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Enantiornithes

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gizzard and rely solely on strong stomachal acids. An example was discovered with what was suspected to be gastroliths in the what would have been the fossil's stomach, re-opening the discussion of the use of gastroliths by Enantiornithes. X-ray and scanning microscope inspection of the rocks determined that they were actually chalcedony crystals, and not gastroliths.
1412:, which includes all living birds as a subset. This means that Enantiornithes were a successful branch of avialan evolution, but one that diversified entirely separately from the lineage leading to modern birds. One study has however found that the shared sternal anatomy was acquired independently and such a relationship needs to be reexamined. 1192:. Juvenile specimens can be identified by a combination of factors: rough texture of their bone tips indicating portions which were still made of cartilage at the time of death, relatively small breastbones, large skulls and eyes, and bones which had not yet fused to one another. Some hatchling specimens have been given formal names, including " 491:(the combined upper foot and ankle bone) is correct, but Walker did not use this reasoning in his original paper. Walker never described the fusion of the tarsometatarsus as opposite, but rather as "Only partial". Also, it is not certain that Enantiornithes had triosseal canals, since no fossil preserves this feature. 1424:
findings impossible. Because it is often unfeasible for other scientists to study each specimen in person given the worldwide distribution of the Enantiornithes, and due to the many uninformative descriptions which have been published on possibly important specimens, many of these specimens become "functional
929:, evolved alongside a short, triangular pygostyle, like the ones in modern birds, rather than the long, rod- or dagger-shaped pygostyles in more primitive avialans like the Enantiornithes. Instead of a feather fan, most Enantiornithes had a pair of long specialized pinfeathers similar to those of the extinct 1106:
A study on paravian digestive systems indicates that known Enantiornithes lacked a crop and a gizzard, didn't use gastroliths and didn't eject pellets. This is considered at odds with the high diversity of diets that their different teeth and skull shapes imply, though some modern birds have lost the
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seems unable to support the muscles that control the modern tail feathers involved in flight. Though some basal Enantiornithes exhibit ancestral flight apparatuses, by the end of the Mesozoic many Enantiornithes had several features convergent with the Neornithes including a deeply keeled sternum, a
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is the most extensively studied family in terms of diet due to their rather unusual rostral anatomy, with long jaws and few teeth arranged at the jaw ends. They have variously been interpreted as piscivores, probers akin to shorebirds and as arboreal bark-probers. A 2022 study however does find them
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outlined a number of criticisms against the prevailing practices of scientists failing to describe many specimens in enough detail for others to evaluate thoroughly. Some species have been described based on specimens which are held in private collections, making further study or review of previous
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deposits dated to 99 million years ago and include hatchlings described in 2017 and 2018, as well as isolated body parts such as wings and feet. These amber remains are among the most well-preserved of any mesozoic dinosaur. Fossils of this clade have been found in both inland and marine sediments,
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Enantiornithes taxonomy is difficult to evaluate, and as a result few clades within the group are consistently found by phylogenetic analyses. Most Enantiornithes are not included in any specific family, and as such are listed here. Many of these have been considered Euenantiornithes, although the
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remains preserved this way (a few isolated feathers are otherwise known, unassigned to any species), and one of the most exquisitely preserved dinosaurian fossils known. The preserved wings show variations in feather pigment and prove that Enantiornithes had fully modern feathers, including barbs,
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Sanz, José L.; Chiappe, Luis M.; Pérez-Moreno, Bernardino P.; Moratalla, José J.; HernÑndez-Carrasquilla, Francisco; Buscalioni, Angela D.; Ortega, Francisco; Poyato-Ariza, Francisco J.; Rasskin-Gutman, Diego; Martı́nez-Delclòs, Xavier (June 6, 1997). "A Nestling Bird from the Lower Cretaceous of
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You, Hai-lu; Lamanna, Matthew C.; Harris, Jerald D.; Chiappe, Luis M.; O'Connor, Jingmai; Ji, Shu-an; LΓΌ, Jun-chang; Yuan, Chong-xi; Li, Da-qing; Zhang, Xing; Lacovara, Kenneth J.; Dodson, Peter; Ji, Qiang (16 June 2006). "A Nearly Modern Amphibious Bird from the Early Cretaceous of Northwestern
331:. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over eighty species of Enantiornithes have been named, but some names represent only single bones, so it is likely that not all are valid. The Enantiornithes became extinct at the 1133:
in 2001 included the remains of four hatchling skeletons of three different species of Enantiornithes. They are substantially complete, very tightly associated, and show surface pitting of the bones that indicates partial digestion. The authors concluded that this association was a regurgitated
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O’Connor, Jingmai K.; Zhou, Zhonghe; Zhang, Fucheng (28 February 2011). "A reappraisal of Boluochia zhengi (Aves: Enantiornithes) and a discussion of intraclade diversity in the Jehol avifauna, China". Journal of Systematic Palaeontology. 9 (1): 51–63. doi:10.1080/14772019.2010.512614. S2CID
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As a very large group of birds, the Enantiornithes displayed a high diversity of different body plans based on differences in ecology and feeding, reflected in an equal diversity of wing forms, many paralleling adaptions to different lifestyles seen in modern birds. In general, the wings of
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Although the vast majority of histology studies and known remains of Enantiornithes point to superprecociality being the norm, one specimen, MPCM-LH-26189, seems to represent an altricial juvenile, implying that like modern birds Enantiornithes explored multiple reproductive strategies.
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Lianhai Hou; LuisM. Chiappe; Fucheng Zhang; Cheng-Ming Chuong (2004). "New Early Cretaceous fossil from China documents a novel trophic specialization for Mesozoic birds". Naturwissenschaften. 91 (1): 22–25. Bibcode:2004NW.....91...22H. doi:10.1007/s00114-003-0489-1. PMC 4382005. PMID
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is subjected to two differing definitions of varying inclusiveness. The more inclusive definition, which follows Cau & Arduini (2008), is used here. Avisaurids were a long-lasting and widespread family of Enantiornithes, which are mainly distinguished by specific features of their
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Morschhauser, E. M.; Varricchio, D.J.; Gao, C.; Liu, J.; Wang, Z.; Cheng, X. & Meng, Q. (2009). "Anatomy of the Early Cretaceous bird Rapaxavis pani, a new species from Liaoning Province, China". Journal of Vertebrate Paleontology. 29 (2): 545–554. doi:10.1671/039.029.0210. S2CID
9271:: Two different wings from hatchling specimens which were described in 2015. They attracted a significant amount of media attention upon their description. They were preserved in exceptional details due to having been trapped within Burmese amber for approximately 99 million years. 1202:
and colleagues considered the practice of naming new species based on juveniles detrimental to the study of Enantiornithes, because it is nearly impossible to determine which adult species a given juvenile specimen belongs to, making any species with a hatchling holotype a
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As a group, the Enantiornithes are often referred to as "enantiornithines" in literature. However, several scientists have noted that this is incorrect, because following the standard rules for forming the names of animal groups, it implies reference only to the subfamily
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Cambra-Moo, Oscar; Buscalioni, Ángela Delgado; Cubo, Jorge; Castanet, Jacques; Loth, Marie-Madeleine; de Margerie, Emmanuel; de Ricqlès, Armand (2006). "Histological observations of Enantiornithine bone (Saurischia, Aves) from the Lower Cretaceous of Las Hoyas (Spain)".
9216:. This specimen was originally considered to be a tiny adult, but later found to be a hatchling. Other specimens have henceforth been assigned to the genus. Due to a lack of distinguishing feature, many paleontologists have considered this genus an undiagnostic 10964:
O'Connor, Jingmai K.; Li, Da-Qing; Lamanna, Matthew C.; Wang, Min; Harris, Jerald D.; Atterholt, Jessie; You, Hai-Lu (30 December 2015). "A new Early Cretaceous enantiornithine (Aves, Ornithothoraces) from northwestern China with elaborate tail ornamentation".
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Bell, Alyssa K.; Chiappe, Luis M.; Erickson, Gregory M.; Suzuki, Shigeru; Watabe, Mahito; Barsbold, Rinchen; Tsogtbaatar, K. (February 2010). "Description and ecologic analysis of Hollanda luceria, a Late Cretaceous bird from the Gobi Desert (Mongolia)".
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birds, which are known to reach adult size quickly). Studies of the rate of bone growth in a variety of Enantiornithes has shown that smaller species tended to grow faster than larger ones, the opposite of the pattern seen in more primitive species like
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was a family of large early Enantiornithes. They had numerous small teeth and numerous primitive features which are lost in most other Enantiornithes. Mostly known from the early Cretaceous of China, with putative Late Cretaceous taxa from Madagascar.
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O'Connor, Jingmai K.; Wang, Xuri; Chiappe, Luis M.; Gao, Chunling; Meng, Qingjin; Cheng, Xiaodong; Liu, Jinyuan (12 March 2009). "Phylogenetic support for a specialized clade of Cretaceous enantiornithine birds with information from a new species".
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Harris, Jerald D.; Lamanna, Matthew C.; You, Hai-lu; Ji, Shu-an; Ji, Qiang (18 May 2006). "A second enantiornithean (Aves: Ornithothoraces) wing from the Early Cretaceous Xiagou Formation near Changma, Gansu Province, People's Republic of China".
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However, several studies have shown that they were efficient flyers, like modern birds, possessing a similarly complex nervous system and wing feather ligaments. Additionally, the lack of a complex tail appears to not have been very relevant for
847:. While most Enantiornithes had claws on at least some of their fingers, many species had shortened hands, a highly mobile shoulder joint, and proportional changes in the wing bones similar to modern birds. Like modern birds, Enantiornithes had 751:
Given their wide range of habitats and diets, the cranial morphology of Enantiornithes varied considerably between species. Skulls of Enantiornithes combined a unique suite of primitive and advanced features. As in more primitive avialans like
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A 2020 study on a juvenile's feathers further stresses the ontological similarities to modern megapodes, but cautions several differences such as the arboreal nature of most Enantiornithes as opposed to the terrestrial lifestyle of megapodes.
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in Enantiornithes must have been a function of tail shortening, not the development of a modern tail feather anatomy. These scientists suggested that a fan of tail feathers and the associated musculature needed to control them, known as the
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O'Connor, Patrick M.; Turner, Alan H.; Groenke, Joseph R.; Felice, Ryan N.; Rogers, Raymond R.; Krause, David W.; Rahantarisoa, Lydia J. (10 December 2020). "Late Cretaceous bird from Madagascar reveals unique development of beaks".
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were large but geologically short-lived early Enantiornithes, with long, hooked talons and robust teeth with curved tips. They may have been equivalent to birds of prey, although this interpretation is open to much debate. The
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Many fossils of Enantiornithes are very fragmentary, and some species are only known from a piece of a single bone. Almost all specimens that are complete, in full articulation, and with soft tissue preservation are known from
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Xing, Lida; O'Connor, Jingmai K.; McKellar, Ryan C.; Chiappe, Luis M.; Tseng, Kuowei; Li, Gang; Bai, Ming (September 2017). "A mid-Cretaceous enantiornithine (Aves) hatchling preserved in Burmese amber with unusual plumage".
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and other birds. The sap would have fossilized and become amber. However, more recently it has been suggested that the sap moved post-mortem, hence not representing true stomachal contents. Combined with the putative fish
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studies have recovered Enantiornithes as a monophyletic group distinct from the modern birds and their closest relatives. The 2002 phylogenetic analysis by Clarke and Norell, though, reduced the number of Enantiornithes
700:(with the latter species being described as similar in size to modern turkeys,) although at least a few larger species may have also existed, including a potentially crane-sized species known only from footprints in the 1309:
Traditionally, they have been considered inferior flyers, due to the shoulder girdle anatomy being assumed to be more primitive and unable to support a ground-based launching mechanism, as well as due to the absence of
1248:, this small species did not reach adult size for a long time, probably several years. Other studies have all supported the view that growth to adult size was slow, as it is in living precocial birds (as opposed to 1120:
due to its larger size, poorly preserved skull and unusual pedal anatomy), being too small for specialised carnivory and herbivory; the atypical rostrum is tentatively speculated to be unrelated to feeding ecology.
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tarsometatarsals (ankle bones). The largest and most advanced members of the group survived in North and South America up until the end of the Cretaceous, yet are very fragmentary compared to some earlier taxa.
2085:, 2022, and includes most named taxa and recovers several previously-named clades. Letters on branches indicate the positions of "wildcard" taxa, those which have been recovered in multiple disparate positions. 12719:
Wang, Li; O'Connor, Zhou; You (2015). "Second species of enantiornithine bird from the Lower Cretaceous Changma Basin, northwestern China with implications for the taxonomic diversity of the Changma avifauna".
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Zelenkov, Nikita V.; Averianov, Alexander O. (13 June 2015). "A historical specimen of enantiornithine bird from the Early Cretaceous of Mongolia representing a new taxon with a specialized neck morphology".
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Xing, Lida; McKellar, Ryan C.; Wang, Min; Bai, Ming; O'Connor, Jingmai K.; Benton, Michael J.; Zhang, Jianping; Wang, Yan; Tseng, Kuowei; Lockley, Martin G.; Li, Gang; Zhang, Weiwei; Xu, Xing (28 June 2016).
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Houde, Peter W. (1988). "Paleognathous Birds from the Early Tertiary of the Northern Hemisphere". Publications of the Nuttall Ornithological Club (Cambridge Massachusetts, USA: Nuttall Ornithological Club)
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Sanz, JosΓ© L.; Chiappe, Luis M.; FernΓ‘dez-Jalvo, Yolanda; Ortega, Francisco; SΓ‘nchez-ChillΓ³n, BegoΓ±a; Poyato-Ariza1, Francisco J.; PΓ©rez-Moreno, Bernardino P. (February 2001). "An early Cretaceous pellet".
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The birds are so named because, among many distinctive features, there is a unique formation of the triosseal canal and the metatarsals are fused proximally to distally, the opposite of that in modern birds
9838:НовыС ΠΏΡ‚ΠΈΡ†Ρ‹ ΠΌΠ΅Π»-ΠΏΠ°Π»Π΅ΠΎΠ³Π΅Π½Π° Π‘Π‘Π‘Π  ΠΈ Π½Π΅ΠΊΠΎΡ‚ΠΎΡ€Ρ‹Π΅ замСчания ΠΏΠΎ истории возникновСния ΠΈ ΡΠ²ΠΎΠ»ΡŽΡ†ΠΈΠΈ класса (New Cretaceous-Paleogene birds of USSR and some remarks about history of the origin and evolution of class) 912:(2006) surveyed all fossils of Enantiornithes then known and concluded that none had preserved tail feathers that formed a lift-generating fan, as in modern birds. They found that all avialans outside of 609:). The widespread occurrence of this group suggests that at least some Enantiornithes were able to cross oceans under their own power; they are the first known avialan lineage with a global distribution. 13460:
Wang, Yan; Wang, Min; O'Connor, Jingmai K.; Wang, Xiaoli; Zheng, Xiaoting; Zhang, Xiaomei (11 January 2016). "A new Jehol enantiornithine bird with three-dimensional preservation and ovarian follicles".
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Because many Enantiornithes lacked complex tails and possessed radically different wing anatomy compared to modern birds, they have been the subject of several studies testing their flight capabilities.
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Hu, Dongyu; Liu, Ying; Li, Jinhua; Xu, Xing; Hou, Lianhai (July 2015). "Yuanjiawaornis viriosus, gen. et sp. nov., a large enantiornithine bird from the Lower Cretaceous of western Liaoning, China".
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patterns of development in birds of today. In other words, Enantiornithes probably hatched from the egg already well developed and ready to run, forage, and possibly even fly at just a few days old.
9277:: A partial corpse of an Enantiornithes hatchling also preserved in Burmese amber. Although indeterminate, it attracted even more media attention than the two wings upon its description in 2017. 982:, suggesting that this might have been the ancestral condition, with pinfeathers being a feature evolved several times in early avialans for display purposes. Another species of Enantiornithes, 12423:
Chiappe, Luis M.; Suzuki, Shigeru; Dyke, Gareth J.; Watabe, Mahito; Tsogtbaatar, K.; Barsbold, Rinchen (January 2007). "A new Enantiornithine bird from the Late Cretaceous of the Gobi desert".
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Hu, Dongyu; Xu, Xing; Hou, Lianhai; Sullivan, Corwin (2012). "A New Enantiornithine Bird from the Lower Cretaceous of Western Liaoning, China, and Its Implications for Early Avian Evolution".
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in many anatomical features of the flight apparatus, but a sternal keel is absent in the basal-most members, only a single basal taxon appears to have had a triosseal canal, and their robust
390:, giving the entire group its name. Since the 1990s, many more complete specimens of Enantiornithes have been discovered, and it was determined that a few previously described "birds" (e.g. 11905:
FernΓ‘ndez, Mariela S.; GarcΓ­a, Rodolfo A.; Fiorelli, Lucas; Scolaro, Alejandro; Salvador, Rodrigo B.; Cotaro, Carlos N.; Kaiser, Gary W.; Dyke, Gareth J.; Farke, Andrew A. (17 April 2013).
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and western edge of prehistoric Asia suggest a worldwide distribution of this group or in the relatively warm regions, at least. Enantiornithes have been found on every continent except
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One of the first Jehol biota Enantiornithes described. Known from many species, although some are now placed into their own genera. May have had a similar appearance and lifestyle to a
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A few specimens preserve actual stomach contents. Unfortunately, none of these preserve the skull, so direct correlation between their known diet and snout/tooth shape cannot be made.
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in Spain, including both slab and counter-slab components. Its 2018 description revealed how various features developed in Enantiornithes as they aged. Such features include the
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Although only known from a few bones, this genus is the namesake of Enantiornithes. It was also one of the largest and last representative of the group prior to their extinction
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O'Connor, Jingmai K.; Falk, Amanda; Wang, Min; Zheng, Xiao-Ting (2020). "First report of immature feathers in juvenile Enantiornithes from the Early Cretaceous Jehol avifauna".
9249:: Briefly described in 2004, this Yixian Enantiornithes specimen had advanced pennaceous feathers on its legs, similar to (albeit shorter than) those of other paravians such as 9625:"The coracoscapular joint of neornithine birdsβ€”extensive homoplasy in a widely neglected articular surface of the avian pectoral girdle and its possible functional correlates" 7161:
was a family of long-snouted early Cretaceous Enantiornithes, with teeth only at the tips of the snout. They are generally considered to be fairly basal members of the group.
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is often found to be the most primitive or basal member of the Enantiornithes, Euenantiornithes may be an extremely inclusive group, made up of all Enantiornithes except for
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Enantiornithes appear to have included waders, swimmers, granivores, insectivores, fishers, and raptors. The vast majority of Enantiornithes were small, between the size of a
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Walker was not clear on his reasons for giving this name in the etymology section of his paper, and this ambiguity led to some confusion among later researchers. For example,
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It has been speculated that superprecociality in Enantiornithes might have prevented them from developing specialised toe arrangements seen in modern birds like zygodactyly.
948:, was initially interpreted as having at least four long tail feathers that overlapped each other and might have formed a lift-generating surface similar to the tail fans of 804:(holes in the side of the head) merged into the orbits as in modern birds due to the postorbitals either not being present or not being long enough to divide the openings. A 466:(the primary bone of the shoulder girdle in vertebrates other than mammals) that is the reverse of that of modern birds. Specifically, in the Enantiornithes, the scapula is 10544:
Wang, Min; Hu, Han; Li, Zhiheng (21 August 2015). "A new small enantiornithine bird from the Jehol Biota, with implications for early evolution of avian skull morphology".
11807:"Palaeohistological inferences of resting metabolic rates in Concornis and Iberomesornis (Enantiornithes, Ornithothoraces) from the Lower Cretaceous of las Hoyas (Spain)" 10179:
Zhang, Zihui; Chiappe, Luis M.; Han, Gang; Chinsamy, Anusuya (2013). "A large bird from the Early Cretaceous of China: new information on the skull of enantiornithines".
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Hu, Han; O'Connor, Jingmai K. (14 November 2016). "First species of Enantiornithes from Sihedang elucidates skeletal development in Early Cretaceous enantiornithines".
13348:"A New Species of Pengornithidae (Aves: Enantiornithes) from the Lower Cretaceous of China Suggests a Specialized Scansorial Habitat Previously Unknown in Early Birds" 851:, or "bastard wings", small forward-pointing arrangements of feathers on the first digit that granted higher maneuverability in the air and aided in precise landings. 1462:
Enantiornithes systematics are highly provisional and notoriously difficult to study, due to their small size and the fact that Enantiornithes tend to be extremely
9845: 879:, however differ by the feathers being shorter, more disorganized (they do not clearly form a wing) and only extend down to the ankle rather than along the foot. 9904:
Xing, Lida; O'Connor, Jingmai K.; McKellar, Ryan C.; Chiappe, Luis M.; Bai, Ming; Tseng, Kuowei; Zhang, Jie; Yang, Haidong; Fang, Jun; Li, Gang (February 2018).
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Kurochkin, E. N.; Chatterjee, S.; Mikhailov, K. E. (19 December 2013). "An embryonic enantiornithine bird and associated eggs from the cretaceous of Mongolia".
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Kurochkin, E. N.; Chatterjee, S.; Mikhailov, K. E. (19 December 2013). "An embryonic enantiornithine bird and associated eggs from the cretaceous of Mongolia".
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Kurochkin, E. N.; Chatterjee, S.; Mikhailov, K. E. (December 2013). "An embryonic enantiornithine bird and associated eggs from the cretaceous of Mongolia".
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Herrera, Gerardo Álvarez; Agnolín, Federico; Rozadilla, SebastiÑn; Lo Coco, Gastón E.; Manabe, Makoto; Tsuihiji, Takanobu; Novas, Fernando E. (2022-12-16).
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Dyke, Gareth; Vremir, MΓ‘tyΓ‘s; Kaiser, Gary; Naish, Darren (June 2012). "A drowned Mesozoic bird breeding colony from the Late Cretaceous of Transylvania".
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Perhaps the most fundamental and characteristic difference between the Enantiornithes and all other birds is in the nature of the articulation between the
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Wang, Min; Zhou, Zhonghe (12 April 2017). "A morphological study of the first known piscivorous enantiornithine bird from the Early Cretaceous of China".
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Chiappe, Luis M.; Bo, Zhao; O'Connor, Jingmai K.; Chunling, Gao; Xuri, Wang; Habib, Michael; Marugan-Lobon, Jesus; Qingjin, Meng; Xiaodong, Cheng (2014).
1224:, these finds demonstrate that hatchling Enantiornithes had the skeletal ossification, well-developed wing feathers, and large brain which correlate with 1033:
Given the wide diversity of skull shape among Enantiornithes, many different dietary specializations must have been present among the group. Some, like
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barbules, and hooklets, and a modern arrangement of wing feather including long flight feathers, short coverts, a large alula and an undercoat of down.
499:. Following the naming conventions used for modern birds as well as extinct groups, it has been pointed out that the correct term is "enantiornithean". 14138: 6066: 7709:
The oldest known member of the family, and one of the oldest putative genera of Enantiornithes known. Possessed extremely well-preserved tail ribbons
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The first Enantiornithes to be discovered were incorrectly referred to modern bird groups. For example, the first known species of Enantiornithes,
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Wang, M.; O'Connor, J. K.; Zhou, Z. (2014). "A new robust enantiornithine bird from the Lower Cretaceous of China with scansorial adaptations".
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However, further discoveries showed that at least among basal Enantiornithes, tail anatomy was more complex than previously thought. One genus,
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and dish-shaped at this joint, and the coracoid is convex. In modern birds, the coracoscapular joint has a concave coracoid and convex scapula.
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Known from a disarticulated skeleton preserved overlying a piece of stomach content composed of fish bones, which may have been its last meal
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suggest that they were predators of small to medium-sized vertebrates, but their robust teeth instead suggest a diet of hard-shelled animals.
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and in non-avialan dinosaurs. Some analyses have interpreted the bone histology to indicate that Enantiornithes may not have had fully avian
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O'Connor, Jingmai K.; Chiappe, Luis M. (28 February 2011). "A revision of enantiornithine (Aves: Ornithothoraces) skull morphology".
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Xing, Lida; O'Connor, Jingmai K.; Chiappe, Luis M.; McKellar, Ryan C.; Carroll, Nathan; Hu, Han; Bai, Ming; Lei, Fumin (2019-07-22).
4183: 12629:
Wang, Xuri; Cau, Andrea; Luo, Xiaoling; KundrΓ‘t, Martin; Wu, Wensheng; Ju, Shubin; Guo, Zhen; Liu, Yichuan; Ji, Qiang (2022-02-11).
12024:
Kaye, Thomas G.; Pittman, Michael; MarugΓ‘n-LobΓ³n, JesΓΊs; MartΓ­n-Abad, Hugo; Sanz, JosΓ© Luis; Buscalioni, Angela D. (21 March 2019).
5516:
One of the first genera of Enantiornithes known from decent remains. Also one of the oldest and most primitive members of the group
1244:
bones showed a growth pattern different from modern birds; although growth was rapid for a few weeks after hatching, probably until
13581: 5374:
since its discovery in 1977 until it received a formal description in 2015. Possessed unique neck vertebrae and a primitive palate
13032:
Liu, Di; Chiappe, Luis M.; Serrano, Francisco; Habib, Michael; Zhang, Yuguang; Meng, Qinjing; Shawkey, Matthew (11 October 2017).
12991:
Wang, Xiaoli; Clark, Alexander D.; O'Connor, Jingmai K.; Zhang, Xiangyu; Wang, Xing; Zheng, Xiaoting; Zhou, Zhonghe (2024-02-27).
11698:
Dinosaur incubation periods directly determined from growth-line counts in embryonic teeth show reptilian-grade development - PNAS
5171:
The proportions of the wings of this genus of Enantiornithes as well as the presence of an alula suggest that it was a good flier
4918:
Developed a massive snout with only a single tooth, despite retaining a "primitive" skull arrangement in contrast to modern birds
920:) with preserved tail feathers had only short coverts or elongated paired tail plumes. They suggested that the development of the 15222: 10076:
Xing, Lida; McKellar, Ryan C.; O'Connor, Jingmai K. (June 2020). "An unusually large bird wing in mid-Cretaceous Burmese amber".
9179:
due to having been described from the same specimen. Despite having been described in 1999, 5 years prior to the description of
10757:"Insight into the evolution of avian flight from a new clade of Early Cretaceous ornithurines from China and the morphology of 826: 13126:
Wang, Min; Zhou, Zhonghe; Xu, Guanghui (7 January 2014). "The first enantiornithine bird from the Upper Cretaceous of China".
8099:
A possible member of the family, known from a well-preserved skeleton complete with structures believed to be developing eggs
1415:
Enantiornithes classification and taxonomy has historically been complicated by a number of factors. In 2010, paleontologists
1238:
Analyses of Enantiornithes bone histology have been conducted to determine the growth rates of these animals. A 2006 study of
9677: 9607: 9458: 9433: 8442:
A toothless advanced genus of Enantiornithes, possessing a robust beak which convergently evolved with those of modern birds
12869:
Hartman, Scott; Mortimer, Mickey; Wahl, William R.; Lomax, Dean R.; Lippincott, Jessica; Lovelace, David M. (10 July 2019).
