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Axolotl

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including eyes and parts of the brain—restoring these alien organs to full functionality. In some cases, axolotls have been known to repair a damaged limb, as well as regenerating an additional one, ending up with an extra appendage that makes them attractive to pet owners as a novelty. In metamorphosed individuals, however, the ability to regenerate is greatly diminished. The axolotl is therefore used as a model for the development of limbs in vertebrates. There are three basic requirements for regeneration of the limb: the wound epithelium, nerve signaling, and the presence of cells from the different limb axes. A wound epidermis is quickly formed by the cells to cover up the site of the wound. In the following days, the cells of the wound epidermis divide and grow quickly forming a blastema, which means the wound is ready to heal and undergo patterning to form the new limb.
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extinction, something that they are already in danger of. Studies have found indicators of a low interpopulation gene flow and higher rates of genetic drift. These are likely the result of multiple “bottleneck” incidents in which events that kill off several individuals of a population occur and sharply reduce the genetic diversity of the remaining population. The offspring produced after bottleneck events have a greater risk of showing decreased fitness and are often less capable of adaptation down the line. Multiple bottleneck events can have disastrous effects on a population. Studies have also found high rates of relatedness that are indicative of inbreeding. Inbreeding can be especially harmful as it can cause an increase in the presence of deleterious, or harmful, genes within a population. The detection of introgressed tiger salamander (
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Ecologica (LRE) in the Universidad Nacional Autonoma de Mexico (UNAM) has built up a population of more than 100 captive-bred individuals. These axolotls are mostly used for research by the lab but plans of a semi-artificial wetland inside the university have been established and the goal is to establish a viable population of axolotls within it. Studies have shown that captive-bred axolotls that are raised in a semi-natural environment can catch prey, survive in the wild, and have moderate success in escaping predators. These captive-bred individuals can be introduced into unpolluted bodies of water or back into Lake Xochimilco to establish or re-establish a wild population.
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oxygen-poor environment that is not well-suited for axolotls. In addition, the intensive use of pesticides from agriculture around Lake Xochimilco causes run off into the lake and a reduction of habitat quality for axolotls. The pesticides used contain chemical compounds that studies show to sharply increase mortality in axolotl embryos and larvae. Of the surviving embryo and larvae, there is also an increase of morphological, behavior, and activity abnormalities.
114: 1004:, though the axolotl differs in its longer toes. Among hobbyists, the process of artificially inducing metamorphosis can often result in death during or even following a successful attempt, and so casual hobbyists are generally discouraged from attempting to induce metamorphosis in pet axolotls. Morphed pet axolotls should be given solid footholds in their enclosure to satisfy their need for land. They should not be given live animals as food. 680: 1225:
can be harmful for amphibians and reptiles. Gravel (common in aquarium use) should not be used, and is recommended that any sand consists of smooth particles with a grain size of under 1mm. One guide to axolotl care for laboratories notes that bowel obstructions are a common cause of death, and recommends that no items with a diameter below 3 cm (or approximately the size of the animal's head) should be available to the animal.
629: 566: 550: 699:. Lake Chalco no longer exists, having been drained as a flood control measure, and Lake Xochimilco remains a remnant of its former self, existing mainly as canals. The water temperature in Xochimilco rarely rises above 20 Â°C (68 Â°F), although it may fall to 6–7 Â°C (43–45 Â°F) in the winter, and perhaps lower. 1224:
Substrates are another important consideration for captive axolotls, as axolotls (like other amphibians and reptiles) tend to ingest bedding material together with food and are commonly prone to gastrointestinal obstruction and foreign body ingestion. Some common substrates used for animal enclosures
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In addition, there is wide individual variability in the size, frequency, and intensity of the gold speckling, and at least one variant develops a black and white piebald appearance upon reaching maturity. Because pet breeders frequently cross the variant colors, double homozygous mutants are common
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organisms, lower temperatures result in slower metabolism and a very unhealthily reduced appetite. Temperatures at approximately 16 Â°C (61 Â°F) to 18 Â°C (64 Â°F) are suggested for captive axolotls to ensure sufficient food intake; stress resulting from more than a day's exposure to
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In the absence of induced metamorphosis, larval axolotls start absorbing iodide into their thyroid glands at 30 days postfertilization. Larval axolotls do produce thyroid hormone from iodide, but the amount appears highly variable. Adult axolotls do not produce thyroid hormone unless metamorphism is
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Another factor that threatens the native axolotl population is the introduction of invasive species such as the Nile tilapia and common carp. These invasive fish species threaten axolotl populations by eating their eggs or young and by out-competing them for natural resources. The presence of these
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limbs, gills and parts of their eyes and brains. Notably, their ability to regenerate declines with age, but it does not disappear. Axolotls keep modestly growing throughout their life and some consider this trait to be a direct contributor to their regenerative abilities. Further research has been
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Lake Xochimilco has poor water quality, caused by the region's aquaculture and agriculture demands. It is also maintained by inputs of only partially treated wastewater. Water quality tests reveal a low nitrogen-phosphorus ratio and a high concentration of chlorophyll a, which are indicative of an
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families in which it seems to be a survival mechanism, in aquatic environments only of mountain and hill, with little food and, in particular, with little iodine. In this way, salamanders can reproduce and survive in the form of a smaller larval stage, which is aquatic and requires a lower quality
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Caballero-Pérez, Juan; Espinal-Centeno, Annie; Falcon, Francisco; García-Ortega, Luis F.; Curiel-Quesada, Everardo; Cruz-Hernåndez, Andrés; Bako, Laszlo; Chen, Xuemei; Martínez, Octavio; Alberto Arteaga-Våzquez, Mario; Herrera-Estrella, Luis (January 2018). "Transcriptional landscapes of Axolotl
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Axolotls are only native to the Mexican Central Valley. Although the native axolotl population once extended through most of the lakes and wetlands that make up this region, the native habitat is now limited to Lake Xochimilco as a result of the expansion of Mexico City. Lake Xochimilco is not a
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Surveys in 1998, 2003, and 2008 found 6,000, 1,000, and 100 axolotls per square kilometer in its Lake Xochimilco habitat, respectively. A four-month-long search in 2013, however, turned up no surviving individuals in the wild. Just a month later, two wild ones were spotted in a network of canals
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in the pet trade, especially white/pink animals with pink eyes that are double homozygous mutants for both the albino and leucistic trait. Axolotls also have some limited ability to alter their color to provide better camouflage by changing the relative size and thickness of their melanophores.
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With such a small native population, there is a large loss of genetic diversity. This lack of genetic diversity can be dangerous for the remaining population, causing an increase in inbreeding and a decrease in general fitness and adaptive potential. It ultimately raises the axolotl's risk for
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An axolotl undergoing metamorphosis experiences a number of physiological changes that help them adapt to life on land. These include increased muscle tone in limbs, the absorption of gills and fins into the body, the development of eyelids, and a reduction in the skin's permeability to water,
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of entire lost appendages in a period of months, and, in certain cases, more vital structures, such as tail, limb, central nervous system, and tissues of the eye and heart. They can even restore less vital parts of their brains. They can also readily accept transplants from other individuals,
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There has been little improvement in the conditions of the lake or the population of native axolotls. Many scientists are focusing their conservation efforts on translocation of captive-bred individuals into new habitats or reintroduction into Lake Xochimilco. The Laboratorio de Restauracion
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and a caudal fin extending from behind the head to the vent. External gills are usually lost when salamander species mature into adulthood, although the axolotl maintains this feature. This is due to their neoteny evolution, where axolotls are much more aquatic than other salamander species.
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The axolotl's body has the capacity to go through metamorphosis if given the necessary hormone, but axolotls do not produce it, and must be exposed to it from an external source, after which an axolotl undergoes an artificially-induced metamorphosis and begins living on land. In laboratory
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A sexually mature adult axolotl, at age 18–27 months, ranges in length from 15 to 45 cm (6 to 18 in), although a size close to 23 cm (9 in) is most common and greater than 30 cm (12 in) is rare. Axolotls possess features typical of salamander larvae, including
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Björklund, N.K. (1993). Small is beautiful: economical axolotl colony maintenance with natural spawnings as if axolotls mattered. In: Handbook on Practical Methods. Ed.: G.M. Malacinski & S.T. Duhon. Bloomington, Department of Biology, Indiana University:
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Woodcock, M. Ryan; Vaughn-Wolfe, Jennifer; Elias, Alexandra; Kump, D. Kevin; Kendall, Katharina Denise; Timoshevskaya, Nataliya; Timoshevskiy, Vladimir; Perry, Dustin W.; Smith, Jeramiah J.; Spiewak, Jessica E.; Parichy, David M.; Voss, S. Randal (2017-01-31).
