Knowledge

Masked booby

Source 📝

31: 1285: 1350: 1370: 532: 1540: 1079: 1468:, a bird flings its head vigorously. The male may also parade in front of the female, walking with an exaggerated high-stepping gait and intermittently tucking his head in his breast, after collecting nesting material and before the pair begins laying. The male presents small sticks and debris as nesting material in a gesture of symbolic nest-building, which leads to copulation. Afterwards, the pair engages in more symbolic nest-building. The twigs and debris are cleared away later as none is actually used in adorning the nest while in use. 1300:, are anywhere from 180 to 3,700 m (590 to 12,140 ft) deep. On these landforms, masked boobies select sites of generally flat, bare or exposed open ground that lie above the high-tide level with access to the ocean. During the breeding season, the species remains near the colony. At other times, juveniles and some adults disperse widely, though some remain at the colony year-round. Most (but not all) birds return to breed at the colony of their birth; once they begin breeding at a site, they will return there annually. 1382: 87: 852: 1309:
colony on Lord Howe Island numbered in the thousands at the time of the island's discovery in 1788, but has declined to under 500 pairs—mostly on offshore islets with the remainder on two hard-to-access headlands—by 2005. Hunting by humans is thought to have played a role; although rats were introduced to the island in 1918, there has been no evidence they are able to kill chicks or eggs—possibly due to the size of the adult boobies. The masked booby was first recorded breeding on
220: 62: 1025:, in 1988, thought to have become extinct due to Polynesian and then European seafarers and settlers. However, the paleoecologist Richard Holdaway and colleagues cast doubt on the distinctness of the fossil taxon in 2001, and a 2010 review by the New Zealand biologist Tammy Steeves and colleagues of the fossil material and DNA found the two overlapped considerably, and hence the extinct and living entities were found to be the same taxon, now known as 1491:
to eight days between the laying of each egg. Occasionally nests with three eggs are reported; these are probably due to an egg from another nest rolling downhill into the nest. The eggs have an average size of 64 mm × 45 mm (2.5 in × 1.8 in) and weigh 75 g (2.6 oz). They are incubated by both adults for 45 days. Parents incubate the eggs by resting on their
1552:
fish. It generally swallows its catch underwater. Fieldwork at Clipperton Island showed that masked boobies flew on average to 103 km (64 mi) from their colony, with a maximum range of 242 km (150 mi), while feeding their chicks. They did not rest at sea at night, though part of their return trip was at night time for longer expeditions. The masked booby forages with the
277:. It has a typical sulid body shape, with a long pointed yellowish bill, long neck, aerodynamic body, long slender wings and pointed tail. The adult is bright white with black wings, a black tail and a dark face mask; at 75–85 cm (30–33 in) long, it is the largest species of booby. The sexes have similar plumage. This species ranges across tropical oceans, except in the eastern 1479:, breeding takes place year-round, with egg-laying peaking from September to early November, while on nearby Raine Island birds begin laying in or after August, likely peaking September to early November. Eggs are laid between May and September on Lord Howe Island, and early July to early January (peaking in September) on Phillip Island. 554:(IOC). The species has also been called the masked gannet, blue-faced booby, white booby (for its plumage), and whistling booby (for its distinctive call). The Australian ornithologist Doug Dorward promoted the name "white booby" as he felt the blue coloration of its face was less prominent than that of the 1444:, in which the booby shakes its head from side to side or up and down and ruffles its head feathers to make its head look bigger and facial markings more prominent. It may cock its tail and hold its wings up away from its body. Neighbouring boobies may escalate by jabbing and lunging at each other. In the 1551:
The masked booby is a spectacular diver, plunging vertically or near-vertically from heights of anywhere from 12 to 100 m (40 to 330 ft)—but more commonly 15 to 35 m (50 to 115 ft)—above the water into the ocean at high speed, to depths of up to 3 m (9.8 ft) in search of
1486:
can be any time from January to early July, peaking in February and March. On Clipperton Island, egg-laying peaks in November to coordinate with peak fish productivity of the surrounding waters in January (for growing chicks). Masked boobies lay at any time in the Caribbean, peaking between March and
1490:
The nest is a cleared area 0.75 to 1 m (2 ft 6 in to 3 ft 3 in) in diameter, within which is a clearly demarcated 25 to 30 cm (10 to 12 in) shallow (1–2 cm (0.4–0.8 in) deep) depression. A clutch of two chalky white eggs is laid, with an interval of five
1313:
off Norfolk Island in 1908, with devegetation by feral animals creating the open ground preferred by the species. By 2007, an estimated 300 pairs were breeding over the island, though the island flora's regeneration after the removal of feral animals might begin to limit suitable nesting sites. In
1858:
in 1982 and 1983 negatively impacted breeding on Christmas Island as the higher water temperatures reduced food supply. Where usually 1500 pairs nested, no young were observed over this period; 50–60 pairs were observed breeding in October 1983. The Australian government has rated both subspecies
1499:
and wrapping their webbed feet over the eggs, with the outermost toes resting on the ground. Their feet are more vascular at this time. When first hatched, the chicks are about 10 cm (3.9 in) long and weigh around 40–60 g (1.4–2.1 oz), with a sparse covering of white down over
1103:
body shape, with a long pointed bill, long neck, aerodynamic body, long slender wings and pointed tail. The adult is bright white with dark wings and a dark tail. The sexes have similar plumage with no seasonal variation, but females are on average slightly heavier and larger than males. The bare
1162:
The juvenile is a streaked or mottled grey-brown on the head and upperparts, with a whitish neck collar. The wings are dark brown and underparts are white. Its bill is yellowish, face is blue-grey and iris a dark brown. Older immature birds have a broader white collar and rump, and more and more
1308:
in the 1890s. These pigs preyed on the crabs that ate the vegetation. After the elimination of pigs in 1964, the crab population rose and vegetation largely disappeared. This was beneficial to the boobies, as they prefer open ground. Clipperton is on a narrow ridge surrounded by deep water. The
1400:
The masked booby generally flies at least 7 m (23 ft) in height, and at speeds of up to 70 km/h (43 mph). It alternates between gliding and active flying with strong periodic wingbeats. It is often encountered alone, or in a small group when returning to its colony.
1850:", a congenital deformity of one or both wings resulting in flightlessness. This coincided with a season of high nestling mortality that was likely related to low numbers of yellowfin tuna due to possible overfishing at a crucial time in the breeding season. The warm phase ( 1814:
around 1000 years ago. The two species subsequently vanished from the Turks and Caicos Islands. A booby yielded around 1–2 kg (2–5 lb) of meat. European sailors in the area also caught and ate tame boobies. Masked booby young and eggs were eaten by the crew of
1230:. In the Indian Ocean it ranges from the coastlines of the Arabian Peninsula and East Africa across to Sumatra and Western Australia, though it is not found off the coast of the Indian subcontinent. Off the Western Australian coastline it is found as far south as the 1012:
had noted that this bird had dark brown rather than pale irises in 1930, but it was not until 1990 that it was formally investigated by R. M. O'Brien and J. Davies and found to also have longer wings than other populations. They classified it as a new subspecies:
3759: 1798:) as they pass through booby colonies and dig their own nests in large numbers. Rats prey on eggs and young of many seabirds, though the size of masked boobies probably prevents direct predation. On Clipperton Island, rats prey on the crab that eats vegetation. 611:
of the former, should be classified as a separate species. Molecular evidence suggests they most likely diverged between 0.8 and 1.1 million years ago. Complex water currents in the eastern Pacific may have established an environmental barrier leading to
1303:
The largest masked booby colony is on Clipperton Island in the eastern Pacific Ocean, a desert atoll southwest of Mexico. In 2003, 112,000 birds were counted, having recovered from 150 individuals in 1958. The population had suffered from the introduction of
840:
sequences shows that populations in the Indian and Pacific Oceans greatly expanded around 180,000 years ago, and that these became separated from Atlantic populations around 115,000 years ago. Furthermore, within each ocean, there is evidence of reduced
308:
to defend its nest. Potential and mated pairs engage in courtship and greeting displays. The female lays two chalky white eggs in a shallow depression on flat ground away from vegetation. The chicks are born featherless, but are soon covered in white
3700: 1846:), which drove their prey item—flying fish—to the surface, facilitating predation by boobies. It may be that overfishing of tuna adversely impacts the availability of fish there. In 2005, 508 young masked boobies at the colony suffered from " 1234:. In the Pacific, it ranges from Brisbane eastwards. It is found in the Caribbean and Atlantic Ocean south to Ascension Island. In the eastern Pacific off the coast of Colombia and Ecuador, the masked booby is replaced by the Nazca booby. A 1169:
The adult masked booby is distinguished from the related Nazca booby by its yellow rather than orange bill, larger size and less distinctive sexual dimorphism. The latter nests on steep cliffs rather than flat ground. The white
835:
change in size across the masked booby's range. Birds in the Atlantic are the smallest, with the size increasing westwards though the Pacific to the Indian Ocean, where the largest individuals are found. Genetic analysis using
1455:
when a female approaches or leaves his territory. In this display, he paces slowly with his neck and bill pointed upwards—between vertical and 45 degrees—with wings partly raised and whistling faintly with an open bill. In a
351:, on its voyage around the world undertaken between August 1822 and March 1825. In the multi-volume publication by Duperrey about the voyage, Lesson authored the ornithological sections. In his 1829 account of the visit to 3726: 1098:
The largest species of booby, the masked booby ranges from 75 to 85 cm (30 to 33 in) long, with a 160–170 cm (63–67 in) wingspan and 1.2–2.2 kg (2.6–4.9 lb) weight. It has a typical
4489:
Pitman, Robert L.; Ballance, Lisa T.; Bost, Charles A. (2012). "Incidence of Wing Deformities ('Angel Wing') Among Masked Boobies at Clipperton Island: Life History Consequences and Insight into Etiology".
2127: 3655: 1292:
Breeding colonies are located on remote islands, atolls and cays. Lord Howe Island is the southernmost colony. Deep water nearby is important for feeding. As an example, waters around
3072:
Rull, I.L.; NicolĂĄs, L.; Neri-Vera, N.; ArgĂĄez, V.; MartĂ­nez, M.; Torres, R. (2016). "Assortative mating by multiple skin color traits in a seabird with cryptic sexual dichromatism".
3798:
Tunnell, John W.; Chapman, Brian R. (2000). Seabirds of the Campeche Bank Islands, southeastern Gulf of Mexico (Report). Atoll Research Bulletin No. 482. Washington DC: Smithsonian.
1112:
is pale yellow with a greenish tinge, sometimes greyish at the base. Conical in shape, the bill is longer than the head and tapers to a slightly downcurved tip. Backward-pointing
1163:
white feathers on the head until the head is wholly white by 14 to 15 months of age. Full adult plumage is acquired three to four months before the bird turns three years old.
1408:(off Norfolk Island) in September 1979 was recovered and released after being caught in fishing gear 24 years and 9.9 months later some 713 km (443 mi) away off the 1132:
are brown-black. The inner webs of the secondaries are white at the base. The underwing is white except for the brown-black flight-feathers that are not covered by the white
2507:
Orta, J.; Jutglar, F.; Garcia, E.F.J.; Kirwan, G.M.; Boesman, P. (2017). del Hoyo, Josep; Elliott, Andrew; Sargatal, Jordi; Christie, David A.; de Juana, Eduardo (eds.).
1867:
are at risk from rising sea levels, and the rising sea temperatures are calculated to reduce food productivity, which may impact on breeding success of both subspecies.
1523:
Although two eggs are often laid, the younger chick almost always perishes within a few days. This has been observed widely across the species' range. Dorward suspected
1448:, a bird tucks the tip of its bill into its chest, possibly positioned to avoid injury to others. This posture is used against intruders or as advertising for a mate. 1166:
The masked booby is usually silent at sea, but is noisy at the nesting colonies. The main call of male birds is a descending whistle; that of females is a loud honk.