12175:"Soft-tissue and dermal arrangement in the wing of an Early Cretaceous bird: Implications for the evolution of avian flight" 8688:
Although incomplete, its skeleton possesses three-dimensional preservation. Possibly flightless due to its wing proportions
13405:
Chiappe, Luis M.; Qingjin, Meng; Serrano, Francisco; Sigurdsen, Trond; Min, Wang; Bell, Alyssa; Di, Liu (25 October 2019).
11005:"A bizarre Early Cretaceous enantiornithine bird with unique crural feathers and an ornithuromorph plough-shaped pygostyle" 9243:
of the skeleton, while later studies reported a squamosal bone present in the specimen but unknown in other Enantiornithes.
8637:
The eponymous avisaurid, as well as one of the largest members of the family. Originally considered a non-avialan dinosaur
7422:
Like other longipterygids, it possessed a thin snout which may have been used for probing for invertebrates in mud or bark
1397:, but in several respects they were more primitive than modern birds, perhaps following an intermediate evolutionary path. 12611: 10406:
Wang, Min; Zhou, Zhonghe; Xu, Guanghui (2014-01-01). "The first enantiornithine bird from the Upper Cretaceous of China".
1326:
also lacked complex tail feathers but were good flyers, and they appear to have been capable of a ground based launching.
9836: 9351:"Insights into the evolution of rachis dominated tail feathers from a new basal enantiornithine (Aves: Ornithothoraces)" 1072:
preserved corpuscles of amber among the fossilized bones, suggesting that this animal fed on tree sap, much like modern
16476: 12026:"Fully fledged enantiornithine hatchling revealed by Laser-Stimulated Fluorescence supports precocial nesting behavior" 7806:
The first pengornithid discovered, and also one of the largest members of the Enantiornithes known from decent remains
6531:
Small but fully mature at the time of its death. One of only a few Chinese Enantiornithes dated to the Late Cretaceous
814:. The presence of these primitive features of the skull would have rendered the Enantiornithes capable of only limited 11178:"The evolution of the modern avian digestive system: insights from paravian fossils from the Yanliao and Jehol biotas" 10224:"Insight into diversity, body size and morphological evolution from the largest Early Cretaceous enantiornithine bird" 12173:
Navalón, Guillermo; MarugÑn-Lobón, Jesús; Chiappe, Luis M.; Luis Sanz, José; Buscalioni, Ángela D. (6 October 2015).
10460:
Carvalho, Ismar; Novas, Fernando; Agnolin, Federico; Isasi, Marcelo; Freitas, Francisco; Andrade, JosΓ© (2015-06-05).
332: 8048:
Adapted for flapping, rather than soaring, flight. Its describers suggest paraphyly or polyphyly of Bohaiornithidae
6104:
A small member of the group which possessed several primitive and derived features compared to other Enantiornithes
1400:
A consensus of scientific analyses indicates that Enantiornithes is one of two major groups within the larger group
11285:"Diet of Mesozoic toothed birds (Longipterygidae) inferred from quantitative analysis of extant avian diet proxies" 11232: 10019:
Xing, Lida; McKellar, Ryan C.; O'Connor, Jingmai K.; Bai, Ming; Tseng, Kuowei; Chiappe, Luis M. (30 January 2019).
9818: 6679:
One disarticulated skeleton from this genus possesses well-preserved bones of the skull, including a quadratojugal
5313:
Originally identified as an ornithuromorph but since reinterpreted as a genus of Enantiornithes closely related to
843:, and displayed some features related to flight similar to those found in the lineage leading to modern birds, the 762:(bones of the snout tip) and most species had toothy jaws rather than toothless beaks. Only a few species, such as 14705: 11067:(Aves: Enantiornithes) from the Lower Cretaceous of Spain and a Reexamination of its Phylogenetic Relationships". 9808:
Discovery of a Cretaceous bird, apparently ancestral to the orders Coraciiformes and Piciformes (Aves: Carinatae)
1235:
Findings suggests Enantiornithes, especially the toothed species, had a longer incubation time than modern birds.
14766: 13554: 11805:
Cubo, Jorge; Buscalioni, Angela D.; Legendre, Lucas J.; Bourdon, Estelle; Sanz, Jose L.; Ricqlès, Armand (2021).
11502:
Zhou, Zhonghe; Zhang, Fucheng (October 22, 2004). "A Precocial Avian Embryo from the Lower Cretaceous of China".
11090:
Dalla Vecchia, Fabio M.; Chiappe, Luis M. (2003). "First avian skeleton from the Mesozoic of northern Gondwana".
10581:"A Comparative Morphological Study of the Jugal and Quadratojugal in Early Birds and Their Dinosaurian Relatives" 5670:
So well preserved that its flight pattern could be reconstructed using the proportions of its feathers and wings
17: 12798:
DalsΓ€tt, J.; Ericson, P. G.; Zhou, Z. (2015). "A New Enantiornithes (Aves) from the Early Cretaceous of China".
12582: 11103: 10365:
Martin, Anthony J.; Vickers-Rich, Patricia; Rich, Thomas H.; Hall, Michael; Angielczyk, Kenneth (January 2014).
7107: 6960: 6016: 5535: 438:
in his landmark paper which established the group. In his paper, Walker explained what he meant by "opposite":
270: 12124: 11757: 11563: 9598:
Hope, Sylvia (2002). "The Mesozoic Radiation of Neornithes". In Chiappe, Luis M.; Witmer, Lawrence M. (eds.).
9388: 673:, however display considerable variation in size with some species. The largest species in this clade include 12871:"A new paravian dinosaur from the Late Jurassic of North America supports a late acquisition of avian flight" 12143: 9225:"Wasaibpanchi": A supposed member of Enantiornithes from Pakistan; the describing paper is of dubious status. 7139:
A large, toothless genus of Enantiornithes, one of the most-well preserved members from the Late Cretaceous.
4327:
Had an unusual ornithuromorph-like pygostyle and brush-like thigh feathers. One of the oldest Enantiornithes
4030: 654: 14711: 12366:"Avian Wing Proportions and Flight Styles: First Step towards Predicting the Flight Modes of Mesozoic Birds" 10920:
O'Connor, Jingmai K.; Wang, Xiaoli; Zheng, Xiaoting; Hu, Han; Zhang, Xiaomei; Zhou, Zhonghe (January 2016).
13574: 12631:"A new bohaiornithid-like bird from the Lower Cretaceous of China fills a gap in enantiornithine disparity" 10661:
Becker, Rachel (28 June 2016). "Bird wings trapped in amber are a fossil first from the age of dinosaurs".
4969:
Possessed an elaborate set of tail feathers, unlike the paired ribbon-like feathers of most Enantiornithes
152: 11782:
Chiappe, L.M. (1995). "The phylogenetic position of the Cretaceous birds of Argentina: Enantiornithes and
9422:"Skeletal Morphology and Systematics of the Cretaceous Euenantiornithes (Ornithothoraces: Enantiornithes)" 6200:
Known from a juvenile specimen which depicts how various features developed in Enantiornithes as they age
4482:
A genus of Enantiornithes from the Changma basin, an area which is unusually dominated by ornithuromorphs
1134:
pellet and, from the details of the digestion and the size, that the hatchlings were swallowed whole by a
16355: 15212: 13411:-like bird from the Early Cretaceous of China: enantiornithine interrelationships and flight performance" 13214:"New enantiornithine bird from the uppermost Cretaceous (Maastrichtian) of southern Patagonia, Argentina" 8905:
Although closely related to some of the largest avisaurids, members of this genus were very small birds
4430: 565:
in age, and it is believed that the Enantiornithes became extinct at the same time as their non-avialan
14994: 10922:"An Enantiornithine with a Fan-Shaped Tail, and the Evolution of the Rectricial Complex in Early Birds" 9047: 8819: 8373:
Some members of the group are obscure or poorly described and may be synonymous with its type species,
1007: 800:. In modern birds these bones are assimilated into the cranium. Some Enantiornithes may have had their 13213: 12992: 7852:
Possesses an elaborate "pintail" tail fan longer than its body, which may have had a display function
5837: 1267:. However a 2021 study rejects the idea that they had less endothermic metabolisms than modern birds. 834:
scan of an amber-encased wing attributable to Enantiornithes showing rachises, skin, muscle and claws.
16466: 16461: 14909: 14899: 14733: 12307:"The most complete enantiornithine from North America and a phylogenetic analysis of the Avisauridae" 11449: 10367:"Oldest known avian footprints from Australia: Eumeralla Formation (Albian), Dinosaur Cove, Victoria" 10316:"The most complete enantiornithine from North America and a phylogenetic analysis of the Avisauridae" 9519: 8738:
beads, however this was later determined as an artefact of preservation and not an indicator of diet
5267:
A poorly known genus of Enantiornithes, but evidently a large and late-surviving member of the group
1498: 712:). Among the smallest described specimens are unnamed hatchlings, although the holotype specimens of 12573: 11866: 11426:
Mikhailov, Konstantin E. (1996). "New Genera of Fossil Eggs from the Upper Cretaceous of Mongolia".
10021:"A fully feathered enantiornithine foot and wing fragment preserved in mid-Cretaceous Burmese amber" 16411: 15235: 15217: 13678: 12993:"First Edentulous Enantiornithine (Aves: Ornithothoraces) from the Lower Cretaceous Jehol Avifauna" 9305:: Another corpse of an indeterminate hatchling preserved in Burmese amber. Described in early 2018. 3835: 1087:
turning out to be fish excrement, the strange stomachal contents of some species turning out to be
1039:, had large, robust jaws suitable for eating hard-shelled invertebrates. The short, blunt teeth of 8489:
Poorly known and described from a skeleton which has now been lost. May have been synonymous with
7524:
Acquired multiple tail feathers which may have been capable of generating lift as in modern birds
4281:
A very well-preserved South American member of the group, complete with ribbon-like tail feathers
3698:
controversy behind this name means that it is not used consistently in studies of Enantiornithes.
15240: 13567: 6632: 14914: 12956:
Wang, M.; Liu, D. (2015). "Taxonomical reappraisal of Cathayornithidae (Aves: Enantiornithes)".
11708: 7426: 4772: 1830: 1800: 524:, may have been the earliest known member of Enantiornithes according to Agnolin et al. (2017). 16438: 16373: 16331: 15207: 15019: 14969: 13034:"Flight aerodynamics in enantiornithines: Information from a new Chinese Early Cretaceous bird" 12568: 11861: 11401: 8757: 435: 12144:"New basal beaked ornithurine bird found from the lower cretaceous of Western Liaoning, China" 11552:"Juvenile Birds from the Early Cretaceous of China: Implications for Enantiornithine Ontogeny" 11283:
Miller, Case Vincent; Pittman, Michael; Wang, Xiaoli; Zheng, Xiaoting; Bright, Jen A. (2022).
9421: 3944: 16433: 15044: 11993:"Exploring the Ecomorphology of Two Cretaceous Enantiornithines With Unique Pedal Morphology" 11380: 9208: 9143: 8929: 8641: 8415: 7277: 6992:
The southernmost known member of Enantiornithes and one of the youngest members of the group
5343: 5286: 4026:
that suggests a feeding specialization similar to hummingbirds, honeyeaters, and woodpeckers
1467: 1194: 831: 369:. The Enantiornithes were first recognized as a distinct lineage, or "subclass" of birds, by 11445: 9515: 6292:
Fragmentary, but may have been a seabird because remains from this genus have been found as
5033:
One of the oldest North American avialans found, albeit known only from fragmentary remains
16420: 15672: 13732: 13513: 13470: 13359: 13304: 13268: 13178: 13135: 13045: 12843: 12764: 12729: 12642: 12479: 12432: 12377: 12243: 12186: 12037: 11918: 11853: 11730: 11664: 11621: 11344: 11142: 11016: 10821: 10708: 10505: 10415: 10378: 10280: 10188: 10085: 10032: 9972: 9917: 9875: 9739: 9703: 9564: 9187:
that the latter name is considered to take priority by most authors. As a result, the name
9088: 9042: 8988: 8924: 8873: 8814: 8752: 8702: 8656: 8597: 8592: 8506: 8456: 8410: 8323: 8264: 8213: 8164: 8113: 8067: 8016: 7961: 7915: 7820: 7774: 7723: 7677: 7626: 7543: 7492: 7441: 7390: 7339: 7293: 7240: 7194: 7052: 7006: 6943: 6910: 6855: 6809: 6750: 6693: 6647: 6596: 6550: 6495: 6481:
Seemingly had four ribbon-like tail feathers instead of only two as in most Enantiornithes
6448: 6411: 6406: 6360: 6310: 6260: 6214: 6168: 6118: 6072: 6011: 5962: 5913: 5854: 5801: 5748: 5689: 5638: 5581: 5484: 5438: 5388: 5338: 5281: 5235: 5185: 5139: 5093: 5047: 4988: 4937: 4886: 4833: 4787: 4736: 4684: 4634: 4588: 4542: 4496: 4450: 4397: 4346: 4295: 4249: 4198: 4144: 4091: 4045: 3990: 3939: 3888: 3830: 3784: 3731: 1501:, this definition of Euenantiornithes was severely criticized by some researchers, such as 1342:
with a short hypocleidium, and ulnar quill knobs that indicate increased aerial abilities.
1078: 323:, also known as enantiornithines or enantiornitheans in literature, are a group of extinct 12125:"A new basal beaked ornithurine bird from the Lower Cretaceous of Western Liaoning, China" 11551: 9906:"A flattened enantiornithine in mid-Cretaceous Burmese amber: morphology and preservation" 5125:
Had robust bones, including feet and claws which may have been adapted for climbing trees
4819:
Known only from a tarsometatarsus which shares some features with those of Enantiornithes
3924: 8: 15202: 15094: 15034: 14979: 14759: 12997: 11806: 11756:
O'Connor, Jingmai K.; Wang, Min; Zheng, Xiao-Ting; Wang, Xiao-Li; Zhou, Zhong-He (2014).
9817:. Vol. 27. Smithsonian Contributions to Paleobiology. pp. 67–73. Archived from 9288: 9204:" is an informally-named bird from North Korea. It may not be a member of Enantiornithes. 9074:
Possessed long wings and a reverse hallux, indicating good flight and perching abilities
8661: 8512: 8462: 8328: 8219: 8169: 8118: 8072: 7966: 7825: 7779: 7728: 7682: 7631: 7548: 7446: 7344: 7298: 7245: 7199: 7092: 7011: 6965: 6915: 6860: 6814: 6652: 6601: 6555: 6454: 6365: 6219: 6154:
Holotype was a small osteologically immature female preserved with medullary bone tissue
6077: 5967: 5833:
A large and long-legged member of the group, proportionally similar to modern shorebirds
5753: 5540: 5443: 5393: 5190: 5098: 4891: 4689: 4351: 4300: 4149: 3995: 3789: 1416: 777: 705: 701: 13517: 13474: 13363: 13308: 13272: 13182: 13139: 13049: 12847: 12768: 12733: 12646: 12483: 12436: 12381: 12247: 12190: 12041: 11922: 11857: 11734: 11697: 11668: 11625: 11348: 11146: 11020: 10825: 10712: 10509: 10419: 10382: 10284: 10192: 10089: 10036: 9976: 9921: 9879: 9743: 9707: 9568: 4528:
Only known from wing bones. May be synonymous with other Lecho formation Enantiornithes
4382: 1470:
have hitherto been very reluctant to suggest delimitations of clades of Enantiornithes.
15985: 15049: 15009: 14974: 14305: 13705: 13529: 13486: 13437: 13406: 13382: 13347: 13328: 13241: 13194: 13151: 13108: 13068: 13033: 12973: 12938: 12897: 12870: 12815: 12780: 12668: 12586: 12559:
Clarke, Julia A.; Norell, Mark A. (2002). "The Morphology and Phylogenetic Position of
12448: 12400: 12365: 12364:
Wang, Xia; McGowan, Alistair J.; Dyke, Gareth J.; Turvey, Samuel T. (7 December 2011).
12341: 12306: 12287: 12207: 12174: 12105: 12101: 12058: 12025: 11941: 11906: 11887: 11826: 11680: 11637: 11575: 11527: 11368: 11311: 11284: 11158: 11115: 11107: 11037: 11004: 10982: 10890: 10859: 10837: 10785: 10756: 10734: 10674: 10610: 10561: 10521: 10439: 10342: 10315: 10296: 10248: 10223: 10204: 10158: 10101: 10053: 10020: 9993: 9960: 9763: 9666: 9646: 9580: 9534: 9497: 7879: 6755: 6265: 6246:
Most similar to bohaiornithids but also has features of other groups of Enantiornithes
5859: 5694: 5586: 4993: 4838: 4096: 3736: 805: 801: 638: 147: 12468:"Insight into the early evolution of the avian sternum from juvenile enantiornithines" 12082: 1282:. Evidence from nesting sites shows that Enantiornithes buried their eggs like modern 780:, and dentaries (the main toothed bones of the lower jaw) without forked rear tips. A 16338: 16111: 16058: 16008: 15151: 15141: 15084: 14884: 14879: 14819: 13884: 13442: 13387: 13332: 13320: 13245: 13233: 13073: 13014: 12902: 12819: 12688:"A new genus and species of enantiornithine bird from the Early Cretaceous of Brazil" 12672: 12660: 12544: 12497: 12405: 12346: 12328: 12279: 12271: 12212: 12063: 11977: 11946: 11879: 11830: 11519: 11360: 11316: 11162: 11119: 11042: 10943: 10895: 10841: 10790: 10776: 10726: 10602: 10462:"A new genus and species of enantiornithine bird from the Early Cretaceous of Brazil" 10461: 10431: 10347: 10253: 10239: 10162: 10150: 10142: 10105: 10058: 9998: 9935: 9755: 9673: 9650: 9603: 9454: 9429: 9163: 5470:
May have been a very maneuverable flier due to the structure of its hand and sternum
4340: 4254: 1270:
Evidence of colonial nesting has been found in Enantiornithes, in sediments from the
1164: 764: 681: 666: 513: 353: 13533: 13490: 13198: 13155: 13112: 12977: 12942: 12784: 12590: 12452: 12109: 11684: 11641: 11579: 11531: 11210: 10986: 10678: 10565: 10525: 10443: 10300: 10208: 9767: 9538: 9501: 15977: 15119: 14791: 13849: 13521: 13478: 13432: 13422: 13377: 13367: 13312: 13276: 13225: 13186: 13143: 13100: 13063: 13053: 13006: 12965: 12930: 12892: 12882: 12851: 12807: 12772: 12737: 12699: 12650: 12578: 12539: 12487: 12440: 12395: 12385: 12336: 12318: 12291: 12261: 12251: 12202: 12194: 12097: 12053: 12045: 12004: 11973: 11936: 11926: 11891: 11871: 11818: 11738: 11672: 11629: 11567: 11559: 11511: 11484: 11352: 11306: 11296: 11189: 11150: 11099: 11072: 11032: 11024: 10974: 10933: 10885: 10875: 10833: 10829: 10780: 10772: 10738: 10716: 10666: 10641: 10614: 10592: 10553: 10513: 10473: 10423: 10386: 10337: 10327: 10288: 10243: 10235: 10196: 10132: 10093: 10048: 10040: 9988: 9980: 9925: 9883: 9789: 9747: 9711: 9636: 9584: 9572: 9555:
Walker, C.A. (1981). "New subclass of birds from the Cretaceous of South America".
9520:"Preliminary note on the palaeognathous bird from the Upper Cretaceous of Mongolia" 9487: 9362: 9213: 9159: 8997: 8516: 8466: 8332: 8273: 8269: 8223: 8173: 8122: 8076: 8025: 8021: 7970: 7924: 7829: 7783: 7732: 7686: 7635: 7552: 7501: 7497: 7450: 7399: 7395: 7348: 7302: 7249: 7203: 7015: 6919: 6864: 6818: 6702: 6698: 6656: 6605: 6559: 6458: 6415: 6369: 6319: 6269: 6223: 6177: 6173: 6127: 6123: 6081: 5971: 5922: 5918: 5757: 5647: 5643: 5544: 5493: 5447: 5397: 5347: 5275: 5240: 5194: 5148: 5144: 5102: 4946: 4942: 4796: 4745: 4741: 4693: 4643: 4459: 4455: 4406: 4402: 4355: 4304: 4258: 4207: 4153: 4039: 3999: 3897: 3893: 3793: 2297: 1375:, anatomically modern birds), and place Enantiornithes in the more inclusive group 1189: 936: 869:
One fossil of Enantiornithes shows wing-like feather tufts on its legs, similar to
785: 617: 570: 496: 377: 40: 13482: 13280: 13229: 13190: 13104: 13010: 12934: 12855: 12741: 12466:
Zheng, Xiaoting; Wang, Xiaoli; O'Connor, Jingmai; Zhou, Zhonghe (9 October 2012).
11488: 11372: 10978: 10557: 10314:
Atterholt, Jessie; Hutchison, J. Howard; O'Connor, Jingmai K. (13 November 2018).
10097: 9793: 9474:
O'Connor, Jingmai K.; Chiappe, Luis M.; Gao, Chunling; Zhao, Bo (September 2011).
8734:
Was once believed to have fed on tree sap as it was preserved in association with
8246:
The first known member of the family, although not considered a close relative of
8001: 7755:
Proposed to have a woodpecker-like lifestyle due to features of the foot and tail
6725:
Some of this genus's remains include well-preserved hindlimbs. the species names,
1513: 1363:
Some researchers classify Enantiornithes, along with the true birds, in the class
1045:
were likely used to feed on soft-bodied arthropods. The strongly hooked talons of
732: 16209: 16037: 15507: 15029: 15014: 14952: 14851: 14779: 14457: 14403: 14396: 14380: 14187: 14088: 13842: 13663: 13647: 13372: 13147: 13058: 12969: 12776: 12704: 12687: 12615: 12390: 11931: 11154: 11063:
Sanz, JosΓ© L.; Chiappe, Luis M.; Buscalioni, Angela D. (1995). "The Osteology of
10517: 10478: 10427: 10292: 10200: 9097: 9093: 9051: 8933: 8882: 8823: 8761: 8711: 8665: 8601: 8419: 7870: 7287: 7234: 7158: 7116: 7112: 7085: 7061: 7057: 6969: 6759: 6504: 6442: 6058:
Although known only from humeri, this genus was large and lived in a broad range
6024: 6020: 5863: 5810: 5806: 5742: 5698: 5590: 5290: 5244: 5056: 4997: 4895: 4842: 4597: 4593: 4551: 4505: 4501: 4100: 4051: 3984: 3948: 3862:
One of the first Enantiornithes known. Once thought to be an ancient relative of
3839: 3740: 3271: 2205: 1741: 1616: 1527: 1447: 1409: 1401: 1352: 1271: 1110: 1046: 962: 898: 844: 815: 773: 744: 594: 487: 370: 327:("birds" in the broad sense), the most abundant and diverse group known from the 301: 291: 249: 133: 44: 8052: 6392:
Many soft tissue details of specimens from this genus were revealed by UV light
467: 373:
in 1981. Walker made this discovery based on some partial remains from the late
16319: 16249: 16201: 16135: 16074: 16029: 16016: 15890: 15853: 15704: 15478: 15315: 15069: 15064: 15059: 15039: 14919: 14894: 14889: 14871: 14752: 14608: 14533: 14501: 14446: 14425: 14332: 14280: 14124: 13877: 12608: 12230:
Brusatte, Stephen L.; O'Connor, Jingmai K.; Jarvis, Erich D. (5 October 2015).
12049: 11742: 11301: 10044: 9930: 9905: 9641: 9624: 8993: 8909: 8696: 8539:
A small and short-snouted genus of Enantiornithes which may be synonymous with
8158: 7589: 7537: 7384: 7000: 5956: 4792: 4289: 3691: 3669: 3639: 3609: 3225: 2942: 2249: 2104: 1595: 1420: 1387: 1330: 1323: 1264: 1229: 1116: 1068: 990: 973: 931: 893: 781: 687: 675: 336: 306: 296: 138: 13525: 13346:
Hu, Han; O'Connor, Jingmai K.; Zhou, Zhonghe; Farke, Andrew A. (3 June 2015).
13316: 12444: 12256: 12231: 11875: 11676: 11633: 10938: 10921: 10670: 10646: 10629: 10137: 10120: 7662: 6942:
The hand of this genus was similar to that of ornithuromorphs, likely through
1599: 16455: 16396: 16257: 16241: 16164: 16156: 15990: 15788: 15638: 15525: 15483: 15283: 15275: 15197: 15161: 15079: 15003: 14831: 14686: 14355: 14314: 14287: 14109: 13947: 13940: 13891: 13237: 13018: 12664: 12332: 12275: 12009: 11992: 10435: 10146: 9887: 9387:
Wang, Min; Zhou, Zhong-He; O'Connor, Jingmai K.; Zelenkov, Nikita V. (2014).
9284: 9101: 9082: 8951: 8841: 8779: 8619: 8605: 7717: 7120: 7065: 7046: 6973: 6028: 5907: 5814: 5478: 5248: 5052: 5015: 4899: 4730: 4678: 4601: 4547: 4509: 4391: 3679: 3312: 3024: 2680: 2598: 2497: 2219: 1919: 1795: 1649: 1485: 1381: 1275: 1177: 976:, had a fan of tail feathers similar to that of more primitive avialans like 855: 769: 754: 658: 623: 536: 527:
Birds with confidently identified characteristics of Enantiornithes found in
474: 412: 392: 82: 12811: 11709:
The origin of the bird's beak: new insights from dinosaur incubation periods
11515: 11133:
O'Connor, Jingmai K. (1 January 2019). "The trophic habits of early birds".
9751: 1896: 16343: 16087: 15932: 15748: 15586: 15534: 15406: 15367: 15136: 15124: 15114: 15104: 14801: 14679: 14494: 14073: 14045: 14038: 13982: 13912: 13856: 13446: 13391: 13324: 13077: 12906: 12501: 12409: 12350: 12283: 12216: 12067: 11950: 11883: 11523: 11364: 11320: 11046: 11003:
Wang, Min; O'Connor, Jingmai K.; Pan, Yanhong; Zhou, Zhonghe (2017-01-31).
10947: 10899: 10794: 10730: 10606: 10351: 10257: 10154: 10062: 10002: 9939: 9784:
Rautian, 1978 a fossil feather from the early Late Jurassic of Kazakhstan.
