2351: 2348: 2345: 2337: 2336: 2333: 2330: 2327: 2324: 2321:Partula remota 2316: 2315: 2312: 2309: 2306: 2303: 2295: 2294: 2291: 2288: 2285: 2282: 2274: 2273: 2270: 2267: 2264: 2261: 2253: 2252: 2249: 2246: 2243: 2240: 2232: 2231: 2228: 2225: 2222: 2219: 2211: 2210: 2207: 2204: 2201: 2198: 2190: 2189: 2186: 2183: 2180: 2177: 2169: 2168: 2165: 2162: 2159: 2156: 2148: 2147: 2140:Espiritu Santo 2136: 2133: 2130: 2127: 2119: 2118: 2115: 2114:Misidentified 2112: 2109: 2106: 2098: 2097: 2086: 2083: 2080: 2077: 2069: 2068: 2065: 2062: 2059: 2056:Partula nodosa 2051: 2050: 2047: 2044: 2041: 2033: 2032: 2029: 2026: 2023: 2015: 2014: 2011: 2008: 2005: 1997: 1996: 1993: 1990: 1987: 1984: 1981:Partula micans 1976: 1975: 1972: 1969: 1966: 1963: 1960:Partula meyeri 1955: 1954: 1951: 1948: 1945: 1942: 1934: 1933: 1930: 1927: 1924: 1921: 1913: 1912: 1905: 1902: 1899: 1896: 1888: 1887: 1880: 1877: 1874: 1871: 1863: 1862: 1859: 1856: 1853: 1850: 1842: 1841: 1838: 1835: 1832: 1829: 1826:Partula lirata 1821: 1820: 1817: 1814: 1811: 1808: 1800: 1799: 1796: 1793: 1790: 1787: 1779: 1778: 1775: 1772: 1769: 1766: 1763:Partula leefei 1758: 1757: 1754: 1751: 1748: 1745: 1737: 1736: 1721: 1718: 1715: 1712: 1704: 1703: 1696: 1693: 1690: 1687: 1679: 1678: 1675: 1672: 1669: 1666: 1658: 1657: 1650: 1647: 1644: 1641: 1633: 1632: 1623: 1620: 1617: 1614: 1606: 1605: 1598: 1595: 1592: 1587: 1579: 1578: 1575: 1572: 1569: 1561: 1560: 1557: 1554: 1551: 1548: 1540: 1539: 1536: 1533: 1530: 1527: 1524:Partula grisea 1519: 1518: 1495: 1492: 1489: 1486: 1478: 1477: 1474: 1471: 1466: 1458: 1457: 1454: 1451: 1446: 1441: 1433: 1432: 1429: 1426: 1423: 1420: 1412: 1411: 1408: 1405: 1402: 1399: 1391: 1390: 1387: 1384: 1381: 1378: 1370: 1369: 1366: 1363: 1360: 1357: 1349: 1348: 1345: 1342: 1339: 1336: 1328: 1327: 1324: 1321: 1318: 1315: 1307: 1306: 1303: 1300: 1297: 1292: 1284: 1283: 1280: 1279:Misidentified 1277: 1274: 1271: 1263: 1262: 1255: 1252: 1249: 1246: 1238: 1237: 1234: 1231: 1228: 1225: 1217: 1216: 1213: 1210: 1207: 1199: 1198: 1195:Bellona Island 1191:Rennell Island 1187: 1184: 1181: 1178: 1170: 1169: 1160: 1157: 1154: 1151: 1148:Partula cootei 1143: 1142: 1139: 1136: 1133: 1130: 1122: 1121: 1118: 1115: 1112: 1109: 1101: 1100: 1097: 1094: 1091: 1088: 1080: 1079: 1076: 1073: 1070: 1067: 1059: 1058: 1055: 1052: 1049: 1046: 1038: 1037: 1034: 1031: 1028: 1025: 1017: 1016: 1013:Torres Islands 1005: 1002: 999: 994: 986: 985: 982: 979: 974: 971:Partula atilis 966: 965: 960:Threatened by 958: 955: 952: 949: 941: 940: 937: 934: 931: 926: 923:Partula arguta 918: 917: 910: 907: 904: 899: 891: 890: 887: 882: 877: 872: 866: 865: 862: 857: 854: 845: 842: 760: 757: 741:Henry Crampton 701:in the family 626: 625: 623: 622: 614: 606: 597: 588: 580: 564: 555: 525: 516: 500: 484: 476: 468: 460: 452: 444: 428: 409: 396:Partula (Echo) 393: 377: 369: 353: 345: 337: 329: 321: 313: 305: 299: 293: 290: 280: 279: 278: 275: 274: 268: 267: 253: 249: 248: 243: 239: 238: 233: 229: 228: 223: 219: 218: 213: 209: 208: 203: 199: 198: 193: 189: 188: 183: 179: 178: 173: 169: 168: 155: 154: 146: 145: 137: 136: 124: 123: 38: 36: 29: 15: 9: 6: 4: 3: 2: 6022: 6011: 6008: 6006: 6003: 6001: 5998: 5996: 5993: 5992: 5990: 5973: 5968: 5964: 5960: 5955: 5951: 5947: 5942: 5938: 5934: 5929: 5925: 5921: 5916: 5912: 5908: 5903: 5899: 5895: 5890: 5886: 5882: 5877: 5873: 5869: 5864: 5860: 5856: 5851: 5847: 5842: 5836: 5832: 5827: 5821: 5817: 5816: 5814: 5812: 5808: 5804: 5799: 5793: 5792:Partula Pages 5789: 5788: 5778: 5774: 5769: 5764: 5760: 5756: 5752: 5747: 5744: 5743: 5738: 5735: 5734: 5729: 5727:, 228: 1-311. 