4609: 3305: 3266: 2069: 4467: 3980: 3938: 3227: 3188: 2119: 512:. Sundevall's binomial name was followed as Lesson's 1829 record did not sufficiently describe the species; however, in 1911, the Australian amateur ornithologist 1766: 1460:
display, one bird stares at another of the opposite sex; this generally leads to other displays. Pairs engage in a (mostly) gentler form of jabbing display, and
1069:, but neither is usually considered distinct; the American biologist Robert Pitman and colleagues found no consistent differences between these three subspecies. 1508:, their eyes are open at birth. Their down thickens as they age, and the chicks are quite fluffy by week 5–6. The primaries and rectrices appear by week 8, and 3860:
Clapp, Roger; Robertson, William B. Jr (1986). "Nesting of the Masked Booby on the Dry Tortugas, Florida: The first record for the contiguous United States".
2956: 1776: 4167: 2350: 1428:
relationships with many pairs remaining together over multiple breeding seasons. Highly territorial when nesting, single males and mated pairs engage in
3784: 3725:
Ratcliffe, Norman; Bella, Mike; Pelembe, Tara; Boyle, Dave; Benjamin, Raymond; White, Richard; Godley, Brendan; Stevenson, Jim; Sanders, Sarah (2010).
2830:
Steeves, Tammy E.; Holdaway, Richard N.; Hale, Marie L.; McLay, Emma; McAllan, Ian A. W.; Christian, Margaret; Hauber, Mark E.; Bunce, Michael (2010).
3907: 1520:
at around 120 days (17 weeks) of age. After leaving the nest, young birds are dependent on their parents for 3–4 weeks before dispersing out to sea.
516:
pointed out that although Lesson's 1829 account did not describe the bird, his 1831 account did, and thus predated Sundevall by six years, and hence
3602: 4533:
Schreiber, Ralph W.; Schreiber, Elizabeth Anne (1984). "Central Pacific Seabirds and the El Niño Southern Oscillation: 1982 to 1983 Perspectives".
4294: 5786: 4721: 1756:) prey on eggs and young. On some islands such as Ascension and Saint Helena, feral cats have been a threat to masked boobies. The tick species 1033:. Fieldwork in the Kermadec Islands indicates the bills of adults are bright yellow, and that adult males had brighter yellow feet than females. 1831: 326: 1516:
appear by week 10. They begin losing their down from week 12 onwards, until they are wholly covered by juvenile plumage by week 15 or 16, and
5838: 2741: 2522: 5992: 4410: 3438: 3409: 3835: 3519: 2623: 2712: 1424:
The masked booby begins breeding by around four or five years of age, though can occasionally do so at three years old. Adults form
1412:, New Caledonia in July 2004. The longest distance travelled is 3,152 km (1,959 mi); a bird tagged at Raine Island in the 5750: 1264:
During the monsoon season (midyear), the masked booby is an occasional vagrant along the western coast of India, with records from
3487:"Foraging strategy of masked boobies from the largest colony in the world: relationship to environmental conditions and fisheries" 1475:, egg-laying takes place from January to July, peaking in June, with juvenile birds from April to December. On Moulter Cay in the 1108:
is described either as black or blue-black. It contrasts with the white plumage and gives a mask-like appearance. The bill of the
5799: 2883: 4387: 4348: 3344: 2254: 2197: 2027: 1990: 1954: 551: 2658: 2297: 5897: 4715: 4685: 4603: 4461: 3974: 3299: 3260: 3221: 3182: 2617: 2228: 2063: 6047: 5804: 3679:"Breeding seabird populations in Brazilian oceanic islands: historical review, update and a call for census standardization" 6027: 4762: 2428:
Patterson, S.A.; Morris-Pocock, J.A.; Friesen, V.L (2011). "A multilocus phylogeny of the Sulidae (Aves: Pelecaniformes)".
1061:
in the Pacific Ocean off the coast of Central America. Birds of the latter two locations have been separated as subspecies
4708:
Handbook of Australian, New Zealand & Antarctic Birds. Volume 1: Ratites to ducks; Part B, Australian pelican to ducks
313:. The second chick born generally does not survive and is killed by its elder sibling. These birds are spectacular plunge 304:
far from the mainland and close to deep water required for foraging. Territorial when breeding, the masked booby performs
6037: 3930: 2394: 1855: 6032: 6022: 5698: 508: 6017: 1440:
as other boobies fly overhead, stretching their necks out and forward. More direct trespassers are confronted with a
1436:—making a short flight and holding his wings in a 'V' shape and making a call as he lands. The mated pair engages in 1139:
The subspecies differ slightly in size and sometimes also in the colour of the irises, bill, legs and feet. The race
3149: 1838:, though the population worldwide is decreasing. At Clipperton Island, the colony was benefitted by the presence of 6042: 6012: 5949: 5729: 2933: 2549:"A role for nonphysical barriers to gene flow in the diversification of a highly vagile seabird, the masked booby ( 5843: 1579:, up to 28 cm (11 in) long (rarely up to 41 cm (16 in)) form the bulk of its diet, along with 6002: 5742: 5711: 1310: 525: 4136: 2319: 2474:
Paleornithology of St. Helena Island, South Atlantic Ocean. Smithsonian contributions to paleobiology.Number 23
1405: 1159:, during the breeding season, the leg colour of male birds contains more yellow-red than those of the females. 4232: 3395:"Noteworthy bird records for Micronesia, with a summary of raptor sightings in the Mariana Islands, 1988–1999" 2934:"Geographic variation and reassessment of species limits in the "Masked" Boobies of the Eastern Pacific Ocean" 2609:
Birds of the Horn of Africa: Ethiopia, Eritrea, Djibouti, Somalia, and Socotra – Revised and Expanded Edition
1409: 1451:
There are several displays related to the establishment and maintenance of pair-bonding. The male initiates
5560: 3551:
Priddel, David; Hutton, Ian; Olson, Samantha; Wheeler, Robert (2005). "Breeding biology of Masked Boobies (
3213:
Atlantic Seabird Photo Journal: Off New York, New Jersey, Delaware and Maryland Coasts to Canyons 1967–2006
1860: 343: 5856: 5602: 3903: 1200: 607:). The masked and Nazca boobies were divergent enough to indicate that the latter, formerly regarded as a 5636: 5584: 3617: 1510: 1432:
to mark their ground against neighbours and interlopers. The male advertises his territory to females by
5641: 6052: 4667: 4255: 2382: 1911: 1726: 1553: 1258: 268: 197: 5954: 5876: 5615: 5551: 5205: 4696: 1493: 1416:
in December 1981 was picked up and released at Phillip Island (off Norfolk Island) in December 1986.
6007: 5851: 5340: 5235: 4755: 3727:"The eradication of feral cats from Ascension Island and its subsequent recolonization by seabirds" 5830: 591:) to be each other's closest relatives, their lineage diverging from a line that gave rise to the 5941: 4848: 4677: 4671: 2320:"Seventeenth supplement to the American Ornithologists' Union check-list of North American Birds" 1704: 1472: 1250: 86: 5928: 4091:
Breeding Biology of the Blue-faced Booby (Sula dactylatra personata) on Green Island, Kure Atoll
321:. The species faces few threats; although its population is declining, it is considered to be a 5791: 5623: 5513: 5215: 3453: 3394: 1608: 348: 5817: 4593: 4029:
Woodward, Paul W. (1972). "The natural history of Kure Atoll, Northwestern Hawaiian Islands".
3820: 3393:
Wiles, G.J.; Worthington, D.J.; Beck, Jr., R.E.; Pratt, H.D.; Aguon, C.F.; Pyle, R.L. (2000).
3289: 3250: 2882:
Ismar, Stefanie M. H.; Baird, Karen; Patel, Selina; Millar, Craig D.; Hauber, Mark E. (2010).
2768: 2092: 2053: 1461: 1314:
2006, two pairs nested in a brown booby colony on Morros del PotosĂ­ (White Friars Rocks) near
5923: 5421: 5302: 5276: 5210: 4858: 4853: 4451: 4095: 3964: 3778: 3486: 3211: 3172: 2607: 1592: 1359: 1057:
Breeds in the central and western Pacific and around Australia, as well as off Mexico and on
886: 521: 322: 181: 5915: 4425: 2832:"Merging ancient and modern DNA: extinct seabird taxon rediscovered in the North Tasman Sea" 2478: 5685: 5628: 5571: 5390: 5350: 5320: 4740: 4636:
Dorward, Douglas Fyffe (1962). "Comparative biology of the White Booby and the Brown Booby
4542: 3869: 3564: 3501: 2564: 2509: 2437: 1624: 1182:) has a more wholly black upperwing, and a longer neck and tail and larger head, while the 1121: 837: 616:. Subfossil bones 14,000 years old belonging to the species have been found in deposits on 416: 356: 1539: 8: 5345: 5312: 5111: 5015: 4748: 2612:. Princeton Field Guides. Princeton, New Jersey: Princeton University Press. p. 44. 1888: 1648: 1616: 1544: 1429: 1425: 1325:
off the coast of Brazil, Ascension Island in the south Atlantic, and five islands of the
1284: 1231: 1227: 1223: 1191: 1133: 1129: 1117: 1109: 367: 305: 51: 5765: 5576: 4546: 3873: 3568: 3505: 2568: 2441: 1369: 1125: 5589: 5488: 5240: 4868: 4655: 4574: 4558: 4515: 4507: 4286: 4159: 3885: 3751: 3580: 3089: 2948: 2856: 2831: 2588: 2342: 2193: 1816: 1672: 1600: 1561: 1413: 1349: 1297: 1235: 81: 2902: 30: 5910: 5659: 5463: 5307: 5261: 5185: 4985: 4942: 4863: 4711: 4681: 4599: 4566: 4457: 4290: 4099: 4089: 3970: 3678: 3643: 3367: 3295: 3256: 3217: 3178: 2861: 2815: 2798: 2693: 2613: 2580: 2576: 2453: 2224: 2059: 1811: 1528: 1175: 1058: 751: 592: 580: 540: 5866: 4578: 4519: 4221:"An ecological comparison of oceanic seabird communities of the South Pacific Ocean" 3755: 3601:
Priddel, David; Carlile, Nicholas; Evans, Olwyn; Evans, Beryl; McCoy, Honey (2010).
3584: 3093: 2799:"Avifauna from the Emily Bay settlement site, Norfolk Island: a preliminary account" 2592: 2472: 5997: 5664: 5416: 5266: 5220: 5138: 5042: 4962: 4895: 4793: 4651: 4550: 4499: 4278: 4151: 4046: 4038: 3877: 3799: 3741: 3690: 3572: 3509: 3081: 2851: 2843: 2810: 2572: 2548: 2445: 2334: 2289: 2189: 1906: 1791: 1749: 1390: 1273: 1091: 1005: 1001: 882: 637: 555: 371: 352: 1214:), though adults of that species have clearly demarcated brown and white plumage. 415:. "Black fingers" refers to the splayed wingtips in flight. The Swedish zoologist 5967: 5884: 5651: 5271: 5230: 5101: 5005: 4771: 4554: 4132: 2794: 1982:
Voyage autour du Monde, Exécuté par Ordre du Roi, Sur la Corvette de Sa Majesté,
1760:
was described parasitising nesting blue-faced boobies off the Arabian coast. The
1712: 1584: 1239: 1171: 909: 832: 513: 5716: 4371: 4320: 3928: 3328: 2449: 2248: 2177: 2021: 1980: 1934: 1337:
in the Gulf of Mexico over 1984 and 1985; 19 pairs were recorded there in 1998.