9759: 9316: 9218: 9036: 8301: 8107: 6734: 6730: 5683: 5432: 5041: 4536: 4085: 4075: 3208: 2854: 2416: 1873: 1451: 1426: 1205: 1199: 517: 328: 10121:"A New Enantiornithine Bird with Unusual Pedal Proportions Found in Amber" 9492: 9475: 7856: 5945:
Originally believed to be an ornithuran, but now considered a relative of
4922: 4234: 1620: 16405: 16148: 15995: 15801: 15780: 15769: 15734: 15601: 15591: 15549: 15440: 15307: 15109: 14934: 14929: 14861: 14846: 14814: 14633: 14592: 14569: 14508: 14439: 14418: 14369: 14341: 14321: 14224: 14173: 14116: 14010: 14003: 13975: 13961: 13954: 13926: 13905: 13863: 13717: 9349:
Wang, X.; O'Connor, J. K.; Zheng, X.; Wang, M.; Hu, H.; Zhou, Z. (2014).
9292: 9251: 9193: 8555: 8207: 7994: 7814: 7759: 7690: 7620: 7528: 7486: 7333: 6787: 6609: 6590: 6373: 6354: 6343: 6304: 6293: 5896: 5179: 5133: 4982: 4880: 4827: 4718: 4582: 4490: 4308: 4138: 3933: 3867: 3629: 3599: 3384: 3085: 2983: 2642: 2351: 2176: 2128: 1967: 1825: 1578: 1573: 1502: 1456: 1368: 1319: 1059: 1035: 1013: 968: 944: 875: 790: 646: 605: 589: 579: 398: 386: 286: 57: 12655: 12630: 11028: 9984: 7143: 5733: 5323: 5079:
Had an enlarged outer toe that may have been an adaptation for perching
16265: 16127: 16066: 16045: 15877: 15861: 15796: 15741: 15713: 15645: 15610: 15488: 15336: 15171: 15146: 14904: 14622: 14615: 14542: 14473: 14466: 14266: 14131: 14102: 14059: 13968: 13933: 13898: 13629: 13620: 13427: 12887: 12492: 12467: 12323: 12266: 11822: 11194: 11177: 11111: 10391: 10366: 10332: 9296: 9257: 9170: 8982: 8808: 8715: 8404: 7955: 7909: 6904: 6781: 6005: 5885: 5848: 5725:
One of the many fragmentary Bissekty Enantiornithes, known only from a
5720: 5612: 5575: 5060: 5001: 4931: 4913: 4864: 4628: 4555: 4444: 4122: 3873: 3863: 3762: 3360: 3285: 2884: 2820: 2547: 2530: 2480: 2375: 1914: 1891: 1699: 1540: 1520:
in 2015, updated from a previous data set created by Jingmai O'Connor.
1439: 1405: 1372: 1255: 1214: 1099: 1093: 1063: 1054: 1019: 984: 949: 917: 913: 759: 738: 657:). Extraordinary remains of Enantiornithes have also been preserved in 562: 558: 521: 374: 358: 210: 102: 67: 14744: 12305:
Atterholt, Jessie; Hutchison, J. Howard; O'Connor, Jingmai K. (2018).
12198: 11176:
O'Connor, Jingmai K.; Zhou, Zhonghe; Smith, Andrew (6 November 2019).
10880: 9367: 9350: 5520: 4973: 4670:, a specialized type of feather which controls air flow over the wing 1286:, which is consistent with their inferred superprecocial adaptations. 839:
Enantiornithes were advanced compared to more primitive avialans like
16225: 16180: 15969: 15762: 15679: 15663: 15652: 15562: 15498: 15457: 15156: 14957: 14944: 14924: 14824: 14665: 14651: 14600: 14557: 14522: 14432: 14410: 14389: 14362: 14273: 14216: 14166: 14159: 14145: 14080: 14052: 14017: 13996: 13870: 13828: 13814: 13690: 13638: 11571: 11356: 11076: 10597: 10580: 9576: 9476:"Anatomy of the Early Cretaceous Enantiornithine Bird Rapaxavis pani" 9324: 9312: 9240: 9231:: A juvenile of an indeterminate specimen of Enantiornithes from the 9115: 9069: 9055: 9001: 8937: 8900: 8886: 8867: 8827: 8765: 8669: 8586: 8500: 8423: 8317: 8149: 8029: 7875: 7768: 7435: 7188: 7079: 6987: 6954: 6767: 6512: 6323: 6287: 6254: 6208: 6131: 6112: 6053: 5891: 5871: 5828: 5706: 5598: 5529: 5497: 5489: 5371: 5294: 5229: 5152: 4850: 4800: 4781: 4647: 4639: 4615: 4523: 4243: 4211: 4203: 4192: 4108: 4055: 3952: 3843: 3824: 3748: 3725: 3589: 3258: 3133: 2956: 2904: 2713: 2663: 2399: 2266: 1962: 1855: 1463: 1393: 1334: 1283: 1260: 1249: 1240: 1225: 1220: 1185: 1158: 1135: 1073: 1041: 1025: 978: 921: 902: 796: 719: 634: 599: 549: 540: 532: 508: 381: 223: 164: 107: 51: 16367: 12519: 12172: 11593:
ElΕΌanowski, Andrzej (1995). "Cretaceous birds and avian phylogeny".
10864:: insights into the aerodynamics and diet of a basal ornithuromorph" 10721: 10696: 9715: 9323:
from various smaller bones, and the fusion of tail vertebrae into a
9142:: Described in 2013 from embryo specimens within eggshells from the 1477:, was defined by Chiappe (2002) as comprising all species closer to 873:. The leg feathers are also reminiscent of the four-winged dinosaur 16390: 16193: 15923: 15898: 15844: 15755: 15727: 15617: 15581: 15573: 15555: 15450: 15435: 15419: 15166: 15129: 14962: 14549: 14515: 14259: 14238: 14194: 14095: 14066: 14031: 13919: 13614: 12023: 9844:(in Russian). Vol. 197. USSR Academy of scienses. p. 82. 9212:: Described in 1999 from a specimen of Enantiornithes found in the 9201: 9147: 8683: 8650: 8450: 8437: 8258: 6849: 6786:
One of many fragmentary Bissekty Enantiornithes, known only from a
6763: 6687: 6508: 6489: 5890:
One of many fragmentary Bissekty Enantiornithes, known only from a
5867: 5795: 5702: 5632: 5618: 5594: 5382: 5365: 5315: 5308: 5262: 4870: 4846: 4128: 4127:
One of many fragmentary Bissekty Enantiornithes, known only from a
4104: 3768: 3744: 3619: 3401: 3109: 2629: 2324: 2159: 1944: 1674: 1479: 1347: 1311: 1245: 1147: 1143: 1139: 862: 859: 713: 670: 650: 566: 554: 455: 447: 404: 340: 197: 184: 97: 92: 77: 72: 62: 10630:"Downsized Dinosaurs: The Evolutionary Transition to Modern Birds" 9780:
Agnolin, F.L., Rozadilla, S., and Ismar de Souza Carvalho (2017).
9028:, although most closely resembles avisaurids among sampled groups 6887:
One of the first Jehol biota Enantiornithes described. Similar to
4574:
Known from a foot preserved in amber with an elongated middle toe
16425: 16103: 15956: 15835: 15720: 15686: 15624: 15445: 15299: 15291: 14984: 14856: 14841: 14672: 14658: 14294: 14245: 13835: 13559: 11211:"Bone digestion and intestinal morphology of the Bearded Vulture" 9320: 9025: 8973: 8918: 8853: 8791: 8746: 8729: 8631: 8359: 8308: 7101: 6794: 6744: 5726: 5617:
One of many fragmentary Bissekty Enantiornithes, known only from
5074: 4869:
One of many fragmentary Bissekty Enantiornithes, known only from
4569: 4331: 3966: 3920:
One specimen from this genus died with an unlaid egg in its body
3882: 3767:
One of many fragmentary Bissekty Enantiornithes, known only from
2757: 2737: 1745: 1376: 1339: 1279: 1188:. An embryo, still curled in its egg, has been reported from the 1088: 709: 692: 443: 366: 362: 324: 236: 124: 112: 87: 11719: 11473: 9961:"Mummified precocial bird wings in mid-Cretaceous Burmese amber" 4378:
Originally mistakenly believed to have possessed a pointed beak
16233: 16172: 15948: 15940: 15869: 15809: 15693: 15631: 15514: 15181: 15176: 15054: 13608: 13549: 13293: 11450:"Embryonic bird skeletons from the late Cretaceous of Mongolia" 11333: 10118: 9903: 9864: 9131: 9019: 9005: 8520: 8470: 8336: 8277: 8227: 8177: 8126: 8080: 7974: 7928: 7833: 7787: 7736: 7639: 7556: 7505: 7454: 7403: 7352: 7306: 7253: 7226:
Originally mistakenly believed to have possessed a hooked beak
7207: 7019: 6923: 6868: 6822: 6706: 6660: 6563: 6462: 6273: 6227: 6181: 6135: 6085: 6042: 5975: 5926: 5761: 5651: 5548: 5451: 5401: 5351: 5198: 5106: 4950: 4814: 4749: 4697: 4463: 4434: 4410: 4359: 4276: 4262: 4157: 4069: 4003: 3971:
Fragmentary but unique in the structure of its tarsometatarsus
3901: 3857: 3797: 3659: 1509:
a good example of a poor choice in a phylogenetic definition".
1181: 957: 808:, which in modern birds is fused to the jugal, is preserved in 544: 528: 454:
This refers to an anatomical feature – the articulation of the
174: 13404: 13211: 12304: 11904: 10755:
Clarke, Julia A.; Zhou, Zhonghe; Zhang, Fucheng (March 2006).
10313: 3975: 16119: 15906: 15469: 11804: 11233:"Short Crystal: Quartz and the Fossilized Bird | GeoRarities" 10857: 9291:. A 2016 analysis of its feathering found elongated putative 9236: 9183:, the description was so poor compared to the description of 8735: 8534: 8484: 8355:
A large member of the family with a well-preserved braincase
8350: 8291: 8241: 8191: 8140: 8094: 8043: 7988: 7942: 7847: 7801: 7750: 7704: 7653: 7570: 7519: 7468: 7417: 7366: 7320: 7267: 7221: 7134: 7033: 6937: 6882: 6836: 6720: 6674: 6623: 6577: 6526: 6476: 6429: 6387: 6337: 6241: 6195: 6149: 6099: 5989: 5940: 5775: 5665: 5562: 5511: 5465: 5415: 5212: 5166: 5120: 5027: 4964: 4763: 4711: 4667: 4661: 4477: 4424: 4373: 4322: 4225: 4177: 4171: 4023: 4017: 3915: 3811: 1506: 1443: 848: 642: 584: 574: 429: 419: 13503: 11654: 11611: 11402:"Classification of fossil eggshells of amniotic vertebrates" 9451:
Glorified Dinosaurs: The Origin and Early Evolution of Birds
9386: 8250:
until a few years later. Preserves a large postorbital bone
7997:, although later these were found to be mineral concretions 7477: 7038:
One of the largest Enantiornithes known from decent remains
6895: 1544: 887: 784:
is preserved in an indeterminate juvenile specimen, while a
458:
bones – which has a concave-convex socket joint between the
16095: 15462: 15425: 15395: 14809: 14775: 12583:
10.1206/0003-0082(2002)387<0001:TMAPPO>2.0.CO;2
12123:
Zhou, Shuang; Zhou, Zhong-He; O'Connor, Jingmai K. (2012).
11104:
10.1671/0272-4634(2002)022[0856:FASFTM]2.0.CO;2
10018: 8963: 6535: 5674: 1364: 1114:
most likely to be generalistic insectivores (sans possibly
1011:
Life reconstruction of enantiornithine birds feeding (left
768:, were fully toothless and had beaks. They also had simple 12990: 12686:
Carvalho; Novas; AgnolΓ­n; Isasi; Freitas; Andrade (2015).
12229: 11991:
Clark, Alexander D.; O'Connor, Jingmai K. (15 June 2021).
10810: 10364: 9473: 7473:
Specialized for perching due to the structure of its feet
6628:
One of the oldest and most primitive members of the group
5424:, although a 2015 study considered it to be a valid genus 4768:
Well-preserved but inconsistent in phylogenetic placement
818:(the ability to move the jaw independent of the cranium). 569:
relatives. The earliest known Enantiornithes are from the
16217: 15074: 14836: 13287: 12832: 12685: 12513: 12511: 12465: 11963: 11755: 11564:
10.1206/0003-0082(2007)3594[1:JBFTEC]2.0.CO;2
11475:
Spain: Implications for Avian Skull and Neck Evolution".
11083: 10963: 10697:"Palaeontology: Leg feathers in an Early Cretaceous bird" 10459: 10222:
Zhou, Zhonghe; Clarke, Julia; Zhang, Fucheng (May 2008).
9834: 9348: 6346:, a skin flap on the shoulder which forms part of a wing 5567:
The earliest known enantiornithine with a toothless beak
854:
Several wings with preserved feathers have been found in
662:
suggesting that they were an ecologically diverse group.
13459: 12868: 10806: 10804: 10491: 10489: 10075: 9415: 9413: 9411: 9409: 9295:, suggesting that a large portion of its feathering was 994:, had a modern-looking pygostyle but lacked a tail fan. 960:-dominated tail feathers similar to feathers present in 13345: 12422: 11545: 11543: 11541: 11002: 10919: 10178: 9728: 4722:, although now considered to be more distantly related 411:
The name "Enantiornithes" means "opposite birds", from
13031: 12508: 11549: 11282: 9024:
Possesses a myriad of features from various groups in
7658:
Possessed a fan-shaped tail composed of many feathers
758:, they retained several separate cranial bones, small 724:
are comparable in size to small tits or hummingbirds.
450:, where the 'normal' condition is completely reversed. 12552: 12363: 11843: 11175: 11062: 10801: 10486: 9406: 7878:
of this group is doubtful, and it may actually be an
561:. Fossils attributable to this group are exclusively 12949: 12754: 12122: 11758:"The histology of two female Early Cretaceous birds" 11538: 11089: 7371:
The most common and well-known member of the family
7325:
May have had a thorn-like structure on its forehead
6841:
Unusually, the premaxillae of this genus were fused
15366: 12718: 12081:Padian, Kevin; Chiappe, Luis M. (11 January 2007). 10495: 10455: 10453: 9957: 9602:. University of California Press. pp. 339–88. 9428:. University of California Press. pp. 240–67. 9424:. In Chiappe, Luis M.; Witmer, Lawrence M. (eds.). 1212:Together with hatchling specimens of the Mongolian 1146:avialans were prey animals, and that some Mesozoic 12797: 12619:. Version 1.0, 2005-NOV- 7. Retrieved 2006-OCT-02. 11713: 10654: 9692: 9665: 9158:(a set of controversial paleontologists including 7993:Originally considered to have been preserved with 4230:One of the most complete Las Hoyas Enantiornithes 12919: 12628: 11990: 11135:Palaeogeography, Palaeoclimatology, Palaeoecology 10750: 10748: 9382: 9380: 9378: 9150:. These specimens were very similar to embryonic 8969:The most complete known North American avisaurid 16453: 12517: 11550:Chiappe, Luis M.; Shu'an, Ji; Qiang, Ji (2007). 10450: 4716:Once considered to be a basal close relative of 12602: 12600: 11058: 11056: 10853: 10851: 10754: 10270: 10221: 9804: 5621:. One species was once considered to belong to 13662: 13339: 13252: 13090: 11605: 10915: 10913: 10911: 10909: 10745: 10695:Zhang, Fucheng; Zhou, Zhonghe (October 2004). 10539: 10537: 10535: 10215: 10174: 10172: 9953: 9951: 9949: 9860: 9858: 9774: 9686: 9375: 5420:Once considered to be dubious or a species of 14760: 13798: 13575: 12080: 11208: 10959: 10957: 10860:"A new specimen of the Early Cretaceous bird 10690: 10688: 9419: 9344: 9342: 9340: 5894:fragment. Originally considered to belong to 15274: 12748: 12597: 12558: 11327: 11132: 11053: 10848: 9600:Mesozoic Birds: Above the Heads of Dinosaurs 9550: 9548: 9453:. Hoboken, New Jersey: John Wiley and Sons. 9426:Mesozoic Birds: Above the Heads of Dinosaurs 9132:Dubious genera and notable unnamed specimens 8561: 8379: 7884: 7595: 7276:, although that genus name is occupied by a 7163: 5836: 3700: 1097:being random mineral precipitates, only the 13453: 11385:: CS1 maint: numeric names: authors list ( 10906: 10621: 10532: 10169: 9946: 9855: 9722: 9420:Chiappe, Luis M.; Walker, Cyril A. (2002). 1142:dinosaur. This was the first evidence that 14767: 14753: 13582: 13568: 13497: 13125: 12712: 12116: 11592: 11444: 10954: 10685: 10405: 9514: 9337: 8859:Would have lived in a coastal environment 4381: 4029: 123: 13436: 13426: 13381: 13371: 13258: 13205: 13067: 13057: 12984: 12896: 12886: 12703: 12679: 12654: 12572: 12543: 12491: 12399: 12389: 12340: 12322: 12265: 12255: 12232:"The Origin and Diversification of Birds" 12206: 12083:"The origin and early evolution of birds" 12057: 12008: 11940: 11930: 11865: 11501: 11425: 11399: 11310: 11300: 11209:Houston, David C.; Copsey, J. A. (1994). 11193: 11036: 10937: 10889: 10879: 10784: 10720: 10694: 10645: 10596: 10477: 10390: 10341: 10331: 10247: 10136: 10052: 9992: 9929: 9640: 9545: 9491: 9366: 9355:Biological Journal of the Linnean Society 9311:: A partial skeleton of a hatchling from 4429:Was well-adapted for climbing due to its 1497:itself. Despite being in accordance with 1379:. Enantiornithes were more advanced than 1150:regurgitated pellets like owls do today. 13168: 12913: 12826: 12518:O'Connor, Jingmai; Dyke, Gareth (2010). 10543: 9663: 8304:on its teeth, unique among fossil birds 1371:definition of Aves (which only includes 1176:Known fossils of Enantiornithes include 1157: 1006: 886: 825: 731: 616: 14774: 12955: 12563:from the Late Cretaceous of Mongolia". 11781: 10627: 9448: 1516:below was found by an analysis by Wang 346: 14: 16454: 11788:Courier Forschungsinstitut Senckenberg 11595:Courier Forschungsinstitut Senckenberg 10660: 10578: 9554: 9154:specimens, although the describers of 8145:A fairly typical member of the family 6891:but usually considered to be distinct 5221:with characteristically slender bones 1438:Enantiornithes is the sister group to 1404:. The other ornithothoracine group is 1066:preserved in its digestive tract, and 952:, though a later study indicates that 704:(and possibly also represented in the 16372: 16371: 15259: 14748: 13601: 13563: 10998: 10996: 10014: 10012: 9899: 9897: 7947:Had a short but robust manual ungual 1322:as a whole - some extinct birds like 1129:A fossil from Spain reported by Sanz 16326: 9672:. New Haven: Yale University Press. 9622: 9597: 384:, which he assigned to a new genus, 16350: 13093:Journal of Systematic Palaeontology 12958:Journal of Systematic Palaeontology 12923:Journal of Systematic Palaeontology 12425:Journal of Systematic Palaeontology 10579:Wang, Min; Hu, Han (January 2017). 10546:Journal of Systematic Palaeontology 10498:Journal of Systematic Palaeontology 9851:from the original on 27 March 2019. 9239:. Its 2006 description studied the 1351:appears to have become secondarily 502: 434:) "birds" . The name was coined by 24: 16472:Early Cretaceous first appearances 13589: 13463:Journal of Vertebrate Paleontology 13171:Journal of Vertebrate Paleontology 13128:Journal of Vertebrate Paleontology 12757:Journal of Vertebrate Paleontology 12102:10.1111/j.1469-185x.1997.tb00024.x 11997:Frontiers in Ecology and Evolution 11092:Journal of Vertebrate Paleontology 10993: 10967:Journal of Vertebrate Paleontology 10814:Journal of Vertebrate Paleontology 10408:Journal of Vertebrate Paleontology 10273:Journal of Vertebrate Paleontology 10181:Journal of Vertebrate Paleontology 10009: 9894: 9835:L. A. Nesov, A. A. Yarkov (1989). 9696:Canadian Journal of Earth Sciences 7865: 7152: 1367:. Others use the more restrictive 1103:displays actual stomach contents. 25: 16488: 15228:Late Quaternary prehistoric birds 13543: 11400:Mikhailov, Konstantin E. (1991). 10634:Evolution: Education and Outreach 9668:The Origin and Evolution of Birds 8368: 7583: 3690: 3685: 2081:The cladogram below is from Wang 1358: 1263:, instead having an intermediate 1058:was found to have the remains of 772:, a complete bar separating each 736:A reconstruction of the skull of 628:a species of early Enantiornithes 597:of North and South America (e.g. 16349: 16337: 16325: 16314: 16313: 16204:(New World vultures and condors) 14729: 14728: 14710: 14704: 13548: 13398: 12545:10.3853/j.0067-1975.62.2010.1540 12532:Records of the Australian Museum 11978:10.19615/j.cnki.1000-3118.190823 10777:10.1111/j.1469-7580.2006.00534.x 10240:10.1111/j.1469-7580.2008.00880.x 9108: 9062: 9012: 8972: 8956: 8944: 8908: 8893: 8846: 8834: 8784: 8772: 8722: 8676: 8640: 8624: 8612: 8527: 8477: 8430: 8358: 8343: 8307: 8284: 8234: 8184: 8148: 8133: 8087: 8051: 8036: 8000: 7981: 7935: 7855: 7840: 7794: 7758: 7743: 7697: 7661: 7646: 7563: 7527: 7512: 7476: 7461: 7425: 7410: 7374: 7359: 7313: 7260: 7214: 7142: 7127: 7091: 7072: 7026: 6980: 6930: 6894: 6875: 6829: 6793: 6774: 6713: 6667: 6631: 6616: 6570: 6534: 6519: 6469: 6422: 6380: 6330: 6280: 6234: 6188: 6142: 6092: 6046: 6035: 5982: 5933: 5878: 5821: 5768: 5732: 5713: 5673: 5658: 5605: 5555: 5519: 5504: 5458: 5408: 5358: 5322: 5301: 5255: 5205: 5159: 5113: 5067: 5020: 5008: 4972: 4957: 4921: 4906: 4857: 4807: 4771: 4756: 4704: 4666:Preserves feathers including an 4654: 4608: 4562: 4516: 4470: 4417: 4366: 4330: 4315: 4269: 4233: 4218: 4182: 4164: 4115: 4062: 4010: 3974: 3959: 3923: 3908: 3872: 3850: 3804: 3755: 1966: 1918: 1895: 1829: 1799: 1744: 1619: 1598: 1577: 1543: 1433: 1091:and the supposed gastroliths of 151: 55: 13162: 13119: 13084: 13025: 12862: 12791: 12622: 12459: 12416: 12357: 12298: 12223: 12166: 12156: 12074: 12017: 11984: 11957: 11898: 11837: 11798: 11775: 11749: 11702: 11691: 11648: 11586: 11495: 11467: 11438: 11419: 11393: 11276: 11266: 11256: 11246: 11225: 11202: 11169: 11126: 10572: 10399: 10358: 10307: 10264: 10112: 10069: 9828: 9798: 9657: 7084:Had a large and unusually wide 1505:, who called it "a ill-defined 1473:One such delineation named the 1153: 916:(the clade they referred to as 858:. These are the first complete 15260: 11215:The Journal of Raptor Research 10834:10.1080/02724634.2009.10010371 9616: 9591: 9508: 9467: 9442: 8550: 6946:. A large member of the group 612: 357:, was originally considered a 13: 1: 13483:10.1080/02724634.2015.1054496 13281:10.1016/j.cretres.2015.02.013 13230:10.1016/j.cretres.2022.105452 13191:10.1080/02724634.2017.1278702 13105:10.1080/14772019.2016.1246111 13011:10.1016/j.cretres.2024.105867 12935:10.1080/14772019.2015.1051146 12856:10.1016/j.cretres.2009.09.001 12742:10.1016/j.cretres.2015.01.008 12609:TaxonSearch: Stem Archosauria 11489:10.1126/science.276.5318.1543 11409:Acta Palaeontologica Polonica 10979:10.1080/02724634.2015.1054035 10558:10.1080/14772019.2015.1073801 10098:10.1016/j.cretres.2020.104412 9794:10.1080/08912963.2017.1413102 9480:Acta Palaeontologica Polonica 9331: 7575:A large member of the family 333:Cretaceous–Paleogene boundary 13373:10.1371/journal.pone.0126791 13148:10.1080/02724634.2013.794814 13059:10.1371/journal.pone.0184637 12970:10.1080/14772019.2014.994087 12777:10.1080/02724634.2013.812101 12705:10.1590/23174889201500020001 12692:Brazilian Journal of Geology 12391:10.1371/journal.pone.0028672 11932:10.1371/journal.pone.0061030 11155:10.1016/j.palaeo.2018.03.006 10518:10.1080/14772019.2010.526639 10479:10.1590/23174889201500020001 10466:Brazilian Journal of Geology 10428:10.1080/02724634.2013.794814 10293:10.1080/02724634.2012.652321 10201:10.1080/02724634.2013.762708 9283:: An indeterminate juvenile 4074:May have been the size of a 1124: 408:) were also Enantiornithes. 