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The axolotl is carnivorous, consuming small prey such as mollusks, worms, insects, other arthropods, and small fish in the wild. Axolotls locate food by smell, and will "snap" at any potential meal, sucking the food into their stomachs with vacuum force.
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studies due to the presence of a mutant gene that causes heart failure in embryos. Since the embryos survive almost to hatching with no heart function, the defect is very observable. The axolotl is also considered an ideal animal model for the study of
625:(gulping air from the surface) may also be used to provide oxygen to their lungs. Buccal pumping can occur in a two-stroke manner that pumps air from the mouth to the lungs, and with four-stroke that reverses this pathway with compression forces. 644:
Axolotls have four pigmentation genes; when mutated, they create different color variants. The normal wild-type animal is brown or tan with gold speckles and an olive undertone. The five most common mutant colors are listed below.
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This animal was X-rayed several times as part of a research project over a period of two years. It was a normal healthy adult (26.3 cm; 159.5 gm) at the beginning of the project and lived several more years after the project
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specimens were not included in the original shipment. Vilem Laufberger in Prague used thyroid hormone injections to induce an axolotl to grow into a terrestrial adult salamander. The experiment was repeated by Englishman
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species has also been shown to change the behavior of axolotls, causing them to be less active to avoid predation. This reduction in activity greatly impacts the axolotls foraging and mating opportunities.
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allowing the axolotl to stay more easily hydrated when on land. The lungs of an axolotl, though present alongside gills after reaching non-metamorphosed adulthood, develop further during metamorphosis.
414: 795:. There are also mutations affecting other organ systems some of which are not well characterized and others that are. The genetics of the color variants of the axolotl have also been widely studied. 1026:
was surprised when, instead of the axolotl, he found in the vivarium a new species, similar to the salamander. This discovery was the starting point of research about neoteny. It is not certain that
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Safi, Rachid; Bertrand, Stéphanie; Marchand, Oriane; Duffraisse, Marilyne; de Luze, Amaury; Vanacker, Jean-Marc; Maraninchi, Marie; Margotat, Alain; Demeneix, Barbara; Laudet, Vincent (2004-02-01).
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Kulbisky, Gordon P; Rickey, Daniel W; Reed, Martin H; Björklund, Natalie; Gordon, Richard (1999). "The axolotl as an animal model for the comparison of 3-D ultrasound with plain film radiography".
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originate behind their heads and are used to move oxygenated water. The external gill rami are lined with filaments (fimbriae) to increase surface area for gas exchange. Four-gill slits lined with
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based on experiments conducted at the University of Manitoba axolotl colony,. As there is no conclusive evidence pointing to gastrolith use, gravel should be avoided due to the high risk of
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lower temperatures may quickly lead to disease and death, and temperatures higher than 24 Â°C (75 Â°F) may lead to metabolic rate increase, also causing stress and eventually death.
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In animals with functioning thyroid glands, iodine in the form of iodide is selectively gathered into the colloid of the thyroid. Inside the colloid, iodide is reduced to elemental iodine (I
436: 3972:"And then we also found out that axolotls are endangered in the real world, and we think it's good to add endangered animals to Minecraft to create awareness about that." - Agnes Larsson 3747: 4058: 1067:, they quickly begin metamorphosis and transform into bigger terrestrial adults, with higher dietary requirements. In fact, in some high mountain lakes there live dwarf forms of 2129: 522:
conducted to examine their heart as a model of human single ventricle and excessive trabeculation. Axolotls were also sold as food in Mexican markets and were a staple in the
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Meyer, Sophie; Lauridsen, Henrik; Pedersen, Kathrine; Andersson, Sofie Amalie; van Ooij, Pim; Willems, Tineke; Berger, Rolf M. F.; Ebels, Tjark; Jensen, Bjarke (2022-11-28).
3782: 847:, it encodes a similar number of proteins, namely 23,251 (the human genome encodes about 20,000 proteins). The size difference is mostly explained by a large fraction of 735:. Their habitat is like that of most neotenic species—a high-altitude body of water surrounded by a risky terrestrial environment. These conditions are thought to favor 2555:
Nowoshilow, Sergej; Schloissnig, Siegfried; Fei, Ji-Feng; Dahl, Andreas; Pang, Andy W. C.; Pippel, Martin; Winkler, Sylke; Hastie, Alex R.; Young, George (2018-01-24).
2496:"Modulating the immune response and the pericardial environment with LPS or prednisolone in the axolotl does not change the regenerative capacity of cryoinjured hearts" 991:, which does not react with proteins, does not trigger metamorphosis. It does speed up the rate of metamorphosis, once it has been triggered by thyroid hormone extract. 3727: 1541: 1127:) DNA in the laboratory axolotl population raises further concerns about the suitability of the captive population as an ark for potential reintroduction purposes. 1063:
and quantity of food compared to the big adult, which is terrestrial. If the salamander larvae ingest a sufficient amount of iodine, directly or indirectly through
1780: 1186:, is harmful to axolotls. A single axolotl typically requires a 150-litre (40-US-gallon) tank. Axolotls spend the majority of the time at the bottom of the tank. 514: 2734:
Demircan, Turan; Ovezmyradov, Guvanch; Yıldırım, Berna; Keskin, Ä°lknur; Ä°lhan, AyƟe Elif; Fesçioğlu, Ece Cana; ÖztĂŒrk, GĂŒrkan; Yıldırım, SĂŒleyman (2018-07-20).
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Respiration and metabolism of embryonic vertebrates: Satellite Symposium of the 29th International Congress of Physiological Sciences, Sydney, Australia, 1983
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Wainwright, P. C.; Sanford, C. P.; Reilly, S. M.; Lauder, G. V. (1989). "Evolution of motor patterns: aquatic feeding in salamanders and ray-finned fishes".
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Masselink, Wouter, and Elly M. Tanaka. "Toward Whole Tissue Imaging of Axolotl Regeneration." Developmental Dynamics, vol. 250, no. 6, 2020, pp. 800–806.,
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in order for the axolotl to go through metamorphosis; therefore, it keeps its gills and lives in water all its life, even after it becomes an adult and is
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Their heads are wide, and their eyes are lidless. Their limbs are underdeveloped and possess long, thin digits. Males are identified by their swollen
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is the cause of complete neoteny in laboratory and pet axolotls. The genes responsible have been narrowed down to a small chromosomal region called
4506: 710:. The axolotl is currently on the International Union for Conservation of Nature's annual Red List of threatened species. Non-native fish, such as 718:, have also recently been introduced to the waters. These new fish have been eating the axolotls' young, as well as their primary source of food. 675:
Lake Xochimilco, Mexico City (Amanecer en Xochimilco). The native habitat of axolotls is important to the study of preservation and conservation.
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are other neotenic salamanders, although unlike axolotls, they cannot be induced to metamorphose by an injection of iodine or thyroxine hormone.
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The genes responsible for neoteny in laboratory animals may have been identified; however, they are not linked in wild populations, suggesting
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spp.), fully aquatic salamanders from a different family that are not closely related to the axolotl but bear a superficial resemblance.
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Pedersen, Katherine; Rasmussen, Rikke Kongsgaard; Dittrich, Anita; Pedersen, Michael; Lauridsen, Henrik (April 17, 2020).
1617: 1323: 2239:"A review of the theories of vertebrate neurulation and their relationship to the mechanics of neural tube birth defects" 17: 3517: 4511: 4501: 3316:"Identification of Mutant Genes and Introgressed Tiger Salamander DNA in the Laboratory Axolotl, Ambystoma mexicanum" 2805: 2001:
Frost, Sally K.; Briggs, Fran; Malacinski, George M. (1984). "A color atlas of pigment genes in the Mexican axolotl (
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In captivity, axolotls eat a variety of readily available foods, including trout and salmon pellets, frozen or live
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are hidden underneath the external gills, which prevent food from entering and allow particles to filter through.
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Robles-Mendoza, C.; GarcĂ­a-Basilio, C.; Cram-Heydrich, S.; HernĂĄndez-Quiroz, M.; Vanegas-PĂ©rez, C. (2009-02-01).
1279: 621:, during which their rakers interlock to close the gill slits. External gills are used for respiration, although 3860: 4221: 739:. However, a terrestrial population of Mexican tiger salamanders occupies and breeds in the axolotl's habitat. 4213: 2625: 1045:
Many other species within the axolotl's genus are also either entirely neotenic or have neotenic populations.