5737: 5724: 5483: 5478: 5473: 5281: 5225: 5190: 5106: 4947: 4642: 2646: 2277: 1839: 1680: 1572:
often harass the species until they disgorge their catch and steal their food.
1330: 1249:
waters off the eastern seaboard of the United States, with single records from
1206:
and a grey rather than yellowish bill. The juvenile masked booby resembles the
1105: 1018: 992: 860: 771: 600: 531: 359:, Lesson mentioned encountering masked boobies, and in a footnote proposed the 4042: 3746: 3085: 2386: 2017: 1976: 1078: 932:
with yellow feet. The subspecies name is derived from the Ancient Greek words
338: 5986: 5812: 5703: 5536: 5457: 5447: 5442: 5370: 5176: 4663: 3821:"Status of seabird nesting populations on Arrecife Alacranes, Gulf of Mexico" 1897: 1835: 1790:
among boobies. On Raine Island and Pandora Cay, nests have been destroyed by
1696: 1640: 1632: 1471:
Breeding takes place at different times of year throughout its range. On the
1326: 1009: 924:. He noted its black mask and blue-grey feet to be distinct from Sundevall's 890: 375: 360: 71: 66: 1245:
In the Atlantic, Caribbean birds occasionally wander north to warm southern
5437: 5427: 5381: 5200: 4570: 2865: 2847: 2584: 2457: 1688: 1527:
on Ascension Island. Siblicide has been observed in the Nazca booby on the
1334: 1293: 1254: 917: 848:
Four subspecies are recognized by the International Ornithologists' Union.
617: 310: 43: 4103: 2293: 431:
September 1827. The species name was derived from the Ancient Greek words
5936: 5889: 5825: 5773: 5545: 5468: 5452: 5331: 5010: 4967: 4957: 4839: 4789: 4411:"Seabird Islands No. 43/1 – Raine Island, Great Barrier Reef, Queensland" 4137:"The role of parents in sibilicidal brood reduction of two booby species" 3146:"Rare Tropical Booby fills up on Northwest Seafood, Catches Flight South" 3145: 1741: 1576: 1569: 1322: 1315: 1269: 1246: 1207: 1201: 1183: 715: 664: 584: 576: 567: 547: 318: 314: 297: 282: 273: 4511: 3603:"A review of the seabirds of Phillip Island in the Norfolk Island Group" 1511: 1065:, and the north west Australian population has been named as subspecies 845:
between populations that does not correspond with any physical barrier.
5962: 5755: 5432: 5401: 5252: 4775: 4562: 4282: 4163: 3889: 3819:
Morales-Vera, T.E.; Ruz-Rosado, F.D.; Velarde, E.; Keith, E.O. (2017).
3695: 2999: 2952: 2740:
van Tets, G.F.; Meredith, C.W.; Fullagar, P.J.; Davidson, P.M. (1988).
2346: 2244: 2173: 1989:(in French). Vol. 1, Part 2. Paris: Arthus Bertrand. p. 494. 1847: 1781: 1664: 1580: 1505: 1483: 1381: 1305: 1087: 613: 608: 479: 290: 219: 138: 4788: 4050: 3804: 3514: 1851: 1017:. Meanwhile, large prehistoric specimens known from the Lord Howe and 271:
in 1831, the masked booby is one of six species of booby in the genus
5410: 5195: 5134: 4503: 4260:
on mainland St Helena, South Atlantic, in the presence of feral cats
3644:"A nesting record of the Masked Booby from Guerrero, Southern Mexico" 3642:
Castillo-Guerrero, Alfredo; Ceyca, Juan Pablo; Mellink, Eric (2007).
2647:"List of birds observed and collected during a voyage in the Red Sea" 1761: 1656: 1524: 1501: 1476: 1113: 961: 939: 878: 842: 460: 438: 384: 98: 5778: 5507: 4155: 3881: 3576: 2338: 1780:
have also been recorded as parasites, the latter possibly spreading
1494: 5677: 5530: 4220: 3011: 1771: 486:
in 1846 from Australia, the species name being the Latin adjective
424: 278: 118: 4710:. Melbourne, Victoria: Oxford University Press. pp. 763–772. 4453:
Talking Taino: Caribbean Natural History from a Native Perspective
4254:
Bolton, M.; Watt, R.; Fowler, E.; Henry, L.; Clingham, E. (2011).
3937:. Australian Government Department of the Environment and Energy. 3818: 3485:
Weimerskirch, Henri; Le Corre, Matthieu; Bost, Charles A. (2008).
1807: 5690: 5610: 5361: 5162: 5096: 5073: 4919: 4891: 4825: 4202: 4200: 4012: 4010: 3177:. Princeton, New Jersey: Princeton University Press. p. 51. 3128: 3126: 3124: 3111: 3109: 3107: 3105: 3103: 3055: 3053: 3040: 3038: 2989: 2987: 2985: 2983: 2981: 2979: 2977: 2547:
Steeves, Tammy E.; Anderson, David J.; Friesen, Vicki L. (2005).
1786: 1100: 317:, plunging into the ocean at high speed in search of prey—mainly 264: 252: 148: 5902: 5037: 2739: 5597: 5087: 5038: 4996: 1583:. Species eaten include various species of flying fish such as 1517: 1265: 851: 260: 108: 4197: 4185: 4007: 3995: 3392: 3288:
Peterson, Roger Tory; Mountfort, Guy; Hollom, P.A.S. (2001) .
3121: 3100: 3050: 3035: 3023: 2974: 2884:"Morphology of the recently re-classified Tasman Masked Booby 2599: 2007:
containing page 494 was published in 1829 (See Cretella, 2010)
1882: 1880: 5672: 5293: 5151: 5062: 5046: 4933: 4908: 4814: 4797: 4115: 4113: 3641: 2427: 1721: 1222:
The masked booby is found across tropical oceans between the
955: 933: 921: 913: 908:
Breeds in the western Indian Ocean. The German ornithologist
565:
The masked booby is one of six species of booby in the genus
454: 432: 378: 301: 256: 158: 4770: 2278:"On some necessary alterations in the nomenclature of birds" 1147:
has dark brown irises and dark grey-green legs and the race
528:
followed in the 17th supplement to their checklist in 1920.
4256:"Re-colonisation and successful breeding of masked boobies 3437:
Pitman, Robert L.; Ballance, Lisa T.; Bost, Charly (2005).
2673: 1877: 1288:
Breeding sites in areas relatively free of vegetation, Oahu
1136:. The legs are yellow-orange or olive. The iris is yellow. 885:. There is significant genetic divergence between birds on 128: 5133: 4110: 4057: 3969:. Sarasota, Florida: Pineapple Press Inc. pp. 95–97. 3724: 3071: 2154: 2142: 1253:
in New Jersey and New York. There are summer records from
281:
and eastern Pacific. In the latter, it is replaced by the
4890: 3929:
Australian Bird & Bat Banding Scheme (ABBBS) (2018).
3550: 3252:
Birds of Maryland, Delaware, and the District of Columbia
2253:. Vol. 2. London: self-published. pp. 506–507. 1404:
Regarding the masked booby's longevity, a bird tagged at
826: 4598:. Collingwood, Victoria: Csiro Publishing. p. 111. 3600: 3484: 2829: 2606:
Redman, Nigel; Stevenson, Terry; Fanshawe, John (2016).
2178:"Descriptions of eleven new species of Australian birds" 2058:. Collingwood, Victoria: Csiro Publishing. p. 151. 1143:
has an orange-yellow bill and olive-grey legs, the race
4456:. Tuscaloosa: University of Alabama Press. p. 44. 3931:"ABBBS Database Search: Sula dactylatra (Masked booby)" 3287: 3216:. Bloomington IN: Xlibris Corporation. pp. 75–76. 2605: 2506: 1531:, and is assumed to occur in the masked booby as well. 4341:
Proceedings of the Entomological Society of Washington
4069: 2546: 2409: 1886: 4253: 3676: 3439:"Clipperton Island: pigsty, rat hole and booby prize" 3432: 3430: 2881: 5317: 4094:. Cambridge, MA: Nuttall Ornithology Club. pp.  3294:. New York: Houghton Mifflin Harcourt. p. 241. 4595:
Climate Change Adaptation Plan for Australian Birds
2387:"Hamerkop, Shoebill, pelicans, boobies, cormorants" 1151:has olive to blueish-grey legs. For the subspecies 4532: 4488: 3677:Mancini, P.L.; Serafini, P.P.; Bugoni, L. (2016). 3436: 3427: 2508: 2223:(5th ed.). London: Cassell Ltd. p. 442. 1935:"The complete collation and dating of the section 1912:10.2305/IUCN.UK.2018-2.RLTS.T22736173A132666363.en 4450:Keegan, William F.; Carlson, Lisabeth A. (2008). 4318: 4219:Ainley, David G.; Boekelheide, Robert J. (1984). 4218: 3966:Common Coastal Birds of Florida and the Caribbean 3480: 3478: 3476: 3474: 2750:, and a description of an extinct new species of 2691: 1810:ate masked and red-footed boobies that nested on 985:van Tets, Meredith, Fullagar & Davidson, 1988 5984: 3326: 3291:A Field Guide to the Birds of Britain and Europe 3174:The Birds of New Jersey: Status and Distribution 2932:Pitman, Robert L.; Jehl, Joseph R. Jnr. (1998). 2792: 2470: 2026:(in French). Paris: F.G. Levrault. p. 601. 1198:) have a buff-yellow crown, shorter tail, white 1174:of the red-footed booby is similar but smaller. 546:"Masked booby" has been designated the official 4694: 4591: 4206: 4191: 4016: 4001: 3906:. National Audubon Society. 12 September 2016. 3255:. Baltimore, Maryland: JHU Press. p. 365. 