7: 16268:(woodpeckers and relatives) 6342:Preserves impressions of a 5994:May have been a synonym of 1442:, and together they form a 485:Feduccia's point about the 10: 16493: 16130:(cormorants and relatives) 15223:Extinct species since 1500 12050:10.1038/s41598-019-41423-7 11743:10.1016/j.crpv.2005.12.018 11302:10.1186/s12915-022-01294-3 10045:10.1038/s41598-018-37427-4 9931:10.1016/j.scib.2018.01.019 9642:10.1007/s00435-021-00528-2 9048:Bajo de la Carpa Formation 8820:Mooreville Chalk Formation 997: 593:) and the latest from the 430: 420: 27:Extinct clade of dinosaurs 16477:Maastrichtian extinctions 16380: 16309: 16260:(kingfishers and rollers) 16192: 16147: 16114:(albatrosses and petrels) 16086: 16057: 16028: 16007: 15998:(swifts and hummingbirds) 15980:(nightjars and relatives) 15968: 15931: 15922: 15889: 15852: 15843: 15834: 15830: 15804:(pheasants and relatives) 15779: 15703: 15662: 15600: 15572: 15548: 15524: 15497: 15418: 15405: 15362: 15335: 15331: 15270: 15266: 15255: 15251: 15190: 15093: 14993: 14943: 14870: 14800: 14787: 14724: 14702: 14632: 14587: 14568: 14532: 14456: 14379: 14331: 14304: 13809: 13794: 13775: 13743: 13725: 13716: 13698: 13689: 13671: 13658: 13597: 13526:10.1134/s0031030113110087 13317:10.1038/s41586-020-2945-x 12565:American Museum Novitates 12445:10.1017/S1477201906001969 12257:10.1016/j.cub.2015.08.003 11876:10.1007/s00114-012-0917-1 11784:Patagopteryx deferrariisi 11677:10.1134/S0031030113110087 11634:10.1134/S0031030113110087 11556:American Museum Novitates 11069:American Museum Novitates 10939:10.1016/j.cub.2015.11.036 10862:Hongshanornis longicresta 10671:10.1038/nature.2016.20162 10647:10.1007/s12052-009-0133-4 10628:Chiappe, Luis M. (2009). 10138:10.1016/j.cub.2019.05.077 9810:[File size 70 MB] 9449:Chiappe, Luis M. (2007). 9120:A very close relative of 8879:Las Curtiembres Formation 8708:Ouadi al Gabour Formation 6017:GrΓ¨s Γ  Reptiles Formation 3398: 3381: 3374: 3357: 3350: 3333: 3326: 3309: 3302: 3282: 3275: 3255: 3248: 3222: 3205: 3198: 3191: 3147: 3130: 3123: 3106: 3099: 3082: 3075: 3068: 3021: 3004: 2997: 2980: 2973: 2953: 2946: 2936: 2901: 2881: 2871: 2851: 2841: 2834: 2817: 2810: 2754: 2734: 2727: 2710: 2703: 2677: 2660: 2653: 2646: 2626: 2619: 2612: 2595: 2588: 2544: 2527: 2520: 2494: 2477: 2470: 2460: 2413: 2396: 2389: 2372: 2365: 2348: 2341: 2321: 2311: 2301: 2263: 2246: 2236: 2216: 2209: 2199: 2173: 2156: 2149: 2142: 2125: 2118: 2101: 2094: 1958: 1940: 1933: 1910: 1887: 1869: 1851: 1844: 1821: 1814: 1791: 1784: 1766: 1759: 1738: 1731: 1713: 1695: 1688: 1670: 1663: 1645: 1638: 1613: 1592: 1569: 1562: 1538: 1531: 1499:phylogenetic nomenclature 1450:(though see above). Most 1300: 283: 278: 148:Scientific classification 146: 131: 122: 34: 16159:(seriemas and relatives) 16138:(pelicans and relatives) 14579:Cau & Arduini, 2008) 12010:10.3389/fevo.2021.654156 9888:10.1016/j.gr.2017.06.001 9805:Pierce Brodkorb (1976). 3836:La Bocana Roja Formation 1468:Phylogenetic taxonomists 1329:Enantiornithes resemble 956:was more likely to have 727: 339:and all other non-avian 16252:(hornbills and hoopoes) 16167:(falcons and relatives) 13506:Paleontological Journal 12812:10.1111/1755-6724.12270 12635:Journal of Paleontology 12528:(Aves: Enantiornithes)" 11846:Die Naturwissenschaften 11657:Paleontological Journal 11614:Paleontological Journal 11516:10.1126/science.1100000 11457:Palaeontologia Polonica 11428:Paleontological Journal 9752:10.1126/science.1126377 9664:Feduccia, Alan (1996). 9527:Palaeontologia Polonica 9166:) consider it distinct. 5217:A possible relative of 1002: 882: 821: 16236:(trogons and quetzals) 16048:(cranes and relatives) 15318:(emus and cassowaries) 15208:Glossary of bird terms 15020:Confuciusornithiformes 12132:Vertebrata PalAsiatica 11966:Vertebrata PalAsiatica 11765:Vertebrata PalAsiatica 11723:Comptes Rendus Palevol 9396:Vertebrata PalAsiatica 8758:Two Medicine Formation 8511:Qiaotou member of the 8461:Qiaotou member of the 8218:Qiaotou member of the 6453:Qiaotou member of the 5841: 4022:Possessed an enlarged 1173: 1030: 905: 835: 748: 715:Parvavis chuxiongensis 629: 621:A life restoration of 539:of South America, and 483: 452: 436:Cyril Alexander Walker 16434:Paleobiology Database 16077:(kagu and sunbittern) 16040:(gulls and relatives) 15045:Songlingornithiformes 15010:Omnivoropterygiformes 13557:at Wikimedia Commons 12800:Acta Geologica Sinica 12472:Nature Communications 11009:Nature Communications 10585:The Anatomical Record 9965:Nature Communications 9493:10.4202/app.2010.0047 9209:Liaoxiornis delicatus 9144:Barun Goyot Formation 8930:Kaiparowits Formation 8416:Barun Goyot Formation 5840: 5344:Andaikhudag Formation 5287:Barun Goyot Formation 1195:Liaoxiornis delicatus 1161: 1010: 890: 829: 776:(eye hole) from each 735: 620: 479: 462:(shoulder blade) and 440: 132:Fossil specimen of a 15899:Phoenicopteriformes 13005:(in press): 105867. 12606:Sereno, P.C. (2005) 12150:. February 13, 2012. 10131:(14): 2396–2401.e2. 8598:Hell Creek Formation 8296:A close relative of 8196:A close relative of 6944:convergent evolution 6412:Yijinholuo Formation 5780:Possibly related to 347:Discovery and naming 15870:Mesitornithiformes 15864:(doves and pigeons) 15198:Families and orders 15060:Ichthyornithiformes 15035:Patagopterygiformes 14202:Piscivorenantiornis 13518:2013PalJ...47.1252K 13475:2016JVPal..36E4496W 13364:2015PLoSO..1026791H 13309:2020Natur.588..272O 13273:2015CrRes..55..210H 13261:Cretaceous Research 13218:Cretaceous Research 13183:2017JVPal..37E8702W 13140:2014JVPal..34..135W 13050:2017PLoSO..1284637L 12998:Cretaceous Research 12848:2010CrRes..31...16B 12836:Cretaceous Research 12769:2014JVPal..34..657W 12734:2015CrRes..55...56W 12722:Cretaceous Research 12656:10.1017/jpa.2022.12 12647:2022JPal...96..961W 12526:Cathayornis yandica 12520:"A Reassessment of 12484:2012NatCo...3.1116Z 12437:2007JSPal...5..193C 12382:2011PLoSO...628672W 12248:2015CBio...25.R888B 12191:2015NatSR...514864N 12042:2019NatSR...9.5006K 11923:2013PLoSO...861030F 11858:2012NW.....99..435D 11735:2006CRPal...5..685C 11669:2013PalJ...47.1252K 11626:2013PalJ...47.1252K 11446:ElΕΌanowski, Andrzej 11349:2001Natur.409..998S 11147:2019PPP...513..178O 11065:Concornis lacustris 11029:10.1038/ncomms14141 11021:2017NatCo...814141W 10826:2009JVPal..29..188O 10759:Yixianornis grabaui 10713:2004Natur.431..925Z 10510:2011JSPal...9..135O 10420:2014JVPal..34..135W 10383:2014Palgy..57....7M 10285:2012JVPal..32..639H 10193:2013JVPal..33.1176Z 10090:2020CrRes.11004412X 10078:Cretaceous Research 10037:2019NatSR...9..927X 9985:10.1038/ncomms12089 9977:2016NatCo...712089X 9922:2018SciBu..63..235X 9880:2017GondR..49..264X 9824:on 19 October 2014. 9744:2006Sci...312.1640Y 9738:(5780): 1640–1643. 9708:2006CaJES..43..547H 9569:1981Natur.292...51W 9516:Elzanowski, Andrzej 9289:Jiufotang Formation 9139:Gobipipus reshetovi 8662:Djadochta Formation 8513:Huajiying Formation 8463:Huajiying Formation 8329:Jiufotang Formation 8220:Huajiying Formation 8170:Jiufotang Formation 8119:Jiufotang Formation 8073:Jiufotang Formation 7967:Jiufotang Formation 7921:Longjiang Formation 7826:Jiufotang Formation 7780:Jiufotang Formation 7729:Jiufotang Formation 7683:Huajiying Formation 7632:Jiufotang Formation 7549:Jiufotang Formation 7447:Jiufotang Formation 7345:Jiufotang Formation 7299:Jiufotang Formation 7246:Jiufotang Formation 7200:Jiufotang Formation 7012:Jiufotang Formation 6966:Chorrillo Formation 6916:Jiufotang Formation 6861:Jiufotang Formation 6815:Jiufotang Formation 6653:Jiufotang Formation 6602:Huajiying Formation 6556:Jiufotang Formation 6545:Piscivorenantiornis 6501:Jiangdihe Formation 6455:Huajiying Formation 6366:Huajiying Formation 6220:Jiufotang Formation 6078:Jiufotang Formation 5968:Jiufotang Formation 5754:Jiufotang Formation 5541:Jiufotang Formation 5444:Jiufotang Formation 5394:Jiufotang Formation 5370:Considered a small 5191:Jiufotang Formation 5099:Jiufotang Formation 4892:Maevarano Formation 4690:Jiufotang Formation 4352:Jiufotang Formation 4301:Huajiying Formation 4150:Jiufotang Formation 4050:Unnamed formation ( 3996:Jiufotang Formation 3945:CsehbΓ‘nya Formation 3790:Jiufotang Formation 1162:Fossilized eggs of 1084:Piscivorenantiornis 935:and certain extant 901:) with a preserved 891:Fossil skeleton of 778:antorbital fenestra 721:Cratoavis cearensis 706:Wonthaggi Formation 702:Eumeralla Formation 16212:(eagles and hawks) 16112:Procellariiformes 16017:Opisthocomiformes 15986:Steatornithiformes 15213:List by population 15050:Hongshanornithidae 14975:Evolution of birds 14306:Iberomesornithidae 13428:10.7717/peerj.7846 12888:10.7717/peerj.7247 12614:2007-02-19 at the 12522:Sinornis santensis 12493:10.1038/ncomms2104 12324:10.7717/peerj.5910 12179:Scientific Reports 12090:Biological Reviews 12030:Scientific Reports 11823:10.1111/pala.12583 11343:(6823): 998–1000. 11195:10.1111/pala.12453 10765:Journal of Anatomy 10392:10.1111/pala.12082 10333:10.7717/peerj.5910 10228:Journal of Anatomy 10025:Scientific Reports 9786:Historical Biology 7880:evolutionary grade 7272:Originally called 6756:Bissekty Formation 6266:Toolebuc Formation 5860:Bissekty Formation 5842: 5695:Bissekty Formation 5587:Bissekty Formation 4994:Woodbine Formation 4839:Bissekty Formation 4433:, like those of a 4097:Bissekty Formation 3737:Bissekty Formation 1174: 1031: 906: 836: 806:quadratojugal bone 802:temporal fenestrae 749: 630: 16449: 16448: 16374:Taxon identifiers 16365: 16364: 16305: 16304: 16301: 16300: 16297: 16296: 16293: 16292: 16289: 16288: 16285: 16284: 16281: 16280: 16277: 16276: 16098:(loons or divers) 16067:Phaethontiformes 16059:Phaethontimorphae 16009:Opisthocomiformes 15978:Caprimulgiformes 15918: 15917: 15907:Podicipediformes 15826: 15825: 15822: 15821: 15818: 15817: 15544: 15543: 15327: 15326: 15284:Struthioniformes 15152:Waterfowl hunting 15085:Gastornithiformes 15080:Aepyornithiformes 15070:Lithornithiformes 14742: 14741: 14720: 14719: 14700: 14699: 14696: 14695: 13885:Cuspirostrisornis 13790: 13789: 13786: 13785: 13771: 13770: 13767: 13766: 13763: 13762: 13553:Media related to 13512:(11): 1252–1269. 13303:(7837): 272–276. 12561:Apsaravis ukhaana 12199:10.1038/srep14864 11663:(11): 1252–1269. 11620:(11): 1252–1269. 10881:10.7717/peerj.234 9868:Gondwana Research 9782:Praeornis sharovi 9679:978-0-300-06460-5 9623:Mayr, G. (2021). 9609:978-0-520-20094-4 9460:978-0-471-24723-4 9435:978-0-520-20094-4 9368:10.1111/bij.12313 9309:MPCM-LH-26189 a/b 9164:Sankar Chatterjee 9129: 9128: 8799:Avisaurus gloriae 8548: 8547: 8375:Gobipteryx minuta 8366: 8365: 7863: 7862: 7581: 7580: 7150: 7149: 6729:, is named after 5782:Cuspirostrisornis 4431:heterodactyl feet 4341:Cuspirostrisornis 4255:Santana Formation 3578: 3577: 3566: 3565: 3557: 3556: 3548: 3547: 3539: 3538: 3530: 3529: 3521: 3520: 3512: 3511: 3503: 3502: 3494: 3493: 3485: 3484: 3476: 3475: 3467: 3466: 3458: 3457: 3449: 3448: 3440: 3439: 3431: 3430: 3422: 3421: 3413: 3412: 3237: 3236: 3180: 3179: 3171: 3170: 3162: 3161: 3054: 3053: 3045: 3044: 3036: 3035: 2925: 2924: 2916: 2915: 2799: 2798: 2787: 2786: 2778: 2777: 2769: 2768: 2692: 2691: 2577: 2576: 2568: 2567: 2559: 2558: 2509: 2508: 2446: 2445: 2437: 2436: 2428: 2427: 2287: 2286: 2278: 2277: 2188: 2187: 2078: 2077: 2069: 2068: 2060: 2059: 2051: 2050: 2042: 2041: 2033: 2032: 2024: 2023: 2015: 2014: 2006: 2005: 1997: 1996: 1988: 1987: 1979: 1978: 1314:in many species. 1165:Gobipteryx minuta 937:birds-of-paradise 765:Gobipteryx minuta 682:Xiangornis shenmi 354:Gobipteryx minuta 317: 316: 274: 16:(Redirected from 16484: 16467:Cretaceous birds 16462:Enantiornitheans 16442: 16441: 16429: 16428: 16416: 16415: 16414: 16401: 16400: 16399: 16369: 16368: 16353: 16352: 16341: 16329: 16328: 16317: 16316: 16269: 16261: 16253: 16245: 16242:Leptosomiformes 16237: 16229: 16221: 16213: 16210:Accipitriformes 16205: 16184: 16183:(perching birds) 16176: 16168: 16160: 16139: 16131: 16123: 16115: 16107: 16104:Sphenisciformes 16099: 16078: 16070: 16049: 16041: 16038:Charadriiformes 16020: 15999: 15981: 15960: 15952: 15949:Musophagiformes 15944: 15929: 15928: 15910: 15902: 15881: 15873: 15865: 15850: 15849: 15841: 15840: 15832: 15831: 15805: 15792: 15570: 15569: 15566: 15559: 15472: 15465: 15429: 15416: 15415: 15412: 15403: 15402: 15399: 15364: 15363: 15360: 15359: 15333: 15332: 15319: 15311: 15303: 15295: 15287: 15272: 15271: 15268: 15267: 15257: 15256: 15253: 15252: 15120:Bird collections 15075:Dinornithiformes 14980:Darwin's finches 14970:Origin of flight 14910:Seabird breeding 14900:Sexual selection 14769: 14762: 14755: 14746: 14745: 14732: 14731: 14715: 14714: 14708: 14644: 14643: 14585: 14584: 14581: 14580: 14139:Microenantiornis 13850:Castignovolucris 13807: 13806: 13796: 13795: 13723: 13722: 13696: 13695: 13669: 13668: 13660: 13659: 13653: 13652: 13599: 13598: 13584: 13577: 13570: 13561: 13560: 13552: 13538: 13537: 13501: 13495: 13494: 13457: 13451: 13450: 13440: 13430: 13402: 13396: 13395: 13385: 13375: 13343: 13337: 13336: 13291: 13285: 13284: 13256: 13250: 13249: 13209: 13203: 13202: 13166: 13160: 13159: 13123: 13117: 13116: 13088: 13082: 13081: 13071: 13061: 13044:(10): e0184637. 13029: 13023: 13022: 12988: 12982: 12981: 12953: 12947: 12946: 12917: 12911: 12910: 12900: 12890: 12866: 12860: 12859: 12830: 12824: 12823: 12795: 12789: 12788: 12752: 12746: 12745: 12716: 12710: 12709: 12707: 12683: 12677: 12676: 12658: 12626: 12620: 12604: 12595: 12594: 12576: 12556: 12550: 12549: 12547: 12515: 12506: 12505: 12495: 12463: 12457: 12456: 12420: 12414: 12413: 12403: 12393: 12361: 12355: 12354: 12344: 12326: 12302: 12296: 12295: 12269: 12259: 12242:(19): R888–898. 12227: 12221: 12220: 12210: 12170: 12164: 12160: 12154: 12151: 12139: 12129: 12120: 12114: 12113: 12087: 12078: 12072: 12071: 12061: 12021: 12015: 12014: 12012: 11988: 11982: 11981: 11961: 11955: 11954: 11944: 11934: 11902: 11896: 11895: 11869: 11841: 11835: 11834: 11802: 11796: 11795: 11779: 11773: 11772: 11762: 11753: 11747: 11746: 11717: 11711: 11706: 11700: 11695: 11689: 11688: 11652: 11646: 11645: 11609: 11603: 11602: 11590: 11584: 11583: 11547: 11536: 11535: 11499: 11493: 11492: 11483:(5318): 1543–6. 11471: 11465: 11464: 11454: 11442: 11436: 11435: 11423: 11417: 11416: 11406: 11397: 11391: 11390: 11384: 11376: 11357:10.1038/35059172 11331: 11325: 11324: 11314: 11304: 11280: 11274: 11270: 11264: 11260: 11254: 11250: 11244: 11243: 11241: 11240: 11229: 11223: 11222: 11206: 11200: 11199: 11197: 11173: 11167: 11166: 11130: 11124: 11123: 11087: 11081: 11080: 11060: 11051: 11050: 11040: 11000: 10991: 10990: 10961: 10952: 10951: 10941: 10917: 10904: 10903: 10893: 10883: 10855: 10846: 10845: 10808: 10799: 10798: 10788: 10752: 10743: 10742: 10724: 10692: 10683: 10682: 10658: 10652: 10651: 10649: 10625: 10619: 10618: 10600: 10598:10.1002/ar.23446 10576: 10570: 10569: 10541: 10530: 10529: 10493: 10484: 10483: 10481: 10457: 10448: 10447: 10403: 10397: 10396: 10394: 10362: 10356: 10355: 10345: 10335: 10311: 10305: 10304: 10268: 10262: 10261: 10251: 10219: 10213: 10212: 10176: 10167: 10166: 10140: 10116: 10110: 10109: 10073: 10067: 10066: 10056: 10016: 10007: 10006: 9996: 9955: 9944: 9943: 9933: 9910:Science Bulletin 9901: 9892: 9891: 9862: 9853: 9852: 9850: 9843: 9832: 9826: 9825: 9823: 9816: 9802: 9796: 9778: 9772: 9771: 9726: 9720: 9719: 9690: 9684: 9683: 9671: 9661: 9655: 9654: 9644: 9620: 9614: 9613: 9595: 9589: 9588: 9577:10.1038/292051a0 9552: 9543: 9542: 9524: 9512: 9506: 9505: 9495: 9471: 9465: 9464: 9446: 9440: 9439: 9417: 9404: 9403: 9393: 9384: 9373: 9372: 9370: 9346: 9214:Yixian Formation 9191:is considered a 9160:Evgeny Kurochkin 9114: 9112: 9111: 9068: 9066: 9065: 9018: 9016: 9015: 8998:Early Cretaceous 8976: 8962: 8960: 8959: 8950: 8948: 8947: 8912: 8899: 8897: 8896: 8852: 8850: 8849: 8840: 8838: 8837: 8797:A new genus for 8790: 8788: 8787: 8778: 8776: 8775: 8728: 8726: 8725: 8682: 8680: 8679: 8644: 8630: 8628: 8627: 8618: 8616: 8615: 8562: 8533: 8531: 8530: 8517:Early Cretaceous 8483: 8481: 8480: 8467:Early Cretaceous 8436: 8434: 8433: 8380: 8362: 8349: 8347: 8346: 8333:Early Cretaceous 8311: 8290: 8288: 8287: 8274:Early Cretaceous 8270:Yixian Formation 8240: 8238: 8237: 8224:Early Cretaceous 8190: 8188: 8187: 8174:Early Cretaceous 8152: 8139: 8137: 8136: 8123:Early Cretaceous 8093: 8091: 8090: 8077:Early Cretaceous 8055: 8042: 8040: 8039: 8026:Early Cretaceous 8022:Yixian Formation 8004: 7987: 7985: 7984: 7971:Early Cretaceous 7941: 7939: 7938: 7925:Early Cretaceous 7885: 7859: 7846: 7844: 7843: 7830:Early Cretaceous 7800: 7798: 7797: 7784:Early Cretaceous 7762: 7749: 7747: 7746: 7733:Early Cretaceous 7703: 7701: 7700: 7687:Early Cretaceous 7665: 7652: 7650: 7649: 7636:Early Cretaceous 7596: 7569: 7567: 7566: 7553:Early Cretaceous 7531: 7518: 7516: 7515: 7502:Early Cretaceous 7498:Yixian Formation 7480: 7467: 7465: 7464: 7451:Early Cretaceous 7429: 7416: 7414: 7413: 7400:Early Cretaceous 7396:Yixian Formation 7378: 7365: 7363: 7362: 7349:Early Cretaceous 7319: 7317: 7316: 7303:Early Cretaceous 7266: 7264: 7263: 7250:Early Cretaceous 7220: 7218: 7217: 7204:Early Cretaceous 7164: 7146: 7133: 7131: 7130: 7095: 7078: 7076: 7075: 7032: 7030: 7029: 7016:Early Cretaceous 6986: 6984: 6983: 6936: 6934: 6933: 6920:Early Cretaceous 6898: 6881: 6879: 6878: 6865:Early Cretaceous 6835: 6833: 6832: 6819:Early Cretaceous 6797: 6780: 6778: 6777: 6719: 6717: 6716: 6703:Early Cretaceous 6699:Xiagou Formation 6673: 6671: 6670: 6657:Early Cretaceous 6635: 6622: 6620: 6619: 6606:Early