5726: 5723: 5719: 5716: 5715: 5703: 5701: 5697: 5693: 5689: 5684: 5682: 5673: 5669: 5663: 5655: 5648: 5642: 5627: 5623: 5616: 5602: 5598: 5591: 5575: 5569: 5567: 5551: 5545: 5539: 5534: 5532: 5524: 5520: 5516: 5510: 5503: 5497: 5491: 5486: 5479: 5473: 5466: 5463: 5460:(June 1821). 5459: 5454: 5450: 5447: 5435: 5434: 5427: 5422: 5419: 5418: 5411: 5406: 5403: 5402: 5395: 5390: 5387: 5386: 5385:Partula gibba 5379: 5374: 5373: 5364: 5363: 5356: 5351: 5348: 5347: 5340: 5335: 5332: 5331: 5324: 5319: 5318: 5309: 5308: 5307:Partula varia 5301: 5296: 5293: 5292: 5291:Partula rosea 5285: 5280: 5279: 5270: 5269: 5262: 5257: 5254: 5253: 5246: 5241: 5238: 5237: 5230: 5225: 5222: 5221: 5214: 5209: 5206: 5205: 5198: 5193: 5192: 5183: 5182: 5175: 5170: 5167: 5166: 5159: 5154: 5151: 5150: 5143: 5138: 5135: 5134: 5127: 5122: 5119: 5118: 5117:Partula clara 5111: 5106: 5103: 5102: 5095: 5090: 5089: 5078: 5076: 5072: 5071: 5066: 5062: 5061: 5056: 5042: 5041: 5033: 5032: 5024: 5023: 5015: 5014: 5011: 5010: 5009: 5002: 5001: 4995: 4994: 4991: 4990: 4987: 4986: 4985: 4978: 4977: 4971: 4970: 4966: 4965: 4962: 4961: 4953: 4952: 4944: 4943: 4935: 4934: 4926: 4925: 4922: 4921: 4920: 4913: 4912: 4909: 4908: 4905: 4904: 4903: 4896: 4895: 4889: 4888: 4884: 4883: 4880: 4879: 4876: 4875: 4874: 4867: 4866: 4860: 4859: 4856: 4855: 4852: 4851: 4850: 4843: 4842: 4839: 4838: 4835: 4834: 4833: 4826: 4825: 4822: 4821: 4813: 4812: 4804: 4803: 4800: 4799: 4798: 4791: 4790: 4787: 4786: 4783: 4782: 4781: 4774: 4773: 4767: 4766: 4763: 4762: 4759: 4758: 4757: 4750: 4749: 4743: 4742: 4736: 4735: 4731: 4730: 4727: 4726: 4718: 4717: 4714: 4713: 4712: 4705: 4704: 4701: 4700: 4697: 4696: 4695: 4688: 4687: 4684: 4683: 4680: 4679: 4678: 4671: 4670: 4667: 4666: 4658: 4657: 4654: 4653: 4652: 4645: 4644: 4641: 4640: 4632: 4631: 4623: 4622: 4614: 4613: 4610: 4609: 4607: 4600: 4599: 4596: 4595: 4592: 4591: 4589: 4582: 4581: 4575: 4574: 4571: 4570: 4562: 4561: 4553: 4552: 4549: 4548: 4546: 4545:P. otaheitana 4539: 4538: 4532: 4531: 4528: 4527: 4524: 4522: 4515: 4514: 4508: 4507: 4501: 4500: 4497: 4496: 4488: 4487: 4479: 4478: 4470: 4469: 4461: 4460: 4457: 4456: 4454: 4453:P. otaheitana 4447: 4446: 4440: 4439: 4436: 4435: 4432: 4431: 4430: 4429:P. otaheitana 4423: 4422: 4416: 4415: 4412: 4411: 4408: 4407: 4406: 4399: 4398: 4392: 4391: 4388: 4387: 4384: 4383: 4382: 4375: 4374: 4371: 4370: 4367: 4366: 4365: 4364:P. otaheitana 4358: 4357: 4351: 4350: 4344: 4343: 4337: 4336: 4333: 4332: 4324: 4323: 4320: 4319: 4318: 4311: 4310: 4307: 4306: 4303: 4302: 4301: 4294: 4293: 4290: 4289: 4286: 4285: 4284: 4277: 4276: 4273: 4272: 4269: 4268: 4267: 4260: 4259: 4256: 4255: 4247: 4246: 4243: 4242: 4241: 4234: 4233: 4230: 4229: 4226: 4225: 4224: 4217: 4216: 4210: 4209: 4206: 4205: 4197: 4196: 4188: 4187: 4179: 4178: 4175: 4174: 4173: 4166: 4165: 4159: 4158: 4155: 4154: 