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that may be responsible for limb regeneration. Although the axolotl genome is about 10 times as large as the
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The feature of the axolotl that attracts most attention is its healing ability: the axolotl does not heal by
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and tube formation; the axolotl's neural tube, unlike the frog's, is not hidden under a layer of superficial
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large body of water, but rather a small series of artificial channels, small lakes, and temporary wetlands.
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was published in 2018 and was the largest animal genome completed at the time. It revealed species-specific
4496: 3174:"Organophosphorus pesticides effect on early stages of the axolotl Ambystoma mexicanum (Amphibia: Caudata)" 2977:
Krylov, O. A. (January 1961). "The role of haloids (bromine and iodine) in the metamorphosis of amphibia".
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Ingram, W. R. (1 December 1928). "Metamorphosis of the Colorado Axolotl by Injection of Inorganic Iodine".
3132: 1343:' trend of adding endangered species to the game to raise awareness. They were also added to its spin-off 1313: 4322: 4244: 934: 894: 4249: 4103: 971:
precursor in thyroxine biosynthesis, causes metamorphosis in axolotls that have their thyroids removed.
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It is believed that during limb generation, axolotls have a different system to regulate their internal
1595: 979:, triggers metamorphosis when injected. This is because diiodotyrosine and thyroxine is produced when I 421: 4024:"Axo got a new style just in time for @maisie_williams & @reubenSelby's hand-picked Locker Bundle" 3173: 1327:(1999), is directly based on an axolotl. The looks of the dragons Toothless and The Night Fury in the 602:
lined with papillae, while females are noticeable for their wider bodies full of eggs. Three pairs of
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A review of gastrolith function with implications for fossil vertebrates and a revised classification
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Dvoskin, Samuel (May 1947). "The Thyroxine-Like Action of Elemental Iodine in the Rat and Chick1".
2339: 1400: 1329: 1199: 1001: 671: 63: 3810: 3229:"Response of a native endangered axolotl, Ambystoma mexicanum (Amphibia), to exotic fish predator" 2433:
Goodwin, James W.; Pinto, Alexander R.; Rosenthal, Nadia A. (June 4, 2013). Olson, Eric N. (ed.).
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Parra-Plea, G; Zamudio, K.R.; Recuero, E.; Aguilar-=Miguel, X.; Huaxuz, D.; Zambrano, L. (2011).
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Albino: pale pink or white, with red eyes, which is more common in axolotls than other species.
533:), which are widespread in much of North America and occasionally become paedomorphic, or with 518: 3050: 4400: 2302: 855:
sizes (22,759 bp) which are 13, 16 and 25 times that observed in human (1,750 bp),
506: 250: 4392: 4187: 3381:"The potential of temporary shelters to increase survival of the endangered Mexican axolotl" 3051:"The history of the oldest self-sustaining laboratory animal: 150 years of axolotl research" 2182: 1145: 4428: 4270: 4182: 4027: 3598: 3392: 3327: 3185: 3062: 2834: 2751: 2570: 2448: 2187: 1727: 913: 4353: 3961: 2080:"Lake Xochimilco, Borough of Xochimilco in southern MĂ©xico City, 162 L ‱ Biotope Aquarium" 1228:
There is some evidence that axolotls might seek out appropriately-sized gravel for use as
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conditions, metamorphosis is reliably induced by administering either the thyroid hormone
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in the wild, with a decreasing population of around 50 to 1,000 adult individuals, by the
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reacts with proteins other than thyroglobulin. If given in a bath instead of injected, I
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Axolotls may be confused with the larval stage of the closely related tiger salamander (
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Pietsch, Paul; Schneider, Carl W. (1985). "Vision and the skin camouflage reactions of
2018: 1862:, Perspectives in vertebrate science, Dordrecht: Springer Netherlands, pp. 55–69, 1756: 1715: 1696: 1345: 1150: 1018: 723: 299: 150: 3544: 3197: 3133:"Water quality in Lake Xochimilco, Mexico: zooplankton indicators and Vibrio cholerae" 1841:
San Francisco Examiner (San Francisco, California) 7 August 1887, page 9, authored by
1305: 1161: 1023: 517:(CITES). Axolotls are used extensively in scientific research due to their ability to 4387: 4257: 3698: 3659: 3628: 3548: 3435: 3412: 3361: 3343: 3291: 3260: 3248: 3209: 3201: 3154: 3086: 3078: 2959: 2862: 2857: 2822: 2801: 2777: 2716: 2606: 2598: 2593: 2523: 2500: 2476: 2415: 2356: 2319: 2252: 2219: 2053: 2049: 1966: 1918: 1908: 1871: 1761: 1743: 1700: 1688: 1680: 1660: 1291: 1106: 1046: 1028: 909:. Neoteny is the term for reaching sexual maturity without undergoing metamorphosis. 778:, which allows viewing of the full development of a vertebrate. Axolotls are used in 3710: 3299: 2514: 2495: 2368: 2337:
Roy, S; Gatien, S (November 2008). "Regeneration in axolotls: a model to aim for!".
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Alcaraz, Guillermina; LĂłpez-Portela, Xarini; Robles-Mendoza, Cecilia (2015-07-01).
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Crowner, Anne; Khatri, Shivam; Blichmann, Dana; Voss, S. Randal (12 April 2019).
2298: 1370: 1355:. An anthropomorphic Axolotl named Axo was also added as a purchasable outfit in 1206: 1071:
that are caused by deficiencies in food and, in particular, iodine, which causes
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provides limited support (BLAST, JBrowse tracks, genome download) for Axolotls.
3655: 3339: 3007: 2763: 2649:: Its Biology and Developmental Genetics, and Its Autonomous Cell-lethal Genes" 2440:
Proceedings of the National Academy of Sciences of the United States of America
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teeth, which develop during metamorphosis. The primary method of feeding is by
603: 591: 467: 217: 3244: 3149: 1922: 1855: 1317:(2002), take some visual inspiration from axolotls. Additionally, the Pokemon 4445: 4348: 4049: 3439: 3347: 3252: 3205: 3158: 3082: 3055:
Journal of Experimental Zoology Part B: Molecular and Developmental Evolution
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The axolotl is a popular exotic pet like its relative, the tiger salamander (
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Clayton, Leigh Ann; Gore, Stacey R. (2007). "Amphibian Emergency Medicine".
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A captive leucistic axolotl, perhaps the most well known form of the axolotl
4158: 3702: 3663: 3552: 3365: 3213: 3090: 2963: 2781: 2720: 2665: 2644: 2610: 2480: 2419: 2360: 2323: 2285: 1970: 1902: 1827: 1806: 1765: 1692: 1481: 1267: 1259: 1174: 844: 820: 788: 614: 470:. The species was originally found in several lakes underlying what is now 4379: 4236: 3811:"Mexico's axolotl, a cartoon hero and genetic marvel, fights for survival" 3459: 2955: 2866: 2256: 2223: 2057: 636: 82: 4296: 4195: 4167: 4009:"Fortnite v13.40 Leaked Skins: Axo, Castaway Jonesy, Crustina & More" 3694: 3428:"Mexico wants you to adopt an axolotl, the amphibian that never grows up" 2557:"The axolotl genome and the evolution of key tissue formation regulators" 1961: 1940: 1245: 1229: 1218: 1189: 1064: 1013: 784: 707: 692: 607: 479: 471: 3074: 2583: 2556: 1945:), a Neotenic Amphibian, Expresses Functional Thyroid Hormone Receptors" 762: 4288: 4200: 3273: 2990: 2901: 1390: 1059: 816: 792: 715: 706:
The wild population has been put under heavy pressure by the growth of
490: 486:, leading to the destruction of much of the axolotl's natural habitat. 451: 207: 2880:
Swingle, W. W. (November 1923). "Iodine and Amphibian Metamorphosis".
2736:"Experimentally induced metamorphosis in highly regenerative axolotl ( 2400: 2215: 1012:
Six adult axolotls (including a leucistic specimen) were shipped from
658:
Melanoid: all black or dark blue with no gold speckling or olive tone.
4301: 3404: 1842: 1335: 1333:
movies are based on axolotls. They were also added to the video game
1283: 1210: 1154: 1072: 1068: 930: 902: 832: 570: 459: 197: 167: 4129: 3587:"The Aquarium Trade as an Invasion Pathway in the Pacific Northwest" 3567: 2893: 2382:
Vieira, Warren A.; Wells, Kaylee M.; McCusker, Catherine D. (2020).
679: 511:
International Union for Conservation of Nature and Natural Resources
4152: 3585:
Strecker, Angela L.; Campbell, Philip M.; Olden, Julian D. (2011).