Cretaceous 6576: 6574: 6573: 6560:Early Cretaceous 6538: 6525: 6523: 6522: 6475: 6473: 6472: 6459:Early Cretaceous 6428: 6426: 6425: 6416:Early Cretaceous 6386: 6384: 6383: 6370:Early Cretaceous 6336: 6334: 6333: 6320:Early Cretaceous 6286: 6284: 6283: 6270:Early Cretaceous 6240: 6238: 6237: 6224:Early Cretaceous 6194: 6192: 6191: 6178:Early Cretaceous 6174:Yixian Formation 6148: 6146: 6145: 6128:Early Cretaceous 6124:Yixian Formation 6098: 6096: 6095: 6082:Early Cretaceous 6067:Microenantiornis 6052: 6050: 6049: 6041: 6039: 6038: 5988: 5986: 5985: 5972:Early Cretaceous 5939: 5937: 5936: 5923:Early Cretaceous 5919:Yixian Formation 5884: 5882: 5881: 5827: 5825: 5824: 5784:or a synonym of 5774: 5772: 5771: 5758:Early Cretaceous 5736: 5719: 5717: 5716: 5677: 5664: 5662: 5661: 5648:Early Cretaceous 5644:Yixian Formation 5611: 5609: 5608: 5561: 5559: 5558: 5545:Early Cretaceous 5523: 5510: 5508: 5507: 5494:Early Cretaceous 5464: 5462: 5461: 5448:Early Cretaceous 5414: 5412: 5411: 5398:Early Cretaceous 5364: 5362: 5361: 5348:Early Cretaceous 5326: 5307: 5305: 5304: 5261: 5259: 5258: 5241:Nemegt Formation 5211: 5209: 5208: 5195:Early Cretaceous 5165: 5163: 5162: 5149:Early Cretaceous 5145:Yixian Formation 5119: 5117: 5116: 5103:Early Cretaceous 5073: 5071: 5070: 5026: 5024: 5023: 5014: 5012: 5011: 4976: 4963: 4961: 4960: 4947:Early Cretaceous 4943:Xiagou Formation 4925: 4912: 4910: 4909: 4863: 4861: 4860: 4813: 4811: 4810: 4797:Early Cretaceous 4775: 4762: 4760: 4759: 4746:Early Cretaceous 4742:Yixian Formation 4710: 4708: 4707: 4694:Early Cretaceous 4660: 4658: 4657: 4644:Early Cretaceous 4614: 4612: 4611: 4568: 4566: 4565: 4522: 4520: 4519: 4476: 4474: 4473: 4460:Early Cretaceous 4456:Xiagou Formation 4423: 4421: 4420: 4407:Early Cretaceous 4403:Yixian Formation 4385: 4372: 4370: 4369: 4356:Early Cretaceous 4334: 4321: 4319: 4318: 4305:Early Cretaceous 4275: 4273: 4272: 4259:Early Cretaceous 4237: 4224: 4222: 4221: 4208:Early Cretaceous 4186: 4170: 4168: 4167: 4154:Early Cretaceous 4121: 4119: 4118: 4068: 4066: 4065: 4040:Castignovolucris 4033: 4016: 4014: 4013: 4000:Early Cretaceous 3978: 3965: 3963: 3962: 3927: 3914: 3912: 3911: 3898:Early Cretaceous 3894:Xiagou Formation 3876: 3856: 3854: 3853: 3810: 3808: 3807: 3794:Early Cretaceous 3761: 3759: 3758: 3701: 3581:Key to letters: 3377: 3376: 3353: 3352: 3329: 3328: 3305: 3304: 3278: 3277: 3251: 3250: 3201: 3200: 3194: 3193: 3126: 3125: 3102: 3101: 3078: 3077: 3071: 3070: 3000: 2999: 2976: 2975: 2949: 2948: 2939: 2938: 2874: 2873: 2844: 2843: 2837: 2836: 2813: 2812: 2730: 2729: 2706: 2705: 2656: 2655: 2649: 2648: 2622: 2621: 2615: 2614: 2591: 2590: 2523: 2522: 2473: 2472: 2463: 2462: 2392: 2391: 2368: 2367: 2344: 2343: 2314: 2313: 2304: 2303: 2298:Euenantiornithes 2239: 2238: 2212: 2211: 2202: 2201: 2152: 2151: 2145: 2144: 2121: 2120: 2097: 2096: 2089: 2088: 1970: 1936: 1935: 1922: 1899: 1847: 1846: 1833: 1817: 1816: 1803: 1787: 1786: 1762: 1761: 1748: 1734: 1733: 1691: 1690: 1666: 1665: 1641: 1640: 1635:Euenantiornithes 1623: 1602: 1581: 1565: 1564: 1547: 1534: 1533: 1524: 1523: 1475:Euenantiornithes 1417:Jingmai O'Connor 1190:Yixian Formation 788:is preserved in 571:Early Cretaceous 503:Origin and range 497:Enantiornithinae 477:stated in 1996: 433: 432: 423: 422: 269: 264: 156: 155: 127: 117: 54: 41:Early Cretaceous 38:Temporal range: 32: 31: 21: 16492: 16491: 16487: 16486: 16485: 16483: 16482: 16481: 16452: 16451: 16450: 16445: 16437: 16432: 16424: 16419: 16410: 16409: 16404: 16395: 16394: 16389: 16376: 16366: 16361: 16273: 16267: 16259: 16251: 16250:Bucerotiformes 16244:(cuckoo-roller) 16243: 16235: 16227: 16219: 16211: 16203: 16202:Cathartiformes 16188: 16182: 16174: 16173:Psittaciformes 16166: 16158: 16143: 16137: 16136:Pelecaniformes 16129: 16121: 16113: 16105: 16097: 16082: 16076: 16075:Eurypygiformes 16068: 16053: 16047: 16039: 16024: 16018: 16003: 15997: 15979: 15964: 15958: 15950: 15942: 15914: 15908: 15900: 15885: 15879: 15878:Pterocliformes 15871: 15863: 15814: 15803: 15790: 15775: 15699: 15658: 15596: 15561: 15560: 15553: 15552: 15540: 15520: 15493: 15470: 15463: 15423: 15422: 15410: 15409: 15393: 15392: 15389: 15387: 15385: 15383: 15381: 15379: 15377: 15375: 15373: 15371: 15369: 15354: 15352: 15350: 15348: 15346: 15344: 15342: 15340: 15338: 15323: 15317: 15316:Casuariiformes 15309: 15308:Apterygiformes 15301: 15293: 15285: 15262: 15247: 15218:Lists by region 15186: 15096: 15089: 15030:Chaoyangiformes 15015:Jeholornithidae 14989: 14953:Origin of birds 14939: 14920:Brood parasites 14866: 14796: 14783: 14773: 14743: 14738: 14716: 14709: 14692: 14637: 14636: 14628: 14574: 14573: 14572: 14564: 14528: 14458:Bohaiornithidae 14452: 14404:Dapingfangornis 14397:Camptodontornis 14381:Longipterygidae 14375: 14327: 14300: 14188:Paraprotopteryx 14153:Monoenantiornis 14089:Largirostrornis 13843:Brevirostruavis 13803: 13782: 13759: 13739: 13712: 13685: 13679:Avemetatarsalia 13664:Avemetatarsalia 13654: 13648:Ornithothoraces 13604: 13603: 13593: 13588: 13546: 13541: 13502: 13498: 13469:(2): e1054496. 13458: 13454: 13403: 13399: 13358:(6): e0126791. 13344: 13340: 13292: 13288: 13257: 13253: 13210: 13206: 13177:(2): e1278702. 13167: 13163: 13124: 13120: 13099:(11): 909–926. 13089: 13085: 13030: 13026: 12989: 12985: 12954: 12950: 12918: 12914: 12867: 12863: 12831: 12827: 12796: 12792: 12753: 12749: 12717: 12713: 12684: 12680: 12627: 12623: 12616:Wayback Machine 12605: 12598: 12574:10.1.1.693.8475 12557: 12553: 12516: 12509: 12464: 12460: 12421: 12417: 12362: 12358: 12303: 12299: 12236:Current Biology 12228: 12224: 12171: 12167: 12161: 12157: 12142: 12127: 12121: 12117: 12085: 12079: 12075: 12022: 12018: 11989: 11985: 11962: 11958: 11903: 11899: 11867:10.1.1.394.9006 11842: 11838: 11803: 11799: 11780: 11776: 11760: 11754: 11750: 11718: 11714: 11707: 11703: 11696: 11692: 11653: 11649: 11610: 11606: 11591: 11587: 11548: 11539: 11500: 11496: 11472: 11468: 11452: 11443: 11439: 11424: 11420: 11404: 11398: 11394: 11378: 11377: 11332: 11328: 11281: 11277: 11271: 11267: 11261: 11257: 11251: 11247: 11238: 11236: 11231: 11230: 11226: 11207: 11203: 11174: 11170: 11131: 11127: 11088: 11084: 11061: 11054: 11001: 10994: 10973:(1): e1054035. 10962: 10955: 10926:Current Biology 10918: 10907: 10856: 10849: 10809: 10802: 10753: 10746: 10722:10.1038/431925a 10693: 10686: 10659: 10655: 10626: 10622: 10577: 10573: 10542: 10533: 10494: 10487: 10458: 10451: 10404: 10400: 10363: 10359: 10312: 10308: 10269: 10265: 10220: 10216: 10177: 10170: 10125:Current Biology 10117: 10113: 10074: 10070: 10017: 10010: 9956: 9947: 9902: 9895: 9863: 9856: 9848: 9841: 9833: 9829: 9821: 9814: 9803: 9799: 9779: 9775: 9727: 9723: 9716:10.1139/e06-007 9691: 9687: 9680: 9662: 9658: 9621: 9617: 9610: 9596: 9592: 9553: 9546: 9522: 9513: 9509: 9472: 9468: 9461: 9447: 9443: 9436: 9418: 9407: 9391: 9385: 9376: 9347: 9338: 9334: 9197:("naked name"). 9175:: A synonym of 9134: 9109: 9107: 9098:Late Cretaceous 9094:Lecho Formation 9063: 9061: 9052:Late Cretaceous 9013: 9011: 8957: 8955: 8945: 8943: 8934:Late Cretaceous 8894: 8892: 8883:Late Cretaceous 8847: 8845: 8835: 8833: 8824:Late Cretaceous 8785: 8783: 8773: 8771: 8762:Late Cretaceous 8723: 8721: 8712:Late Cretaceous 8677: 8675: 8666:Late Cretaceous 8625: 8623: 8613: 8611: 8602:Late Cretaceous 8553: 8528: 8526: 8478: 8476: 8431: 8429: 8420:Late Cretaceous 8371: 8344: 8342: 8285: 8283: 8235: 8233: 8185: 8183: 8134: 8132: 8088: 8086: 8037: 8035: 7982: 7980: 7936: 7934: 7868: 7866:Bohaiornithidae 7841: 7839: 7795: 7793: 7744: 7742: 7698: 7696: 7647: 7645: 7586: 7564: 7562: 7513: 7511: 7462: 7460: 7411: 7409: 7360: 7358: 7314: 7312: 7288:Dapingfangornis 7261: 7259: 7235:Camptodontornis 7215: 7213: 7159:Longipterygidae 7155: 7153:Longipterygidae 7128: 7126: 7117:Late Cretaceous 7113:Qiupa Formation 7086:tarsometatarsal 7073: 7071: 7062:Late Cretaceous 7058:Lecho Formation 7027: 7025: 6981: 6979: 6970:Late Cretaceous 6931: 6929: 6876: 6874: 6830: 6828: 6775: 6773: 6760:Late Cretaceous 6714: 6712: 6668: 6666: 6617: 6615: 6571: 6569: 6520: 6518: 6505:Late Cretaceous 6470: 6468: 6443:Paraprotopteryx 6423: 6421: 6381: 6379: 6331: 6329: 6281: 6279: 6235: 6233: 6189: 6187: 6163:Monoenantiornis 6143: 6141: 6093: 6091: 6047: 6045: 6036: 6034: 6025:Late Cretaceous 6021:Lecho Formation 5983: 5981: 5934: 5932: 5879: 5877: 5864:Late Cretaceous 5822: 5820: 5811:Late Cretaceous 5807:Lecho Formation 5769: 5767: 5743:Largirostrornis 5714: 5712: 5699:Late Cretaceous 5659: 5657: 5606: 5604: 5591:Late Cretaceous 5556: 5554: 5505: 5503: 5459: 5457: 5409: 5407: 5359: 5357: 5302: 5300: 5291:Late Cretaceous 5256: 5254: 5245:Late Cretaceous 5206: 5204: 5160: 5158: 5114: 5112: 5068: 5066: 5057:Late Cretaceous 5021: 5019: 5009: 5007: 4998:Late Cretaceous 4958: 4956: 4907: 4905: 4896:Late Cretaceous 4858: 4856: 4843:Late Cretaceous 4808: 4806: 4757: 4755: 4705: 4703: 4655: 4653: 4609: 4607: 4598:Late Cretaceous 4594:Lecho Formation 4563: 4561: 4552:Late Cretaceous 4517: 4515: 4506:Late Cretaceous 4502:Lecho Formation 4471: 4469: 4418: 4416: 4367: 4365: 4316: 4314: 4270: 4268: 4219: 4217: 4165: 4163: 4116: 4114: 4101:Late Cretaceous 4063: 4061: 4052:Late Cretaceous 4011: 4009: 3985:Brevirostruavis 3960: 3958: 3949:Late Cretaceous 3909: 3907: 3851: 3849: 3840:Late Cretaceous 3805: 3803: 3756: 3754: 3741:Late Cretaceous 3695: 3688: 3673: 3663: 3653: 3643: 3633: 3623: 3613: 3603: 3593: 3579: 3567: 3558: 3549: 3540: 3531: 3522: 3513: 3504: 3495: 3486: 3477: 3468: 3459: 3450: 3441: 3432: 3423: 3414: 3272:Bohaiornithidae 3238: 3181: 3172: 3163: 3055: 3046: 3037: 2926: 2917: 2800: 2788: 2779: 2770: 2693: 2578: 2569: 2560: 2510: 2447: 2438: 2429: 2288: 2279: 2206:Longipterygidae 2189: 2092:Enantiornithes 2079: 2070: 2061: 2052: 2043: 2034: 2025: 2016: 2007: 1998: 1989: 1980: 1742:Bohaiornithidae 1617:Longipterygidae 1528:Ornithothoraces 1448:Ornithothoraces 1436: 1410:Ornithuromorpha 1402:Ornithothoraces 1361: 1331:Ornithuromorphs 1303: 1272:Late Cretaceous 1156: 1127: 1111:Longipterygidae 1047:Bohaiornithidae 1005: 1000: 963:Paraprotopteryx 885: 845:Ornithuromorpha 824: 816:cranial kinesis 730: 615: 595:Late Cretaceous 505: 488:tarsometatarsus 424:) "opposite" + 380:of what is now 371:Cyril A. Walker 349: 302:Longipterygidae 292:Bohaiornithidae 268: 262: 250:Ornithothoraces 150: 118: 116: 115: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 49: 48: 45:Late Cretaceous 39: 36: 28: 23: 22: 18:Enantiornithine 15: 12: 11: 5: 16490: 16480: 16479: 16474: 16469: 16464: 16447: 16446: 16444: 16443: 16430: 16417: 16412:Enantiornithes 16402: 16386: 16384: 16382:Enantiornithes 16378: 16377: 16363: 16362: 16360: 16359: 16347: 16335: 16323: 16310: 16307: 16306: 16303: 16302: 16299: 16298: 16295: 16294: 16291: 16290: 16287: 16286: 16283: 16282: 16279: 16278: 16275: 16274: 16272: 16271: 16263: 16258:Coraciiformes 16255: 16247: 16239: 16234:Trogoniformes 16231: 16223: 16215: 16207: 16198: 16196: 16190: 16189: 16187: 16186: 16181:Passeriformes 16178: 16170: 16165:Falconiformes 16162: 16157:Cariamiformes 16153: 16151: 16145: 16144: 16142: 16141: 16133: 16125: 16120:Ciconiiformes 16117: 16109: 16101: 16092: 16090: 16084: 16083: 16081: 16080: 16072: 16063: 16061: 16055: 16054: 16052: 16051: 16043: 16034: 16032: 16030:Cursorimorphae 16026: 16025: 16023: 16022: 16013: 16011: 16005: 16004: 16002: 16001: 15993: 15988: 15983: 15974: 15972: 15966: 15965: 15963: 15962: 15954: 15946: 15937: 15935: 15926: 15920: 15919: 15916: 15915: 15913: 15912: 15904: 15895: 15893: 15891:Mirandornithes 15887: 15886: 15884: 15883: 15875: 15867: 15862:Columbiformes 15858: 15856: 15854:Columbimorphae 15847: 15838: 15828: 15827: 15824: 15823: 15820: 15819: 15816: 15815: 15813: 15812: 15807: 15799: 15794: 15789:Meleagridinae 15785: 15783: 15777: 15776: 15774: 15773: 15766: 15759: 15752: 15745: 15738: 15731: 15724: 15717: 15709: 15707: 15705:Odontophoridae 15701: 15700: 15698: 15697: 15690: 15683: 15676: 15668: 15666: 15660: 15659: 15657: 15656: 15649: 15642: 15635: 15628: 15621: 15614: 15606: 15604: 15598: 15597: 15595: 15594: 15589: 15584: 15578: 15576: 15567: 15546: 15545: 15542: 15541: 15539: 15538: 15530: 15528: 15522: 15521: 15519: 15518: 15511: 15503: 15501: 15495: 15494: 15492: 15491: 15486: 15484:Stictonettinae 15481: 15479:Dendrocygninae 15476: 15475: 15474: 15467: 15455: 15454: 15453: 15448: 15443: 15432: 15430: 15413: 15400: 15357: 15329: 15328: 15325: 15324: 15322: 15321: 15313: 15305: 15297: 15289: 15280: 15278: 15264: 15263: 15249: 15248: 15246: 15245: 15244: 15243: 15238: 15232:Notable birds 15230: 15225: 15220: 15215: 15210: 15205: 15200: 15194: 15192: 15188: 15187: 15185: 15184: 15182:Egg collecting 15179: 15174: 15169: 15164: 15159: 15154: 15149: 15144: 15139: 15134: 15133: 15132: 15122: 15117: 15112: 15107: 15101: 15099: 15091: 15090: 15088: 15087: 15082: 15077: 15072: 15067: 15065:Hesperornithes 15062: 15057: 15052: 15047: 15042: 15040:Ambiortiformes 15037: 15032: 15027: 15025:Enantiornithes 15022: 15017: 15012: 15007: 14999: 14997: 14991: 14990: 14988: 14987: 14982: 14977: 14972: 14967: 14966: 14965: 14960: 14949: 14947: 14941: 14940: 14938: 14937: 14932: 14927: 14922: 14917: 14912: 14907: 14902: 14897: 14892: 14887: 14882: 14876: 14874: 14868: 14867: 14865: 14864: 14859: 14854: 14849: 14844: 14839: 14834: 14829: 14828: 14827: 14817: 14812: 14806: 14804: 14798: 14797: 14795: 14794: 14788: 14785: 14784: 14772: 14771: 14764: 14757: 14749: 14740: 14739: 14737: 14736: 14725: 14722: 14721: 14718: 14717: 14703: 14701: 14698: 14697: 14694: 14693: 14691: 14690: 14683: 14676: 14669: 14662: 14655: 14647: 14645: 14642:Chiappe, 1992) 14630: 14629: 14627: 14626: 14619: 14612: 14609:Enantiophoenix 14605: 14597: 14588: 14582: 14566: 14565: 14563: 14562: 14554: 14546: 14538: 14536: 14534:Gobipterygidae 14530: 14529: 14527: 14526: 14519: 14512: 14505: 14502:Parabohaiornis 14498: 14491: 14484: 14477: 14470: 14462: 14460: 14454: 14453: 14451: 14450: 14447:Shengjingornis 14443: 14436: 14429: 14426:Longirostravis 14422: 14415: 14407: 14400: 14393: 14385: 14383: 14377: 14376: 14374: 14373: 14366: 14359: 14352: 14345: 14337: 14335: 14333:Pengornithidae 14329: 14328: 14326: 14325: 14318: 14310: 14308: 14302: 14301: 14299: 14298: 14291: 14284: 14281:Yuanjiawaornis 14277: 14270: 14263: 14256: 14249: 14242: 14235: 14228: 14221: 14213: 14205: 14198: 14191: 14184: 14177: 14170: 14163: 14156: 14149: 14142: 14135: 14128: 14125:Longchengornis 14121: 14113: 14106: 14099: 14092: 14085: 14077: 14070: 14063: 14056: 14049: 14042: 14035: 14028: 14021: 14014: 14007: 14000: 13993: 13986: 13979: 13972: 13965: 13958: 13951: 13944: 13937: 13930: 13923: 13916: 13909: 13902: 13895: 13888: 13881: 13878:Cruralispennia 13874: 13867: 13860: 13853: 13846: 13839: 13832: 13825: 13822:Alethoalaornis 13818: 13810: 13804: 13801:Enantiornithes 13799: 13792: 13791: 13788: 13787: 13784: 13783: 13779:Enantiornithes 13776: 13773: 13772: 13769: 13768: 13765: 13764: 13761: 13760: 13758: 13757: 13756: 13755: 13749: 13747: 13745:Enantiornithes 13741: 13740: 13738: 13737: 13736: 13735: 13726: 13720: 13714: 13713: 13711: 13710: 13709: 13708: 13699: 13693: 13687: 13686: 13684: 13683: 13682: 13681: 13672: 13666: 13656: 13655: 13651: 13650: 13641: 13632: 13623: 13617: 13611: 13595: 13594: 13591:Enantiornithes 13587: 13586: 13579: 13572: 13564: 13555:Enantiornithes 13545: 13544:External links 13542: 13540: 13539: 13496: 13452: 13397: 13338: 13286: 13251: 13204: 13161: 13134:(1): 135–145. 13118: 13083: 13024: 12983: 12948: 12929:(4): 319–338. 12912: 12861: 12825: 12806:(2): 801–807. 12790: 12763:(3): 657–671. 12747: 12711: 12698:(2): 161–171. 12678: 12641:(4): 961–976. 12621: 12596: 12567:(3387): 1–46. 12551: 12507: 12458: 12431:(2): 193–208. 12415: 12376:(12): e28672. 12356: 12297: 12222: 12165: 12155: 12153: 12152: 12115: 12073: 12016: 11983: 11956: 11897: 11836: 11797: 11774: 11748: 11712: 11701: 11690: 11647: 11604: 11585: 11558:(3594): 1–46. 11537: 11494: 11466: 11437: 11418: 11392: 11326: 11275: 11265: 11255: 11245: 11224: 11201: 11168: 11125: 11082: 11071:(3133): 1–23. 11052: 10992: 10953: 10932:(1): 114–119. 10905: 10847: 10820:(1): 188–204. 10800: 10771:(3): 287–308. 10744: 10684: 10653: 10620: 10571: 10552:(6): 481–497. 10531: 10504:(1): 135–157. 10485: 10472:(2): 161–171. 10449: 10414:(1): 135–145. 10398: 10357: 10306: 10263: 10214: 10187:(5): 1176–89. 10168: 10111: 10068: 10008: 9945: 9916:(4): 235–243. 9893: 9854: 9827: 9797: 9773: 9721: 9702:(5): 547–554. 9685: 9678: 9656: 9615: 9608: 9590: 9563:(5818): 51–3. 9544: 9507: 9486:(3): 463–475. 9466: 9459: 9441: 9434: 9405: 9374: 9361:(3): 805–819. 