4151: 4150: 4149: 4142: 4141: 4135: 4134: 4131: 4130: 4122: 4121: 4113: 4112: 4104: 4103: 4100: 4099: 4098: 4091: 4090: 4084: 4083: 4080: 4079: 4076: 4075: 4074: 4067: 4066: 4063: 4062: 4059: 4058: 4057: 4050: 4049: 4043: 4042: 4039: 4038: 4035: 4034: 4033: 4026: 4025: 4019: 4018: 4012: 4011: 4008: 4007: 3999: 3998: 3995: 3994: 3993: 3986: 3985: 3982: 3981: 3978: 3977: 3975: 3968: 3967: 3964: 3963: 3960: 3959: 3958: 3951: 3950: 3947: 3946: 3943: 3941: 3940: 3933: 3932: 3929: 3928: 3925: 3923: 3922: 3915: 3914: 3911: 3910: 3902: 3901: 3893: 3892: 3884: 3883: 3880: 3879: 3878: 3871: 3870: 3867: 3866: 3863: 3862: 3860: 3859:P. otaheitana 3853: 3852: 3846: 3845: 3842: 3841: 3833: 3832: 3829: 3828: 3826: 3825:P. otaheitana 3819: 3818: 3815: 3814: 3811: 3810: 3808: 3807:P. otaheitana 3801: 3800: 3797: 3796: 3793: 3792: 3790: 3789:P. otaheitana 3783: 3782: 3776: 3775: 3772: 3771: 3768: 3767: 3766: 3759: 3758: 3752: 3751: 3748: 3747: 3744: 3742: 3735: 3734: 3728: 3727: 3724: 3723: 3715: 3714: 3711: 3710: 3709: 3702: 3701: 3698: 3697: 3694: 3693: 3692: 3685: 3684: 3681: 3680: 3672: 3671: 3668: 3667: 3666: 3659: 3658: 3655: 3654: 3651: 3650: 3649: 3642: 3641: 3635: 3634: 3631: 3630: 3622: 3621: 3618: 3617: 3616: 3609: 3608: 3605: 3604: 3601: 3600: 3599: 3592: 3591: 3588: 3587: 3584: 3583: 3582: 3575: 3574: 3571: 3570: 3567: 3566: 3565: 3558: 3557: 3554: 3553: 3550: 3549: 3548: 3541: 3540: 3537: 3536: 3533: 3532: 3531: 3524: 3523: 3520: 3519: 3516: 3515: 3514: 3507: 3506: 3500: 3499: 3493: 3492: 3489: 3488: 3480: 3479: 3471: 3470: 3462: 3461: 3458: 3457: 3456: 3455:P. hebe bella 3449: 3448: 3442: 3441: 3438: 3437: 3434: 3433: 3432: 3425: 3424: 3421: 3420: 3412: 3411: 3403: 3402: 3399: 3398: 3397: 3390: 3389: 3383: 3382: 3379: 3378: 3375: 3374: 3373: 3366: 3365: 3359: 3358: 3352: 3351: 3348: 3347: 3339: 3338: 3335: 3334: 3333: 3326: 3325: 3322: 3321: 3318: 3317: 3316: 3309: 3308: 3305: 3304: 3301: 3300: 3299: 3292: 3291: 3288: 3287: 3284: 3283: 3282: 3275: 3274: 3268: 3267: 3264: 3263: 3255: 3254: 3246: 3245: 3242: 3241: 3240: 3233: 3232: 3226: 3225: 3222: 3221: 3218: 3217: 3216: 3209: 3208: 3205: 3204: 3196: 3195: 3192: 3191: 3190: 3183: 3182: 3179: 3178: 3175: 3174: 3173: 3166: 3165: 3162: 3161: 3158: 3157: 3156: 3149: 3148: 3145: 3144: 3141: 3140: 3139: 3132: 3131: 3125: 3124: 3118: 3117: 3111: 3110: 3104: 3103: 3097: 3096: 3090: 3089: 3085: 3084: 3078: 3077: 3071: 3070: 3066: 3065: 3061: 3059: 3055: 3045: 3044: 3043: 3038: 3034: 3033: 3032: 3027: 3026: 3021: 3017: 3016: 3015: 3010: 3009: 3004: 3000: 2999: 2998: 2993: 2992: 2987: 2983: 2982: 2981: 2976: 2975: 2970: 2966: 2965: 2964: 2959: 2958: 2953: 2952: 2947: 2946: 2941: 2937: 2936: 2934: 2930: 2925: 2924: 2919: 2918: 2913: 2911: 2907: 2902: 2898: 2897: 2896: 2891: 2890: 2885: 2881: 2880: 2879: 2875: 2871: 2867: 2862: 2861: 2856: 2852: 2851: 2850: 2849:P. otaheitana 2845: 2844: 2839: 2838: 2833: 2832: 2827: 2826: 2821: 2820: 2815: 2811: 2803: 2801: 2797: 2790: 2786: 2782: 2781: 2780: 2775: 2774: 2770: 2766: 2765: 2764: 2759: 2757: 2753: 2748: 2747: 2742: 2740: 2736: 2731: 2730: 2726: 2722: 2721: 2720: 2715: 2714: 2709: 2708: 2704: 2700: 2699: 2698: 2693: 2692: 2687: 2686: 2681: 2680: 2675: 2674: 2669: 2668: 2664: 