3380: 2537: 1431: 1380: 1275: 1183: 1179: 1076: 1051: 534: 187: 3276:"Conservation genetics of threatened Mexican axolotls (Ambystoma)" 3226: 2823:"The role of thyroid hormone in zebrafish and axolotl development" 1000:
An axolotl that has gone through metamorphosis resembles an adult
4031: 3966: 2733: 2493: 2435:"Macrophages are required for adult salamander limb regeneration" 1854:
McIndoe, Rosemary; Smith, D. G. (1984), Seymour, Roger S. (ed.),
1278:
on its back. It was recognized as "Bank Note of the Year" by the
1252: 1214: 1091: 951: 898: 882: 872: 851:, but such repeated elements also contribute to increased median 736: 628: 498: 448: 4275: 2795: 1520:
Amphibian Species of the World: an Online Reference. Version 6.0
1110:
Lake Xochimilco is the only native habitat left for the axolotl.
4208: 4068: 3677:
Gresens, Jill (2004). "An Introduction to the Mexican Axolotl (
3131:
Nandini, Sarma; GarcĂ­a, Pedro Ramirez; Sarma, S. S. S. (2016).
1996: 1994: 1713: 1318: 1308: 1263: 1248: 988: 852: 836: 775: 599: 565: 177: 4109: 2384:"Advancements to the Axolotl Model for Regeneration and Aging" 2130:"Endangered 'water monster' Axolotl found in Mexico City lake" 1985:"18 Types of Axolotl Colors You Can Own (Axolotl Color Guide)" 1938: 1427: 1425: 403: 4413: 3745:
Gastroliths – How I Learned to Stop Worrying and Love Gravel.
3460:"Axolotls – Requirements & Water Conditions in Captivity" 3312: 2201: 1298: 1271: 886: 856: 523: 502: 140: 134: 2554: 1991: 1781:"Mythic Salamander Faces Crucial Test: Survival in the Wild" 881:
which are unable to live on dry land, often being dubbed as
549: 3836:"Mexican axolotl will be the new image of the 50 peso bill" 3644:
Veterinary Clinics of North America: Exotic Animal Practice
3530: 2036:
larvae: the effects of eye transplants and brain lesions".
1422: 1287: 804: 466:. Instead of taking to the land, adults remain aquatic and 391: 4124: 3378: 2682: 1221:, but care should be taken as fish may contain parasites. 787:
closure due to the similarities between human and axolotl
497:, as well as the introduction of invasive species such as 397: 1395: 482:. These lakes were drained by Spanish settlers after the 382: 4099:
Mystical amphibian venerated by Aztecs nears extinction
3049:
Reiß, Christian; Olsson, Lennart; Hoßfeld, Uwe (2015).
515:
Convention on International Trade in Endangered Species
3385:
Aquatic Conservation: Marine and Freshwater Ecosystems
2177: 2175: 3584: 2432: 2000: 1904:
Vertebrates: comparative anatomy, function, evolution
1856:"Functional morphology of gills in larval amphibians" 1202:, are often added to the water to prevent infection. 388: 385: 3379:
Ramos, A.G.; Mena-Gonzalez, H.; Zambrano, L (2021).
400: 394: 3962:""Minecraft Live: Caves & Cliffs - First Look"" 2381: 2172: 379: 3130: 3008:"Axolotls - Metamorphosed & Tiger Salamanders" 2270:Armstrong, John B. (1985). "The axolotl mutants". 1618:"Are Axolotls Endangered? You Need To Be Careful
" 770:Today, the axolotl is still used in research as a 493:due to urbanization in Mexico City and consequent 4120:. Vol. 20 (11th ed.). 1911. p. 63. 3048: 2244:Journal of Embryology and Experimental Morphology 1534: 1086: 721:Axolotls are members of the tiger salamander, or 4443: 2794:Chapter 49, "Synthesis of Thyroid Hormones" in: 2330: 2155:"Mexico City's 'water monster' nears extinction" 2031: 1615: 1509: 687:The axolotl is native only to the freshwater of 462:in that it reaches adulthood without undergoing 2827:Proceedings of the National Academy of Sciences 2105:"Mexico's 'water monster' may have disappeared" 1658: 3805: 3803: 3801: 3799: 2740:) under constant diet restructures microbiota" 1779:Tickell, Sofia Castello Y. (30 October 2012). 1165:Axolotl in a pet store in Melbourne, Australia 513:(IUCN) and is listed under Appendix II of the 2979:Bulletin of Experimental Biology and Medicine 2096: 1562: 1457:10.2305/IUCN.UK.2020-3.RLTS.T1095A53947343.en 1258:They appear in the works of Mexican muralist 4482:Critically endangered fauna of North America 4021: 1853: 1644:(Television series). Dartmoor, England, UK: 1432:IUCN SSC Amphibian Specialist Group (2020). 1022:in Paris in 1863. Unaware of their neoteny, 4452:IUCN Red List critically endangered species 3796: 3641: 3631:. Washington, D.C.: National Academy Press. 2814: 2796:Walter F. Boron; Emile L. Boulpaep (2012). 1798: 1593: 23: 4472:Animals bred for albinism on a large scale 4084:Follow the Eggs and Hatchlings (2nd Batch) 3734:Acta Palaeontologica Polonica 52 (1): 1–16 2678: 2676: 2642: 2183:"Ambystoma mexicanum (Salamandra ajolote)" 1804: 920:, which contains several candidate genes. 751: 731:, along with all other Mexican species of 112: 81: 24: 4074:Follow the Eggs, Hatchlings and Juveniles 3959: 3355: 3148: 2856: 2846: 2788: 2771: 2710: 2700: 2664: 2617: 2592: 2582: 2513: 2470: 2460: 2409: 2399: 2336: 2269: 2102: 1960: 1826: 1755: 1455: 1270:banknote featuring an axolotl along with 4457:Amphibians described in the 18th century 4104:The animal that’s everywhere and nowhere 3622: 3620: 3524: 1659:Sandoval-GuzmĂĄn, Tatiana (August 2023). 1634: 1239: 1188: 1160: 1144: 1105: 761: 678: 670: 635: 627: 576: 564: 556: 548: 32:This is an accepted version of this page 4507:Taxa named by Frederick Polydore Nodder 3775:"Common Disease Conditions in Axolotls" 3676: 3670: 3103: 2941: 2879: 2673: 1900: 1778: 1505: 1503: 1501: 1476: 1474: 1040: 640:Axolotls displaying variations in color 14: 4527:Species endangered by invasive species 4444: 2976: 2914: 2236: 666: 428: 4477:Critically endangered biota of Mexico 4134: 4133: 3861:"Billete de 50 pesos de la familia G" 3626: 3617: 3002: 3000: 2821:Brown, Donald D. (25 November 1997). 2820: 2550: 2548: 2546: 1934: 1932: 1896: 1894: 1892: 1805:Malacinski, George M. (Spring 1978). 1648:. 2009-11-11. Event occurs at 00:25. 1157:, with less pigmentation than normal. 877:Most amphibians begin their lives as 649:Leucistic: pale pink with black eyes. 561:Face of a common or wild type axolotl 3425: 2643:Malacinski, George M. (1978-05-01). 