9335: 9333: 9330: 9329: 9328: 9306: 9300: 9278: 9272: 9262: 9244: 9226: 9223: 9205: 9198: 9167: 9133: 9130: 9127: 9126: 9124: 9118: 9105: 9091: 9086: 9078: 9077: 9075: 9072: 9059: 9045: 9040: 9032: 9031: 9029: 9022: 9009: 8994:Ilek Formation 8991: 8986: 8978: 8977: 8970: 8967: 8941: 8927: 8922: 8914: 8913: 8906: 8903: 8890: 8876: 8871: 8863: 8862: 8860: 8857: 8831: 8817: 8812: 8804: 8803: 8801: 8795: 8769: 8755: 8750: 8742: 8741: 8739: 8732: 8719: 8705: 8700: 8697:Enantiophoenix 8692: 8691: 8689: 8686: 8673: 8659: 8654: 8646: 8645: 8638: 8635: 8609: 8595: 8590: 8582: 8581: 8578: 8575: 8572: 8569: 8566: 8552: 8549: 8546: 8545: 8543: 8537: 8524: 8509: 8504: 8496: 8495: 8493: 8487: 8474: 8459: 8454: 8446: 8445: 8443: 8440: 8427: 8413: 8408: 8400: 8399: 8396: 8393: 8390: 8387: 8384: 8370: 8369:Gobipterygidae 8367: 8364: 8363: 8356: 8353: 8340: 8326: 8321: 8313: 8312: 8305: 8294: 8281: 8267: 8262: 8254: 8253: 8251: 8244: 8231: 8216: 8211: 8203: 8202: 8200: 8194: 8181: 8167: 8162: 8159:Parabohaiornis 8154: 8153: 8146: 8143: 8130: 8116: 8111: 8103: 8102: 8100: 8097: 8084: 8070: 8065: 8057: 8056: 8049: 8046: 8033: 8019: 8014: 8006: 8005: 7998: 7991: 7978: 7964: 7959: 7951: 7950: 7948: 7945: 7932: 7918: 7913: 7905: 7904: 7901: 7898: 7895: 7892: 7889: 7871:Bohaiornithids 7867: 7864: 7861: 7860: 7853: 7850: 7837: 7823: 7818: 7810: 7809: 7807: 7804: 7791: 7777: 7772: 7764: 7763: 7756: 7753: 7740: 7726: 7721: 7713: 7712: 7710: 7707: 7694: 7680: 7675: 7667: 7666: 7659: 7656: 7643: 7629: 7624: 7616: 7615: 7612: 7609: 7606: 7603: 7600: 7590:Pengornithidae 7585: 7584:Pengornithidae 7582: 7579: 7578: 7576: 7573: 7560: 7546: 7541: 7538:Shengjingornis 7533: 7532: 7525: 7522: 7509: 7495: 7490: 7482: 7481: 7474: 7471: 7458: 7444: 7439: 7431: 7430: 7423: 7420: 7407: 7393: 7388: 7385:Longirostravis 7380: 7379: 7372: 7369: 7356: 7342: 7337: 7329: 7328: 7326: 7323: 7310: 7296: 7291: 7283: 7282: 7280: 7270: 7257: 7243: 7238: 7230: 7229: 7227: 7224: 7211: 7197: 7192: 7184: 7183: 7180: 7177: 7174: 7171: 7168: 7154: 7151: 7148: 7147: 7140: 7137: 7124: 7110: 7105: 7097: 7096: 7089: 7082: 7069: 7055: 7050: 7042: 7041: 7039: 7036: 7023: 7009: 7004: 7001:Yuanjiawaornis 6996: 6995: 6993: 6990: 6977: 6963: 6958: 6950: 6949: 6947: 6940: 6927: 6913: 6908: 6900: 6899: 6892: 6885: 6872: 6858: 6853: 6845: 6844: 6842: 6839: 6826: 6812: 6807: 6799: 6798: 6791: 6784: 6771: 6753: 6748: 6740: 6739: 6737: 6733:from the band 6723: 6710: 6696: 6691: 6683: 6682: 6680: 6677: 6664: 6650: 6645: 6637: 6636: 6629: 6626: 6613: 6599: 6594: 6586: 6585: 6583: 6580: 6567: 6553: 6548: 6540: 6539: 6532: 6529: 6516: 6498: 6493: 6485: 6484: 6482: 6479: 6466: 6451: 6446: 6438: 6437: 6435: 6432: 6419: 6409: 6404: 6396: 6395: 6393: 6390: 6377: 6363: 6358: 6350: 6349: 6347: 6340: 6327: 6313: 6308: 6300: 6299: 6297: 6290: 6277: 6263: 6258: 6250: 6249: 6247: 6244: 6231: 6217: 6212: 6204: 6203: 6201: 6198: 6185: 6171: 6166: 6158: 6157: 6155: 6152: 6139: 6121: 6116: 6108: 6107: 6105: 6102: 6089: 6075: 6070: 6062: 6061: 6059: 6056: 6032: 6014: 6009: 6001: 6000: 5998: 5992: 5979: 5965: 5960: 5957:Longchengornis 5952: 5951: 5949: 5943: 5930: 5916: 5911: 5903: 5902: 5900: 5888: 5875: 5857: 5852: 5844: 5843: 5834: 5831: 5818: 5804: 5799: 5791: 5790: 5788: 5778: 5765: 5751: 5746: 5738: 5737: 5730: 5723: 5710: 5692: 5687: 5679: 5678: 5671: 5668: 5655: 5641: 5636: 5628: 5627: 5625: 5615: 5602: 5584: 5579: 5571: 5570: 5568: 5565: 5552: 5538: 5533: 5525: 5524: 5517: 5514: 5501: 5487: 5482: 5474: 5473: 5471: 5468: 5455: 5441: 5436: 5428: 5427: 5425: 5418: 5405: 5391: 5386: 5378: 5377: 5375: 5368: 5355: 5341: 5336: 5328: 5327: 5320: 5311: 5298: 5284: 5279: 5271: 5270: 5268: 5265: 5252: 5238: 5233: 5225: 5224: 5222: 5215: 5202: 5188: 5183: 5175: 5174: 5172: 5169: 5156: 5142: 5137: 5129: 5128: 5126: 5123: 5110: 5096: 5091: 5083: 5082: 5080: 5077: 5064: 5050: 5045: 5037: 5036: 5034: 5031: 5005: 4991: 4986: 4978: 4977: 4970: 4967: 4954: 4940: 4935: 4927: 4926: 4919: 4916: 4903: 4889: 4884: 4876: 4875: 4873: 4867: 4854: 4836: 4831: 4823: 4822: 4820: 4817: 4804: 4793:Ilek Formation 4790: 4785: 4777: 4776: 4769: 4766: 4753: 4739: 4734: 4726: 4725: 4723: 4714: 4701: 4687: 4682: 4674: 4673: 4671: 4664: 4651: 4637: 4632: 4624: 4623: 4621: 4618: 4605: 4591: 4586: 4578: 4577: 4575: 4572: 4559: 4545: 4540: 4532: 4531: 4529: 4526: 4513: 4499: 4494: 4486: 4485: 4483: 4480: 4467: 4453: 4448: 4440: 4439: 4437: 4427: 4414: 4400: 4395: 4387: 4386: 4379: 4376: 4363: 4349: 4344: 4336: 4335: 4328: 4325: 4312: 4298: 4293: 4290:Cruralispennia 4285: 4284: 4282: 4279: 4266: 4252: 4247: 4239: 4238: 4231: 4228: 4215: 4201: 4196: 4188: 4187: 4180: 4174: 4161: 4147: 4142: 4134: 4133: 4131: 4125: 4112: 4094: 4089: 4081: 4080: 4078: 4072: 4059: 4048: 4043: 4035: 4034: 4027: 4020: 4007: 3993: 3988: 3980: 3979: 3972: 3969: 3956: 3942: 3937: 3929: 3928: 3921: 3918: 3905: 3891: 3886: 3878: 3877: 3870: 3860: 3847: 3833: 3828: 3820: 3819: 3817: 3814: 3801: 3787: 3782: 3779:Alethoalaornis 3774: 3773: 3771: 3765: 3752: 3734: 3729: 3721: 3720: 3717: 3714: 3711: 3708: 3705: 3694: 3692:Incertae sedis 3689: 3687: 3686:List of genera 3684: 3670:Yuanjiawaornis 3640:Parabohaiornis 3610:Enantiophoenix 3576: 3575: 3569: 3568: 3564: 3563: 3560: 3559: 3555: 3554: 3551: 3550: 3546: 3545: 3542: 3541: 3537: 3536: 3533: 3532: 3528: 3527: 3524: 3523: 3519: 3518: 3515: 3514: 3510: 3509: 3506: 3505: 3501: 3500: 3497: 3496: 3492: 3491: 3488: 3487: 3483: 3482: 3479: 3478: 3474: 3473: 3470: 3469: 3465: 3464: 3461: 3460: 3456: 3455: 3452: 3451: 3447: 3446: 3443: 3442: 3438: 3437: 3434: 3433: 3429: 3428: 3425: 3424: 3420: 3419: 3416: 3415: 3411: 3410: 3407: 3406: 3397: 3394: 3393: 3390: 3389: 3380: 3375: 3373: 3370: 3369: 3366: 3365: 3356: 3351: 3349: 3346: 3345: 3342: 3341: 3332: 3327: 3325: 3322: 3321: 3318: 3317: 3308: 3303: 3301: 3295: 3294: 3291: 3290: 3281: 3276: 3274: 3268: 3267: 3264: 3263: 3254: 3249: 3247: 3244: 3243: 3240: 3239: 3235: 3234: 3231: 3230: 3226:Shengjingornis 3221: 3218: 3217: 3214: 3213: 3204: 3199: 3197: 3192: 3190: 3187: 3186: 3183: 3182: 3178: 3177: 3174: 3173: 3169: 3168: 3165: 3164: 3160: 3159: 3156: 3155: 3146: 3143: 3142: 3139: 3138: 3129: 3124: 3122: 3119: 3118: 3115: 3114: 3105: 3100: 3098: 3095: 3094: 3091: 3090: 3081: 3076: 3074: 3069: 3067: 3064: 3063: 3057: 3056: 3052: 3051: 3048: 3047: 3043: 3042: 3039: 3038: 3034: 3033: 3030: 3029: 3020: 3017: 3016: 3013: 3012: 3003: 2998: 2996: 2993: 2992: 2989: 2988: 2979: 2974: 2972: 2966: 2965: 2962: 2961: 2952: 2947: 2945: 2943:Pengornithidae 2937: 2935: 2932: 2931: 2928: 2927: 2923: 2922: 2919: 2918: 2914: 2913: 2910: 2909: 2900: 2894: 2893: 2890: 2889: 2880: 2872: 2870: 2864: 2863: 2860: 2859: 2850: 2842: 2840: 2835: 2833: 2830: 2829: 2826: 2825: 2816: 2811: 2809: 2806: 2805: 2802: 2801: 2797: 2796: 2790: 2789: 2785: 2784: 2781: 2780: 2776: 2775: 2772: 2771: 2767: 2766: 2763: 2762: 2753: 2747: 2746: 2743: 2742: 2733: 2728: 2726: 2723: 2722: 2719: 2718: 2709: 2704: 2702: 2699: 2698: 2695: 2694: 2690: 2689: 2686: 2685: 2676: 2673: 2672: 2669: 2668: 2659: 2654: 2652: 2647: 2645: 2639: 2638: 2635: 2634: 2625: 2620: 2618: 2613: 2611: 2608: 2607: 2604: 2603: 2594: 2589: 2587: 2584: 2583: 2580: 2579: 2575: 2574: 2571: 2570: 2566: 2565: 2562: 2561: 2557: 2556: 2553: 2552: 2543: 2540: 2539: 2536: 2535: 2526: 2521: 2519: 2516: 2515: 2512: 2511: 2507: 2506: 2503: 2502: 2493: 2490: 2489: 2486: 2485: 2476: 2471: 2469: 2461: 2459: 2453: 2452: 2449: 2448: 2444: 2443: 2440: 2439: 2435: 2434: 2431: 2430: 2426: 2425: 2422: 2421: 2412: 2409: 2408: 2405: 2404: 2395: 2390: 2388: 2385: 2384: 2381: 2380: 2371: 2366: 2364: 2361: 2360: 2357: 2356: 2347: 2342: 2340: 2334: 2333: 2330: 2329: 2320: 2312: 2310: 2302: 2300: 2294: 2293: 2290: 2289: 2285: 2284: 2281: 2280: 2276: 2275: 2272: 2271: 2262: 2259: 2258: 2255: 2254: 2250:Longirostravis 2245: 2237: 2235: 2229: 2228: 2225: 2224: 2215: 2210: 2208: 2200: 2198: 2195: 2194: 2191: 2190: 2186: 2185: 2182: 2181: 2172: 2169: 2168: 2165: 2164: 2155: 2150: 2148: 2143: 2141: 2138: 2137: 2134: 2133: 2124: 2119: 2117: 2114: 2113: 2110: 2109: 2105:Cruralispennia 2100: 2095: 2093: 2087: 2076: 2075: 2072: 2071: 2067: 2066: 2063: 2062: 2058: 2057: 2054: 2053: 2049: 2048: 2045: 2044: 2040: 2039: 2036: 2035: 2031: 2030: 2027: 2026: 2022: 2021: 2018: 2017: 2013: 2012: 2009: 2008: 2004: 2003: 2000: 1999: 1995: 1994: 1991: 1990: 1986: 1985: 1982: 1981: 1977: 1976: 1973: 1972: 1957: 1954: 1953: 1950: 1949: 1939: 1934: 1932: 1929: 1928: 1925: 1924: 1909: 1906: 1905: 1902: 1901: 1886: 1883: 1882: 1879: 1878: 1868: 1865: 1864: 1861: 1860: 1850: 1845: 1843: 1840: 1839: 1836: 1835: 1820: 1815: 1813: 1810: 1809: 1806: 1805: 1790: 1785: 1783: 1780: 1779: 1776: 1775: 1765: 1760: 1758: 1755: 1754: 1751: 1750: 1737: 1732: 1730: 1727: 1726: 1723: 1722: 1712: 1709: 1708: 1705: 1704: 1694: 1689: 1687: 1684: 1683: 1680: 1679: 1669: 1664: 1662: 1659: 1658: 1655: 1654: 1644: 1639: 1637: 1630: 1629: 1626: 1625: 1612: 1609: 1608: 1605: 1604: 1596:Pengornithidae 1591: 1588: 1587: 1584: 1583: 1568: 1563: 1561: 1559:Enantiornithes 1554: 1553: 1550: 1549: 1537: 1532: 1530: 1522: 1459:to just four. 1457:autapomorphies 1435: 1432: 1388:Confuciusornis 1360: 1359:Classification 1357: 1302: 1299: 1265:metabolic rate 1230:superprecocial 1155: 1152: 1126: 1123: 1117:Shengjingornis 1069:Enantiophoenix 1004: 1001: 999: 996: 991:Cruralispennia 972:, a primitive 932:Confuciusornis 927:rectrical bulb 894:Rapaxavis pani 884: 881: 823: 820: 782:squamosal bone 770:quadrate bones 729: 726: 694:Mirarce eatoni 688:Zhouornis hani 676:Pengornis houi 614: 611: 504: 501: 348: 345: 337:Hesperornithes 321:Enantiornithes 315: 314: 310: 309: 307:Pengornithidae 304: 299: 297:Gobipterygidae 294: 289: 281: 280: 276: 275: 266:Enantiornithes 260: 253: 252: 247: 240: 239: 234: 227: 226: 221: 214: 213: 208: 201: 200: 195: 188: 187: 182: 178: 177: 172: 168: 167: 162: 158: 157: 144: 143: 139:Zhouornis hani 129: 128: 120: 119: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 37: 35:Enantiornithes 26: 9: 6: 4: 3: 2: 16489: 16478: 16475: 16473: 16470: 16468: 16465: 16463: 16460: 16459: 16457: 16440: 16435: 16431: 16427: 16422: 16418: 16413: 16407: 16403: 16398: 16392: 16388: 16387: 16385: 16383: 16379: 16375: 16370: 16358: 16357: 16348: 16346: 16345: 16340: 16336: 16334: 16333: 16324: 16322: 16321: 16312: 16311: 16308: 16270: 16264: 16262: 16256: 16254: 16248: 16246: 16240: 16238: 16232: 16230: 16224: 16222: 16218:Strigiformes 16216: 16214: 16208: 16206: 16200: 16199: 16197: 16195: 16191: 16185: 16179: 16177: 16171: 16169: 16163: 16161: 16155: 16154: 16152: 16150: 16146: 16140: 16134: 16132: 16126: 16124: 16118: 16116: 16110: 16108: 16102: 16100: 16094: 16093: 16091: 16089: 16085: 16079: 16073: 16071: 16069:(tropicbirds) 16065: 16064: 16062: 16060: 16056: 16050: 16044: 16042: 16036: 16035: 16033: 16031: 16027: 16021: 16015: 16014: 16012: 16010: 16006: 16000: 15994: 15992: 15991:Podargiformes 15989: 15987: 15984: 15982: 15976: 15975: 15973: 15971: 15967: 15961: 15955: 15953: 15947: 15945: 15941:Cuculiformes 15939: 15938: 15936: 15934: 15930: 15927: 15925: 15921: 15911: 15905: 15903: 15897: 15896: 15894: 15892: 15888: 15882: 15876: 15874: 15868: 15866: 15860: 15859: 15857: 15855: 15851: 15848: 15846: 15842: 15839: 15837: 15833: 15829: 15811: 15808: 15806: 15800: 15798: 15795: 15793: 15787: 15786: 15784: 15782: 15778: 15772: 15771: 15767: 15765: 15764: 15760: 15758: 15757: 15753: 15751: 15750: 15746: 15744: 15743: 15739: 15737: 15736: 15732: 15730: 15729: 15725: 15723: 15722: 15718: 15716: 15715: 15711: 15710: 15708: 15706: 15702: 15696: 15695: 15691: 15689: 15688: 15684: 15682: 15681: 15677: 15675: 15674: 15670: 15669: 15667: 15665: 15661: 15655: 15654: 15650: 15648: 15647: 15643: 15641: 15640: 15639:Macrocephalon 15636: 15634: 15633: 15629: 15627: 15626: 15622: 15620: 15619: 15615: 15613: 15612: 15608: 15607: 15605: 15603: 15599: 15593: 15590: 15588: 15585: 15583: 15580: 15579: 15577: 15575: 15571: 15568: 15564: 15557: 15551: 15547: 15537: 15536: 15532: 15531: 15529: 15527: 15526:Anseranatidae 15523: 15517: 15516: 15512: 15510: 15509: 15505: 15504: 15502: 15500: 15496: 15490: 15487: 15485: 15482: 15480: 15477: 15473: 15468: 15466: 15461: 15460: 15459: 15456: 15452: 15449: 15447: 15444: 15442: 15439: 15438: 15437: 15434: 15433: 15431: 15427: 15421: 15417: 15414: 15408: 15404: 15401: 15397: 15391: 15365: 15361: 15358: 15356: 15334: 15330: 15320: 15314: 15312: 15306: 15304: 15300:Tinamiformes 15298: 15296: 15290: 15288: 15282: 15281: 15279: 15277: 15276:Palaeognathae 15273: 15269: 15265: 15258: 15254: 15250: 15242: 15239: 15237: 15234: 15233: 15231: 15229: 15226: 15224: 15221: 15219: 15216: 15214: 15211: 15209: 15206: 15204: 15201: 15199: 15196: 15195: 15193: 15189: 15183: 15180: 15178: 15175: 15173: 15170: 15168: 15165: 15163: 15162:Pigeon racing 15160: 15158: 15155: 15153: 15150: 15148: 15145: 15143: 15140: 15138: 15135: 15131: 15128: 15127: 15126: 15123: 15121: 15118: 15116: 15113: 15111: 15108: 15106: 15103: 15102: 15100: 15098: 15092: 15086: 15083: 15081: 15078: 15076: 15073: 15071: 15068: 15066: 15063: 15061: 15058: 15056: 15053: 15051: 15048: 15046: 15043: 15041: 15038: 15036: 15033: 15031: 15028: 15026: 15023: 15021: 15018: 15016: 15013: 15011: 15008: 15006: 15005: 15004:Archaeopteryx 15001: 15000: 14998: 14996: 14992: 14986: 14983: 14981: 14978: 14976: 14973: 14971: 14968: 14964: 14961: 14959: 14956: 14955: 14954: 14951: 14950: 14948: 14946: 14942: 14936: 14933: 14931: 14928: 14926: 14923: 14921: 14918: 14916: 14913: 14911: 14908: 14906: 14903: 14901: 14898: 14896: 14893: 14891: 14888: 14886: 14883: 14881: 14878: 14877: 14875: 14873: 14869: 14863: 14860: 14858: 14855: 14853: 14850: 14848: 14845: 14843: 14840: 14838: 14835: 14833: 14830: 14826: 14823: 14822: 14821: 14818: 14816: 14813: 14811: 14808: 14807: 14805: 14803: 14799: 14793: 14790: 14789: 14786: 14781: 14777: 14770: 14765: 14763: 14758: 14756: 14751: 14750: 14747: 14735: 14727: 14726: 14723: 14713: 14707: 14689: 14688: 14687:Soroavisaurus 14684: 14682: 14681: 14677: 14675: 14674: 14670: 14668: 14667: 14663: 14661: 14660: 14656: 14654: 14653: 14649: 14648: 14646: 14641: 14635: 14631: 14625: 14624: 14620: 14618: 14617: 14613: 14611: 14610: 14606: 14603: 14602: 14598: 14595: 14594: 14590: 14589: 14586: 14583: 14578: 14571: 14567: 14560: 14559: 14555: 14552: 14551: 14547: 14545: 14544: 14540: 14539: 14537: 14535: 14531: 14525: 14524: 14520: 14518: 14517: 14513: 14511: 14510: 14506: 14504: 14503: 14499: 14497: 14496: 14492: 14490: 14489: 14485: 14483: 14482: 14478: 14476: 14475: 14471: 14469: 14468: 14464: 14463: 14461: 14459: 14455: 14449: 14448: 14444: 14442: 14441: 14437: 14435: 14434: 14430: 14428: 14427: 14423: 14421: 14420: 14416: 14413: 14412: 14408: 14406: 14405: 14401: 14399: 14398: 14394: 14392: 14391: 14387: 14386: 14384: 14382: 14378: 14372: 14371: 14367: 14365: 14364: 14360: 14358: 14357: 14356:Parapengornis 14353: 14351: 14350: 14346: 14344: 14343: 14339: 14338: 14336: 14334: 14330: 14324: 14323: 14319: 14317: 14316: 14315:Iberomesornis 14312: 14311: 14309: 14307: 14303: 14297: 14296: 14292: 14290: 14289: 14288:Yungavolucris 14285: 14283: 14282: 14278: 14276: 14275: 14271: 14269: 14268: 14264: 14262: 14261: 14257: 14255: 14254: 14250: 14248: 14247: 14243: 14241: 14240: 14236: 14234: 14233: 14229: 14227: 14226: 14222: 14219: 14218: 14214: 14211: 14210: 14206: 14204: 14203: 14199: 14197: 14196: 14192: 14190: 14189: 14185: 14183: 14182: 14178: 14176: 14175: 14171: 14169: 14168: 14164: 14162: 14161: 14157: 14155: 14154: 14150: 14148: 14147: 14143: 14141: 14140: 14136: 14134: 14133: 14129: 14127: 14126: 14122: 14119: 14118: 14114: 14112: 14111: 14110:Liaoningornis 14107: 14105: 14104: 14100: 14098: 14097: 14093: 14091: 14090: 14086: 14083: 14082: 14078: 14076: 14075: 14071: 14069: 14068: 14064: 14062: 14061: 14057: 14055: 14054: 14050: 14048: 14047: 14043: 14041: 14040: 14036: 14034: 14033: 14029: 14027: 14026: 14022: 14020: 14019: 14015: 14013: 14012: 14008: 14006: 14005: 14001: 13999: 13998: 13994: 13992: 13991: 13987: 13985: 13984: 13980: 13978: 13977: 13973: 13971: 13970: 13966: 13964: 13963: 13959: 13957: 13956: 13952: 13950: 13949: 13948:Eoenantiornis 13945: 13943: 13942: 13941:Eocathayornis 13938: 13936: 13935: 13931: 13929: 13928: 13924: 13922: 13921: 13917: 13915: 13914: 13910: 13908: 13907: 13903: 13901: 13900: 13896: 13894: 13893: 13892:Dalingheornis 13889: 13887: 13886: 13882: 13880: 13879: 13875: 13873: 13872: 13868: 13866: 13865: 13861: 13859: 13858: 13854: 13852: 13851: 13847: 13845: 13844: 13840: 13838: 13837: 13833: 13831: 13830: 13826: 13824: 13823: 13819: 13817: 13816: 13812: 13811: 13808: 13805: 13802: 13797: 13793: 13781: 13780: 13774: 13753: 13752: 13751: 13750: 13748: 13746: 13742: 13734: 13730: 13729: 13728: 13727: 13724: 13721: 13719: 13715: 13707: 13703: 13702: 13701: 13700: 13697: 13694: 13692: 13688: 13680: 13676: 13675: 13674: 13673: 13670: 13667: 13665: 13661: 13657: 13649: 13645: 13642: 13640: 13636: 13633: 13631: 13627: 13624: 13622: 13618: 13616: 13612: 13610: 13606: 13605: 13600: 13596: 13592: 13585: 13580: 13578: 13573: 13571: 13566: 13565: 13562: 13558: 13556: 13551: 13535: 13531: 13527: 13523: 13519: 13515: 13511: 13507: 13500: 13492: 13488: 13484: 13480: 13476: 13472: 13468: 13464: 13456: 13448: 13444: 13439: 13434: 13429: 13424: 13420: 13416: 13412: 13410: 13401: 13393: 13389: 13384: 13379: 13374: 13369: 13365: 13361: 13357: 13353: 13349: 13342: 13334: 13330: 13326: 13322: 13318: 13314: 13310: 13306: 13302: 13298: 13290: 13282: 13278: 13274: 13270: 13266: 13262: 13255: 13247: 13243: 13239: 13235: 13231: 13227: 13223: 13219: 13215: 13208: 13200: 13196: 13192: 13188: 13184: 13180: 13176: 13172: 13165: 13157: 13153: 13149: 13145: 13141: 13137: 13133: 13129: 13122: 13114: 13110: 13106: 13102: 13098: 13094: 13087: 13079: 13075: 13070: 13065: 13060: 13055: 13051: 13047: 13043: 13039: 13035: 13028: 13020: 13016: 13012: 13008: 13004: 13000: 12999: 12994: 12987: 12979: 12975: 12971: 12967: 12963: 12959: 12952: 12944: 12940: 12936: 12932: 12928: 12924: 12916: 12908: 12904: 12899: 12894: 12889: 12884: 12880: 12876: 12872: 12865: 12857: 12853: 12849: 12845: 12841: 12837: 12829: 12821: 12817: 12813: 12809: 12805: 12801: 12794: 12786: 12782: 12778: 12774: 12770: 12766: 12762: 12758: 12751: 12743: 12739: 12735: 12731: 12727: 12723: 12715: 12706: 12701: 12697: 12693: 12689: 12682: 12674: 12670: 12666: 12662: 12657: 12652: 12648: 12644: 12640: 12636: 12632: 12625: 12618: 12617: 12613: 12610: 12603: 12601: 12592: 12588: 12584: 12580: 12575: 12570: 12566: 12562: 12555: 12546: 12541: 12537: 12533: 12529: 12527: 12523: 12514: 12512: 12503: 12499: 12494: 12489: 12485: 12481: 12477: 12473: 12469: 12462: 12454: 12450: 12446: 12442: 12438: 12434: 12430: 12426: 12419: 12411: 12407: 12402: 12397: 12392: 12387: 12383: 12379: 12375: 12371: 12367: 12360: 12352: 12348: 12343: 12338: 12334: 12330: 12325: 12320: 12316: 12312: 12308: 12301: 12293: 12289: 12285: 12281: 12277: 12273: 12268: 12263: 12258: 12253: 12249: 12245: 12241: 12237: 12233: 12226: 12218: 12214: 12209: 12204: 12200: 12196: 12192: 12188: 12184: 12180: 12176: 12169: 12159: 12149: 12145: 12141: 12140: 12137: 12133: 12126: 12119: 12111: 12107: 12103: 12099: 12095: 12091: 12084: 12077: 12069: 12065: 12060: 12055: 12051: 12047: 12043: 12039: 12035: 12031: 12027: 12020: 12011: 12006: 12002: 11998: 11994: 11987: 11979: 11975: 11971: 11967: 11960: 11952: 11948: 11943: 11938: 11933: 11928: 11924: 11920: 11917:(4): e61030. 11916: 11912: 11908: 11901: 11893: 11889: 11885: 11881: 11877: 11873: 11868: 11863: 11859: 11855: 11852:(6): 435–42. 11851: 11847: 11840: 11832: 11828: 11824: 11820: 11816: 11812: 11811:Palaeontology 11808: 11801: 11793: 11789: 11785: 11778: 11770: 11766: 11759: 11752: 11744: 11740: 11736: 11732: 11729:(5): 685–91. 11728: 11724: 11716: 11710: 11705: 11699: 11694: 11686: 11682: 11678: 11674: 11670: 11666: 11662: 11658: 11651: 11643: 11639: 11635: 11631: 11627: 11623: 11619: 11615: 11608: 11600: 11596: 11589: 11581: 11577: 11573: 11569: 11565: 11561: 11557: 11553: 11546: 11544: 11542: 11533: 11529: 11525: 11521: 11517: 11513: 11510:(5696): 653. 11509: 11505: 11498: 11490: 11486: 11482: 11478: 11470: 11462: 11458: 11451: 11447: 11441: 11433: 11429: 11422: 11415:(2): 193–238. 11414: 11410: 11403: 11396: 11388: 11382: 11374: 11370: 11366: 11362: 11358: 11354: 11350: 11346: 11342: 11338: 11330: 11322: 11318: 11313: 11308: 11303: 11298: 11294: 11290: 11286: 11279: 11269: 11259: 11249: 11234: 11228: 11220: 11216: 11212: 11205: 11196: 11191: 11187: 11183: 11182:Palaeontology 11179: 11172: 11164: 11160: 11156: 11152: 11148: 11144: 11140: 11136: 11129: 11121: 11117: 11113: 11109: 11105: 11101: 11098:(4): 856–60. 11097: 11093: 11086: 11078: 11074: 11070: 11066: 11059: 11057: 11048: 11044: 11039: 11034: 11030: 11026: 11022: 11018: 11014: 11010: 11006: 10999: 10997: 10988: 10984: 10980: 10976: 10972: 10968: 10960: 10958: 10949: 10945: 10940: 10935: 10931: 10927: 10923: 10916: 10914: 10912: 10910: 10901: 10897: 10892: 10887: 10882: 10877: 10873: 10869: 10865: 10863: 10854: 10852: 10843: 10839: 10835: 10831: 10827: 10823: 10819: 10815: 10807: 10805: 10796: 10792: 10787: 10782: 10778: 10774: 10770: 10766: 10762: 10760: 10751: 10749: 10740: 10736: 10732: 10728: 10723: 10718: 10714: 10710: 10707:(7011): 925. 10706: 10702: 10698: 10691: 10689: 10680: 10676: 10672: 10668: 10664: 10657: 10648: 10643: 10640:(2): 248–56. 10639: 10635: 10631: 10624: 10616: 10612: 10608: 10604: 10599: 10594: 10590: 10586: 10582: 10575: 10567: 10563: 10559: 10555: 10551: 10547: 10540: 10538: 10536: 10527: 10523: 10519: 10515: 10511: 10507: 10503: 10499: 10492: 10490: 10480: 10475: 10471: 10467: 10463: 10456: 10454: 10445: 10441: 10437: 10433: 10429: 10425: 10421: 10417: 10413: 10409: 10402: 10393: 10388: 10384: 10380: 10376: 10372: 10371:Palaeontology 10368: 10361: 10353: 10349: 10344: 10339: 10334: 10329: 10325: 10321: 10317: 10310: 10302: 10298: 10294: 10290: 10286: 10282: 10279:(3): 639–45. 