3132: 3115: 3059: 3044: 3029: 3017: 3005: 2993: 2754:from Lord Howe and Norfolk Islands, Tasman Sea" 2182:Proceedings of the Zoological Society of London 3859: 3797: 3783:: CS1 maint: DOI inactive as of August 2024 ( 3471: 2003:The title page gives the year as 1828 but the 1834:(IUCN) lists the masked booby as a species of 1832:International Union for Conservation of Nature 327:International Union for Conservation of Nature 4756: 4449: 3337:Journal of the Bombay Natural History Society 2927: 2925: 2923: 2698:from Lord Howe, Norfolk and Kermadec Islands" 2120:"NRM 569854 Dysporus cyanops Sundevall, 1838" 1986:, pendant les annĂ©es 1822, 1823, 1824 et 1825 4662: 4380:Bulletin of the British Ornithologists' Club 4333:n. sp., a parasite of the blue-faced booby ( 3720: 3718: 3596: 3594: 3546: 3544: 3542: 3540: 3281: 2735: 2733: 2679: 2542: 2540: 2160: 2148: 2023:TraitĂ© d'Ornithologie, ou Tableau MĂ©thodique 1321:Major nesting areas in the Atlantic include 1261:, as well as waters off the coast of Spain. 4592:Garnett, Stephen; Franklin, Donald (2014). 4526: 4369: 2803:Records of the Australian Museum Supplement 2381: 2172: 1801: 1217: 1021:had been classified as a separate species, 487: 400: 267:. First described by the French naturalist 4763: 4749: 4695:Marchant, S.; Higgins, P.G., eds. (1990). 4022: 3958: 3956: 2931: 2920: 2377: 2375: 2373: 2371: 2051: 2047: 2045: 1736: 1482:In the northern hemisphere, egg-laying on 478:The English ornithologist and bird artist 218: 60: 29: 3803: 3745: 3715: 3694: 3591: 3537: 3513: 3148:. Oregon Coast Aquarium. 7 October 2015. 2855: 2814: 2730: 2537: 2090: 1910: 1419: 4131: 4028: 2877: 2875: 2644: 2519:Handbook of the Birds of the World Alive 1932: 1538: 1380: 1368: 1348: 1283: 1077: 850: 530: 4635: 4119: 4075: 4063: 3953: 3365: 3248: 2415: 2393:. International Ornithologists' Union. 2368: 2275: 2218: 2055:Australian Bird Names: A Complete Guide 2042: 1979:(1829). Duperrey, Louis Isidore (ed.). 1926: 1863:. The low-lying colonies of subspecies 5985: 4087: 3962: 2502: 2500: 2498: 2496: 2494: 2492: 2490: 2488: 2016: 1975: 1825: 1758:Ornithodoros (Alectorobius) muesebecki 827:Intraspecific variation and subspecies 5512: 5511: 5132: 5036: 4889: 4787: 4744: 3333:from the shores of coastal Karnataka" 3209: 3170: 2872: 2430:Molecular Phylogenetics and Evolution 2243: 2166: 2101:Physiographiska SĂ€llskapets Tidskrift 5877:f8dc48e9-337c-43b9-b9fd-399cd8558da3 5743:7b667bc9-c1e5-48d6-9a2e-ad0ac4863bbe 5460:(imperial cormorant, blue-eyed shag) 4612:from the original on 30 October 2023 4470:from the original on 30 October 2023 4408: 3983:from the original on 30 October 2023 3941:from the original on 30 October 2023 3910:from the original on 8 February 2022 3765:from the original on 2 February 2017 3308:from the original on 30 October 2023 3269:from the original on 30 October 2023 3230:from the original on 30 October 2023 3191:from the original on 30 October 2023 2962:from the original on 26 January 2021 2661:from the original on 5 December 2018 2626:from the original on 7 February 2023 2525:from the original on 30 October 2023 2356:from the original on 22 October 2021 2300:from the original on 5 December 2018 2200:from the original on 5 December 2018 2117: 2072:from the original on 30 October 2023 1993:from the original on 10 October 2018 1279: 881:and some Atlantic islands including 427:collected in the Atlantic Ocean on 6 289:), which was formerly regarded as a 5993:IUCN Red List least concern species 4372:"Piroplasmosis in the masked booby 4235:from the original on 11 August 2020 3555:) on Lord Howe Island, Australia". 3525:from the original on 10 August 2020 3372:) along the western coast of India" 2521:. Barcelona, Spain: Lynx Edicions. 2485: 2477:. Smithsonian Institution. p.  2052:Gray, Jeannie; Fraser, Ian (2013). 1898:IUCN Red List of Threatened Species 1333:. The species attempted to nest at 552:International Ornithologists' Union 13: 4656:10.1111/j.1474-919X.1962.tb07244.x 4390:from the original on 20 March 2022 4370:Peirce, M.A.; Feare, C.J. (1978). 4351:from the original on 15 April 2019 4337:) on Hasikiya Island, Arabian Sea" 3841:from the original on 13 April 2021 3703:from the original on 28 March 2021 3658:from the original on 28 March 2021 3415:from the original on 22 April 2018 3347:from the original on 28 March 2021 2742:"Osteological differences between 2718:from the original on 27 April 2018 2694:"A new subspecies of Masked Booby 2692:O'Brien, R.M.; Davies, J. (1990). 2250:Handbook to the Birds of Australia 2194:10.1111/j.1469-7998.1846.tb00135.x 1859:occurring in Australian waters as 920:off the coast of Somalia near the 509:Handbook to the Birds of Australia 14: 6064: 4492:The Wilson Journal of Ornithology 3683:Revista Brasileira de Ornitologia 3368:"Recent records of masked booby ( 2888:breeding on the Kermadec Islands" 2397:from the original on 12 June 2018 2130:from the original on 6 March 2019 1500:their grey to pinkish-grey skin. 1104:skin around the face, throat and 370:of the masked booby in 1831. The 366:. Lesson subsequently provided a 4300:from the original on 2 June 2019 4173:from the original on 28 May 2019 3152:from the original on 1 June 2019 2816:10.3853/j.0812-7387.27.2001.1343 2577:10.1111/j.1365-294X.2005.02713.x 2257:from the original on 29 May 2019 2030:from the original on 30 May 2019 1957:from the original on 30 May 2019 341:was a member of the crew on the 85: 4727:from the original on 7 May 2019 4585: 4482: 4443: 4402: 4376:in the Amirantes, Indian Ocean" 4363: 4312: 4247: 4212: 4125: 4081: 3922: 3896: 3853: 3812: 3791: 3670: 3635: 3386: 3359: 3320: 3242: 3203: 3164: 3138: 3065: 2823: 2786: 2685: 2638: 2464: 2421: 2385:; Donsker, David, eds. (2017). 2312: 2269: 2237: 2212: 1887:BirdLife International (2018). 1746:Chroicocephalus novaehollandiae 1272:states. It is a vagrant to the 1029:as this name has priority over 928:with a blue face, and Lesson's 4628: 3494:Marine Ecology Progress Series 3329:"First report of masked booby 3171:Boyle, William J. Jr. (2011). 2111: 2091:Sundevall, Carl Jakob (1838). 2084: 2010: 1969: 1073: 1: 4676:(Abridged ed.). Oxford: 3935:Bird and bat banding database 1870: 1008:. The New Zealand naturalist 889:off Ascension and those from 526:American Ornithological Union 4859:Christmas Island frigatebird 4555:10.1126/science.225.4663.713 3904:"Dry Tortugas National Park" 3452:(2): 193–194. Archived from 2901:(2): 105–109. Archived from 1861:vulnerable to climate change 1856:El Niño–Southern Oscillation 1340: 962: 940: 461: 439: 385: 7: 6028:Birds of the Atlantic Ocean 5160: 5071: 4917: 4823: 4207:Marchant & Higgins 1990 4192:Marchant & Higgins 1990 4088:Kepler, Cameron B. (1969). 4017:Marchant & Higgins 1990 4002:Marchant & Higgins 1990 3750:(inactive 27 August 2024). 3376:Newsletter for Birdwatchers 3133:Marchant & Higgins 1990 3116:Marchant & Higgins 1990 3060:Marchant & Higgins 1990 3045:Marchant & Higgins 1990 3030:Marchant & Higgins 1990 3018:Marchant & Higgins 1990 3006:Marchant & Higgins 1990 2994:Marchant & Higgins 1990 2450:10.1016/j.ympev.2010.11.021 2391:World Bird List Version 7.1 1046:S. d. californica 332: 300:, generally on islands and 10: 6069: 6038:Birds of the Pacific Ocean 4424:(3): 73–77. Archived from 4319:Hoogstraal, Harry (1969). 3249:Beehler, Bruce M. (2019). 2221:Cassell's Latin Dictionary 2124:Maturhistoriska riksmuseet 1933:Cretella, Massimo (2010). 1731:Sthenoteuthis oualaniensis 1719:) and mullet of the genus 1554:white-bellied storm petrel 1534: 1259:Worcester County, Maryland 956: 934: 855:A breeding pair of subsp. 455: 433: 379: 6048:Taxa named by RenĂ© Lesson 6033:Birds of the Indian Ocean 6023:Birds of Ascension Island 5520: 5399: 5379: 5359: 5329: 5291: 5250: 5174: 5159: 5150: 5146: 5128: 5085: 5070: 5061: 5057: 5032: 4994: 4976: 4931: 4916: 4907: 4903: 4885: 4837: 4822: 4813: 4809: 4783: 4043:10.5479/si.00775630.164.1 3963:Nellis, David W. (2001). 3747:10.1017/S003060530999069X 3616:: 113–127. Archived from 3327:Madhyastha, N.A. (1987). 3086:10.1007/s10336-016-1352-4 2471:Olson, Storrs L. (1975). 2276:Mathews, Gregory (1911). 1905:: e.T22736173A132666363. 1605:Parexocoetus brachypterus 1547:, are a common prey item. 1543:Flying fish, such as the 1031:S. d. fullagari 1015:S. d. fullagari 989:S. d. fullagari 769: 749: 742: 713: 696: 689: 682: 662: 655: 635: 628: 502:. Gould adopted the name 419:described the species as 226: 217: 210: 205: 187: 180: 82:Scientific classification 80: 58: 49: 37: 28: 23: 6018:Birds of the Middle East 5341:Double-crested cormorant 5236:White-breasted cormorant 4374:Sula dactylatra melanops 4335:Sula dactylatra melanops 4225:Studies in Avian Biology 3008:, pp. 763, 771–772. 