1522:. American Museum of Natural History 1498: 1471: 937:. The former is more commonly used. 344:DumĂ©ril, Bibron, and DumĂ©ril, 1854 3772: 3518:"Axolotls: Keeping a Water Monster" 3515: 3106:"Iodine and Evolution. DIMI-Marche" 2800:(2nd ed.). Elsevier/Saunders. 2623: 1596:"Axolotl verges on wild extinction" 1443:IUCN Red List of Threatened Species 889:, they go through a process called 56: 3533:Ultrasound in Medicine and Biology 3042: 2997: 2636: 2543: 2072: 2019:10.1111/j.1432-0436.1984.tb01393.x 1929: 1889: 1616:PetAquariums.com (22 April 2020). 975:, which contains both iodide and I 57: 4538: 4041: 4022:@FortniteGame (August 11, 2021). 3565: 3484:"Caudata Culture Species Entry – 3198:10.1016/j.chemosphere.2008.09.087 2917:Experimental Biology and Medicine 1572:. Aquarium Online. Archived from 1266:released a new design for its 50- 1058:Neoteny has been observed in all 489:As of 2020, the axolotl was near 3939:International Astronomical Union 3426:PaĂșl, MarĂ­a Luisa (2023-12-01). 3292:10.1111/j.1469-1795.2011.00488.x 2538:https://doi.org/10.1002/dvdy.282 1542:"Mexican Walking Fish, Axolotls 1324:PokĂ©mon Gold, Silver and Crystal 923: 375: 154: 4522:Species endangered by pollution 4094:University of KY Axolotl Colony 4015: 4001: 3977: 3953: 3926: 3902: 3878: 3853: 3828: 3785:from the original on 2020-08-04 3766: 3756: 3737: 3717: 3635: 3578: 3559: 3509: 3476: 3452: 3419: 3372: 3306: 3267: 3220: 3165: 3124: 3097: 3024: 2970: 2935: 2908: 2873: 2727: 2530: 2515:10.1096/fasebj.2020.34.s1.04015 2487: 2426: 2375: 2292: 2263: 2230: 2195: 2157:. November 2008. Archived from 2147: 2122: 2025: 1977: 1847: 1835: 1772: 1707: 1642:Weird Creatures with Nick Baker 1280:International Bank Note Society 1244:The species is named after the 1134: 967:, an analogue of the iodinated 835:long sequence of the axolotl's 798: 652:Xanthic: grey, with black eyes. 3886:"Banknote of 2021 Nominations" 3611:10.1577/03632415.2011.10389070 1652: 1609: 1587: 1087:Online Model Organism Database 766:Leucistic axolotl in captivity 544: 13: 1: 3960:Minecraft (October 3, 2020). 3545:10.1016/s0301-5629(99)00040-x 2204:Brain, Behavior and Evolution 1416: 1349:in 2022 and are available in 1311:and its evolutions, added in 1301:, was named AxĂłlotl in 2019. 863:(906 bp), respectively. 613:Axolotls have barely visible 4487:Endemic amphibians of Mexico 4467:Animal testing on amphibians 4079:Mating Dance and Laying Eggs 3773:Loh, Richmond (2015-05-15). 2624:Ley, Willy (February 1968). 2103:Stevenson, M. (2014-01-28). 2050:10.1016/0006-8993(85)90772-3 484:conquest of the Aztec Empire 7: 3985:"The Guardian Battle 21180" 2353:10.1016/j.exger.2008.09.003 2316:10.1016/j.ydbio.2017.08.022 1901:Kardong, Kenneth V (2019). 1868:10.1007/978-94-009-6536-2_4 1364: 987:has no effect on axolotls. 935:thyroid-stimulating hormone 895:thyroid-stimulating hormone 604:external gill stalks (rami) 284:IUCN range of the axolotl. 10: 4543: 3656:10.1016/j.cvex.2007.02.004 3340:10.1038/s41598-017-00059-1 3032:"Transition & Feeding" 2764:10.1038/s41598-018-29373-y 2689:Frontiers in Endocrinology 2626:"Epitaph for a Lonely Olm" 1740:10.1038/s41598-022-24442-9 1677:10.1038/s41592-023-01961-5 1594:Matt Walker (2009-08-26). 1570:"Axolotols (Walking Fish)" 1138: 1097: 1007: 897:, which is needed for the 870: 866: 755: 61: 4512:Taxa named by George Shaw 4142: 3914:www.nameexoworlds.iau.org 3245:10.1007/s10750-015-2194-4 3150:10.4081/jlimnol.2015.1213 2594:21.11116/0000-0003-F659-4 1907:. McGraw-Hill Education. 1510:Frost, Darrel R. (2018). 1386:Lake Patzcuaro salamander 1339:in 2020. It is following 1314:PokĂ©mon Ruby and Sapphire 950:), which reacts with the 826: 703:leading from Xochimilco. 305: 298: 283: 279: 256: 249: 151:Scientific classification 149: 132: 110: 101: 89: 80: 75: 4502:Regenerative biomedicine 4089:Indiana U Axolotl Colony 2929:10.3181/00379727-26-4212 2848:10.1073/pnas.94.24.13011 2702:10.3389/fendo.2019.00237 2628:. For Your Information. 2340:Experimental Gerontology 2300:(Ambystoma mexicanum)". 1516:(Shaw and Nodder, 1798)" 1401:Regenerative biomedicine 1330:How to Train Your Dragon 1217:. Axolotls can also eat 1083:, as it does in humans. 1002:plateau tiger salamander 64:Axolotl (disambiguation) 39:latest accepted revision 4117:EncyclopĂŠdia Britannica 4059:considered for deletion 2882:The Biological Bulletin 2462:10.1073/pnas.1300290110 2251:(Supplement): 229–255. 1376:Barred tiger salamander 752:Use as a model organism 742: 454:closely related to the 425: 312:Shaw and Nodder, 1798 2645:"The Mexican Axolotl, 2630:Galaxy Science Fiction 2286:10.1002/dvg.1020060102 2273:Developmental Genetics 1807:"The Mexican Axolotl, 1406:Texas blind salamander 1358:Fortnite Battle Royale 1195: 1166: 1158: 1111: 807:and is capable of the 767: 684: 676: 641: 633: 586: 574: 562: 554: 458:. It is unusual among 4401:Paleobiology Database 3627:Pough, F. H. (1992). 3568:"Health and Diseases" 3280:American Conservation 2956:10.1210/endo-40-5-334 2303:Developmental Biology 1240:Cultural significance 1200:Holtfreter's solution 1192: 1164: 1148: 1109: 765: 682: 674: 639: 631: 580: 568: 560: 552: 507:critically endangered 430:[aːˈʃoːloːtÉŹ] 119:Critically Endangered 59:Species of salamander 3695:10.1038/laban1004-41 3137:Journal of Limnology 3104:Venturi, S. (2004). 2666:10.1093/icb/18.2.195 2188:Animal Diversity Web 1962:10.1210/en.2003-0913 1828:10.1093/icb/18.2.195 1482:"Appendices | CITES" 1450:: e.T1095A53947343. 1182:, commonly added to 1041:In other salamanders 914:artificial selection 859:(1,469 bp) and 849:repetitive sequences 349:Amblystoma weismanni 62:For other uses, see 4497:Population genetics 4227:Ambystoma-mexicanum 4214:ambystoma-mexicanum 4188:Ambystoma_mexicanum 4174:Ambystoma mexicanum 4144:Ambystoma mexicanum 3942:. December 17, 2019 3679:Ambystoma mexicanum 3603:2011Fish...36...74S 3486:Ambystoma mexicanum 3397:2021ACMFE..31.1535R 3332:2017NatSR...7....6W 3190:2009Chmsp..74..703R 3075:10.1002/jez.b.22617 3067:2015JEZB..324..393R 2839:1997PNAS...9413011B 2833:(24): 13011–13016. 