1471:
1477:
642:
2027:
231:
213:
2057:
2033:
25:
557:
544:, at a distance of 380 kilometres from the holotype, in Alberta in the Dinosaur Provincial Park, on the north bank of the Red Deer River, one kilometre east of Happy Jack's Cabin, in a layer of the lower Dinosaur Park Formation, two metres above the
536:, was found by Triebold, in layer of the Judith River Formation dating from the middle Campanian, about seventy-seven million years old. It consists of a partial right squamosal, possibly of a subadult individual. It was acquired by the
567:
was a medium-sized ceratopsian, reaching 4 metres (13 ft) in length and 1 metric ton (1.1 short tons) in body mass. The holotype squamosal has a length of 793 millimetres (31.2 in; 2.602 ft).
540:. The second specimen, UALVP 54559, was referred. It too is a partial right squamosal but slightly larger and probably of an older individual. It was found by Susan Owen-Kagen, a preparator at the
583:. The squamosal possesses a rear branch that is narrowed and rod-shaped instead of being blunt, triangular and tapering, the normal chasmosaurine form, or broadly rounded as with
548:. Associated with UALVP 54559, remains of a brow horn core were discovered but these were not referred because the connection with the squamosal was considered too uncertain.
501:, a reference to the similarity of the neck shield to the winged helmet of the messenger of the gods, with ~ceratops, "horn face", a usual suffix in ceratopian names. The
596:
on the frill rim. Based on the narrow form of the squamosal, it was deduced that the inner shield bones, the parietals, were very wide and pierced by enormous parietal
626:
shape and later triangular squamosals, the describers rejected this hypothesis because juvenile centrosaurines do not show such a transitional phase during their
688:
Ryan, M. J.; Evans, D. C.; Currie, P. J.; Loewen, M. A. (2014). "A new chasmosaurine from northern
Laramidia expands frill disparity in ceratopsid dinosaurs".
2173:
2122:
2183:
35:
2208:
2178:
93:
2213:
813:
750:
65:
792:
2223:
1470:
72:
1476:
2050:
655:
615:
79:
50:
2203:
230:
61:
2188:
2218:
2193:
2026:
2163:
2198:
883:
785:
2168:
1001:
525:
502:
2140:
2077:
521:
506:
480:
2127:
2135:
2060:
910:
541:
397:
86:
2032:
2109:
697:
537:
374:
8:
1166:
778:
647:
498:
46:
701:
1886:
1836:
1268:
721:
494:
225:
622:
squamosal suggests it was a transitional form between older rectangular squamosals of
1969:
1907:
1870:
1344:
1231:
756:
746:
713:
510:
725:
1999:
1797:
1622:
1324:
1019:
705:
545:
2100:
1893:
1845:
1804:
1740:
1594:
1296:
1220:
1175:
1049:
1035:
994:
868:
852:
592:
The squamosal carries at least six, and perhaps as many as eight, episquamosals,
450:
125:
1900:
1859:
1790:
1777:
1748:
1733:
1703:
1664:
1650:
1567:
1528:
1407:
1317:
1282:
1275:
1155:
1105:
1028:
1008:
623:
607:
589:. The narrowing is caused by an indentation of the outer rim of the squamosal.
517:
349:
307:
709:
2157:
1978:
1957:
1927:
1918:
1818:
1712:
1657:
1636:
1608:
1601:
1587:
1580:
1386:
1375:
1368:
1310:
1303:
1289:
1247:
1182:
1148:
1119:
1072:
760:
611:
440:
163:
1992:
1943:
1936:
1852:
1825:
1811:
1719:
1692:
1685:
1678:
1629:
1615:
1507:
1494:
1425:
1414:
1399:
1361:
1353:
1335:
1240:
1206:
1189:
1141:
1134:
1090:
895:
843:
717:
576:
487:
336:
291:
212:
1985:
1879:
1762:
1726:
1549:
1261:
1126:
585:
138:
42:
1755:
1671:
1521:
1254:
1042:
976:
963:
922:
801:
593:
443:
323:
183:
148:
610:. It is the oldest chasmosaurine known from Canada, and the first pre-
740:
497:
and Mark Loewen. The generic name combines the name of the Roman god
454:
242:
188:
132:
2071:
24:
2094:
1556:
1542:
1535:
1098:
834:
825:
627:
529:
520:
collected from approximately time-equivalent sections of the upper
446:
275:
262:
178:
173:
158:
153:
143:
556:
2114:
1950:
1112:
1056:
513:, in reference to the similar specimens the species is based on.