2767:: 35–57 . Archived from 2680:Liddell & Scott 1980 2161:Liddell & Scott 1980 2149:Liddell & Scott 1980 1802:Relationship with humans 1754:Gallirallus philippensis 1597:Hirundichthys speculiger 1385:Egg and chick of subsp. 1218:Distribution and habitat 1116:line the mandibles. The 571:. A 2011 genetic study ( 6043:Birds described in 1831 6013:Birds of Norfolk Island 5284:(long-tailed cormorant) 4849:Magnificent frigatebird 4678:Oxford University Press 4673:A Greek-English Lexicon 4031:Atoll Research Bulletin 3553:Sula dactylatra tasmani 2886:Sula dactylatra tasmani 1947:Bollettino Malacologico 1737:Predators and parasites 1727:purpleback flying squid 1709:Euleptorhamphus viridis 1473:Cocos (Keeling) Islands 1238:was rescued in 2015 in 1050:S. d. bedouti 1027:S. d. tasmani 296:Nesting takes place in 6003:Birds of the Caribbean 5413:(Magellanic cormorant) 5243:(Temminck's cormorant) 5216:Little black cormorant 4778:(Phalacrocoraciformes) 3366:Kasambe, Raju (2010). 3074:Journal of Ornithology 2848:10.1098/rsbl.2009.0478 2219:Simpson, D.P. (1979). 2118:Johanson, Kjell Arne. 1623:), other fish such as 1613:Cheilopogon atrisignis 1548: 1420:Breeding and courtship 1393: 1374: 1362: 1289: 1095: 863: 583:showed the masked and 543: 488: 401: 349:Louis Isidore Duperrey 337:The French naturalist 5924:Paleobiology Database 5422:New Zealand king shag 5277:Little pied cormorant 5211:Black-faced cormorant 4854:Ascension frigatebird 4664:Liddell, Henry George 2645:Hartlaub, G. (1859). 2294:10.5962/bhl.part.1688 1822:on Lord Howe Island. 1767:Ornithodoros capensis 1645:Decapterus macarellus 1621:Cheilopogon melanurus 1593:mirrorwing flyingfish 1542: 1384: 1372: 1360:French Frigate Shoals 1352: 1296:, at the edge of the 1287: 1276:north of New Guinea. 1081: 1000:The form breeding on 922:town of the same name 887:Boatswain Bird Island 854: 534: 323:least-concern species 269:RenĂ©-PrimevĂšre Lesson 5738:Fauna Europaea (new) 5391:Red-legged cormorant 5351:Flightless cormorant 5321:Spectacled cormorant 5141:: Phalacrocoracidae) 5045:: Anhingidae · 4796:: Fregatidae · 4640:spp. at Ascension". 3210:Brady, Alan (2009). 3020:, pp. 763, 771. 2333:(3): 439–449. 1920. 2282:Novitates Zoologicae 1677:Cirrhitops fasciatus 1653:Coryphaena equiselis 1625:yellowtail amberjack 1509: 1492: 1318:in southern Mexico. 1199: 838:mtDNA control region 417:Carl Jakob Sundevall 357:South Atlantic Ocean 5346:Neotropic cormorant 5313:Red-faced cormorant 5112:Australasian darter 5016:Australasian gannet 4892:Gannets and boobies 4547:1984Sci...225..713S 4409:King, B.R. (1986). 4122:, pp. 194–197. 4066:, pp. 185–186. 3874:1986ColWa...9..113C 3862:Colonial Waterbirds 3623:on 21 February 2021 3569:2005EmuAO.105..105P 3506:2008MEPS..362..291W 3459:on 19 November 2020 2941:The Wilson Bulletin 2793:Holdaway, Richard; 2774:on 17 February 2021 2569:2005MolEc..14.3877S 2442:2011MolPE..58..181P 1826:Conservation status 1777:Amblyomma loculosum 1661:Coryphaena hippurus 1649:pompano dolphinfish 1617:Atlantic flyingfish 1575:Fish, particularly 1545:Atlantic flyingfish 1357:, Courting display, 1232:Dampier Archipelago 1228:30th parallel south 1224:30th parallel north 1192:Australasian gannet 1110:nominate subspecies 1082:Juveniles of subsp 243:), also called the 52:Conservation status 5489:South Georgia shag 5424:(rough-faced shag) 5303:Brandt's cormorant 5241:Japanese cormorant 4869:Lesser frigatebird 4283:10.61350/sbj.24.60 3828:Marine Ornithology 3696:10.1007/BF03544338 3446:Marine Ornithology 2895:Marine Ornithology 2705:Marine Ornithology 1673:redbarred hawkfish 1637:Katsuwonus pelamis 1601:sailfin flyingfish 1589:Exocoetus volitans 1558:Fregetta grallaria 1549: 1442:yes-no headshaking 1430:agonistic displays 1414:Great Barrier Reef 1394: 1375: 1363: 1298:Great Barrier Reef 1290: 1096: 864: 544: 368:formal description 306:agonistic displays 173:S. dactylatra 6053:Pantropical fauna 5978: 5977: 5911:Open Tree of Life 5514:Taxon identifiers 5505: 5504: 5501: 5500: 5497: 5496: 5464:Heard Island shag 5308:Pelagic cormorant 5262:Crowned cormorant 5186:Socotra cormorant 5170: 5169: 5124: 5123: 5120: 5119: 5081: 5080: 5028: 5027: 5024: 5023: 4943:Blue-footed booby 4927: 4926: 4881: 4880: 4877: 4876: 4864:Great frigatebird 4833: 4832: 4717:978-0-19-553068-1 4687:978-0-19-910207-5 4605:978-0-643-10803-5 4541:(4663): 713–716. 4463:978-0-8173-5508-1 3976:978-1-56164-191-8 3515:10.3354/meps07424 3301:978-0-618-16675-6 3262:978-1-4214-2733-1 3223:978-1-4500-8159-7 3184:978-1-4008-3828-8 2619:978-0-691-17289-7 2563:(12): 3877–3887. 2557:Molecular Ecology 2230:978-0-304-52257-6 2065:978-0-643-10471-6 1844:Thunnus albacares 1812:Grand Turk Island 1792:green sea turtles 1750:buff-banded rails 1669:Kyphosus bigibbus 1609:glider flyingfish 1566:Bulweria bulwerii 1529:Galapagos Islands 1466:oblique headshake 1434:flight circuiting 1398: 1397: 1280:Breeding colonies 1268:, Karnataka, and 1155:and the nominate 1059:Clipperton Island 1043: 986: 903: 872: 859:and their chick, 823: 822: 814: 813: 805: 804: 796: 795: 787: 786: 752:Blue-footed booby 731: 730: 581:mitochondrial DNA 541:Grand Turk Island 293:of masked booby. 231: 230: 227:Range (in green) 75: 6060: 5971: 5970: 5958: 5957: 5945: 5944: 5932: 5931: 5919: 5918: 5906: 5905: 5893: 5892: 5880: 5879: 5870: 5869: 5860: 5859: 5847: 5846: 5834: 5833: 5821: 5820: 5808: 5807: 5795: 5794: 5782: 5781: 5769: 5768: 5759: 5758: 5746: 5745: 5733: 5732: 5720: 5719: 5707: 5706: 5694: 5693: 5681: 5680: 5668: 5667: 5655: 5654: 5645: 5644: 5632: 5631: 5619: 5618: 5616:22D1B4A2F0911712 5606: 5605: 5593: 5592: 5580: 5579: 5567: 5566: 5556: 5555: 5554: 5541: 5540: 5539: 5509: 5508: 5417:Guanay cormorant 5319: 5267:Little cormorant 5221:Indian cormorant 5157: 5156: 5148: 5147: 5130: 5129: 5068: 5067: 5059: 5058: 5034: 5033: 4963:Red-footed booby 4914: 4913: 4905: 4904: 4887: 4886: 4820: 4819: 4811: 4810: 4785: 4784: 4765: 4758: 4751: 4742: 4741: 4736: 4734: 4732: 4726: 4705: 4691: 4659: 4622: 4621: 4619: 4617: 4589: 4583: 4582: 4530: 4524: 4523: 4504:10.1676/11-208.1 4486: 4480: 4479: 4477: 4475: 4447: 4441: 4440: 4438: 4436: 4431:on 15 April 2019 4430: 4415: 4406: 4400: 4399: 4397: 4395: 4367: 4361: 4360: 4358: 4356: 4316: 4310: 4309: 4307: 4305: 4299: 4268: 4251: 4245: 4244: 4242: 4240: 4216: 4210: 4204: 4195: 4189: 4183: 4182: 4180: 4178: 4172: 4141: 4133:Anderson, Atholl 4129: 4123: 4117: 4108: 4107: 4085: 4079: 4073: 4067: 4061: 4055: 4054: 4026: 4020: 4014: 4005: 3999: 3993: 3992: 3990: 3988: 3960: 3951: 3950: 3948: 3946: 3926: 3920: 3919: 3917: 3915: 3900: 3894: 3893: 3857: 3851: 3850: 3848: 3846: 3840: 3825: 3816: 3810: 3809: 3807: 3795: 3789: 3788: 3782: 3774: 3772: 3770: 3764: 3749: 3731: 3722: 3713: 3712: 3710: 3708: 3698: 3674: 3668: 3667: 3665: 3663: 3639: 3633: 3632: 3630: 3628: 3622: 3607: 3598: 3589: 3588: 3548: 3535: 3534: 3532: 3530: 3524: 3517: 3491: 3482: 3469: 3468: 3466: 3464: 3458: 3443: 3434: 3425: 3424: 3422: 3420: 3414: 3399: 3390: 3384: 3383: 3363: 3357: 3356: 3354: 3352: 3324: 3318: 3317: 3315: 3313: 3285: 3279: 3278: 3276: 3274: 3246: 3240: 3239: 3237: 3235: 3207: 3201: 3200: 3198: 3196: 3168: 3162: 3161: 3159: 3157: 3142: 3136: 3130: 3119: 3113: 3098: 3097: 3080:(4): 1049–1062. 