2756:2018NatSR...810974D 2738:Ambystoma mexicanum 2647:Ambystoma mexicanum 2584:10.1038/nature25458 2575:2018Natur.554...50N 2453:2013PNAS..110.9415G 2237:Gordon, R. (1985). 2003:Ambystoma mexicanum 1943:Ambystoma mexicanum 1809:Ambystoma mexicanum 1732:2022NatSR..1220491M 1646:The Science Channel 1544:Ambystoma mexicanum 1514:Ambystoma mexicanum 1436:Ambystoma mexicanum 1361:on August 9, 2020. 1346:Minecraft: Dungeons 819:level and suppress 667:Habitat and ecology 583:Ambystoma mexicanum 444:Ambystoma mexicanum 352:Wiedersheim, 1879 291:Ambystoma mexicanum 260:Ambystoma mexicanum 104:Conservation status 29:Page version status 18:Ambystoma mexicanum 3865:www.banxico.org.mx 3750:2020-09-22 at the 3730:2016-03-04 at the 3320:Scientific Reports 2991:10.1007/BF00796048 2798:Medical Physiology 2744:Scientific Reports 2653:American Zoologist 2632:. pp. 95–104. 2111:. Associated Press 1987:. August 14, 2019. 1815:American Zoologist 1785:The New York Times 1720:Scientific Reports 1196: 1171:Ambystoma tigrinum 1167: 1159: 1151:Vancouver Aquarium 1149:These axolotls at 1112: 1019:Jardin des Plantes 768: 724:Ambystoma tigrinum 685: 677: 642: 634: 587: 575: 563: 555: 505:. It is listed as 341:Siredon Humboldtii 35: 4439: 4438: 4388:Open Tree of Life 4136:Taxon identifiers 3840:The Yucatan Times 3516:Wiegert, Joshua. 2501:The FASEB Journal 2447:(23): 9415–9420. 2401:10.1159/000504294 2216:10.1159/000116519 1914:978-1-259-70091-0 1877:978-94-009-6536-2 1558:on 15 March 2018. 1029:Ambystoma velasci 907:able to reproduce 581:Axolotl's gills ( 422:Classical Nahuatl 367: 366: 361: 353: 345: 337: 329: 321: 317:Siren pisciformis 313: 309:Gyrinus mexicanus 242:A. mexicanum 144: 127: 92:National Aquarium 47:15 September 2024 26: 16:(Redirected from 4534: 4517:Valley of Mexico 4492:Mole salamanders 4432: 4431: 4422: 4421: 4409: 4408: 4396: 4395: 4383: 4382: 4370: 4369: 4357: 4356: 4344: 4343: 4331: 4330: 4318: 4317: 4305: 4304: 4292: 4291: 4279: 4278: 4266: 4265: 4253: 4252: 4240: 4239: 4230: 4229: 4217: 4216: 4204: 4203: 4191: 4190: 4178: 4177: 4176: 4163: 4162: 4161: 4131: 4130: 4121: 4113: 4062: 4036: 4035: 4019: 4013: 4012: 4011:. 5 August 2020. 4005: 3999: 3998: 3996: 3995: 3981: 3975: 3974: 3957: 3951: 3950: 3948: 3947: 3930: 3924: 3923: 3921: 3920: 3910:"Approved names" 3906: 3900: 3899: 3897: 3896: 3882: 3876: 3875: 3873: 3872: 3857: 3851: 3850: 3848: 3847: 3832: 3826: 3825: 3823: 3822: 3807: 3794: 3793: 3791: 3790: 3770: 3764: 3760: 3754: 3741: 3735: 3721: 3715: 3714: 3674: 3668: 3667: 3639: 3633: 3632: 3624: 3615: 3614: 3582: 3576: 3575: 3563: 3557: 3556: 3528: 3522: 3521: 3513: 3507: 3506: 3504: 3503: 3494:. Archived from 3480: 3474: 3473: 3471: 3470: 3456: 3450: 3449: 3447: 3446: 3423: 3417: 3416: 3405:10.1002/aqc.3520 3391:(6): 1535–1542. 3376: 3370: 3369: 3359: 3310: 3304: 3303: 3271: 3265: 3264: 3224: 3218: 3217: 3169: 3163: 3162: 3152: 3128: 3122: 3121: 3119: 3117: 3108:. Archived from 3101: 3095: 3094: 3046: 3040: 3039: 3036:Morphed Axolotls 3028: 3022: 3021: 3019: 3018: 3004: 2995: 2994: 2974: 2968: 2967: 2939: 2933: 2932: 2912: 2906: 2905: 2877: 2871: 2870: 2860: 2850: 2818: 2812: 2811: 2792: 2786: 2785: 2775: 2731: 2725: 2724: 2714: 2704: 2680: 2671: 2670: 2668: 2640: 2634: 2633: 2621: 2615: 2614: 2596: 2586: 2552: 2541: 2534: 2528: 2527: 2517: 2491: 2485: 2484: 2474: 2464: 2430: 2424: 2423: 2413: 2403: 2379: 2373: 2372: 2334: 2328: 2327: 2296: 2290: 2289: 2267: 2261: 2260: 2234: 2228: 2227: 2199: 2193: 2192: 2179: 2170: 2169: 2167: 2166: 2151: 2145: 2144: 2142: 2141: 2126: 2120: 2119: 2117: 2116: 2100: 2094: 2093: 2091: 2090: 2084:Biotope Aquarium 2076: 2070: 2069: 2029: 2023: 2022: 2013:(1–3): 182–188. 1998: 1989: 1988: 1981: 1975: 1974: 1964: 1936: 1927: 1926: 1898: 1887: 1886: 1885: 1884: 1851: 1845: 1839: 1833: 1832: 1830: 1802: 1796: 1795: 1793: 1791: 1776: 1770: 1769: 1759: 1711: 1705: 1704: 1671:(8): 1117–1119. 1656: 1650: 1649: 1638: 1632: 1631: 1629: 1628: 1622:PetAquariums.com 1613: 1607: 1606: 1604: 1603: 1591: 1585: 1584: 1582: 1581: 1576:on 10 April 2013 1566: 1560: 1559: 1557: 1551:. Archived from 1550: 1538: 1532: 1531: 1529: 1527: 1507: 1496: 1495: 1493: 1492: 1478: 1469: 1468: 1466: 1464: 1459: 1429: 1411:Texas salamander 973:Lugol's solution 841:genetic pathways 697:Valley of Mexico 456:tiger salamander 440: 439: 438: 432: 419: 418: 417: 416: 409: 406: 405: 402: 399: 396: 393: 390: 387: 384: 381: 359: 351: 343: 335: 333:Axolotes guttata 327: 319: 311: 288: 262: 159: 158: 138: 121: 116: 115: 96:Washington, D.C. 85: 73: 72: 21: 4542: 4541: 4537: 4536: 4535: 4533: 4532: 4531: 4442: 4441: 4440: 4435: 4427: 4425: 4417: 4412: 4404: 4399: 4391: 4386: 4378: 4375:Observation.org 4373: 4365: 4360: 4352: 4347: 4339: 4334: 4326: 4321: 4313: 4308: 4300: 4295: 4287: 4282: 4274: 4269: 4261: 4256: 4248: 4243: 4235: 4233: 4225: 4220: 4212: 4207: 4199: 4194: 4186: 4181: 4172: 4171: 4166: 4157: 4156: 4151: 4138: 4111:"Axolotl"  4108: 4047: 4044: 4039: 4020: 4016: 4007: 4006: 4002: 3993: 3991: 3983: 3982: 3978: 3958: 3954: 3945: 3943: 3932: 3931: 3927: 3918: 3916: 3908: 3907: 3903: 3894: 3892: 3890:www.theibns.org 3884: 3883: 3879: 3870: 3868: 3859: 3858: 3854: 3845: 3843: 3834: 3833: 3829: 3820: 3818: 3809: 3808: 3797: 3788: 3786: 3771: 3767: 3761: 3757: 3752:Wayback Machine 3742: 3738: 3732:Wayback Machine 3722: 3718: 3675: 3671: 3640: 3636: 3625: 3618: 3583: 3579: 3566:Clare, John P. 3564: 3560: 3529: 3525: 3514: 3510: 3501: 3499: 3492:www.caudata.org 3482: 3481: 3477: 3468: 3466: 3458: 3457: 3453: 3444: 3442: 3432:Washington Post 3424: 3420: 3377: 3373: 3311: 3307: 3272: 3268: 3225: 3221: 3170: 3166: 3129: 3125: 3115: 3113: 3112:on 4 March 2017 3102: 3098: 3047: 3043: 3030: 3029: 3025: 3016: 3014: 3012:www.axolotl.org 3006: 3005: 2998: 2975: 2971: 2940: 2936: 2913: 2909: 2894:10.2307/1536749 2878: 2874: 2819: 2815: 2808: 2793: 2789: 2732: 2728: 2681: 2674: 2641: 2637: 2622: 2618: 2569:(7690): 50–55. 