10278: 10274: 10267: 10259: 10255: 10250: 10245: 10241: 10237: 10234:(5): 565–77. 10233: 10229: 10225: 10218: 10210: 10206: 10202: 10198: 10194: 10190: 10186: 10182: 10175: 10173: 10164: 10160: 10156: 10152: 10148: 10144: 10139: 10134: 10130: 10126: 10122: 10115: 10107: 10103: 10099: 10095: 10091: 10087: 10083: 10079: 10072: 10064: 10060: 10055: 10050: 10046: 10042: 10038: 10034: 10030: 10026: 10022: 10015: 10013: 10004: 10000: 9995: 9990: 9986: 9982: 9978: 9974: 9970: 9966: 9962: 9954: 9952: 9950: 9941: 9937: 9932: 9927: 9923: 9919: 9915: 9911: 9907: 9900: 9898: 9889: 9885: 9881: 9877: 9873: 9869: 9861: 9859: 9847: 9840: 9839: 9831: 9820: 9813: 9812: 9811: 9801: 9795: 9791: 9787: 9783: 9777: 9769: 9765: 9761: 9757: 9753: 9749: 9745: 9741: 9737: 9733: 9725: 9717: 9713: 9709: 9705: 9701: 9697: 9689: 9681: 9675: 9670: 9669: 9660: 9652: 9648: 9643: 9638: 9635:(2): 217–28. 9634: 9630: 9629:Zoomorphology 9626: 9619: 9611: 9605: 9601: 9594: 9586: 9582: 9578: 9574: 9570: 9566: 9562: 9558: 9551: 9549: 9540: 9536: 9532: 9528: 9521: 9517: 9511: 9503: 9499: 9494: 9489: 9485: 9481: 9477: 9470: 9462: 9456: 9452: 9445: 9437: 9431: 9427: 9423: 9416: 9414: 9412: 9410: 9401: 9397: 9390: 9383: 9381: 9379: 9369: 9364: 9360: 9356: 9352: 9345: 9343: 9341: 9336: 9326: 9322: 9318: 9314: 9310: 9307: 9304: 9301: 9298: 9294: 9290: 9286: 9285:bohaiornithid 9282: 9279: 9276: 9273: 9270: 9266: 9263: 9260: 9259: 9254: 9253: 9248: 9245: 9242: 9238: 9235:Formation of 9234: 9230: 9227: 9224: 9221: 9220: 9215: 9211: 9210: 9206: 9203: 9199: 9196: 9195: 9190: 9186: 9182: 9178: 9174: 9173:fengningensis 9172: 9168: 9165: 9161: 9157: 9153: 9149: 9145: 9141: 9140: 9136: 9135: 9125: 9123: 9119: 9117: 9106: 9103: 9102:Maastrichtian 9099: 9095: 9092: 9090: 9087: 9085: 9084: 9083:Soroavisaurus 9080: 9079: 9076: 9073: 9071: 9060: 9057: 9053: 9049: 9046: 9044: 9041: 9039: 9038: 9034: 9033: 9030: 9027: 9023: 9021: 9010: 9007: 9003: 8999: 8995: 8992: 8990: 8987: 8985: 8984: 8980: 8979: 8975: 8971: 8968: 8965: 8953: 8952:United States 8942: 8939: 8935: 8931: 8928: 8926: 8923: 8921: 8920: 8916: 8915: 8911: 8907: 8904: 8902: 8891: 8888: 8884: 8880: 8877: 8875: 8872: 8870: 8869: 8865: 8864: 8861: 8858: 8855: 8843: 8842:United States 8832: 8829: 8825: 8821: 8818: 8816: 8813: 8811: 8810: 8806: 8805: 8802: 8800: 8796: 8793: 8781: 8780:United States 8770: 8767: 8763: 8759: 8756: 8754: 8751: 8749: 8748: 8744: 8743: 8740: 8737: 8733: 8731: 8720: 8717: 8713: 8709: 8706: 8704: 8701: 8699: 8698: 8694: 8693: 8690: 8687: 8685: 8674: 8671: 8667: 8663: 8660: 8658: 8655: 8653: 8652: 8648: 8647: 8643: 8639: 8636: 8633: 8621: 8620:United States 8610: 8607: 8606:Maastrichtian 8603: 8599: 8596: 8594: 8591: 8589: 8588: 8584: 8583: 8579: 8576: 8573: 8570: 8567: 8564: 8563: 8560: 8557: 8544: 8542: 8538: 8536: 8525: 8522: 8518: 8514: 8510: 8508: 8505: 8503: 8502: 8498: 8497: 8494: 8492: 8488: 8486: 8475: 8472: 8468: 8464: 8460: 8458: 8455: 8453: 8452: 8448: 8447: 8444: 8441: 8439: 8428: 8425: 8421: 8417: 8414: 8412: 8409: 8407: 8406: 8402: 8401: 8397: 8394: 8391: 8388: 8385: 8382: 8381: 8378: 8376: 8361: 8357: 8354: 8352: 8341: 8338: 8334: 8330: 8327: 8325: 8322: 8320: 8319: 8315: 8314: 8310: 8306: 8303: 8300:with grooved 8299: 8295: 8293: 8282: 8279: 8275: 8271: 8268: 8266: 8263: 8261: 8260: 8256: 8255: 8252: 8249: 8245: 8243: 8232: 8229: 8225: 8221: 8217: 8215: 8212: 8210: 8209: 8205: 8204: 8201: 8199: 8195: 8193: 8182: 8179: 8175: 8171: 8168: 8166: 8163: 8161: 8160: 8156: 8155: 8151: 8147: 8144: 8142: 8131: 8128: 8124: 8120: 8117: 8115: 8112: 8110: 8109: 8105: 8104: 8101: 8098: 8096: 8085: 8082: 8078: 8074: 8071: 8069: 8066: 8064: 8063: 8059: 8058: 8054: 8050: 8047: 8045: 8034: 8031: 8027: 8023: 8020: 8018: 8015: 8013: 8012: 8008: 8007: 8003: 7999: 7996: 7992: 7990: 7979: 7976: 7972: 7968: 7965: 7963: 7960: 7958: 7957: 7953: 7952: 7949: 7946: 7944: 7933: 7930: 7926: 7922: 7919: 7917: 7914: 7912: 7911: 7907: 7906: 7902: 7899: 7896: 7893: 7890: 7887: 7886: 7883: 7881: 7877: 7872: 7858: 7854: 7851: 7849: 7838: 7835: 7831: 7827: 7824: 7822: 7819: 7817: 7816: 7812: 7811: 7808: 7805: 7803: 7792: 7789: 7785: 7781: 7778: 7776: 7773: 7771: 7770: 7766: 7765: 7761: 7757: 7754: 7752: 7741: 7738: 7734: 7730: 7727: 7725: 7722: 7720: 7719: 7718:Parapengornis 7715: 7714: 7711: 7708: 7706: 7695: 7692: 7688: 7684: 7681: 7679: 7676: 7674: 7673: 7669: 7668: 7664: 7660: 7657: 7655: 7644: 7641: 7637: 7633: 7630: 7628: 7625: 7623: 7622: 7618: 7617: 7613: 7610: 7607: 7604: 7601: 7598: 7597: 7594: 7591: 7577: 7574: 7572: 7561: 7558: 7554: 7550: 7547: 7545: 7542: 7540: 7539: 7535: 7534: 7530: 7526: 7523: 7521: 7510: 7507: 7503: 7499: 7496: 7494: 7491: 7489: 7488: 7484: 7483: 7479: 7475: 7472: 7470: 7459: 7456: 7452: 7448: 7445: 7443: 7440: 7438: 7437: 7433: 7432: 7428: 7424: 7421: 7419: 7408: 7405: 7401: 7397: 7394: 7392: 7389: 7387: 7386: 7382: 7381: 7377: 7373: 7370: 7368: 7357: 7354: 7350: 7346: 7343: 7341: 7338: 7336: 7335: 7331: 7330: 7327: 7324: 7322: 7311: 7308: 7304: 7300: 7297: 7295: 7292: 7290: 7289: 7285: 7284: 7281: 7279: 7275: 7271: 7269: 7258: 7255: 7251: 7247: 7244: 7242: 7239: 7237: 7236: 7232: 7231: 7228: 7225: 7223: 7212: 7209: 7205: 7201: 7198: 7196: 7193: 7191: 7190: 7186: 7185: 7181: 7178: 7175: 7172: 7169: 7166: 7165: 7162: 7160: 7145: 7141: 7138: 7136: 7125: 7122: 7121:Maastrichtian 7118: 7114: 7111: 7109: 7106: 7104: 7103: 7099: 7098: 7094: 7090: 7088:(ankle bone) 7087: 7083: 7081: 7070: 7067: 7066:Maastrichtian 7063: 7059: 7056: 7054: 7051: 7049: 7048: 7047:Yungavolucris 7044: 7043: 7040: 7037: 7035: 7024: 7021: 7017: 7013: 7010: 7008: 7005: 7003: 7002: 6998: 6997: 6994: 6991: 6989: 6978: 6975: 6974:Maastrichtian 6971: 6967: 6964: 6962: 6959: 6957: 6956: 6952: 6951: 6948: 6945: 6941: 6939: 6928: 6925: 6921: 6917: 6914: 6912: 6909: 6907: 6906: 6902: 6901: 6897: 6893: 6890: 6886: 6884: 6873: 6870: 6866: 6862: 6859: 6857: 6854: 6852: 6851: 6847: 6846: 6843: 6840: 6838: 6827: 6824: 6820: 6816: 6813: 6811: 6808: 6806: 6805: 6801: 6800: 6796: 6792: 6789: 6785: 6783: 6772: 6769: 6765: 6761: 6757: 6754: 6752: 6749: 6747: 6746: 6742: 6741: 6738: 6736: 6732: 6728: 6724: 6722: 6711: 6708: 6704: 6700: 6697: 6695: 6692: 6690: 6689: 6685: 6684: 6681: 6678: 6676: 6665: 6662: 6658: 6654: 6651: 6649: 6646: 6644: 6643: 6639: 6638: 6634: 6630: 6627: 6625: 6614: 6611: 6607: 6603: 6600: 6598: 6595: 6593: 6592: 6588: 6587: 6584: 6581: 6579: 6568: 6565: 6561: 6557: 6554: 6552: 6549: 6547: 6546: 6542: 6541: 6537: 6533: 6530: 6528: 6517: 6514: 6510: 6506: 6502: 6499: 6497: 6494: 6492: 6491: 6487: 6486: 6483: 6480: 6478: 6467: 6464: 6460: 6456: 6452: 6450: 6447: 6445: 6444: 6440: 6439: 6436: 6434:Poorly known 6433: 6431: 6420: 6417: 6413: 6410: 6408: 6405: 6403: 6402: 6398: 6397: 6394: 6391: 6389: 6378: 6375: 6371: 6367: 6364: 6362: 6359: 6357: 6356: 6352: 6351: 6348: 6345: 6341: 6339: 6328: 6325: 6321: 6317: 6314: 6312: 6309: 6307: 6306: 6302: 6301: 6298: 6295: 6291: 6289: 6278: 6275: 6271: 6267: 6264: 6262: 6259: 6257: 6256: 6252: 6251: 6248: 6245: 6243: 6232: 6229: 6225: 6221: 6218: 6216: 6213: 6211: 6210: 6206: 6205: 6202: 6199: 6197: 6186: 6183: 6179: 6175: 6172: 6170: 6167: 6165: 6164: 6160: 6159: 6156: 6153: 6151: 6140: 6137: 6133: 6129: 6125: 6122: 6120: 6117: 6115: 6114: 6110: 6109: 6106: 6103: 6101: 6090: 6087: 6083: 6079: 6076: 6074: 6071: 6069: 6068: 6064: 6063: 6060: 6057: 6055: 6044: 6033: 6030: 6029:Maastrichtian 6026: 6022: 6018: 6015: 6013: 6010: 6008: 6007: 6003: 6002: 5999: 5997: 5993: 5991: 5980: 5977: 5973: 5969: 5966: 5964: 5961: 5959: 5958: 5954: 5953: 5950: 5948: 5944: 5942: 5931: 5928: 5924: 5920: 5917: 5915: 5912: 5910: 5909: 5908:Liaoningornis 5905: 5904: 5901: 5899: 5898: 5893: 5889: 5887: 5876: 5873: 5869: 5865: 5861: 5858: 5856: 5853: 5851: 5850: 5846: 5845: 5839: 5835: 5832: 5830: 5819: 5816: 5815:Maastrichtian 5812: 5808: 5805: 5803: 5800: 5798: 5797: 5793: 5792: 5789: 5787: 5783: 5779: 5777: 5766: 5763: 5759: 5755: 5752: 5750: 5747: 5745: 5744: 5740: 5739: 5735: 5731: 5728: 5724: 5722: 5711: 5708: 5704: 5700: 5696: 5693: 5691: 5688: 5686: 5685: 5681: 5680: 5676: 5672: 5669: 5667: 5656: 5653: 5649: 5645: 5642: 5640: 5637: 5635: 5634: 5630: 5629: 5626: 5624: 5620: 5616: 5614: 5603: 5600: 5596: 5592: 5588: 5585: 5583: 5580: 5578: 5577: 5573: 5572: 5569: 5566: 5564: 5553: 5550: 5546: 5542: 5539: 5537: 5534: 5532: 5531: 5527: 5526: 5522: 5518: 5515: 5513: 5502: 5499: 5495: 5491: 5488: 5486: 5483: 5481: 5480: 5479:Iberomesornis 5476: 5475: 5472: 5469: 5467: 5456: 5453: 5449: 5445: 5442: 5440: 5437: 5435: 5434: 5430: 5429: 5426: 5423: 5419: 5417: 5406: 5403: 5399: 5395: 5392: 5390: 5387: 5385: 5384: 5380: 5379: 5376: 5373: 5369: 5367: 5356: 5353: 5349: 5345: 5342: 5340: 5337: 5335: 5334: 5330: 5329: 5325: 5321: 5318: 5317: 5312: 5310: 5299: 5296: 5292: 5288: 5285: 5283: 5280: 5278: 5277: 5273: 5272: 5269: 5266: 5264: 5253: 5250: 5249:Maastrichtian 5246: 5242: 5239: 5237: 5234: 5232: 5231: 5227: 5226: 5223: 5220: 5216: 5214: 5203: 5200: 5196: 5192: 5189: 5187: 5184: 5182: 5181: 5177: 5176: 5173: 5170: 5168: 5157: 5154: 5150: 5146: 5143: 5141: 5138: 5136: 5135: 5131: 5130: 5127: 5124: 5122: 5111: 5108: 5104: 5100: 5097: 5095: 5092: 5090: 5089: 5085: 5084: 5081: 5078: 5076: 5065: 5062: 5058: 5054: 5053:Burmese amber 5051: 5049: 5046: 5044: 5043: 5039: 5038: 5035: 5032: 5029: 5017: 5016:United States 5006: 5003: 4999: 4995: 4992: 4990: 4987: 4985: 4984: 4980: 4979: 4975: 4971: 4968: 4966: 4955: 4952: 4948: 4944: 4941: 4939: 4936: 4934: 4933: 4929: 4928: 4924: 4920: 4917: 4915: 4904: 4901: 4900:Maastrichtian 4897: 4893: 4890: 4888: 4885: 4883: 4882: 4878: 4877: 4874: 4872: 4868: 4866: 4855: 4852: 4848: 4844: 4840: 4837: 4835: 4832: 4830: 4829: 4825: 4824: 4821: 4818: 4816: 4805: 4802: 4798: 4794: 4791: 4789: 4786: 4784: 4783: 4779: 4778: 4774: 4770: 4767: 4765: 4754: 4751: 4747: 4743: 4740: 4738: 4735: 4733: 4732: 4731:Eoenantiornis 4728: 4727: 4724: 4721: 4720: 4715: 4713: 4702: 4699: 4695: 4691: 4688: 4686: 4683: 4681: 4680: 4679:Eocathayornis 4676: 4675: 4672: 4669: 4665: 4663: 4652: 4649: 4645: 4641: 4638: 4636: 4633: 4631: 4630: 4626: 4625: 4622: 4619: 4617: 4606: 4603: 4602:Maastrichtian 4599: 4595: 4592: 4590: 4587: 4585: 4584: 4580: 4579: 4576: 4573: 4571: 4560: 4557: 4553: 4549: 4548:Burmese Amber 4546: 4544: 4541: 4539: 4538: 4534: 4533: 4530: 4527: 4525: 4514: 4511: 4510:Maastrichtian 4507: 4503: 4500: 4498: 4495: 4493: 4492: 4488: 4487: 4484: 4481: 4479: 4468: 4465: 4461: 4457: 4454: 4452: 4449: 4447: 4446: 4442: 4441: 4438: 4436: 4432: 4428: 4426: 4415: 4412: 4408: 4404: 4401: 4399: 4396: 4394: 4393: 4392:Dalingheornis 4389: 4388: 4384: 4380: 4377: 4375: 4364: 4361: 4357: 4353: 4350: 4348: 4345: 4343: 4342: 4338: 4337: 4333: 4329: 4326: 4324: 4313: 4310: 4306: 4302: 4299: 4297: 4294: 4292: 4291: 4287: 4286: 4283: 4280: 4278: 4267: 4264: 4260: 4256: 4253: 4251: 4248: 4246: 4245: 4241: 4240: 4236: 4232: 4229: 4227: 4216: 4213: 4209: 4205: 4202: 4200: 4197: 4195: 4194: 4190: 4189: 4185: 4181: 4179: 4175: 4173: 4162: 4159: 4155: 4151: 4148: 4146: 4143: 4141: 4140: 4136: 4135: 4132: 4130: 4126: 4124: 4113: 4110: 4106: 4102: 4098: 4095: 4093: 4090: 4088: 4087: 4083: 4082: 4079: 4077: 4073: 4071: 4060: 4057: 4053: 4049: 4047: 4044: 4042: 4041: 4037: 4036: 4032: 4028: 4025: 4021: 4019: 4008: 4005: 4001: 3997: 3994: 3992: 3989: 3987: 3986: 3982: 3981: 3977: 3973: 3970: 3968: 3957: 3954: 3950: 3946: 3943: 3941: 3938: 3936: 3935: 3931: 3930: 3926: 3922: 3919: 3917: 3906: 3903: 3899: 3895: 3892: 3890: 3887: 3885: 3884: 3880: 3879: 3875: 3871: 3869: 3865: 3861: 3859: 3848: 3845: 3841: 3837: 3834: 3832: 3829: 3827: 3826: 3822: 3821: 3818: 3816:Poorly known 3815: 3813: 3802: 3799: 3795: 3791: 3788: 3786: 3783: 3781: 3780: 3776: 3775: 3772: 3770: 3766: 3764: 3753: 3750: 3746: 3742: 3738: 3735: 3733: 3730: 3728: 3727: 3723: 3722: 3718: 3715: 3712: 3709: 3706: 3703: 3702: 3699: 3693: 3683: 3682: 3681: 3680:Yungavolucris 3676: 3672: 3671: 3666: 3662: 3661: 3656: 3652: 3651: 3646: 3642: 3641: 3636: 3632: 3631: 3626: 3622: 3621: 3616: 3612: 3611: 3606: 3602: 3601: 3596: 3592: 3591: 3586: 3582: 3574: 3571: 3570: 3562: 3561: 3553: 3552: 3544: 3543: 3535: 3534: 3526: 3525: 3517: 3516: 3508: 3507: 3499: 3498: 3490: 3489: 3481: 3480: 3472: 3471: 3463: 3462: 3454: 3453: 3445: 3444: 3436: 3435: 3427: 3426: 3418: 3417: 3409: 3408: 3405: 3404: 3403: 3396: 3395: 3392: 3391: 3388: 3387: 3386: 3379: 3378: 3372: 3371: 3368: 3367: 3364: 3363: 3362: 3355: 3354: 3348: 3347: 3344: 3343: 3340: 3339: 3338: 3331: 3330: 3324: 3323: 3320: 3319: 3316: 3315: 3314: 3313:Eoenantiornis 3307: 3306: 3300: 3297: 3296: 3293: 3292: 3289: 3288: 3287: 3280: 3279: 3273: 3270: 3269: 3266: 3265: 3262: 3261: 3260: 3253: 3252: 3246: 3245: 3242: 3241: 3233: 3232: 3229: 3228: 3227: 3220: 3219: 3216: 3215: 3212: 3211: 3210: 3203: 3202: 3196: 3195: 3189: 3188: 3185: 3184: 3176: 3175: 3167: 3166: 3158: 3157: 3154: 3153: 3152: 3145: 3144: 3141: 3140: 3137: 3136: 3135: 3128: 3127: 3121: 3120: 3117: 3116: 3113: 3112: 3111: 3104: 3103: 3097: 3096: 3093: 3092: 3089: 3088: 3087: 3080: 3079: 3073: 3072: 3066: 3065: 3062: 3059: 3058: 3050: 3049: 3041: 3040: 3032: 3031: 3028: 3027: 3026: 3025:Parapengornis 3019: 3018: 3015: 3014: 3011: 3010: 3009: 3002: 3001: 2995: 2994: 2991: 2990: 2987: 2986: 2985: 2978: 2977: 2971: 2968: 2967: 2964: 2963: 2960: 2959: 2958: 2951: 2950: 2944: 2941: 2940: 2934: 2933: 2930: 2929: 2921: 2920: 2912: 2911: 2908: 2907: 2906: 2899: 2896: 2895: 2892: 2891: 2888: 2887: 2886: 2879: 2876: 2875: 2869: 2866: 2865: 2862: 2861: 2858: 2857: 2856: 2849: 2846: 2845: 2839: 2838: 2832: 2831: 2828: 2827: 2824: 2823: 2822: 2815: 2814: 2808: 2807: 2804: 2803: 2795: 2792: 2791: 2783: 2782: 2774: 2773: 2765: 2764: 2761: 2760: 2759: 2752: 2749: 2748: 2745: 2744: 2741: 2740: 2739: 2732: 2731: 2725: 2724: 2721: 2720: 2717: 2716: 2715: 2708: 2707: 2701: 2700: 2697: 2696: 2688: 2687: 2684: 2683: 2682: 2681:Soroavisaurus 2675: 2674: 2671: 2670: 2667: 2666: 2665: 2658: 2657: 2651: 2650: 2644: 2641: 2640: 2637: 2636: 2633: 2632: 2631: 2624: 2623: 2617: 2616: 2610: 2609: 2606: 2605: 2602: 2601: 2600: 2599:Eocathayornis 2593: 2592: 2586: 2585: 2582: 2581: 2573: 2572: 2564: 2563: 2555: 2554: 2551: 2550: 2549: 2542: 2541: 2538: 2537: 2534: 2533: 2532: 2525: 2524: 2518: 2517: 2514: 2513: 2505: 2504: 2501: 2500: 2499: 2498:Liaoningornis 2492: 2491: 2488: 2487: 2484: 2483: 2482: 2475: 2474: 2468: 2465: 2464: 2458: 2455: 2454: 2451: 2450: 2442: 2441: 2433: 2432: 2424: 2423: 2420: 2419: 2418: 2411: 2410: 2407: 2406: 2403: 2402: 2401: 2394: 2393: 2387: 2386: 2383: 2382: 2379: 2378: 2377: 2370: 2369: 2363: 2362: 2359: 2358: 2355: 2354: 2353: 2346: 2345: 2339: 2336: 2335: 2332: 2331: 2328: 2327: 2326: 2319: 2316: 2315: 2309: 2306: 2305: 2299: 2296: 2295: 2292: 2291: 2283: 2282: 2274: 2273: 2270: 2269: 2268: 2261: 2260: 2257: 2256: 2253: 2252: 2251: 2244: 2241: 2240: 2234: 2231: 2230: 2227: 2226: 2223: 2222: 2221: 2220:Iberomesornis 2214: 2213: 2207: 2204: 2203: 2197: 2196: 2193: 2192: 2184: 2183: 2180: 2179: 2178: 2171: 2170: 2167: 2166: 2163: 2162: 2161: 2154: 2153: 2147: 2146: 2140: 2139: 2136: 2135: 2132: 2131: 2130: 2123: 2122: 2116: 2115: 2112: 2111: 2108: 2107: 2106: 2099: 2098: 2091: 2090: 2086: 2084: 2074: 2073: 2065: 2064: 2056: 2055: 2047: 2046: 2038: 2037: 2029: 2028: 2020: 2019: 2011: 2010: 2002: 2001: 1993: 1992: 1984: 1983: 1975: 1974: 1971: 1969: 1965: 1964: 1956: 1955: 1952: 1951: 1948: 1947: 1946: 1938: 1937: 1931: 1930: 1927: 1926: 1923: 1921: 1917: 1916: 1908: 1907: 1904: 1903: 1900: 1898: 1894: 1893: 1885: 1884: 1881: 1880: 1877: 1876: 1875: 1867: 1866: 1863: 1862: 1859: 1858: 1857: 1849: 1848: 1842: 1841: 1838: 1837: 1834: 1832: 1828: 1827: 1819: 1818: 1812: 1811: 1808: 1807: 1804: 1802: 1798: 1797: 1796:Eoenantiornis 1789: 1788: 1782: 1781: 1778: 1777: 1774: 1773: 1772: 1764: 1763: 1757: 1756: 1753: 1752: 1749: 1747: 1743: 1736: 1735: 1729: 1728: 1725: 1724: 1721: 1720: 1719: 1711: 1710: 1707: 1706: 1703: 1702: 1701: 1693: 1692: 1686: 1685: 1682: 1681: 1678: 1677: 1676: 1668: 1667: 1661: 1660: 1657: 1656: 1653: 1652: 1651: 1650:Eocathayornis 1643: 1642: 1636: 1632: 1631: 1628: 1627: 1624: 1622: 1618: 1611: 1610: 1607: 1606: 1603: 1601: 1597: 1590: 1589: 1586: 1585: 1582: 1580: 1576: 1575: 1567: 1566: 1560: 1556: 1555: 1552: 1551: 1548: 1546: 1542: 1536: 1535: 1529: 1526: 1525: 1521: 1519: 1515: 1510: 1508: 1504: 1500: 1496: 1495:Iberomesornis 1492: 1491:Iberomesornis 1488: 1487: 1486:Iberomesornis 1482: 1481: 1476: 1471: 1469: 1465: 1460: 1458: 1453: 1449: 1445: 1441: 1434:Relationships 1431: 1429: 1428: 1422: 1418: 1413: 1411: 1407: 1403: 1398: 1396: 1395: 1390: 1389: 1384: 1383: 1382:Archaeopteryx 1378: 1374: 1370: 1366: 1356: 1354: 1350: 1349: 1343: 1341: 1336: 1332: 1327: 1325: 1321: 1315: 1313: 1307: 1298: 1294: 1291: 1287: 1285: 1281: 1277: 1276:Maastrichtian 1273: 1268: 1266: 1262: 1258: 1257: 1251: 1247: 1243: 1242: 1236: 1233: 1231: 1227: 1223: 1222: 1217: 1216: 1210: 1208: 1207: 1201: 1197: 1196: 1191: 1187: 1183: 1179: 1171: 1167: 1166: 1160: 1151: 1149: 1145: 1141: 1137: 1132: 1122: 1119: 1118: 1112: 1108: 1104: 1102: 1101: 1096: 1095: 1090: 1086: 1085: 1080: 1075: 1071: 1070: 1065: 1062:from aquatic 1061: 1057: 1056: 1050: 1048: 1044: 1043: 1038: 1037: 1028: 1027: 1022: 1021: 1016: 1015: 1009: 995: 993: 992: 987: 986: 981: 980: 975: 971: 970: 965: 964: 959: 955: 951: 947: 946: 940: 938: 934: 933: 928: 923: 919: 915: 911: 904: 900: 899:longipterygid 896: 895: 889: 880: 878: 877: 872: 871:Archaeopteryx 867: 864: 861: 857: 856:Burmese amber 852: 850: 846: 842: 841:Archaeopteryx 833: 828: 819: 817: 813: 812: 807: 803: 799: 798: 793: 792: 787: 783: 779: 775: 771: 767: 766: 761: 757: 756: 755:Archaeopteryx 746: 745:bohaiornithid 742: 740: 734: 725: 723: 722: 717: 716: 711: 707: 703: 699: 696: 695: 690: 689: 684: 683: 678: 677: 672: 668: 663: 660: 659:Burmese amber 656: 652: 648: 644: 640: 636: 627: 625: 624:Iberomesornis 619: 610: 608: 607: 602: 601: 596: 592: 591: 586: 582: 581: 576: 572: 568: 564: 560: 556: 552: 551: 546: 542: 538: 537:Maastrichtian 534: 530: 525: 523: 519: 515: 511: 510: 500: 498: 492: 490: 489: 482: 478: 476: 475:Alan Feduccia 471: 469: 465: 461: 457: 451: 449: 445: 439: 437: 427: 417: 414: 413:Ancient Greek 409: 407: 406: 401: 400: 395: 394: 393:Iberomesornis 389: 388: 383: 379: 376: 372: 368: 364: 360: 356: 355: 344: 342: 338: 335:, along with 334: 330: 326: 322: 313: 312:and see text 308: 305: 303: 300: 298: 295: 293: 290: 288: 285: 284: 282: 277: 272: 267: 261: 258: 255: 254: 251: 248: 245: 242: 241: 238: 235: 232: 229: 228: 225: 222: 219: 216: 215: 212: 209: 206: 203: 202: 199: 196: 193: 190: 189: 186: 183: 180: 179: 176: 173: 170: 169: 166: 163: 160: 159: 154: 149: 145: 141: 140: 135: 134:bohaiornithid 130: 126: 121: 114: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 53: 46: 42: 33: 30: 19: 16381: 16354: 16342: 16330: 16318: 16228:(mousebirds) 16226:Coliiformes 16096:Gaviiformes 16088:Aequornithes 15996:Apodiformes 15957:Otidiformes 15933:Otidimorphae 15880:(sandgrouse) 15802:Phasianinae 15768: 15761: 15754: 15749:Odontophorus 15747: 15740: 15733: 15726: 15719: 15712: 15692: 15685: 15678: 15671: 15651: 15644: 15637: 15630: 15623: 15616: 15609: 15587:Oreophasinae 15533: 15513: 15506: 15411:(waterfowls) 15407:Anseriformes 15157:Cockfighting 15142:Conservation 15137:Bird feeding 15125:Birdwatching 15115:Ornithomancy 15055:Gansuiformes 15024: 15002: 14995:Fossil birds 14885:Intelligence 14685: 14680:Neuquenornis 14678: 14671: 14664: 14657: 14650: 14639: 14621: 14614: 14607: 14599: 14591: 14576: 14556: 14548: 14541: 14521: 14514: 14507: 14500: 14495:Longusunguis 14493: 14486: 14479: 14472: 14465: 14445: 14438: 14431: 14424: 14417: 14409: 14402: 14395: 14388: 14368: 14361: 14354: 14347: 14340: 14320: 14313: 14293: 14286: 14279: 14272: 14265: 14258: 14251: 14244: 14237: 14230: 14223: 14215: 14207: 14200: 14193: 14186: 14179: 14172: 14165: 14158: 14151: 14144: 14137: 14130: 14123: 14115: 14108: 14101: 14094: 14087: 14079: 14074:Kizylkumavis 14072: 14065: 14058: 14051: 14046:Ilerdopteryx 14044: 14039:Huoshanornis 14037: 14030: 14023: 14016: 14009: 14002: 13995: 13990:Fortunguavis 13988: 13983:Fortipesavis 13981: 13974: 13967: 13960: 13953: 13946: 13939: 13932: 13925: 13918: 13913:Elektorornis 13911: 13904: 13897: 13890: 13883: 13876: 13869: 13862: 13857:Catenoleimus 13855: 13848: 13841: 13834: 13827: 13820: 13813: 13800: 13778: 13777: 13744: 13643: 13634: 13625: 13590: 13547: 13509: 13505: 13499: 13466: 13462: 13455: 13418: 13414: 13408: 13400: 13355: 13351: 13341: 13300: 13296: 13289: 13264: 13260: 13254: 13221: 13217: 13207: 13174: 13170: 13164: 13131: 13127: 13121: 13096: 13092: 13086: 13041: 13037: 13027: 13002: 12996: 12986: 12961: 12957: 12951: 12926: 12922: 12915: 12878: 12874: 12864: 12842:(1): 16–26. 