3069: 3063: 3057: 3048: 3042: 3033: 3027: 3021: 3015: 3009: 3003: 2997: 2991: 2972: 2971: 2969: 2967: 2961: 2938: 2929: 2918: 2917: 2915: 2913: 2907: 2892: 2879: 2870: 2869: 2859: 2827: 2821: 2820: 2818: 2795:Anderson, Atholl 2790: 2784: 2783: 2781: 2779: 2773: 2758: 2737: 2728: 2727: 2725: 2723: 2717: 2702: 2689: 2683: 2677: 2671: 2670: 2668: 2666: 2642: 2636: 2635: 2633: 2631: 2603: 2597: 2596: 2544: 2535: 2534: 2532: 2530: 2516: 2504: 2483: 2482: 2468: 2462: 2461: 2425: 2419: 2413: 2407: 2406: 2404: 2402: 2379: 2366: 2365: 2363: 2361: 2355: 2324: 2316: 2310: 2309: 2307: 2305: 2273: 2267: 2266: 2264: 2262: 2241: 2235: 2234: 2216: 2210: 2209: 2207: 2205: 2170: 2164: 2158: 2152: 2146: 2140: 2139: 2137: 2135: 2115: 2109: 2108: 2095:Dysporus cyanops 2088: 2082: 2081: 2079: 2077: 2049: 2040: 2039: 2037: 2035: 2014: 2008: 2002: 2000: 1998: 1973: 1967: 1966: 1964: 1962: 1930: 1924: 1923: 1921: 1919: 1914: 1884: 1685:Gempylus serpens 1513: 1496: 1391:Ascension Island 1345: 1344: 1274:Caroline Islands 1212:Sula leucogaster 1203: 1180:Papasula abbotti 1041: 1006:Kermadec Islands 984: 975: 972: 969: 965: 959: 958: 953: 950: 947: 943: 937: 936: 901: 893:off Puerto Rico. 883:Ascension Island 870: 868:S. d. dactylatra 745: 744: 692: 691: 685: 684: 669:Sula leucogaster 658: 657: 638:Red-footed booby 631: 630: 624: 623: 601:Peruvian boobies 556:red-footed booby 518:Sula dactylactra 501: 498: 495: 491: 474: 471: 468: 464: 458: 457: 452: 449: 446: 442: 436: 435: 430: 421:Dysporus cyanops 414: 411: 408: 404: 399:, and the Latin 398: 395: 392: 388: 382: 381: 372:specific epithet 353:Ascension Island 249:blue-faced booby 222: 193: 90: 89: 69: 64: 63: 33: 21: 20: 6068: 6067: 6063: 6062: 6061: 6059: 6058: 6057: 6008:Birds of Hawaii 5983: 5982: 5979: 5974: 5968:Sula-dactylatra 5966: 5961: 5953: 5948: 5940: 5935: 5927: 5922: 5914: 5909: 5901: 5896: 5888: 5885:Observation.org 5883: 5875: 5873: 5865: 5863: 5855: 5850: 5842: 5837: 5829: 5824: 5816: 5811: 5803: 5798: 5790: 5785: 5777: 5772: 5764: 5762: 5754: 5749: 5741: 5736: 5728: 5723: 5715: 5710: 5702: 5697: 5689: 5684: 5676: 5671: 5663: 5658: 5650: 5648: 5640: 5635: 5627: 5622: 5614: 5609: 5603:sula-dactylatra 5601: 5596: 5590:Sula_dactylatra 5588: 5583: 5577:Sula_dactylatra 5575: 5570: 5564: 5559: 5552:Sula dactylatra 5550: 5549: 5544: 5535: 5534: 5529: 5522:Sula dactylatra 5516: 5506: 5493: 5395: 5375: 5355: 5325: 5287: 5272:Pygmy cormorant 5246: 5231:Great cormorant 5166: 5142: 5116: 5102:Oriental darter 5077: 5053: 5020: 5006:Northern gannet 4990: 4972: 4923: 4899: 4873: 4829: 4805: 4779: 4769: 4739: 4730: 4728: 4724: 4718: 4703: 4699:Sula dactylatra 4688: 4631: 4626: 4625: 4615: 4613: 4606: 4590: 4586: 4531: 4527: 4487: 4483: 4473: 4471: 4464: 4448: 4444: 4434: 4432: 4428: 4413: 4407: 4403: 4393: 4391: 4368: 4364: 4354: 4352: 4317: 4313: 4303: 4301: 4297: 4266: 4258:Sula dactylatra 4252: 4248: 4238: 4236: 4217: 4213: 4205: 4198: 4190: 4186: 4176: 4174: 4170: 4156:10.2307/4089018 4139: 4130: 4126: 4118: 4111: 4086: 4082: 4074: 4070: 4062: 4058: 4037:(164): 1–318 . 4027: 4023: 4015: 4008: 4000: 3996: 3986: 3984: 3977: 3961: 3954: 3944: 3942: 3927: 3923: 3913: 3911: 3902: 3901: 3897: 3882:10.2307/1521152 3858: 3854: 3844: 3842: 3838: 3823: 3817: 3813: 3796: 3792: 3776: 3775: 3768: 3766: 3762: 3729: 3723: 3716: 3706: 3704: 3675: 3671: 3661: 3659: 3640: 3636: 3626: 3624: 3620: 3605: 3599: 3592: 3577:10.1071/MU04028 3549: 3538: 3528: 3526: 3522: 3489: 3483: 3472: 3462: 3460: 3456: 3441: 3435: 3428: 3418: 3416: 3412: 3408:(2): 257–284 . 3397: 3391: 3387: 3370:Sula dactylatra 3364: 3360: 3350: 3348: 3325: 3321: 3311: 3309: 3302: 3286: 3282: 3272: 3270: 3263: 3247: 3243: 3233: 3231: 3224: 3208: 3204: 3194: 3192: 3185: 3169: 3165: 3155: 3153: 3144: 3143: 3139: 3131: 3122: 3114: 3101: 3070: 3066: 3058: 3051: 3043: 3036: 3028: 3024: 3016: 3012: 3004: 3000: 2992: 2975: 2965: 2963: 2959: 2936: 2930: 2921: 2911: 2909: 2908:on 22 July 2018 2905: 2890: 2880: 2873: 2836:Biology Letters 2828: 2824: 2791: 2787: 2777: 2775: 2771: 2756: 2738: 2731: 2721: 2719: 2715: 2700: 2696:Sula dactylatra 2690: 2686: 2678: 2674: 2664: 2662: 2657:(4): 337–352 . 2643: 2639: 2629: 2627: 2620: 2604: 2600: 2551:Sula dactylatra 2545: 2538: 2528: 2526: 2512:Sula dactylatra 2510:"Masked booby ( 2505: 2486: 2469: 2465: 2426: 2422: 2414: 2410: 2400: 2398: 2380: 2369: 2359: 2357: 2353: 2339:10.2307/4073271 2322: 2318: 2317: 2313: 2303: 2301: 2274: 2270: 2260: 2258: 2242: 2238: 2231: 2217: 2213: 2203: 2201: 2171: 2167: 2159: 2155: 2147: 2143: 2133: 2131: 2116: 2112: 2089: 2085: 2075: 2073: 2066: 2050: 2043: 2033: 2031: 2015: 2011: 1996: 1994: 1974: 1970: 1960: 1958: 1931: 1927: 1917: 1915: 1891:Sula dactylatra 1885: 1878: 1873: 1828: 1804: 1739: 1713:flat needlefish 1705:ribbon halfbeak 1701:Cololabis saira 1629:Seriola lalandi 1585:blue flyingfish 1562:Bulwer's petrel 1537: 1515: 1498: 1446:pelican posture 1422: 1358: 1343: 1282: 1240:Newport, Oregon 1220: 1205: 1092:Hawaiian island 1076: 1039:S. d. personata 995:): Tasman booby 973: 970: 967: 951: 948: 945: 912:described this 910:Gustav Hartlaub 829: 824: 815: 806: 797: 788: 732: 700:Sula dactylatra 514:Gregory Mathews 499: 496: 493: 472: 469: 466: 450: 447: 444: 428: 423:in 1837 from a 412: 409: 406: 396: 393: 390: 364:Sula dactylatra 347:, captained by 335: 240:Sula dactylatra 201: 195: 191:Sula dactylatra 189: 176: 84: 76: 65: 61: 54: 17: 16:Species of bird 12: 11: 5: 6066: 6056: 6055: 6050: 6045: 6040: 6035: 6030: 6025: 6020: 6015: 6010: 6005: 6000: 5995: 5976: 5975: 5973: 5972: 5959: 5946: 5933: 5920: 5907: 5894: 5881: 5871: 5861: 5848: 5835: 5822: 5809: 5796: 5783: 5770: 5760: 5747: 5734: 5725:Fauna Europaea 5721: 5708: 5695: 5682: 5669: 5656: 5646: 5633: 5620: 5607: 5594: 5581: 5568: 5557: 5542: 5526: 5524: 5518: 5517: 5503: 5502: 5499: 5498: 5495: 5494: 5492: 5491: 5486: 5484:Macquarie shag 5481: 5479:Kerguelen shag 5476: 5474:Antarctic shag 5471: 5466: 5461: 5455: 5450: 5445: 5440: 5435: 5430: 5425: 5419: 5414: 5407: 5405: 5397: 5396: 5394: 5393: 5387: 5385: 5377: 5376: 5374: 5373: 5367: 5365: 5357: 5356: 5354: 5353: 5348: 5343: 5337: 5335: 5327: 5326: 5324: 5323: 5315: 5310: 5305: 5299: 5297: 5289: 5288: 5286: 5285: 5282:Reed cormorant 5279: 5274: 5269: 5264: 5258: 5256: 5248: 5247: 5245: 5244: 5238: 5233: 5228: 5226:Cape cormorant 5223: 5218: 5213: 5208: 5206:Pied cormorant 5203: 5198: 5193: 5191:Bank cormorant 5188: 5182: 5180: 5172: 5171: 5168: 5167: 5154: 5144: 5143: 5126: 5125: 5122: 5121: 5118: 5117: 5115: 5114: 5109: 5107:African darter 5104: 5099: 5093: 5091: 5083: 5082: 5079: 5078: 5065: 5055: 5054: 5030: 5029: 5026: 5025: 5022: 5021: 5019: 5018: 5013: 5008: 5002: 5000: 4992: 4991: 4989: 4988: 4986:Abbott's booby 4982: 4980: 4974: 4973: 4971: 4970: 4965: 4960: 4955: 4950: 4948:Peruvian booby 4945: 4939: 4937: 4929: 4928: 4925: 4924: 4911: 4901: 4900: 4883: 4882: 4879: 4878: 4875: 4874: 4872: 4871: 4866: 4861: 4856: 4851: 4845: 4843: 4835: 4834: 4831: 4830: 4817: 4807: 4806: 4781: 4780: 4768: 4767: 4760: 4753: 4745: 4738: 4737: 4716: 4692: 4686: 4660: 4650:(2): 174–220. 4632: 4630: 4627: 4624: 4623: 4604: 4584: 4525: 4498:(3): 597–602. 4481: 4462: 4442: 4401: 4362: 4311: 4246: 4211: 4209:, p. 766. 4196: 4194:, p. 765. 4184: 4150:(4): 860–869. 4124: 4109: 4080: 4078:, p. 179. 4068: 4056: 4021: 4019:, p. 768. 4006: 4004:, p. 767. 3994: 3975: 3952: 3921: 3895: 3868:(1): 113–116. 3852: 3811: 3790: 3714: 3689:(2): 94–115 . 3669: 3634: 3590: 3563:(2): 105–113. 3536: 3470: 3426: 3385: 3358: 3331:Sula dactylara 3319: 3300: 3280: 3261: 3241: 3222: 3202: 3183: 3163: 3137: 3135:, p. 764. 3120: 3118:, p. 769. 3099: 3064: 3062:, p. 771. 3049: 3047:, p. 770. 3034: 3032:, p. 772. 3022: 3010: 2998: 2996:, p. 763. 2973: 2947:(2): 155–170. 2919: 2871: 2822: 2785: 2729: 2684: 2682:, p. 431. 2672: 2637: 2618: 2598: 2536: 2484: 2463: 2436:(2): 181–191. 2420: 2418:, p. 174. 2408: 2367: 2311: 2268: 2236: 2229: 2211: 2165: 2163:, p. 804. 2153: 2151:, p. 397. 2141: 2126:(in Swedish). 2110: 2103:(in Swedish). 2083: 2064: 2041: 2009: 1968: 1925: 1875: 1874: 1872: 1869: 1840:yellowfin tuna 1827: 1824: 1803: 1800: 1796:Chelonia mydas 1738: 1735: 1717:Ablennes hians 1681:snake mackerel 1536: 1533: 1421: 1418: 1396: 1395: 1377: 1376: 1365: 1364: 1342: 1339: 1331:Gulf of Mexico 1281: 1278: 1219: 1216: 1196:Morus serrator 1188:Morus capensis 1176:Abbott's booby 1075: 1072: 1071: 1070: 1054: 1053: 1035: 1034: 1019:Norfolk Island 997: 996: 993:junior synonym 978: 977: 905: 904: 902:Hartlaub, 1859 899:S. d. melanops 895: 894: 877:Breeds in the 874: 873: 861:Norfolk Island 828: 825: 821: 820: 817: 816: 812: 811: 808: 807: 803: 802: 799: 798: 794: 793: 790: 789: 785: 784: 781: 780: 776:Sula variegata 772:Peruvian booby 768: 765: 764: 761: 760: 748: 743: 741: 738: 737: 734: 733: 729: 728: 725: 724: 712: 709: 708: 705: 704: 698:Masked booby ( 695: 690: 688: 683: 681: 678: 677: 674: 673: 661: 656: 654: 651: 650: 647: 646: 634: 629: 627: 622: 605:Sula variegata 573:depicted below 484:Sula personata 334: 331: 229: 228: 224: 223: 215: 214: 208: 207: 203: 202: 196: 185: 184: 178: 177: 170: 168: 164: 163: 156: 152: 151: 146: 142: 141: 136: 132: 131: 126: 122: 121: 116: 112: 111: 106: 102: 101: 96: 92: 91: 78: 77: 59: 56: 55: 50: 47: 46: 38:Adult, subsp. 