2553: 2544: 2535: 2531: 2492: 2488: 2431: 2427: 2380: 2376: 2335: 2331: 2297: 2293: 2268: 2264: 2235: 2231: 2200: 2196: 2181: 2180: 2173: 2164: 2162: 2153: 2152: 2148: 2139: 2137: 2134:The Independent 2128: 2127: 2123: 2114: 2112: 2101: 2097: 2088: 2086: 2078: 2077: 2073: 2030: 2026: 2007:Differentiation 1999: 1992: 1983: 1982: 1978: 1937: 1930: 1915: 1899: 1890: 1882: 1880: 1878: 1852: 1848: 1840: 1836: 1803: 1799: 1789: 1787: 1777: 1773: 1712: 1708: 1657: 1653: 1640: 1639: 1635: 1626: 1624: 1614: 1610: 1601: 1599: 1592: 1588: 1579: 1577: 1568: 1567: 1563: 1555: 1548: 1540: 1539: 1535: 1525: 1523: 1508: 1499: 1490: 1488: 1480: 1479: 1472: 1462: 1460: 1430: 1423: 1419: 1371:Amphibious fish 1367: 1242: 1198:Salts, such as 1143: 1137: 1100: 1089: 1043: 1024:Auguste DumĂ©ril 1010: 994: 993: 992: 986: 982: 978: 949: 926: 879:aquatic animals 875: 869: 831:The 32 billion 829: 801: 760: 754: 745: 729:species complex 712:African tilapia 689:Lake Xochimilco 669: 547: 495:water pollution 476:Lake Xochimilco 434: 433: 412: 411: 378: 374: 325:Siredon axolotl 294: 289: Axolotl ( 286: 275: 264: 258: 245: 153: 145: 128: 117: 113: 106: 67: 60: 55: 54: 53: 52: 51: 50: 34: 22: 15: 12: 11: 5: 4540: 4530: 4529: 4524: 4519: 4514: 4509: 4504: 4499: 4494: 4489: 4484: 4479: 4474: 4469: 4464: 4459: 4454: 4437: 4436: 4434: 4433: 4423: 4410: 4397: 4384: 4371: 4358: 4345: 4332: 4319: 4306: 4293: 4280: 4267: 4254: 4241: 4231: 4218: 4205: 4192: 4179: 4164: 4148: 4146: 4140: 4139: 4128: 4127: 4122: 4106: 4101: 4096: 4091: 4086: 4081: 4076: 4071: 4065:Ambystomatidae 4043: 4042:External links 4040: 4038: 4037: 4030:) – via 4014: 4000: 3976: 3952: 3925: 3901: 3877: 3852: 3827: 3795: 3765: 3755: 3736: 3716: 3669: 3650:(2): 587–620. 3634: 3616: 3577: 3558: 3539:(6): 969–975. 3523: 3508: 3475: 3451: 3418: 3371: 3305: 3266: 3219: 3184:(5): 703–710. 3164: 3123: 3096: 3061:(5): 393–404. 3041: 3023: 2996: 2985:(1): 724–727. 2969: 2950:(5): 334–352. 2934: 2907: 2888:(5): 229–253. 2872: 2813: 2806: 2787: 2726: 2672: 2659:(2): 195–206. 2635: 2616: 2542: 2529: 2486: 2425: 2394:(3): 212–222. 2374: 2347:(11): 968–73. 2329: 2310:(2): 227–239. 2291: 2262: 2229: 2210:(6): 329–341. 2194: 2171: 2146: 2121: 2095: 2071: 2038:Brain Research 2024: 1990: 1976: 1955:(2): 760–772. 1941:"The Axolotl ( 1928: 1913: 1888: 1876: 1846: 1834: 1821:(2): 195–206. 1797: 1771: 1706: 1665:Nature Methods 1651: 1633: 1608: 1586: 1561: 1533: 1497: 1470: 1420: 1418: 1415: 1414: 1413: 1408: 1403: 1398: 1393: 1388: 1383: 1378: 1373: 1366: 1363: 1352:Lego Minecraft 1341:Mojang Studios 1241: 1238: 1175:poikilothermic 1173:). As for all 1141:Herpetoculture 1136: 1133: 1099: 1096: 1088: 1085: 1081:hypothyroidism 1042: 1039: 1009: 1006: 984: 980: 976: 965:Diiodotyrosine 947: 943:Role of iodine 941: 940: 939: 925: 922: 871:Main article: 868: 865: 828: 825: 800: 797: 772:model organism 758:Model organism 753: 750: 744: 741: 668: 665: 660: 659: 656: 653: 650: 632:Buccal pumping 623:buccal pumping 592:external gills 546: 543: 365: 364: 363: 362: 354: 346: 338: 330: 328:Wagler, 1830 322: 314: 303: 302: 296: 295: 285: 281: 280: 277: 276: 265: 254: 253: 247: 246: 239: 237: 233: 232: 225: 221: 220: 218:Ambystomatidae 215: 211: 210: 205: 201: 200: 195: 191: 190: 185: 181: 180: 175: 171: 170: 165: 161: 160: 147: 146: 133: 130: 129: 111: 108: 107: 102: 99: 98: 87: 86: 78: 77: 58: 36: 30: 27: 25: 9: 6: 4: 3: 2: 4539: 4528: 4525: 4523: 4520: 4518: 4515: 4513: 4510: 4508: 4505: 4503: 4500: 4498: 4495: 4493: 4490: 4488: 4485: 4483: 4480: 4478: 4475: 4473: 4470: 4468: 4465: 4463: 4462:Animal models 4460: 4458: 4455: 4453: 4450: 4449: 4447: 4430: 4424: 4420: 4415: 4411: 4407: 4402: 4398: 4394: 4389: 4385: 4381: 4376: 4372: 4368: 4363: 4359: 4355: 4350: 4346: 4342: 4337: 4333: 4329: 4324: 4320: 4316: 4311: 4307: 4303: 4298: 4294: 4290: 4285: 4281: 4277: 4272: 4268: 4264: 4259: 4255: 4251: 4246: 4242: 4238: 4232: 4228: 4223: 4219: 4215: 4210: 4206: 4202: 4197: 4193: 4189: 4184: 4180: 4175: 4169: 4165: 4160: 4154: 4150: 4149: 4147: 4145: 4141: 4137: 4132: 4126: 4123: 4119: 4118: 4112: 4107: 4105: 4102: 4100: 4097: 4095: 4092: 4090: 4087: 4085: 4082: 4080: 4077: 4075: 4072: 4070: 4066: 4060: 4056: 4055: 4051: 4046: 4045: 4033: 4029: 4025: 4018: 4010: 4004: 3990: 3986: 3980: 3973: 3969: 3968: 3963: 3956: 3941: 3940: 3935: 3929: 3915: 3911: 3905: 3891: 3887: 3881: 3866: 3862: 3856: 3841: 3837: 3831: 3816: 3812: 3806: 3804: 3802: 3800: 3784: 3780: 3776: 3769: 3759: 3753: 3749: 3746: 3740: 3733: 3729: 3726: 3720: 3712: 3708: 3704: 3700: 3696: 3692: 3688: 3684: 3680: 3673: 3665: 3661: 3657: 3653: 3649: 3645: 3638: 3630: 3623: 3621: 3612: 3608: 3604: 3600: 3596: 3592: 3588: 3581: 3573: 3569: 3562: 3554: 3550: 3546: 3542: 3538: 3534: 3527: 3519: 3512: 3498:on 2016-03-15 3497: 3493: 3489: 3487: 3479: 3465: 3461: 3455: 3441: 3437: 3433: 3429: 3422: 3414: 3410: 3406: 3402: 3398: 3394: 3390: 3386: 3382: 3375: 3367: 3363: 3358: 3353: 3349: 3345: 3341: 3337: 3333: 3329: 3325: 3321: 3317: 3309: 3301: 3297: 3293: 3289: 3285: 3281: 3277: 3270: 3262: 3258: 3254: 3250: 3246: 3242: 3238: 3234: 3233:Hydrobiologia 3230: 3223: 3215: 3211: 3207: 3203: 3199: 3195: 3191: 3187: 3183: 3179: 3175: 3168: 3160: 3156: 3151: 3146: 3142: 3138: 3134: 3127: 3111: 3107: 3100: 3092: 3088: 3084: 3080: 3076: 3072: 3068: 3064: 3060: 3056: 3052: 3045: 3037: 3033: 3027: 3013: 3009: 3003: 3001: 2992: 2988: 2984: 2980: 2973: 2965: 2961: 2957: 2953: 2949: 2945: 2944:Endocrinology 2938: 2930: 2926: 2922: 2918: 2911: 2903: 2899: 2895: 2891: 2887: 2883: 2876: 2868: 2864: 2859: 2854: 2849: 2844: 2840: 2836: 2832: 2828: 2824: 2817: 2809: 2807:9781437717532 2803: 2799: 2791: 2783: 2779: 2774: 2769: 2765: 2761: 2757: 2753: 2749: 2745: 2741: 2739: 2730: 2722: 2718: 2713: 2708: 2703: 2698: 2694: 2690: 2686: 2679: 2677: 2667: 2662: 2658: 2654: 2650: 2648: 2639: 2631: 2627: 2620: 2612: 2608: 2604: 2600: 2595: 2590: 2585: 2580: 2576: 2572: 2568: 2564: 2563: 2558: 2551: 2549: 2547: 2539: 2533: 2525: 2521: 2516: 2511: 2507: 2503: 2502: 2497: 2490: 2482: 2478: 2473: 2468: 2463: 2458: 2454: 2450: 2446: 2442: 2441: 2436: 2429: 2421: 2417: 2412: 2407: 2402: 2397: 2393: 2389: 2385: 2378: 2370: 2366: 2362: 2358: 2354: 2350: 2346: 2342: 2341: 2333: 2325: 2321: 2317: 2313: 2309: 