2660: 2659: 2658: 2657:P. otaheitana 2653: 2652: 2647: 2646: 2641: 2640: 2635: 2634: 2629: 2625: 2623: 2622: 2617: 2602: 2599: 2596: 2594: 2593: 2592:Partula varia 2589: 2588: 2584: 2580: 2577: 2574: 2571: 2569: 2568: 2564: 2563: 2559: 2556: 2553: 2550: 2548: 2547: 2543: 2542: 2538: 2535: 2532: 2529: 2527: 2526: 2522: 2521: 2517: 2514: 2511: 2508: 2506: 2505: 2501: 2500: 2496: 2493: 2490: 2488: 2487: 2483: 2482: 2478: 2475: 2472: 2469: 2467: 2466: 2462: 2461: 2457: 2454: 2451: 2448: 2446: 2445: 2441: 2440: 2436: 2433: 2430: 2427: 2425: 2424: 2420: 2419: 2415: 2412: 2409: 2406: 2404: 2403: 2399: 2398: 2394: 2391: 2388: 2385: 2383: 2382: 2378: 2377: 2373: 2370: 2367: 2364: 2362: 2361: 2357: 2356: 2352: 2349: 2346: 2344: 2343: 2342:Partula rosea 2339: 2338: 2334: 2331: 2328: 2325: 2323: 2322: 2318: 2317: 2313: 2310: 2307: 2304: 2302: 2301: 2297: 2296: 2292: 2289: 2286: 2283: 2281: 2280: 2276: 2275: 2271: 2268: 2265: 2262: 2260: 2259: 2255: 2254: 2250: 2247: 2244: 2241: 2239: 2238: 2234: 2233: 2229: 2226: 2223: 2220: 2218: 2217: 2213: 2212: 2208: 2205: 2202: 2199: 2197: 2196: 2192: 2191: 2187: 2184: 2181: 2178: 2176: 2175: 2171: 2170: 2166: 2163: 2160: 2157: 2155: 2154: 2150: 2149: 2145: 2141: 2137: 2134: 2131: 2128: 2126: 2125: 2121: 2120: 2116: 2113: 2110: 2107: 2105: 2104: 2100: 2099: 2095: 2091: 2087: 2084: 2081: 2078: 2076: 2075: 2074:Partula obesa 2071: 2070: 2066: 2063: 2060: 2058: 2057: 2053: 2052: 2048: 2045: 2042: 2040: 2039: 2035: 2034: 2030: 2027: 2024: 2022: 2021: 2017: 2016: 2012: 2009: 2006: 2004: 2003: 1999: 1998: 1994: 1991: 1988: 1985: 1983: 1982: 1978: 1977: 1973: 1970: 1967: 1964: 1962: 1961: 1957: 1956: 1952: 1949: 1946: 1943: 1941: 1940: 1936: 1935: 1931: 1928: 1925: 1922: 1920: 1919: 1915: 1914: 1910: 1906: 1903: 1900: 1897: 1895: 1894: 1893:Partula lutea 1890: 1889: 1885: 1881: 1878: 1875: 1872: 1870: 1869: 1865: 1864: 1860: 1857: 1854: 1851: 1849: 1848: 1844: 1843: 1839: 1836: 1833: 1830: 1828: 1827: 1823: 1822: 1818: 1815: 1812: 1809: 1807: 1806: 1802: 1801: 1797: 1794: 1791: 1788: 1786: 1785: 1781: 1780: 1776: 1773: 1770: 1767: 1765: 1764: 1760: 1759: 1755: 1752: 1749: 1746: 1744: 1743: 1739: 1738: 1734: 1730: 1726: 1722: 1719: 1716: 1713: 1711: 1710: 1706: 1705: 1701: 1697: 1694: 1691: 1688: 1686: 1685: 1681: 1680: 1676: 1673: 1670: 1667: 1665: 1664: 1660: 1659: 1655: 1651: 1648: 1645: 1642: 1640: 1639: 1635: 1634: 1630: 1629: 1628:Partula clara 1624: 1621: 1618: 1615: 1613: 1612: 1608: 1607: 1603: 1599: 1596: 1593: 1591: 1588: 1586: 1585: 1581: 1580: 1576: 1573: 1570: 1568: 1567: 1563: 1562: 1558: 1555: 1552: 1549: 1547: 1546: 1542: 1541: 1537: 1534: 1531: 1528: 1526: 1525: 1521: 1520: 1516: 1513:. Extinct on 1512: 1508: 1504: 1500: 1496: 1493: 1490: 1487: 1485: 1484: 1483:Partula gibba 1480: 1479: 1475: 1472: 1470: 1467: 1465: 1464: 1460: 1459: 1455: 1452: 1450: 1447: 1445: 1442: 1440: 1439: 1435: 1434: 1430: 1427: 1424: 1421: 1419: 1418: 1414: 1413: 1409: 1406: 1403: 1400: 1398: 1397: 1393: 1392: 1388: 1385: 1382: 1379: 1377: 1376: 1372: 1371: 1367: 1364: 1361: 1358: 1356: 1355: 1351: 1350: 1346: 1343: 1340: 1337: 1335: 1334: 1330: 1329: 1325: 1322: 1319: 1316: 1314: 1313: 1309: 1308: 1304: 1301: 1298: 1296: 1293: 1291: 1290: 1286: 