35: 34: 26: 25: 15: 9: 6: 4: 3: 2: 6065: 6054: 6051: 6049: 6046: 6044: 6041: 6039: 6036: 6034: 6031: 6029: 6026: 6024: 6021: 6019: 6016: 6014: 6011: 6009: 6006: 6004: 6001: 5999: 5996: 5994: 5991: 5990: 5988: 5981: 5969: 5964: 5960: 5956: 5951: 5947: 5943: 5938: 5934: 5930: 5925: 5921: 5917: 5912: 5908: 5904: 5899: 5895: 5891: 5886: 5882: 5878: 5872: 5868: 5862: 5858: 5853: 5849: 5845: 5840: 5836: 5832: 5827: 5823: 5819: 5814: 5810: 5806: 5801: 5797: 5793: 5788: 5784: 5780: 5775: 5771: 5767: 5761: 5757: 5752: 5748: 5744: 5739: 5735: 5731: 5726: 5722: 5718: 5713: 5709: 5705: 5700: 5696: 5692: 5687: 5683: 5679: 5674: 5670: 5666: 5661: 5657: 5653: 5647: 5643: 5638: 5634: 5630: 5625: 5621: 5617: 5612: 5608: 5604: 5599: 5595: 5591: 5586: 5582: 5578: 5573: 5569: 5562: 5558: 5553: 5547: 5543: 5538: 5532: 5528: 5527: 5525: 5523: 5519: 5515: 5510: 5490: 5487: 5485: 5482: 5480: 5477: 5475: 5472: 5470: 5467: 5465: 5462: 5459: 5458:Imperial shag 5456: 5454: 5451: 5449: 5448:Campbell shag 5446: 5444: 5443:Auckland shag 5441: 5439: 5436: 5434: 5431: 5429: 5426: 5423: 5420: 5418: 5415: 5412: 5409: 5408: 5406: 5404: 5403: 5398: 5392: 5389: 5388: 5386: 5384: 5383: 5378: 5372: 5371:European shag 5369: 5368: 5366: 5364: 5363: 5358: 5352: 5349: 5347: 5344: 5342: 5339: 5338: 5336: 5334: 5333: 5328: 5322: 5316: 5314: 5311: 5309: 5306: 5304: 5301: 5300: 5298: 5296: 5295: 5290: 5283: 5280: 5278: 5275: 5273: 5270: 5268: 5265: 5263: 5260: 5259: 5257: 5255: 5254: 5249: 5242: 5239: 5237: 5234: 5232: 5229: 5227: 5224: 5222: 5219: 5217: 5214: 5212: 5209: 5207: 5204: 5202: 5199: 5197: 5194: 5192: 5189: 5187: 5184: 5183: 5181: 5179: 5178: 5177:Phalacrocorax 5173: 5165: 5164: 5158: 5155: 5153: 5149: 5145: 5140: 5136: 5131: 5127: 5113: 5110: 5108: 5105: 5103: 5100: 5098: 5095: 5094: 5092: 5090: 5089: 5084: 5076: 5075: 5069: 5066: 5064: 5060: 5056: 5051: 5048: 5044: 5040: 5035: 5031: 5017: 5014: 5012: 5009: 5007: 5004: 5003: 5001: 4999: 4998: 4993: 4987: 4984: 4983: 4981: 4979: 4975: 4969: 4966: 4964: 4961: 4959: 4956: 4954: 4951: 4949: 4946: 4944: 4941: 4940: 4938: 4936: 4935: 4930: 4922: 4921: 4915: 4912: 4910: 4906: 4902: 4897: 4893: 4888: 4884: 4870: 4867: 4865: 4862: 4860: 4857: 4855: 4852: 4850: 4847: 4846: 4844: 4842: 4841: 4836: 4828: 4827: 4821: 4818: 4816: 4812: 4808: 4803: 4799: 4795: 4791: 4786: 4782: 4777: 4773: 4766: 4761: 4759: 4754: 4752: 4747: 4746: 4743: 4723: 4719: 4713: 4709: 4702: 4701:Masked booby" 4700: 4693: 4689: 4683: 4679: 4675: 4674: 4669: 4668:Scott, Robert 4665: 4661: 4657: 4653: 4649: 4645: 4644: 4639: 4634: 4633: 4611: 4607: 4601: 4597: 4596: 4588: 4580: 4576: 4572: 4568: 4564: 4560: 4556: 4552: 4548: 4544: 4540: 4536: 4529: 4521: 4517: 4513: 4509: 4505: 4501: 4497: 4493: 4485: 4469: 4465: 4459: 4455: 4454: 4446: 4427: 4423: 4419: 4412: 4405: 4389: 4385: 4381: 4377: 4375: 4366: 4350: 4346: 4342: 4338: 4336: 4332: 4328: 4324: 4315: 4296: 4292: 4288: 4284: 4280: 4276: 4272: 4265: 4263: 4259: 4250: 4234: 4230: 4226: 4222: 4215: 4208: 4203: 4201: 4193: 4188: 4169: 4165: 4161: 4157: 4153: 4149: 4145: 4138: 4134: 4128: 4121: 4116: 4114: 4105: 4101: 4097: 4093: 4092: 4084: 4077: 4072: 4065: 4060: 4052: 4048: 4044: 4040: 4036: 4032: 4025: 4018: 4013: 4011: 4003: 3998: 3982: 3978: 3972: 3968: 3967: 3959: 3957: 3940: 3936: 3932: 3925: 3909: 3905: 3899: 3891: 3887: 3883: 3879: 3875: 3871: 3867: 3863: 3856: 3837: 3833: 3829: 3822: 3815: 3806: 3801: 3794: 3786: 3780: 3761: 3757: 3753: 3748: 3743: 3739: 3735: 3728: 3721: 3719: 3702: 3697: 3692: 3688: 3684: 3680: 3673: 3657: 3653: 3649: 3648:Western Birds 3645: 3638: 3619: 3615: 3611: 3604: 3597: 3595: 3586: 3582: 3578: 3574: 3570: 3566: 3562: 3558: 3554: 3547: 3545: 3543: 3541: 3521: 3516: 3511: 3507: 3503: 3499: 3495: 3488: 3481: 3479: 3477: 3475: 3455: 3451: 3447: 3440: 3433: 3431: 3411: 3407: 3403: 3396: 3389: 3381: 3377: 3373: 3371: 3362: 3346: 3342: 3338: 3334: 3332: 3323: 3307: 3303: 3297: 3293: 3292: 3284: 3268: 3264: 3258: 3254: 3253: 3245: 3229: 3225: 3219: 3215: 3214: 3206: 3190: 3186: 3180: 3176: 3175: 3167: 3151: 3147: 3141: 3134: 3129: 3127: 3125: 3117: 3112: 3110: 3108: 3106: 3104: 3095: 3091: 3087: 3083: 3079: 3075: 3068: 3061: 3056: 3054: 3046: 3041: 3039: 3031: 3026: 3019: 3014: 3007: 3002: 2995: 2990: 2988: 2986: 2984: 2982: 2980: 2978: 2958: 2954: 2950: 2946: 2942: 2935: 2928: 2926: 2924: 2904: 2900: 2896: 2889: 2887: 2878: 2876: 2867: 2863: 2858: 2853: 2849: 2845: 2841: 2837: 2833: 2826: 2817: 2812: 2808: 2804: 2800: 2796: 2789: 2770: 2766: 2762: 2755: 2753: 2749: 2745: 2736: 2734: 2714: 2710: 2706: 2699: 2697: 2688: 2681: 2676: 2660: 2656: 2652: 2648: 2641: 2625: 2621: 2615: 2611: 2610: 2602: 2594: 2590: 2586: 2582: 2578: 2574: 2570: 2566: 2562: 2558: 2554: 2552: 2543: 2541: 2524: 2520: 2515: 2513: 2503: 2501: 2499: 2497: 2495: 2493: 2491: 2489: 2480: 2476: 2475: 2467: 2459: 2455: 2451: 2447: 2443: 2439: 2435: 2431: 2424: 2417: 2412: 2396: 2392: 2388: 2384: 2378: 2376: 2374: 2372: 2352: 2348: 2344: 2340: 2336: 2332: 2328: 2321: 2315: 2299: 2295: 2291: 2287: 2283: 2279: 2272: 2256: 2252: 2251: 2246: 2240: 2232: 2226: 2222: 2215: 2199: 2195: 2191: 2187: 2183: 2179: 2175: 2169: 2162: 2157: 2150: 2145: 2129: 2125: 2121: 2114: 2106: 2102: 2098: 2096: 2087: 2071: 2067: 2061: 2057: 2056: 2048: 2046: 2029: 2025: 2024: 2019: 2013: 2006: 1992: 1988: 1987: 1983: 1978: 1972: 1956: 1952: 1948: 1944: 1942: 1938: 1929: 1913: 1908: 1904: 1900: 1899: 1894: 1892: 1883: 1881: 1876: 1868: 1866: 1862: 1857: 1853: 1849: 1845: 1841: 1837: 1836:least concern 1833: 1823: 1821: 1820: 1813: 1809: 1799: 1797: 1793: 1789: 1788: 1783: 1782:piroplasmosis 1779: 1778: 1773: 1769: 1768: 1763: 1759: 1755: 1751: 1747: 1743: 1734: 1732: 1728: 1724: 1723: 1718: 1714: 1710: 1706: 1702: 1698: 1697:Pacific saury 1694: 1693:Auxis thazard 1690: 1686: 1682: 1678: 1674: 1670: 1666: 1662: 1658: 1654: 1650: 1646: 1642: 1641:mackerel scad 1638: 1634: 1633:skipjack tuna 1630: 1626: 1622: 1618: 1614: 1610: 1606: 1602: 1598: 1594: 1590: 1586: 1582: 1578: 1573: 1571: 1567: 1563: 1559: 1555: 1546: 1541: 1532: 1530: 1526: 1521: 1519: 1514: 1507: 1503: 1497: 1488: 1485: 1480: 1478: 1474: 1469: 1467: 1463: 1459: 1454: 1449: 1447: 1443: 1439: 1435: 1431: 1427: 1417: 1415: 1411: 1410:Isle of Pines 1407: 1406:Nepean Island 1402: 1392: 1388: 1383: 1379: 1378: 1371: 1367: 1366: 1361: 1356: 1351: 1347: 1346: 1338: 1336: 1332: 1328: 1327:Campeche Bank 1324: 1319: 1317: 1312: 1311:Philip Island 1307: 1301: 1299: 1295: 1286: 1277: 1275: 1271: 1267: 1262: 1260: 1256: 1252: 1248: 1243: 1241: 1237: 1233: 1229: 1225: 1215: 1213: 1209: 1204: 1197: 1193: 1189: 1185: 1181: 1177: 1173: 1167: 1164: 1160: 1158: 1154: 1150: 1146: 1142: 1137: 1135: 1131: 1127: 1123: 1119: 1115: 1111: 1107: 1102: 1093: 1089: 1085: 1080: 1068: 1064: 1060: 1056: 1055: 1051: 1047: 1040: 1037: 1036: 1032: 1028: 1024: 1020: 1016: 1011: 1010:Walter Oliver 1007: 1003: 999: 998: 994: 990: 983: 982:S. d. tasmani 980: 979: 964: 942: 931: 927: 923: 919: 916:in 1859 from 915: 911: 907: 906: 900: 897: 896: 892: 891:Monito Island 888: 884: 880: 876: 875: 869: 866: 865: 862: 858: 853: 849: 846: 844: 839: 834: 819: 818: 810: 809: 801: 800: 792: 791: 783: 782: 779: 777: 773: 767: 766: 763: 762: 759: 757: 756:Sula nebouxii 753: 747: 746: 740: 739: 736: 735: 727: 726: 723: 721: 717: 711: 710: 707: 706: 703: 701: 694: 693: 687: 686: 680: 679: 676: 675: 672: 670: 666: 660: 659: 653: 652: 649: 648: 645: 643: 639: 633: 632: 626: 625: 621: 619: 615: 610: 606: 602: 598: 597:Sula nebouxii 594: 590: 586: 585:Nazca boobies 582: 578: 575:) using both 574: 570: 569: 563: 561: 557: 553: 549: 542: 538: 533: 529: 527: 523: 519: 515: 511: 510: 505: 490: 485: 481: 476: 463: 441: 426: 422: 418: 403: 387: 377: 376:Ancient Greek 374:combines the 373: 369: 365: 362: 361:binomial name 358: 354: 350: 346: 345: 340: 330: 328: 324: 320: 316: 312: 307: 303: 299: 294: 292: 288: 284: 280: 276: 275: 270: 266: 262: 258: 254: 251:, is a large 250: 246: 245:masked gannet 242: 241: 236: 225: 221: 216: 213: 209: 204: 199: 194: 192: 186: 183: 182:Binomial name 179: 175: 174: 169: 166: 165: 162: 161: 157: 154: 153: 150: 147: 144: 143: 140: 137: 134: 133: 130: 127: 124: 123: 120: 117: 114: 113: 110: 107: 104: 103: 100: 97: 94: 93: 88: 83: 79: 73: 68: 67:Least Concern 57: 53: 48: 45: 41: 36: 32: 27: 24:Masked booby 22: 19: 5980: 5867:masked-booby 5766:masked-booby 5521: 5438:Chatham shag 5428:Foveaux shag 5400: 5382:Poikilocarbo 5380: 5360: 5330: 5292: 5251: 5201:Spotted shag 5175: 5161: 5086: 5072: 5049: 4995: 4977: 4953:Masked booby 4952: 4932: 4918: 4838: 4824: 4801: 4790:Frigatebirds 4729:. Retrieved 4707: 4698: 4672: 4647: 4641: 4637: 4614:. Retrieved 4594: 4587: 4538: 4534: 4528: 4495: 4491: 4484: 4472:. Retrieved 4452: 4445: 4433:. Retrieved 4426:the original 4421: 4417: 4404: 4392:. Retrieved 4383: 4379: 4373: 4365: 4353:. Retrieved 4344: 4340: 4334: 4330: 4327:Alectorobius 4326: 4323:Ornithodoros 4322: 4314: 4302:. Retrieved 4274: 4270: 4261: 4257: 4249: 4237:. Retrieved 4228: 4224: 4214: 4187: 4175:. Retrieved 4147: 4143: 4127: 4120:Dorward 1962 4090: 4083: 4076:Dorward 1962 4071: 4064:Dorward 1962 4059: 4034: 4030: 4024: 3997: 3985:. Retrieved 3965: 3943:. Retrieved 3934: 3924: 3912:. Retrieved 3898: 3865: 3861: 3855: 3843:. Retrieved 3831: 3827: 3814: 3793: 3779:cite journal 3767:. Retrieved 3740:(1): 20–29. 