12839: 12835: 12828: 12803: 12799: 12793: 12760: 12756: 12750: 12725: 12721: 12714: 12695: 12691: 12681: 12638: 12634: 12624: 12607: 12564: 12560: 12554: 12535: 12531: 12525: 12521: 12475: 12471: 12461: 12428: 12424: 12418: 12373: 12369: 12359: 12314: 12310: 12300: 12239: 12235: 12225: 12185:(1): 14864. 12182: 12178: 12168: 12158: 12147: 12135: 12131: 12118: 12093: 12089: 12076: 12033: 12029: 12019: 12000: 11996: 11986: 11969: 11965: 11959: 11914: 11910: 11900: 11849: 11845: 11839: 11814: 11810: 11800: 11791: 11787: 11783: 11777: 11771:(1): 112–28. 11768: 11764: 11751: 11726: 11722: 11715: 11704: 11693: 11660: 11656: 11650: 11617: 11613: 11607: 11598: 11594: 11588: 11555: 11507: 11503: 11497: 11480: 11476: 11469: 11460: 11456: 11440: 11431: 11427: 11421: 11412: 11408: 11395: 11381:cite journal 11340: 11336: 11329: 11292: 11288: 11278: 11268: 11258: 11248: 11237:. Retrieved 11235:. 2021-03-12 11227: 11218: 11214: 11204: 11188:(1): 13–27. 11185: 11181: 11171: 11138: 11134: 11128: 11095: 11091: 11085: 11068: 11064: 11012: 11008: 10970: 10966: 10929: 10925: 10871: 10867: 10861: 10817: 10813: 10768: 10764: 10758: 10704: 10700: 10662: 10656: 10637: 10633: 10623: 10591:(1): 62–75. 10588: 10584: 10574: 10549: 10545: 10501: 10497: 10469: 10465: 10411: 10407: 10401: 10374: 10370: 10360: 10323: 10319: 10309: 10276: 10272: 10266: 10231: 10227: 10217: 10184: 10180: 10128: 10124: 10114: 10081: 10077: 10071: 10028: 10024: 9971:(1): 12089. 9968: 9964: 9913: 9909: 9871: 9867: 9837: 9830: 9819:the original 9809: 9807: 9800: 9785: 9781: 9776: 9735: 9731: 9724: 9699: 9695: 9688: 9667: 9659: 9632: 9628: 9618: 9599: 9593: 9560: 9556: 9530: 9526: 9510: 9483: 9479: 9469: 9450: 9444: 9425: 9399: 9395: 9358: 9354: 9317:ossification 9308: 9302: 9280: 9274: 9268: 9264: 9256: 9250: 9247:IVPP V 13939 9246: 9228: 9219:nomen dubium 9217: 9207: 9192: 9188: 9184: 9180: 9176: 9169: 9155: 9151: 9137: 9121: 9081: 9037:Neuquenornis 9035: 8981: 8917: 8866: 8807: 8798: 8745: 8695: 8649: 8585: 8554: 8540: 8499: 8490: 8449: 8403: 8374: 8372: 8316: 8297: 8257: 8247: 8206: 8197: 8157: 8108:Longusunguis 8106: 8060: 8009: 7954: 7908: 7869: 7813: 7767: 7716: 7670: 7619: 7587: 7536: 7485: 7434: 7383: 7332: 7286: 7274:Camptodontus 7273: 7233: 7187: 7156: 7100: 7045: 6999: 6953: 6903: 6888: 6848: 6802: 6790:(shin bone) 6743: 6735:Bad Religion 6731:Greg Graffin 6726: 6686: 6640: 6589: 6543: 6488: 6441: 6399: 6353: 6303: 6296:gut content 6253: 6207: 6161: 6111: 6065: 6004: 5995: 5955: 5946: 5906: 5895: 5847: 5794: 5785: 5781: 5741: 5684:Kizylkumavis 5682: 5631: 5622: 5574: 5528: 5477: 5433:Huoshanornis 5431: 5421: 5381: 5331: 5314: 5274: 5228: 5218: 5178: 5132: 5088:Fortunguavis 5086: 5042:Fortipesavis 5040: 4981: 4930: 4879: 4826: 4780: 4729: 4717: 4677: 4627: 4581: 4537:Elektorornis 4535: 4489: 4443: 4390: 4339: 4288: 4242: 4191: 4137: 4086:Catenoleimus 4084: 4076:Canada goose 4038: 3983: 3932: 3881: 3823: 3777: 3724: 3696: 3678: 3674: 3668: 3664: 3658: 3654: 3648: 3644: 3638: 3634: 3628: 3624: 3618: 3614: 3608: 3604: 3598: 3594: 3588: 3584: 3583: 3580: 3572: 3400: 3399: 3383: 3382: 3359: 3358: 3337:Fortunguavis 3335: 3334: 3311: 3310: 3298: 3284: 3283: 3257: 3256: 3224: 3223: 3209:Huoshanornis 3207: 3206: 3149: 3148: 3132: 3131: 3108: 3107: 3084: 3083: 3060: 3023: 3022: 3006: 3005: 2982: 2981: 2969: 2955: 2954: 2903: 2902: 2897: 2883: 2882: 2877: 2867: 2855:Longusunguis 2853: 2852: 2847: 2819: 2818: 2793: 2756: 2755: 2750: 2736: 2735: 2712: 2711: 2679: 2678: 2662: 2661: 2628: 2627: 2597: 2596: 2546: 2545: 2529: 2528: 2496: 2495: 2479: 2478: 2466: 2456: 2417:Neuquenornis 2415: 2414: 2398: 2397: 2374: 2373: 2350: 2349: 2337: 2323: 2322: 2317: 2307: 2265: 2264: 2248: 2247: 2242: 2232: 2218: 2217: 2175: 2174: 2158: 2157: 2127: 2126: 2103: 2102: 2082: 2080: 1961: 1959: 1943: 1941: 1913: 1911: 1890: 1888: 1874:Neuquenornis 1872: 1870: 1854: 1852: 1824: 1822: 1794: 1792: 1769: 1767: 1739: 1718:Fortunguavis 1716: 1714: 1698: 1696: 1673: 1671: 1648: 1646: 1634: 1614: 1593: 1572: 1570: 1558: 1539: 1517: 1511: 1494: 1490: 1484: 1478: 1474: 1472: 1461: 1452:phylogenetic 1437: 1427:nomina dubia 1425: 1414: 1399: 1392: 1386: 1380: 1362: 1346: 1344: 1328: 1320:avian flight 1316: 1308: 1304: 1295: 1292: 1288: 1269: 1254: 1239: 1237: 1234: 1219: 1213: 1211: 1206:nomen dubium 1204: 1200:Luis Chiappe 1198:"; however, 1193: 1175: 1163: 1154:Life history 1130: 1128: 1115: 1109: 1105: 1098: 1092: 1082: 1067: 1060:exoskeletons 1053: 1051: 1040: 1034: 1032: 1024: 1018: 1012: 989: 983: 977: 974:pengornithid 967: 961: 953: 943: 941: 930: 926: 909: 907: 892: 874: 870: 868: 853: 840: 837: 809: 795: 789: 763: 753: 750: 737: 720: 714: 708:by a single 698: 693: 686: 680: 674: 664: 631: 622: 604: 598: 588: 578: 548: 526: 518:Kimmeridgian 507: 506: 493: 486: 484: 480: 472: 463: 459: 453: 441: 425: 415: 410: 403: 397: 391: 385: 352: 350: 329:Mesozoic era 320: 318: 311: 265: 256: 243: 230: 217: 204: 191: 137: 50:136–66  29: 16406:Wikispecies 16356:WikiProject 16266:Piciformes 16149:Australaves 16128:Suliformes 16046:Gruiformes 15901:(flamingos) 15810:Tetraoninae 15781:Phasianidae 15770:Rhynchortyx 15735:Dactylortyx 15602:Megapodidae 15592:Penelopinae 15550:Galliformes 15292:Rheiformes 15286:(ostriches) 15236:individuals 15110:Ornithology 15097:interaction 14852:Preen gland 14634:Avisauridae 14593:Bauxitornis 14570:Avisauridae 14509:Shenqiornis 14481:Gretcheniao 14440:Shanweiniao 14419:Longipteryx 14370:Yuanchuavis 14349:Eopengornis 14342:Chiappeavis 14322:Noguerornis 14232:Pterygornis 14225:Protopteryx 14174:Orienantius 14117:Liaoxiornis 14011:Gracilornis 14004:Grabauornis 13976:Flexomornis 13962:Falcatakely 13955:Explorornis 13927:Enantiornis 13906:Elbretornis 13864:Cathayornis 13733:Maniraptora 13718:Maniraptora 13267:: 210–219. 12478:(1): 1116. 12267:10161/11144 12096:(1): 1–42. 12036:(1): 5006. 11434:(2): 246–8. 11289:BMC Biology 11221:(2): 73–78. 11141:: 178–195. 10377:(1): 7–19. 9874:: 264–277. 9402:(1): 31–76. 9303:DIP-V-15102 9293:melanosomes 9269:DIP-V-15101 9265:DIP-V-15100 9252:Microraptor 9194:nomen nudum 8556:Avisauridae 8551:Avisauridae 8298:Shenqiornis 8208:Shenqiornis 8011:Gretcheniao 7995:gastroliths 7815:Yuanchuavis 7691:Hauterivian 7672:Eopengornis 7621:Chiappeavis 7487:Shanweiniao 7334:Longipteryx 6889:Cathayornis 6788:tibiotarsus 6727:Q. graffini 6642:Pterygornis 6610:Hauterivian 6591:Protopteryx 6374:Hauterivian 6355:Orienantius 6344:propatagium 6305:Noguerornis 6294:ichthyosaur 5996:Cathayornis 5897:Ichthyornis 5786:Cathayornis 5623:Enantiornis 5422:Cathayornis 5219:Cathayornis 5180:Gracilornis 5134:Grabauornis 4983:Flexomornis 4881:Falcatakely 4828:Explorornis 4719:Cathayornis 4583:Enantiornis 4491:Elbretornis 4309:Hauterivian 4139:Cathayornis 3934:Bauxitornis 3868:woodpeckers 3650:Pterygornis 3630:Longipteryx 3600:Cathayornis 3385:Shenqiornis 3151:Gretcheniao 3086:Grabauornis 3008:Eopengornis 2984:Yuanchuavis 2643:Avisauridae 2352:Enantiornis 2177:Flexomornis 2129:Protopteryx 1826:Cathayornis 1771:Pterygornis 1574:Protopteryx 1503:Paul Sereno 1464:homoplastic 1421:Gareth Dyke 1369:crown group 1170:Dinosaurium 1064:crustaceans 1036:Shenqiornis 1014:Longipteryx 969:Chiappeavis 954:Shanweiniao 945:Shanweiniao 876:Microraptor 811:Pterygornis 791:Shenqiornis 786:postorbital 760:premaxillae 647:Jehol group 613:Description 606:Enantiornis 590:Protopteryx 580:Noguerornis 512:, from the 399:Cathayornis 387:Enantiornis 361:related to 287:Avisauridae 16456:Categories 16106:(penguins) 15959:(bustards) 15797:Perdicinae 15742:Dendrortyx 15714:Callipepla 15646:Megapodius 15611:Aepypodius 15489:Tadorninae 15471:true geese 15302:(tinamous) 15261:Neornithes 15172:Pheasantry 15147:Aviculture 14915:Incubation 14905:Lek mating 14623:Mystiornis 14616:Halimornis 14543:Gobipteryx 14488:Linyiornis 14474:Bohaiornis 14467:Beiguornis 14267:Xiangornis 14209:Platanavis 14132:Martinavis 14103:Lenesornis 14060:Incolornis 13969:Feitianius 13934:Eoalulavis 13899:Dunhuangia 13754:see below↓ 13630:Dinosauria 13621:Sauropsida 13409:Bohaiornis 13224:: 105452. 12138:(1): 9–24. 12003:: 654156. 11295:(1): 101. 11239:2021-05-06 10084:: 104412. 10031:(1): 927. 9332:References 9297:iridescent 9281:CUGB P1202 9258:Anchiornis 9233:El Montsec 9189:Hebeiornis 9171:Hebeiornis 9152:Gobipteryx 8983:Mystiornis 8809:Halimornis 8716:Cenomanian 8571:Formation 8405:Gobipteryx 8389:Formation 8248:Bohaiornis 8198:Bohaiornis 8062:Linyiornis 7956:Bohaiornis 7910:Beiguornis 7894:Formation 7605:Formation 7173:Formation 6905:Xiangornis 6782:Uzbekistan 6316:El Montsec 6006:Martinavis 5947:Eoalulavis 5886:Uzbekistan 5849:Lenesornis 5721:Uzbekistan 5613:Uzbekistan 5576:Incolornis 5061:Cenomanian 5002:Cenomanian 4932:Feitianius 4914:Madagascar 4865:Uzbekistan 4629:Eoalulavis 4556:Cenomanian 4445:Dunhuangia 4123:Uzbekistan 3763:Uzbekistan 3710:Formation 3361:Mystiornis 3286:Bohaiornis 2885:Dunhuangia 2821:Feitianius 2548:Hebeiornis 2531:Gobipteryx 2481:Eoalulavis 2376:Halimornis 1915:Eoalulavis 1892:Gobipteryx 1700:Dunhuangia 1541:Euornithes 1489:. Because 1440:Euornithes 1406:Euornithes 1373:neornithes 1353:flightless 1324:lithornids 1261:endothermy 1256:Jeholornis 1215:Gobipteryx 1186:hatchlings 1148:pan-avians 1100:Eoalulavis 1094:Bohaiornis 1074:sapsuckers 1055:Eoalulavis 1023:and right 1020:Bohaiornis 985:Feitianius 950:Euronithes 918:Ornithurae 914:Euornithes 739:Bohaiornis 563:Cretaceous 559:Antarctica 522:Kazakhstan 375:Cretaceous 359:paleognath 279:Subgroups 211:Saurischia 198:Dinosauria 16175:(parrots) 16019:(hoatzin) 15970:Strisores 15951:(turacos) 15943:(cuckoos) 15872:(mesites) 15791:(turkeys) 15763:Philortyx 15680:Agelastes 15673:Acryllium 15664:Numididae 15653:Talegalla 15563:gamebirds 15556:landfowls 15535:Anseranas 15499:Anhimidae 15458:Anserinae 15241:fictional 14963:dinosaurs 14958:Theropoda 14945:Evolution 14890:Migration 14872:Behaviour 14666:Intiornis 14652:Avisaurus 14601:Concornis 14558:Vescornis 14523:Zhouornis 14433:Rapaxavis 14411:Evgenavis 14390:Boluochia 14363:Pengornis 14274:Yatenavis 14253:Shangyang 14217:Praeornis 14181:Otogornis 14167:Nanantius 14160:Musivavis 14146:Mirusavis 14081:Kuszholia 14053:Imparavis 14018:Gurilynia 13997:Gobipipus 13871:Cratoavis 13829:Alexornis 13815:Abavornis 13706:Theropoda 13691:Theropoda 13639:Theropoda 13607:Kingdom: 13421:: e7846. 13333:227174405 13246:254804249 13238:0195-6671 13019:0195-6671 12881:: e7247. 12820:131170912 12728:: 56–65. 12673:247432530 12665:0022-3360 12569:CiteSeerX 12333:2167-8359 12317:: e5910. 12276:1879-0445 11972:: 24–44. 11862:CiteSeerX 11831:245082389 11572:2246/5890 11463:: 147–79. 11273:84643293. 11263:14740099. 11253:84817636. 11163:133781513 11120:130143737 11077:2246/3667 11015:: 14141. 10842:196607241 10436:0272-4634 10326:: e5910. 10163:195887085 10147:0960-9822 10106:213510021 9651:236397491 9533:: 103–9. 9325:pygostyle 9313:Las Hoyas 9287:from the 9241:histology 9185:Vescornis 9181:Vescornis 9177:Vescornis 9156:Gobipipus 9122:Avisaurus 9116:Argentina 9070:Argentina 9056:Santonian 9002:Barremian 8938:Campanian 8901:Argentina 8887:Campanian 8868:Intiornis 8828:Campanian 8766:Campanian 8670:Campanian 8587:Avisaurus 8574:Location 8501:Vescornis 8491:Vescornis 8424:Campanian 8392:Location 8318:Zhouornis 8030:Barremian 7897:Location 7876:monophyly 7769:Pengornis 7608:Location 7436:Rapaxavis 7189:Boluochia 7176:Location 7080:Argentina 6988:Argentina 6955:Yatenavis 6804:Shangyang 6768:Coniacian 6513:Santonian 6401:Otogornis 6324:Barremian 6288:Australia 6255:Nanantius 6209:Musivavis 6132:Barremian 6113:Mirusavis 6054:Argentina 5892:synsacrum 5872:Coniacian 5829:Argentina 5729:fragment 5707:Coniacian 5619:coracoids 5599:Coniacian 5530:Imparavis 5498:Barremian 5490:Las Hoyas 5372:pterosaur 5295:Campanian 5230:Gurilynia 5153:Barremian 4871:coracoids 4851:Coniacian 4801:Barremian 4782:Evgenavis 4648:Barremian 4640:Las Hoyas 4616:Argentina 4524:Argentina 4244:Cratoavis 4212:Barremian 4204:Las Hoyas 4193:Concornis 4109:Coniacian 4056:Campanian 3953:Santonian 3844:Campanian 3825:Alexornis 3769:coracoids 3749:Coniacian 3726:Abavornis 3713:Location 3590:Boluochia 3259:Zhouornis 3134:Cratoavis 2957:Pengornis 2905:Musivavis 2714:Avisaurus 2664:Intiornis 2400:Concornis 2267:Rapaxavis 1963:Concornis 1856:Vescornis 1514:cladogram 1394:Sapeornis 1345:At least 1335:pygostyle 1312:rectrices 1284:megapodes 1250:altricial 1241:Concornis 1226:precocial 1221:Gobipipus 1138:or small 1136:pterosaur 1125:Predation 1042:Pengornis 1026:Pengornis 1017:, centre 979:Sapeornis 922:pygostyle 903:pygostyle 797:Pengornis 635:Las Hoyas 600:Avisaurus 550:Alexornis 541:Campanian 533:Australia 514:Oxfordian 509:Praeornis 382:Argentina 363:ostriches 341:dinosaurs 224:Theropoda 171:Kingdom: 165:Eukaryota 16426:42331729 16391:Wikidata 16320:Category 16194:Afroaves 16122:(storks) 15924:Passerea 15909:(grebes) 15845:Columbea 15756:Oreortyx 15728:Cyrtonyx 15618:Alectura 15582:Cracinae 15574:Cracidae 15451:Oxyurini 15441:Aythyini 15436:Anatinae 15420:Anatidae 15167:Falconry 15130:big year 14985:Seabirds 14895:Foraging 14842:Feathers 14734:Category 14550:Jibeinia 14516:Sulcavis 14260:Sinornis 14239:Qiliania 14195:Parvavis 14096:Lectavis 14067:Junornis 14032:Houornis 14025:Holbotia 13920:Elsornis 13615:Chordata 13613:Phylum: 13609:Animalia 13534:86747842 13491:85807045 13447:31667014 13392:26039693 13352:PLOS ONE 13325:33239782 13199:89858642 13156:85255564 13113:89551799 13078:29020077 13038:PLOS ONE 12978:86665059 12964:: 1–19. 12943:85784633 12907:31333906 12785:85313872 12612:Archived 12591:52971055 12538:: 7–20. 12502:23047674 12453:85391743 12410:22163324 12370:PLOS ONE 12351:30479894 12284:26439352 12217:26440221 12148:Phys.org 12110:86007060 12068:30899080 11951:23613776 11911:PLOS ONE 11884:22575918 11794:: 55–63. 11685:86747842 11642:86747842 11601:: 37–53. 11580:85871695 11532:34504916 11524:15499011 11448:(1981). 11365:11234054 11321:35550084 11047:28139644 10987:85800831 10948:26748849 10900:24482756 10874:: e234. 10795:16533313 10731:15496911 10679:88601510 10607:28000410 10566:83603202 10526:86503357 10444:85255564 10352:30479894 10301:85942925 10258:18397240 10209:84677039 10155:31303484 10063:30700773 10003:27352215 9940:36659012 9846:Archived 9768:42723583 9760:16778053 9730:China". 9539:53487062 9518:(1974). 9502:55311115 9275:HPG-15-1 9202:Proornis 9148:Mongolia 8684:Mongolia 8651:Elsornis 8541:Jibeinia 8451:Jibeinia 8438:Mongolia 8259:Sulcavis 6850:Sinornis 6764:Turonian 6688:Qiliania 6509:Turonian 6490:Parvavis 5868:Turonian 5796:Lectavis 5703:Turonian 5633:Junornis 5595:Turonian 5383:Houornis 5366:Mongolia 5333:Holbotia 5316:Lectavis 5309:Mongolia 5276:Hollanda 5263:Mongolia 4847:Turonian 4129:coracoid 4105:Turonian 3745:Turonian 3620:Houornis 3402:Sulcavis 3110:Parvavis 2630:Qiliania 2325:Sinornis 2160:Elsornis 1945:Qiliania 1675:Elsornis 1483:than to 1480:Sinornis 1348:Elsornis 1246:fledging 1172:(Prague) 1144:Mesozoic 1140:theropod 863:dinosaur 860:Mesozoic 832:micro-CT 671:starling 651:Liaoning 645:and the 567:dinosaur 555:Mongolia 464:coracoid 456:shoulder 448:coracoid 446:and the 426:ornithes 421:ἐνάντιος 416:enantios 405:Sinornis 325:avialans 185:Chordata 181:Phylum: 175:Animalia 161:Domain: 16397:Q132268 16332:Commons 15836:Neoaves 15721:Colinus 15687:Guttera 15625:Eulipoa 15446:Mergini 15310:(kiwis) 15294:(rheas) 15105:Ringing 14930:Hybrids 14925:Nesting 14880:Singing 14857:Plumage 14832:Dactyly 14802:Anatomy 14792:Outline 14782:: Aves) 14673:Mirarce 14659:Gettyia 14295:Yuornis 14246:Sazavis 13836:Avimaia 13619:Class: 13514:Bibcode 13471:Bibcode 13438:6816414 13383:4454694 13360:Bibcode 13305:Bibcode 13269:Bibcode 13179:Bibcode 13136:Bibcode 13069:5636078 13046:Bibcode 12898:6626525 12844:Bibcode 12765:Bibcode 12730:Bibcode 12643:Bibcode 12480:Bibcode 12433:Bibcode 12401:3233598 12378:Bibcode 12342:6238772 12292:3099017 12244:Bibcode 12208:4594305 12187:Bibcode 12059:6428842 12038:Bibcode 11942:3629076 11919:Bibcode 11892:1396792 11854:Bibcode 11731:Bibcode 11665:Bibcode 11622:Bibcode 11504:Science 11477:Science 11345:Bibcode 11312:9097364 11143:Bibcode 11112:4524284 11038:5290326 11017:Bibcode 10891:3898307 10822:Bibcode 10786:2100246 10739:4322054 10709:Bibcode 10615:3649504 10506:Bibcode 10416:Bibcode 10379:Bibcode 10343:6238772 10281:Bibcode 10249:2409080 10189:Bibcode 10086:Bibcode 10054:6353931 10033:Bibcode 9994:4931330 9973:Bibcode 9918:Bibcode 9876:Bibcode 9740:Bibcode 9732:Science 9704:Bibcode 9585:4340858 9565:Bibcode 9321:sternum 9319:of the 9229:LP-4450 9026:Paraves 8919:Mirarce 8854:Alabama 8792:Montana 8747:Gettyia 8730:Lebanon 8632:Montana 8580:Images 8398:Images 7903:Images 7614:Images 7182:Images 7102:Yuornis 6745:Sazavis 5727:humerus 5075:Myanmar 4570:Myanmar 3967:Hungary 3883:Avimaia 3864:rollers 3719:Images 2758:Mirarce 2738:Gettyia 1446:called 1377:Avialae 1340:furcula 1338:narrow 1280:Romania 1182:embryos 1089:ovaries 1079:pellets 998:Biology 908:Clarke 710:furcula 667:sparrow 468:concave 460:scapula 444:scapula 431:ὄρνιθΡς 367:tinamou 237:Avialae 43:– 16344:Portal 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Index

Enantiornithine
Early Cretaceous
Late Cretaceous
Ma
Preκž’
κž’
O
S
D
C
P
T
J
K
Pg
N

bohaiornithid
Zhouornis hani
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Saurischia
Theropoda
Avialae
Ornithothoraces
Enantiornithes

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