1285: 1281: 1278: 1275: 1272: 1270: 1269: 1265: 1264: 1260: 1256: 1253: 1250: 1247: 1245: 1244: 1240: 1239: 1235: 1232: 1229: 1226: 1224: 1223: 1219: 1218: 1214: 1211: 1208: 1206: 1205: 1201: 1200: 1196: 1192: 1188: 1185: 1182: 1179: 1177: 1176: 1172: 1171: 1167: 1166: 1161: 1158: 1155: 1152: 1150: 1149: 1145: 1144: 1140: 1137: 1134: 1131: 1129: 1128: 1124: 1123: 1119: 1116: 1113: 1110: 1108: 1107: 1103: 1102: 1098: 1095: 1092: 1089: 1087: 1086: 1085:Partula clara 1082: 1081: 1077: 1074: 1071: 1068: 1066: 1065: 1061: 1060: 1056: 1053: 1050: 1047: 1045: 1044: 1040: 1039: 1035: 1032: 1029: 1026: 1024: 1023: 1019: 1018: 1014: 1010: 1006: 1003: 1000: 998: 995: 993: 992: 988: 987: 983: 980: 978: 975: 973: 972: 968: 967: 963: 959: 956: 953: 950: 948: 947: 943: 942: 938: 935: 932: 930: 927: 925: 924: 920: 919: 915: 911: 908: 905: 903: 900: 898: 897: 893: 892: 888: 886: 883: 881: 878: 876: 873: 870: 869: 863: 860: 859: 853: 851: 841: 839: 833: 831: 830:least concern 827: 823: 819: 815: 811: 807: 803: 798: 796: 791: 789: 785: 780: 778: 777: 772: 771: 766: 756: 754: 750: 746: 742: 738: 733: 731: 727: 723: 722:Pacific Ocean 719: 715: 711: 706: 704: 700: 697: 694: 691: 687: 683: 679: 678: 671: 670: 665: 659: 658: 657:Partula gibba 653: 647: 646: 641: 637: 632: 620:Hartman, 1881 618: 615: 612:Iredale, 1941 610: 607: 601: 598: 594:Hartman, 1881 592: 589: 586:Pilsbry, 1909 584: 581: 577:Goodsir, 1845 575: 570:Hartman, 1881 568: 565: 559: 556: 550: 545:Lamarck, 1812 543: 536: 531:Hartman, 1881 529: 526: 520: 517: 513:Lamarck, 1818 511: 506:Hartman, 1881 504: 501: 495: 490:Hartman, 1881 488: 485: 482:Pilsbry, 1909 480: 477: 474:Hartman, 1881 472: 469: 466:Pilsbry, 1909 464: 461: 458:Pilsbry, 1909 456: 453: 450:Pilsbry, 1909 448: 445: 439: 434:Hartman, 1881 432: 429: 426: 422:Mulsant, 1846 420: 415:Hartman, 1881 413: 410: 404: 399:Hartman, 1881 397: 394: 388: 383:Hartman, 1881 381: 378: 375:Pilsbry, 1909 373: 370: 364: 359:Hartman, 1881 357: 354: 351:Hartman, 1881 349: 346: 343:Hartman, 1881 341: 338: 335:Hartman, 1881 333: 330: 327:Pilsbry, 1909 325: 322: 319:Hartman, 1881 317: 314: 311:Hartman, 1881 309: 306: 303:Hartman, 1881 300: 297:Hartman, 1881 294: 291: 288:Hartman, 1881 285: 284: 276: 273: 269: 264: 259: 258: 254: 251: 250: 247: 244: 241: 240: 237: 234: 232:Superfamily: 231: 230: 227: 224: 221: 220: 217: 214: 211: 210: 207: 204: 201: 200: 197: 194: 191: 190: 187: 184: 181: 180: 177: 174: 171: 170: 165: 160: 156: 153: 152: 147: 143: 138: 135: 131: 128: 120: 117: 109: 98: 95: 91: 88: 84: 81: 77: 74: 70: 67: –  66: 62: 61:Find sources: 55: 51: 45: 44: 39:This article 37: 33: 28: 27: 22: 5810: 5758: 5754: 5741: 5732: 5721: 5699: 5695: 5691: 5671: 5662: 5653: 5647:"Tentacle 3" 5641: 5629:. Retrieved 5625: 5615: 5604:. Retrieved 5600: 5590: 5578:. Retrieved 5553:. Retrieved 5544: 5522: 5518: 5509: 5501: 5496: 5485: 5472: 5464: 5461: 5453: 5445: 5431: 5415: 5399: 5383: 5360: 5346:Partula hebe 5344: 5328: 5305: 5289: 5266: 5250: 5234: 5218: 5202: 5179: 5163: 5147: 5131: 5115: 5099: 5068: 5058: 5054: 5053: 5006: 5005: 4982: 4981: 4967: 4917: 4916: 4900: 4899: 4887:Palaopartula 4885: 4871: 4870: 4847: 4846: 4830: 4829: 4795: 4794: 4778: 4777: 4754: 4753: 4732: 4709: 4708: 