2305: 2304: 2295: 2287: 2283: 2279: 2275: 2274: 2266: 2258: 2254: 2250: 2246: 2245: 2240: 2233: 2225: 2221: 2217: 2213: 2209: 2205: 2198: 2190: 2189: 2184: 2178: 2176: 2161:on 2011-07-23 2160: 2156: 2150: 2135: 2131: 2125: 2110: 2106: 2099: 2085: 2081: 2075: 2067: 2063: 2059: 2055: 2051: 2047: 2043: 2039: 2035: 2028: 2020: 2016: 2012: 2008: 2004: 1997: 1995: 1986: 1980: 1972: 1968: 1963: 1958: 1954: 1950: 1949:Endocrinology 1946: 1944: 1935: 1933: 1924: 1920: 1916: 1910: 1906: 1905: 1897: 1895: 1893: 1879: 1873: 1869: 1865: 1861: 1857: 1850: 1844: 1838: 1829: 1824: 1820: 1816: 1812: 1810: 1801: 1786: 1782: 1775: 1767: 1763: 1758: 1753: 1749: 1745: 1741: 1737: 1733: 1729: 1725: 1721: 1717: 1710: 1702: 1698: 1694: 1690: 1686: 1682: 1678: 1674: 1670: 1666: 1662: 1661:"The axolotl" 1655: 1647: 1643: 1637: 1623: 1619: 1612: 1597: 1590: 1575: 1571: 1565: 1554: 1547: 1545: 1537: 1521: 1517: 1515: 1506: 1504: 1502: 1487: 1483: 1477: 1475: 1458: 1453: 1449: 1445: 1444: 1439: 1437: 1428: 1426: 1421: 1412: 1409: 1407: 1404: 1402: 1399: 1397: 1394: 1392: 1389: 1387: 1384: 1382: 1379: 1377: 1374: 1372: 1369: 1368: 1362: 1360: 1359: 1354: 1353: 1348: 1347: 1342: 1338: 1337: 1332: 1331: 1326: 1325: 1320: 1316: 1315: 1310: 1307: 1302: 1300: 1296: 1295:constellation 1293: 1289: 1285: 1281: 1277: 1273: 1269: 1265: 1261: 1256: 1254: 1250: 1247: 1237: 1235: 1231: 1226: 1222: 1220: 1216: 1212: 1208: 1203: 1201: 1191: 1187: 1185: 1181: 1176: 1172: 1163: 1156: 1152: 1147: 1142: 1132: 1128: 1126: 1120: 1116: 1108: 1104: 1095: 1093: 1084: 1082: 1078: 1074: 1070: 1066: 1061: 1056: 1054: 1053: 1048: 1038: 1036: 1035:Julian Huxley 1031: 1030: 1025: 1021: 1020: 1015: 1005: 1003: 998: 990: 974: 970: 969:thyroglobulin 966: 963: 959: 957: 956:thyroglobulin 953: 944: 938: 936: 932: 924:Metamorphosis 921: 919: 915: 910: 908: 904: 900: 896: 892: 891:metamorphosis 888: 884: 880: 874: 864: 862: 858: 854: 850: 846: 842: 838: 834: 824: 822: 818: 813: 810: 806: 796: 794: 790: 786: 781: 777: 773: 764: 759: 749: 740: 738: 734: 730: 726: 725: 719: 717: 713: 709: 704: 700: 698: 694: 690: 681: 673: 664: 657: 654: 651: 648: 647: 646: 638: 630: 626: 624: 620: 616: 611: 609: 605: 601: 596: 593: 584: 579: 572: 569:The speckled 567: 559: 551: 542: 540: 536: 532: 527: 525: 520: 516: 512: 508: 504: 500: 496: 492: 487: 485: 481: 477: 473: 469: 465: 464:metamorphosis 461: 457: 453: 450: 446: 445: 437: 431: 427: 423: 415: 408: 372: 358: 357:Siredon edule 355: 350: 347: 342: 339: 334: 331: 326: 323: 318: 315: 310: 307: 306: 304: 301: 297: 292: 282: 278: 273: 269: 263: 261: 255: 252: 251:Binomial name 248: 244: 243: 238: 235: 234: 231: 230: 226: 223: 222: 219: 216: 213: 212: 209: 206: 203: 202: 199: 196: 193: 192: 189: 186: 183: 182: 179: 176: 173: 172: 169: 166: 163: 162: 157: 152: 148: 142: 136: 131: 125: 120: 109: 105: 100: 97: 93: 88: 84: 79: 74: 71: 69: 65: 48: 44: 40: 33: 28: 19: 4143: 4115: 4052: 4017: 4003: 3992:. Retrieved 3989:www.lego.com 3988: 3979: 3971: 3965: 3955: 3944:. Retrieved 3937: 3928: 3917:. Retrieved 3913: 3904: 3893:. Retrieved 3889: 3880: 3869:. Retrieved 3867:(in Spanish) 3864: 3855: 3844:. Retrieved 3842:. 2020-02-21 3839: 3830: 3819:. Retrieved 3817:. 2018-11-20 3814: 3787:. Retrieved 3778: 3768: 3758: 3739: 3719: 3689:(9): 41–47. 3686: 3682: 3678: 3672: 3647: 3643: 3637: 3597:(2): 74–85. 3594: 3590: 3580: 3571: 3561: 3536: 3532: 3526: 3511: 3500:. Retrieved 3496:the original 3491: 3485: 3478: 3467:. Retrieved 3463: 3454: 3443:. Retrieved 3431: 3421: 3388: 3384: 3374: 3323: 3319: 3308: 3286:(1): 61–72. 3283: 3279: 3269: 3239:(1): 73–80. 3236: 3232: 3222: 3181: 3177: 3167: 3140: 3136: 3126: 3116:25 September 3114:. Retrieved 3110:the original 3099: 3058: 3054: 3044: 3035: 3026: 3015:. Retrieved 3011: 2982: 2978: 2972: 2947: 2943: 2937: 2920: 2916: 2910: 2885: 2881: 2875: 2830: 2826: 2816: 2797: 2790: 2750:(1): 10974. 2747: 2743: 2737: 2729: 2692: 2688: 2656: 2652: 2646: 2638: 2629: 2619: 2566: 2560: 2532: 2505: 2499: 2489: 2444: 2438: 2428: 2391: 2387: 2377: 2344: 2338: 2332: 2307: 2301: 2294: 2277: 2271: 2265: 2248: 2242: 2232: 2207: 2203: 2197: 2186: 2163:. Retrieved 2159:the original 2149: 2138:. Retrieved 2136:. 2014-02-24 2133: 2124: 2113:. Retrieved 2108: 2098: 2087:. Retrieved 2083: 2074: 2044:(1): 37–60. 2041: 2037: 2033: 2027: 2010: 2006: 2002: 1979: 1952: 1948: 1942: 1903: 1881:, retrieved 1859: 1849: 1837: 1818: 1814: 1808: 1800: 1788:. Retrieved 1784: 1774: 1726:(1): 20491. 1723: 1719: 1709: 1668: 1664: 1654: 1641: 1636: 1625:. Retrieved 1621: 1611: 1600:. Retrieved 1589: 1578:. Retrieved 1574:the original 1564: 1553:the original 1543: 1536: 1524:. Retrieved 1519: 1513: 1489:. Retrieved 1485: 1461:. Retrieved 1447: 1441: 1435: 1356: 1350: 1344: 1334: 1328: 1322: 1312: 1303: 1260:Diego Rivera 1257: 1243: 1227: 1223: 1204: 1197: 1170: 1168: 1135:Captive care 1129: 1124: 1121: 1117: 1113: 1101: 1090: 1057: 1050: 1044: 1027: 1017: 1011: 999: 995: 960: 954:residues of 945: 942: 927: 917: 911: 876: 861:Tibetan frog 845:human genome 830: 821:inflammation 814: 809:regeneration 802: 799:Regeneration 789:neural plate 780:heart defect 769: 746: 732: 722: 720: 705: 701: 686: 661: 643: 612: 597: 588: 582: 538: 530: 528: 488: 449:paedomorphic 443: 442: 370: 368: 360:DugĂšs, 1888 356: 348: 340: 336:Owen, 1844 332: 324: 320:Shaw, 1802 316: 308: 290: 259: 257: 241: 240: 228: 70: 68: 46: 37:This is the 31: 4297:iNaturalist 4196:AmphibiaWeb 4168:Wikispecies 4125:xenbase.org 3743:Gordon, N, 3572:axolotl.org 3464:axolotl.org 3178:Chemosphere 2388:Gerontology 2280:(1): 1–25. 1463:12 November 1321:, added in 1262:. 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Index

Ambystoma mexicanum
latest accepted revision
reviewed
Axolotl (disambiguation)

National Aquarium
Washington, D.C.
Conservation status
Critically Endangered
IUCN 3.1
CITES
CITES
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Amphibia
Urodela
Ambystomatidae
Ambystoma
Binomial name
Shaw
Nodder
Synonyms
/ˈéksəlɒtəl/

Classical Nahuatl
[aːˈʃoːloːtÉŹ]

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