462:
458:
433:
193:
168:
819:
252:
770:
436:
479:
614:
ceratopsid to have been collected on both sides of the
493:
was named and described by
Michael Ryan, David Evans,
687:
637:
683:
681:
679:
677:
675:
673:
671:
2155:
465:, United States. It contains a single species,
867:
668:
786:
51:introducing citations to additional sources
1505:
745:. Princeton University Press. p. 293.
2174:Late Cretaceous dinosaurs of North America
793:
779:
211:
555:
41:Relevant discussion may be found on the
516:The species is represented only by two
2156:
742:The Princeton Field Guide to Dinosaurs
2076:
2075:
1492:
961:
812:
774:
738:
18:
13:
14:
2235:
2056:
2055:
2031:
2025:
1493:
1475:
1469:
656:Timeline of ceratopsian research
640:
229:
136:
34:relies largely or entirely on a
23:
2184:Taxa named by Philip J. Currie
962:
800:
732:
551:
1:
2209:Ceratopsians of North America
2179:Fossil taxa described in 2014
661:
505:is that of the constellation
2214:Late Cretaceous ceratopsians
618:. Though the profile of the
575:(2014) established a single
474:
7:
633:
579:, unique derived trait, of
10:
2240:
16:Extinct genus of dinosaurs
2224:Monotypic dinosaur genera
2084:
2045:
2023:
1968:
1926:
1917:
1869:
1835:
1785:
1776:
1702:
1575:
1566:
1516:
1501:
1488:
1466:
1424:
1394:
1385:
1343:
1334:
1230:
1215:
1204:
1165:
1085:
1081:
1070:
1018:
989:
985:
974:
970:
957:
921:
903:
894:
876:
863:
808:
739:Paul, Gregory S. (2016).
710:10.1007/s00114-014-1183-1
403:
396:
226:Scientific classification
224:
219:
210:
118:
509:, named after the twins
1002:Micropachycephalosaurus
526:Dinosaur Park Formation
561:
522:Judith River Formation
491:Mercuriceratops gemini
481:Fergus County, Montana
468:Mercuriceratops gemini
410:Mercuriceratops gemini
2136:Paleobiology Database
559:
542:University of Alberta
2204:Ornithischian genera
538:Royal Ontario Museum
47:improve this article
2189:Dinosaur Park fauna
1167:Archaeoceratopsidae
702:2014NW....101..505R
690:Naturwissenschaften
648:Paleontology portal
2219:Cretaceous Montana
2194:Fossils of Montana
1887:Crittendenceratops
1837:Albertaceratopsini
606:was placed in the
562:
495:Philip John Currie
2164:Fossils of Canada
2151:
2150:
2078:Taxon identifiers
2069:
2068:
2041:
2040:
2021:
2020:
2017:
2016:
2013:
2012:
2009:
2008:
1970:Pachyrhinosaurini
1908:Yehuecauhceratops
1871:Nasutoceratopsini
1772:
1771:
1484:
1483:
1464:
1463:
1460:
1459:
1456:
1455:
1452:
1451:
1448:
1447:
1444:
1443:
1345:Protoceratopsidae
1232:Leptoceratopsidae
1200:
1199:
1066:
1065:
953:
952:
949:
948:
945:
944:
941:
940:
752:978-1-78684-190-2
511:Castor and Pollux
425:
424:
377:
112:
111:
97:
62:"Mercuriceratops"
2231:
2199:Campanian genera
2144:
2143:
2131:
2130:
2118:
2117:
2105:
2104:
2103:
2073:
2072:
2059:
2058:
2036:
2035:
2029:
2000:Pachyrhinosaurus
1924:
1923:
1798:Machairoceratops
1783:
1782:
1623:Coahuilaceratops
1573:
1572:
1514:
1513:
1503:
1502:
1490:
1489:
1479:
1474:
1473:
1392:
1391:
1341:
1340:
1325:Zhuchengceratops
1213:
1212:
1083:
1082:
1079:
1078:
1020:Chaoyangsauridae
987:
986:
983:
982:
972:
971:
959:
958:
901:
900:
874:
873:
865:
864:
858:
857:
810:
809:
795:
788:
781:
772:
771:
765:
764:
736:
730:
729:
685:
650:
645:
644:
643:
616:Canada–US border
546:Oldman Formation
412:
408:
368:
361:
348:
335:
322:
306:
290:
234:
233:
215:
205:
135:
124:Temporal range:
116:
115:
107:
104:
98:
96:
55:
27:
19:
2239:
2238:
2234:
2233:
2232:
2230:
2229:
2228:
2154:
2153:
2152:
2147:
2139:
2134:
2126:
2121:
2113:
2108:
2099:
2098:
2093:
2086:Mercuriceratops
2080:
2070:
2065:
2037:
2030:
2005:
1964:
1913:
1894:Furcatoceratops
1865:
1846:Albertaceratops
1831:
1805:Menefeeceratops
1768:
1741:Tatankaceratops
1698:
1644:Mercuriceratops
1595:Arrhinoceratops
1562:
1510:
1497:
1480:
1468:
1440:
1420:
1381:
1330:
1297:Montanoceratops
1226:
1221:Graciliceratops
1209:
1196:
1176:Archaeoceratops
1161:
1075:
1062:
1050:Xuanhuaceratops
1036:Hualianceratops
1014:
995:Albalophosaurus
979:
966:
937:
917:
890:
884:Avemetatarsalia
869:Avemetatarsalia
859:
853:Marginocephalia
815:
814:
804:
799:
769:
768:
753:
737:
733:
686:
669:
664:
646:
641:
639:
636:
620:Mercuriceratops
604:Mercuriceratops
581:Mercuriceratops
565:Mercuriceratops
554:
518:squamosal bones
477:
451:Late Cretaceous
449:known from the
439:of herbivorous
429:Mercuriceratops
421:
414:
406:
405:
392:
367:
364:Mercuriceratops
359:
346:
333:
320:
304:
288:
228:
206:
204:
203:
202:
201:
196:
191:
186:
181:
176:
171:
166:
161:
156:
151:
146:
141:
130:
129:
126:Late Cretaceous
122:
120:Mercuriceratops
108:
102:
99:
56:
54:
40:
28:
17:
12:
11:
5:
2237:
2227:
2226:
2221:
2216:
2211:
2206:
2201:
2196:
2191:
2186:
2181:
2176:
2171:
2169:Chasmosaurines
2166:
2149:
2148:
2146:
2145:
2132:
2119:
2106:
2090:
2088:
2082:
2081:
2067:
2066:
2064:
2063:
2053:
2046:
2043:
2042:
2039:
2038:
2024:
2022:
2019:
2018:
2015:
2014:
2011:
2010:
2007:
2006:
2004:
2003:
1996:
1989:
1982:
1974:
1972:
1966:
1965:
1963:
1962:
1954:
1947:
1940:
1932:
1930:
1921:
1915:
1914:
1912:
1911:
1904:
1901:Nasutoceratops
1897:
1890:
1883:
1875:
1873:
1867:
1866:
1864:
1863:
1860:Medusaceratops
1856:
1849:
1841:
1839:
1833:
1832:
1830:
1829:
1822:
1815:
1808:
1801:
1794:
1791:Diabloceratops
1786:
1780:
1778:Centrosaurinae
1774:
1773:
1770:
1769:
1767:
1766:
1759:
1752:
1749:Titanoceratops
1745:
1737:
1734:Regaliceratops
1730:
1723:
1716:
1708:
1706:
1704:Triceratopsini
1700:
1699:
1697:
1696:
1689:
1682:
1675:
1668:
1665:Sierraceratops
1661:
1654:
1651:Navajoceratops
1647:
1640:
1633:
1626:
1619:
1612:
1605:
1598:
1591:
1584:
1576:
1570:
1568:Chasmosaurinae
1564:
1563:
1561:
1560:
1553:
1546:
1539:
1532:
1529:Brachyceratops
1525:
1517:
1511:
1506:
1499:
1498:
1486:
1485:
1482:
1481:
1467:
1465:
1462:
1461:
1458:
1457:
1454:
1453:
1450:
1449:
1446:
1445:
1442:
1441:
1439:
1438:
1437:
1436:
1430:
1428:
1422:
1421:
1419:
1418:
1411:
1408:Turanoceratops
1404:
1395:
1389:
1383:
1382:
1380:
1379:
1372:
1365:
1358:
1349:
1347:
1338:
1332:
1331:
1329:
1328:
1321:
1318:Unescoceratops
1314:
1307:
1300:
1293:
1286:
1283:Ischioceratops
1279:
1276:Gryphoceratops
1272:
1265:
1258:
1251:
1244:
1236:
1234:
1228:
1227:
1225:
1224:
1216:
1210:
1205:
1202:
1201:
1198:
1197:
1195:
1194:
1186:
1179:
1171:
1169:
1163:
1162:
1160:
1159:
1156:Sasayamagnomus
1152:
1145:
1138:
1131:
1123:
1116:
1109:
1106:Auroraceratops
1102:
1095:
1086:
1076:
1071:
1068:
1067:
1064:
1063:
1061:
1060:
1053:
1046:
1039:
1032:
1029:Chaoyangsaurus
1024:
1022:
1016:
1015:
1013:
1012:
1009:Psittacosaurus
1005:
998:
990:
980:
975:
968:
967:
955:
954:
951:
950:
947:
946:
943:
942:
939:
938:
936:
935:
934:
933:
927:
925:
919:
918:
916:
915:
914:
913:
904:
898:
892:
891:
889:
888:
887:
886:
877:
871:
861:
860:
856:
855:
846:
837:
828:
822:
806:
805:
798:
797:
790:
783:
775:
767:
766:
751:
731:
696:(6): 505–512.
666:
665:
663:
660:
659:
658:
652:
651:
635:
632:
608:Chasmosaurinae
553:
550:
524:and the lower
476:
473:
423:
422:
415:
401:
400:
394:
393:
389:M. gemini
385:
383:
379:
378:
357:
353:
352:
350:Chasmosaurinae
344:
340:
339:
331:
327:
326:
318:
311:
310:
308:Neornithischia
302:
295:
294:
286:
279:
278:
273:
266:
265:
260:
256:
255:
250:
246:
245:
240:
236:
235:
222:
221:
217:
216:
208:
207:
199:
198:
197:
192:
187:
182:
177:
172:
167:
162:
157:
152:
147:
142:
137:
123:
110:
109:
45:. Please help
31:
29:
22:
15:
9:
6:
4:
3:
2:
2236:
2225:
2222:
2220:
2217:
2215:
2212:
2210:
2207:
2205:
2202:
2200:
2197:
2195:
2192:
2190:
2187:
2185:
2182:
2180:
2177:
2175:
2172:
2170:
2167:
2165:
2162:
2161:
2159:
2142:
2137:
2133:
2129:
2124:
2120:
2116:
2111:
2107:
2102:
2096:
2092:
2091:
2089:
2087:
2083:
2079:
2074:
2062:
2054:
2052:
2048:
2047:
2044:
2034:
2028:
2002:
2001:
1997:
1995:
1994:
1990:
1988:
1987:
1983:
1981:
1980:
1979:Achelousaurus
1976:
1975:
1973:
1971:
1967:
1960:
1959:
1958:Styracosaurus
1955:
1953:
1952:
1948:
1946:
1945:
1941:
1939:
1938:
1934:
1933:
1931:
1929:
1928:Centrosaurini
1925:
1922:
1920:
1919:Eucentrosaura
1916:
1910:
1909:
1905:
1903:
1902:
1898:
1896:
1895:
1891:
1889:
1888:
1884:
1882:
1881:
1877:
1876:
1874:
1872:
1868:
1862:
1861:
1857:
1855:
1854:
1850:
1848:
1847:
1843:
1842:
1840:
1838:
1834:
1828:
1827:
1823:
1821:
1820:
1819:Wendiceratops
1816:
1814:
1813:
1809:
1807:
1806:
1802:
1800:
1799:
1795:
1793:
1792:
1788:
1787:
1784:
1781:
1779:
1775:
1765:
1764:
1760:
1758:
1757:
1753:
1751:
1750:
1746:
1743:
1742:
1738:
1736:
1735:
1731:
1729:
1728:
1724:
1722:
1721:
1717:
1715:
1714:
1713:Eotriceratops
1710:
1709:
1707:
1705:
1701:
1695:
1694:
1690:
1688:
1687:
1683:
1681:
1680:
1676:
1674:
1673:
1669:
1667:
1666:
1662:
1660:
1659:
1658:Pentaceratops
1655:
1653:
1652:
1648:
1646:
1645:
1641:
1639:
1638:
1637:Kosmoceratops
1634:
1632:
1631:
1627:
1625:
1624:
1620:
1618:
1617:
1613:
1611:
1610:
1609:Bravoceratops
1606:
1604:
1603:
1602:Bisticeratops
1599:
1597:
1596:
1592:
1590:
1589:
1588:Anchiceratops
1585:
1583:
1582:
1581:Agujaceratops
1578:
1577:
1574:
1571:
1569:
1565:
1559:
1558:
1554:
1552:
1551:
1547:
1545:
1544:
1540:
1538:
1537:
1533:
1531:
1530:
1526:
1524:
1523:
1519:
1518:
1515:
1512:
1509:
1504:
1500:
1496:
1491:
1487:
1478:
1472:
1434:
1433:
1432:
1431:
1429:
1427:
1423:
1417:
1416:
1412:
1410:
1409:
1405:
1402:
1401:
1397:
1396:
1393:
1390:
1388:
1387:Ceratopsoidea
1384:
1378:
1377:
1376:Protoceratops
1373:
1371:
1370:
1369:Breviceratops
1366:
1364:
1363:
1359:
1356:
1355:
1351:
1350:
1348:
1346:
1342:
1339:
1337:
1333:
1327:
1326:
1322:
1320:
1319:
1315:
1313:
1312:
1311:Udanoceratops
1308:
1306:
1305:
1304:Prenoceratops
1301:
1299:
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1292:
1291:
1290:Leptoceratops
1287:
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1273:
1271:
1270:
1266:
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1259:
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1183:Helioceratops
1180:
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1149:Mosaiceratops
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1132:
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1128:
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1120:Koreaceratops
1117:
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1100:
1096:
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1073:Neoceratopsia
1069:
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624:centrosaurine
621:
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612:Maastrichtian
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503:specific name
500:
496:
492:
489:
486:In 2014, the
484:
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472:
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469:
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461:, Canada and
460:
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441:chasmosaurine
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398:Binomial name
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74:
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67:
64: –
63:
59:
58:Find sources:
52:
48:
44:
38:
37:
36:single source
32:This article
30:
26:
21:
20:
2085:
1998:
1993:Stellasaurus
1991:
1984:
1977:
1956:
1949:
1944:Coronosaurus
1942:
1937:Centrosaurus
1935:
1906:
1899:
1892:
1885:
1878:
1858:
1853:Lokiceratops
1851:
1844:
1826:Xenoceratops
1824:
1817:
1812:Sinoceratops
1810:
1803:
1796:
1789:
1761:
1754:
1747:
1739:
1732:
1725:
1720:Nedoceratops
1718:
1711:
1693:Vagaceratops
1691:
1686:Utahceratops
1684:
1679:Terminocavus
1677:
1670:
1663:
1656:
1649:
1643:
1642:
1635:
1630:Judiceratops
1628:
1621:
1616:Chasmosaurus
1614:
1607:
1600:
1593:
1586:
1579:
1555:
1548:
1541:
1534:
1527:
1520:
1508:Ceratopsidae
1495:Ceratopsidae
1426:Ceratopsidae
1415:Zuniceratops
1413:
1406:
1400:Ajkaceratops
1398:
1374:
1367:
1362:Bagaceratops
1360:
1354:Ajkaceratops
1352:
1336:Coronosauria
1323:
1316:
1309:
1302:
1295:
1288:
1281:
1274:
1267:
1260:
1253:
1246:
1241:Asiaceratops
1239:
1219:
1207:Euceratopsia
1190:Yamaceratops
1188:
1181:
1174:
1154:
1147:
1142:Microceratus
1140:
1135:Liaoceratops
1133:
1125:
1118:
1111:
1104:
1097:
1091:Ajkaceratops
1089:
1055:
1048:
1041:
1034:
1027:
1007:
1000:
993:
911:Ornithischia
896:Ornithischia
848:
844:Ornithischia
839:
830:
741:
734:
693:
689:
619:
603:
602:
597:
591:
584:
580:
577:autapomorphy
572:
570:
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563:
533:
515:
490:
488:type species
485:
478:
467:
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426:
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388:
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370:
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362:
337:Ceratopsidae
314:
298:
292:Ornithischia
282:
269:
220:Restoration
119:
113:
103:October 2021
100:
90:
83:
76:
69:
57:
33:
1986:Einiosaurus
1880:Avaceratops
1763:Triceratops
1727:Ojoceratops
1550:Monoclonius
1262:Ferrisaurus
1127:Kulceratops
586:Triceratops
560:Restoration
552:Description
343:Subfamily:
2158:Categories
2049:See also:
1756:Torosaurus
1672:Spiclypeus
1522:Agathaumas
1435:see below↓
1255:Cerasinops
1043:Stenopelix
977:Ceratopsia
964:Ceratopsia
932:see below↓
923:Ceratopsia
835:Dinosauria
802:Ceratopsia
662:References
594:osteoderms
457:stage) of
444:ceratopsid
324:Ceratopsia
276:Dinosauria
73:newspapers
2101:Q16934814
818:Kingdom:
761:985402380
598:fenestrae
534:ROM 64222
475:Discovery
455:Campanian
382:Species:
249:Kingdom:
243:Eukaryota
43:talk page
2115:42418332
2095:Wikidata
2061:Category
2051:Timeline
1557:Polyonax
1543:Dysganus
1536:Ceratops
1099:Aquilops
826:Chordata
824:Phylum:
820:Animalia
726:13957187
718:24859020
634:See also
628:ontogeny
530:holotype
447:dinosaur
330:Family:
263:Chordata
259:Phylum:
253:Animalia
239:Domain:
131:77
2128:1482557
1951:Spinops
1269:Gremlin
1057:Yinlong
698:Bibcode
499:Mercury
463:Montana
459:Alberta
434:extinct
407:†
386:†
356:Genus:
200:↓
87:scholar
2141:295748
759:
749:
724:
716:
573:et al.
528:. The
507:Gemini
432:is an
420:, 2014
418:et al.
371:et al.
89:
82:
75:
68:
60:
2123:IRMNG
849:Clade
840:Clade
831:Clade
722:S2CID
571:Ryan
437:genus
416:Ryan
369:Ryan
315:Clade
299:Clade
283:Clade
270:Clade
94:JSTOR
80:books
909:see
882:see
757:OCLC
747:ISBN
714:PMID
375:2014
139:PreꞒ
66:news
2110:EoL
1113:Beg
706:doi
694:101
49:by
2160::
2138::
2125::
2112::
2097::
851::
842::
833::
755:.
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704:.
692:.
670:^
630:.
600:.
532:,
483:.
471:.
373:,
317::
301::
285::
272::
189:Pg
133:Ma
128:,
1961:?
1744:?
1403:?
1357:?
1193:?
1130:?
1094:?
794:e
787:t
780:v
763:.
728:.
708::
700::
453:(
360:†
347:†
334:†
321:†
305:†
289:†
194:N
184:K
179:J
174:T
169:P
164:C
159:D
154:S
149:O
144:Ꞓ
105:)
101:(
91:·
84:·
77:·
70:·
53:.
39:.
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