4692: 4691: 4677:P. auraniana 4675: 4674: 4649: 4648: 4604: 4603: 4586: 4585: 4543: 4542: 4518: 4451: 4450: 4427: 4426: 4403: 4402: 4379: 4378: 4362: 4361: 4317:P. similaris 4315: 4314: 4298: 4297: 4283:P. radiolata 4281: 4280: 4264: 4263: 4238: 4237: 4223:P. assimilis 4221: 4220: 4170: 4169: 4146: 4145: 4095: 4094: 4071: 4070: 4056:P. dentifera 4054: 4053: 4032:P. dentifera 4030: 4029: 3992:P. tohiveana 3990: 3989: 3974:P. mirabilis 3972: 3971: 3955: 3954: 3937: 3936: 3919: 3918: 3875: 3874: 3857: 3856: 3823: 3822: 3805: 3804: 3787: 3786: 3763: 3762: 3738: 3706: 3705: 3689: 3688: 3663: 3662: 3646: 3645: 3613: 3612: 3596: 3595: 3579: 3578: 3562: 3561: 3545: 3544: 3528: 3527: 3511: 3510: 3453: 3452: 3429: 3428: 3394: 3393: 3370: 3369: 3330: 3329: 3313: 3312: 3298:P. mirabilis 3296: 3295: 3279: 3278: 3237: 3236: 3213: 3212: 3187: 3186: 3170: 3169: 3153: 3152: 3136: 3135: 3087: 3086: 3067: 3058:monophyletic 3053: 3051: 3042:P. assimilis 3040: 3037:Cook Islands 3036: 3035: 3031:P. similaris 3029: 3025:P. auraniana 3023: 3019: 3018: 3012: 3008:P. cramptoni 3006: 3002: 3001: 2995: 2989: 2985: 2984: 2978: 2972: 2968: 2967: 2963:P. radiolata 2961: 2957:P. lutaensis 2955: 2951:P. langfordi 2949: 2943: 2939: 2938: 2933:P. dentifera 2927: 2921: 2915: 2906:P. garrettii 2904: 2900: 2899: 2893: 2887: 2883: 2882: 2878:P. tohiveana 2870:P. suturalis 2863: 2860:P. mirabilis 2859: 2854: 2853: 2847: 2841: 2835: 2829: 2823: 2817: 2813: 2812: 2809: 2799: 2796:P. garrettii 2795: 2794: 2788: 2779:P. langfordi 2777: 2771: 2768: 2767: 2761: 2756:P. dentifera 2750: 2744: 2735:P. garrettii 2733: 2727: 2724: 2723: 2717: 2711: 2705: 2702: 2701: 2697:P. tohiveana 2695: 2689: 2685:P. suturalis 2683: 2677: 2673:P. mirabilis 2671: 2665: 2662: 2661: 2655: 2649: 2643: 2637: 2631: 2627: 2626: 2619: 2615: 2613: 2590: 2565: 2544: 2523: 2502: 2484: 2463: 2442: 2421: 2400: 2379: 2360:Partula rufa 2358: 2340: 2319: 2298: 2277: 2256: 2235: 2214: 2193: 2172: 2151: 2122: 2101: 2072: 2054: 2036: 2018: 2000: 1979: 1958: 1937: 1916: 1891: 1866: 1845: 1824: 1803: 1782: 1761: 1740: 1707: 1682: 1661: 1636: 1626: 1609: 1601: 1582: 1566:Partula hebe 1564: 1543: 1522: 1497:Persists in 1481: 1461: 1436: 1417:Partula faba 1415: 1394: 1373: 1352: 1331: 1310: 1287: 1266: 1241: 1220: 1202: 1173: 1163: 1146: 1125: 1104: 1083: 1062: 1041: 1020: 989: 969: 962:habitat loss 944: 921: 894: 849: 847: 834: 805: 804:contains 73 799: 792: 783: 781: 774: 768: 762: 745:Yoshio Kondo 743:(along with 734: 709: 707: 676: 675: 674: 667: 655: 643: 639: 616: 608: 603:Clench, 1941 599: 590: 582: 573: 566: 561:Clench, 1941 557: 548: 541: 534: 527: 518: 509: 502: 493: 486: 478: 470: 462: 454: 446: 437: 430: 424: 418: 411: 402: 395: 386: 379: 371: 366:Röding, 1798 362: 355: 347: 339: 331: 323: 315: 307: 256: 255: 222:Infraorder: 149: 133: 127: 112: 103: 93: 86: 79: 72: 60: 48:Please help 43:verification 40: 5902:iNaturalist 5835:Wikispecies 5688:Mayer A. G. 5515:Burch J. B. 5008:Eua zebrina 4984:Eua globosa 3957:P. mooreana 3877:P. producta 3827:sinistrorsa 3809:sinistralis 3691:P. mooreana 3615:P. taeniata 3547:P. mooreana 3138:P. aurantia 2974:P. emersoni 2874:P. taeniata 2866:P. mooreana 2837:P. incrassa 2691:P. taeniata 2679:P. mooreana 2667:P. aurantia 2581:Endemic to 2088:Endemic to 1882:Endemic to 1723:Endemic to 1009:Aore Island 793:Today, the 690:terrestrial 686:land snails 609:Scilistylus 441:Risso, 1826 406:Selys, 1853 236:Pupilloidea 226:Pupilloidei 5989:Categories 5606:2023-05-27 5442:References 4694:P. turneri 4608:intermedia 4405:P. affinis 4240:P. luteola 4172:P. turgida 4073:P. tristis 3939:P. hyalina 3921:P. hyalina 3708:P. tohivea 3332:P. hyalina 3069:Partulidae 2969:Micronesia 2910:P. tristis 2831:P. hyalina 2819:P. affinis 2769:Marianas - 2763:P. turgida 2739:P. tristis 2645:P. hyalina 2633:P. affinis 914:Tahiti Iti 826:vulnerable 822:endangered 737:Polynesian 730:New Guinea 703:Partulidae 538:Oken, 1807 425:; Marianna 246:Partulidae 206:Gastropoda 76:newspapers 5672:d-nb.info 5436:(extinct) 4711:P. lirata 4606:P. nodosa 4590:composita 4588:P. nodosa 4455:rubescens 4097:P. meyeri 3976:propinqua 3942:(Tahiti) 3530:P. exigua 3048:Cladogram 3014:P. micans 2997:P. lirata 2991:P. leefei 2923:P. meyeri 2843:P. nodosa 2725:Raiatea - 2707:P. arguta 2703:Huahine - 2651:P. nodosa 2536:Formerly 1909:Bora Bora 1753:Formerly 1674:Formerly 1654:Mt. Aorai 1494:Formerly 1428:Formerly 1259:Mt. Marau 1138:Formerly 1117:Formerly 1096:Formerly 1033:Formerly 936:Formerly 856:Full list 852:include: 816:, 17 are 812:, 10 are 696:gastropod 693:pulmonate 600:Rennellia 552:Lea, 1833 182:Kingdom: 176:Eukaryota 106:June 2024 5826:Q6296067 5820:Wikidata 5777:17610827 5720:(1916). 5626:Mongabay 5075:turmeric 5070:Dracaena 5065:plantain 5060:Caladium 4266:P. gibba 3791:lignaria 3648:P. clara 3513:P. clara 3396:P. rosea 3372:P. varia 3281:P. clara 2945:P. gibba 2940:Marianas 2895:P. varia 2889:P. rosea 2825:P. clara 2773:P. gibba 2719:P. varia 2713:P. rosea 2663:Moorea - 2639:P. clara 2583:Vanikoro 1515:Aguiguan 871:Species 824:, 2 are 820:, 8 are 770:Achatina 699:mollusks 645:Caladium 591:Pasithea 549:Pasithea 542:Pasithea 535:Pasithea 419:Harmonia 324:Marianna 308:Harmonia 286:Astraea 272:Synonyms 263:Férussac 242:Family: 196:Mollusca 192:Phylum: 186:Animalia 172:Domain: 5920:1349766 5894:4597417 5881:4925260 5841:Partula 5811:Partula 5692:Partula 5580:28 June 5555:28 June 5519:Partula 5502:Partula 5081:Gallery 5055:Partula 5050:Ecology 4734:Samoana 4148:P. faba 3431:P. hebe 3088:Partula 3054:Partula 2980:P. rufa 2917:P. hebe 2901:Raiatea 2884:Huahine 2789:Partula 2746:P. hebe 2729:P. faba 2621:ex situ 2616:Partula 1503:Sarigan 850:Partula 844:Species 838:Mo'orea 810:extinct 806:Partula 788:extinct 784:Partula 759:Decline 716:" and " 710:Partula 677:Partula 640:Partula 617:Sterope 574:Sterope 363:Astraea 295:Clytia 257:Partula 252:Genus: 212:Order: 202:Class: 134:Partula 90:scholar 5972:861628 5959:324349 5946:362162 5933:145553 5775:  5631:17 May 4547:crassa 3861:crassa 2855:Moorea 2814:Tahiti 2628:Tahiti 2094:Futuna 1700:Tahiti 1511:Saipan 889:Notes 765:Tahiti 636:Saipan 510:Oenone 438:Helena 387:Clytia 348:Oenone 332:Matata 316:Helena 265:, 1821 92:  85:  78:  71:  63:  5967:WoRMS 5915:IRMNG 5907:90863 5868:7PBZ7 5650:(PDF) 2132:6-10 2111:5-10 2090:Alofi 1733:Naiau 1729:Cicia 1507:Pagan 682:genus 680:is a 494:Nenia 340:Nenia 301:Echo 97:JSTOR 83:books 5928:NCBI 5889:GBIF 5855:3012 5850:BOLD 5773:PMID 5700:XXVI 5668:"34" 5633:2023 5582:2024 5557:2024 5073:and 5063:and 2986:Fiji 2798:and 2614:The 2603:Yes 2497:Yes 2476:Yes 2455:Yes 2353:Yes 2305:N/A 2287:1-2 2245:4-5 2144:Aore 2142:and 2092:and 2067:Yes 2049:Yes 2031:Yes 2013:Yes 1944:N/A 1884:Rota 1873:N/A 1731:and 1725:Mago 1597:Yes 1594:1-5 1577:Yes 1509:and 1499:Guam 1476:Yes 909:Yes 547:and 403:Echo 69:news 5876:EoL 5863:CoL 5763:doi 5702:(2) 5521:". 4969:Eua 2597:EW 2578:No 2572:CR 2557:No 2551:EX 2530:EX 2509:EX 2491:EW 2470:CR 2449:EW 2434:No 2428:LC 2413:No 2407:EX 2392:No 2386:EX 2371:No 2365:PE 2347:EW 2332:No 2326:EX 2311:No 2290:No 2284:EN 2269:No 2263:EN 2248:No 2242:EN 2227:No 2221:EX 2206:No 2200:EX 2185:No 2179:EX 2164:No 2158:EX 2135:No 2129:VU 2108:EN 2085:No 2079:CR 2061:EW 2043:EW 2025:EW 2007:EW 1992:No 1986:LC 1971:No 1965:PE 1950:No 1929:No 1923:EX 1904:No 1898:EX 1879:No 1858:No 1852:EX 1837:No 1834:11 1831:EN 1816:No 1810:EX 1795:No 1789:EX 1774:No 1768:CR 1747:EX 1720:No 1714:EN 1695:No 1689:PE 1668:EX 1649:No 1643:PE 1622:No 1616:CR 1602:may 1571:EW 1556:No 1550:EX 1535:No 1529:LC 1488:EN 1453:No 1422:EX 1407:No 1401:EX 1386:No 1380:CR 1365:No 1359:EX 1344:No 1338:EX 1323:No 1317:EX 1302:No 1295:N/A 1273:EX 1254:No 1248:PE 1233:No 1227:EX 1215:No 1209:EX 1186:No 1180:CR 1159:No 1153:EX 1132:PE 1111:EX 1093:20 1090:CR 1075:No 1069:EX 1054:No 1048:EX 1027:EX 1004:No 984:No 957:No 951:CR 728:to 705:. 52:by 5991:: 5969:: 5956:: 5943:: 5930:: 5917:: 5904:: 5891:: 5878:: 5865:: 5852:: 5837:: 5822:: 5771:. 5759:17 5757:. 5753:. 5680:^ 5670:. 5652:. 5624:. 5599:. 5565:^ 5530:^ 5465:92 5077:. 4523:) 3743:) 3039:- 3028:, 3022:- 3011:, 3005:- 2994:, 2988:- 2977:, 2971:- 2960:, 2954:, 2948:, 2942:- 2926:, 2920:, 2914:, 2903:- 2892:, 2886:- 2876:, 2872:, 2868:, 2864:, 2857:- 2846:, 2840:, 2834:, 2828:, 2822:, 2816:- 2776:, 2760:, 2749:, 2743:, 2732:, 2716:, 2710:, 2694:, 2688:, 2682:, 2676:, 2670:, 2654:, 2648:, 2642:, 2636:, 2630:- 2600:0 2585:. 2575:1 2554:0 2533:0 2512:0 2494:0 2473:2 2452:0 2431:? 2410:0 2389:0 2368:1 2350:0 2329:0 2308:? 2266:1 2224:0 2203:0 2182:0 2161:0 2082:2 2064:0 2046:0 2028:0 2010:0 1989:? 1968:1 1947:0 1926:0 1901:0 1876:? 1855:0 1813:0 1792:0 1771:1 1750:0 1735:. 1727:, 1717:3 1702:. 1692:1 1671:0 1656:. 1646:1 1631:. 1619:1 1590:VU 1574:0 1553:0 1532:? 1517:. 1505:, 1501:, 1491:4 1473:0 1469:EW 1444:LC 1425:0 1404:0 1383:2 1362:0 1341:0 1320:0 1299:0 1276:0 1251:1 1230:0 1212:0 1197:. 1183:1 1156:0 1135:1 1114:0 1072:0 1051:0 1030:0 1015:. 1001:5 997:EN 981:0 977:EX 954:1 933:0 929:PE 906:1 902:CR 840:. 832:. 790:. 755:. 732:. 688:, 579:) 554:) 540:, 515:) 499:) 443:) 392:) 368:) 5779:. 5765:: 5674:. 5656:. 5635:. 5609:. 5584:. 5559:. 5525:. 5480:. 4519:( 3739:( 2935:) 2931:( 2912:) 2908:( 2758:) 2754:( 2741:) 2737:( 1449:? 885:C 880:P 875:S 119:) 113:( 108:) 104:( 94:· 87:· 80:· 73:· 46:. 23:.

Index

Partula (disambiguation)

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Partula radiolata
Scientific classification
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Eukaryota
Animalia
Mollusca
Gastropoda
Stylommatophora
Pupilloidei
Pupilloidea
Partulidae
Partula
Férussac
Synonyms

Saipan
Caladium

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