3737: 3733: 3705:. Retrieved 3686: 3682: 3672: 3660:. Retrieved 3651: 3647: 3637: 3625:. Retrieved 3618:the original 3613: 3609: 3560: 3556: 3552: 3527:. Retrieved 3497: 3493: 3461:. Retrieved 3454:the original 3449: 3445: 3417:. Retrieved 3405: 3401: 3388: 3379: 3375: 3369: 3361: 3349:. Retrieved 3340: 3336: 3330: 3322: 3310:. Retrieved 3290: 3283: 3271:. Retrieved 3251: 3244: 3232:. Retrieved 3212: 3205: 3193:. Retrieved 3173: 3166: 3154:. Retrieved 3140: 3077: 3073: 3067: 3025: 3013: 3001: 2964:. Retrieved 2944: 2940: 2910:. Retrieved 2903:the original 2898: 2894: 2885: 2842:(1): 94–97. 2839: 2835: 2825: 2806: 2802: 2788: 2776:. Retrieved 2769:the original 2764: 2760: 2751: 2747: 2743: 2720:. Retrieved 2708: 2704: 2695: 2687: 2675: 2663:. Retrieved 2654: 2650: 2640: 2628:. Retrieved 2608: 2601: 2560: 2556: 2550: 2527:. Retrieved 2518: 2511: 2473: 2466: 2433: 2429: 2423: 2416:Dorward 1962 2411: 2399:. Retrieved 2390: 2358:. Retrieved 2330: 2326: 2314: 2302:. Retrieved 2285: 2281: 2271: 2259:. Retrieved 2249: 2239: 2220: 2214: 2202:. Retrieved 2185: 2181: 2168: 2156: 2144: 2132:. Retrieved 2123: 2113: 2104: 2100: 2094: 2086: 2074:. Retrieved 2054: 2032:. Retrieved 2022: 2018:Lesson, RenĂ© 2012: 2004: 1995:. Retrieved 1985: 1981: 1977:Lesson, RenĂ© 1971: 1959:. Retrieved 1950: 1946: 1940: 1936: 1928: 1916:. Retrieved 1902: 1896: 1890: 1864: 1843: 1829: 1818: 1805: 1795: 1785: 1775: 1765: 1757: 1753: 1745: 1742:Silver gulls 1740: 1730: 1720: 1716: 1708: 1700: 1692: 1689:frigate tuna 1684: 1676: 1668: 1660: 1652: 1644: 1636: 1628: 1620: 1612: 1604: 1596: 1588: 1574: 1570:Frigatebirds 1568:) at times. 1565: 1557: 1550: 1522: 1489: 1481: 1470: 1465: 1462:allopreening 1457: 1453:sky-pointing 1452: 1450: 1445: 1441: 1437: 1433: 1423: 1403: 1399: 1386: 1354: 1335:Dry Tortugas 1320: 1302: 1294:Raine Island 1291: 1263: 1255:Delaware Bay 1251:Island Beach 1244: 1221: 1211: 1195: 1187: 1179: 1168: 1165: 1161: 1156: 1152: 1148: 1144: 1140: 1138: 1097: 1083: 1066: 1062: 1049: 1045: 1038: 1030: 1026: 1022: 1014: 988: 981: 929: 925: 918:Maydh Island 898: 871:Lesson, 1831 867: 856: 847: 830: 775: 770: 755: 750: 719: 714: 699: 697: 668: 663: 641: 636: 604: 596: 588: 572: 566: 564: 559: 545: 536: 517: 507: 506:in his 1865 504:Sula cyanops 503: 483: 477: 420: 363: 342: 336: 295: 286: 272: 248: 244: 239: 238: 235:masked booby 234: 232: 211: 190: 188: 172: 171: 159: 44:Midway Atoll 39: 18: 5937:SeaLifeBase 5852:Neotropical 5826:NatureServe 5774:iNaturalist 5546:Wikispecies 5469:Crozet shag 5453:Bounty shag 5332:Nannopterum 5011:Cape gannet 4968:Brown booby 4958:Nazca booby 4629:Cited texts 4347:: 368–374. 4262:Felis catus 3834:: 175–185. 3654:: 229–231. 3500:: 291–302. 3402:Micronesica 3382:(4): 59–60. 2383:Gill, Frank 2245:Gould, John 2174:Gould, John 1984:La Coquille 1953:: 83–103 . 1918:12 November 1581:cephalopods 1577:flying fish 1487:September. 1323:Rocas Atoll 1316:Zihuatanejo 1270:Maharashtra 1247:Gulf Stream 1208:brown booby 1184:Cape gannet 1122:secondaries 1074:Description 1063:californica 1042:Gould, 1846 966:), meaning 944:), meaning 831:There is a 720:Sula granti 716:Nazca booby 665:Brown booby 593:blue-footed 589:Sula granti 548:common name 539:In flight, 465:), meaning 443:), meaning 389:), meaning 344:La Coquille 339:RenĂ© Lesson 319:flying fish 287:Sula granti 283:Nazca booby 206:Subspecies 5987:Categories 5963:Xeno-canto 5433:Otago shag 5402:Leucocarbo 5253:Microcarbo 5135:Cormorants 4898:: Sulidae) 4776:Suliformes 4331:muesebecki 4051:10088/6078 3805:10088/4870 2809:: 85–100. 2711:(1): 1–7. 2630:4 December 2401:31 January 2107:(3): 218 . 1871:References 1848:angel wing 1784:caused by 1725:, and the 1665:brown chub 1506:nidicolous 1484:Kure Atoll 1438:outposting 1426:monogamous 1387:dactylatra 1306:feral pigs 1190:) and the 1157:dactylatra 1114:serrations 1088:Kure Atoll 1044:(includes 1023:S. tasmani 987:(includes 930:dactylatra 618:St. Helena 614:speciation 609:subspecies 537:dactylatra 492:, meaning 482:described 480:John Gould 405:, meaning 291:subspecies 139:Suliformes 5411:Rock shag 5196:Pitt shag 4670:(1980) . 4386:: 38–40. 4291:131226663 4277:: 60–71. 4231:: 2–23 . 2188:: 18–21. 2005:livraison 1865:personata 1854:) of the 1817:HMS  1657:mahi-mahi 1525:siblicide 1512:scapulars 1502:Altricial 1477:Coral Sea 1389:in nest, 1355:personata 1341:Behaviour 1149:personata 1130:rectrices 1118:primaries 1084:personata 1002:Lord Howe 879:Caribbean 843:gene flow 642:Sula sula 560:Sula sula 489:personata 212:See text 167:Species: 105:Kingdom: 99:Eukaryota 40:personata 5831:2.105902 5818:22736173 5792:10190427 5691:45509069 5629:22736173 5624:BirdLife 5531:Wikidata 4978:Papasula 4722:Archived 4610:Archived 4579:40459951 4571:17810291 4520:85187886 4512:23324568 4468:Archived 4388:Archived 4349:Archived 4295:Archived 4233:Archived 4168:Archived 4135:(1995). 3981:Archived 3939:Archived 3908:Archived 3836:Archived 3760:Archived 3756:51902194 3701:Archived 3656:Archived 3610:Notornis 3585:55658891 3520:Archived 3419:26 April 3410:Archived 3345:Archived 3306:Archived 3267:Archived 3228:Archived 3189:Archived 3150:Archived 3094:11376276 2957:Archived 2866:19675005 2797:(2001). 2761:Notornis 2713:Archived 2659:Archived 2651:The Ibis 2624:Archived 2593:25708378 2585:16202102 2523:Archived 2458:21144905 2395:Archived 2351:Archived 2298:Archived 2288:: 9–10. 2255:Archived 2247:(1865). 2198:Archived 2176:(1846). 2128:Archived 2070:Archived 2028:Archived 2020:(1831). 1991:Archived 1955:Archived 1941:Coquille 1937:Zoologie 1770:and the 1464:. In an 1202:humerals 1141:melanops 1126:humerals 1004:and the 620:Island. 522:priority 425:subadult 380:ÎŽÎŹÎșÏ„Ï…Î»ÎżÏ‚ 333:Taxonomy 329:(IUCN). 298:colonies 279:Atlantic 263:family, 145:Family: 119:Chordata 115:Phylum: 109:Animalia 95:Domain: 72:IUCN 3.1 5998:Boobies 5756:2480967 5611:Avibase 5537:Q602084 5362:Gulosus 5163:Species 5097:Anhinga 5088:Anhinga 5074:Species 5050:Anhinga 5039:Darters 4920:Species 4840:Fregata 4826:Species 4802:Fregata 4563:1693159 4543:Bibcode 4535:Science 4435:13 June 4418:Corella 4394:13 June 4355:13 June 4271:Seabird 4239:6 March 4164:4089018 4144:The Auk 3890:1521152 3870:Bibcode 3845:11 June 3565:Bibcode 3502:Bibcode 3463:1 March 3351:13 June 3343:: 434. 2953:4163925 2912:22 July 2857:2817237 2778:3 March 2722:5 March 2665:6 March 2565:Bibcode 2529:2 March 2438:Bibcode 2360:4 March 2347:4073271 2304:4 March 2204:4 March 2134:3 March 1997:4 March 1943:voyage" 1939:of the 1852:El Niño 1787:Babesia 1762:argasid 1535:Feeding 1353:Subsp. 1329:in the 1236:vagrant 1153:tasmani 1145:tasmani 1134:coverts 1090:in the 1067:bedouti 941:mĂ©lanos 935:ÎŒÎ­Î»Î±ÎœÎżÏ‚ 926:cyanops 857:tasmani 577:nuclear 550:by the 535:Subsp. 355:in the 325:by the 265:Sulidae 255:of the 253:seabird 247:or the 155:Genus: 149:Sulidae 135:Order: 125:Class: 70: ( 5955:212599 5929:186230 5916:170197 5903:212599 5874:NZOR: 5864:NZBO: 5857:masboo 5805:174699 5763:GNAB: 5712:EURING 5704:100701 5678:masboo 5652:masboo 5598:ARKive 5565:masboo 5139:family 5043:family 4896:family 4794:family 4731:25 May 4714:  4684:  4616:6 June 4602:  4577:  4569:  4561:  4518:  4510:  4474:6 June 4460:  4304:2 June 4289:  4177:28 May 4162:  4104:248142 4102:  3987:6 June 3973:  3945:1 June 3914:1 June 3888:  3769:2 June 3754:  3707:6 June 3662:4 July 3627:2 June 3583:  3529:31 May 3312:6 June 3298:  3273:6 June 3259:  3234:6 June 3220:  3195:6 June 3181:  3156:1 June 3092:  2966:28 May 2951:  2864:  2854:  2616:  2591:  2583:  2456:  2345:  2261:29 May 2227:  2076:6 June 2062:  2034:30 May 1961:30 May 1819:Supply 1772:ixodid 1748:) and 1615:) and 1560:) and 1518:fledge 1458:gazing 1266:Kerala 1257:, and 954:, and 833:clinal 599:) and 524:. The 497:masked 453:, and 440:kĂșanos 434:ÎșÏÎ±ÎœÎżÏ‚ 429:  394:finger 386:dĂĄktul 315:divers 302:atolls 261:gannet 200:, 1831 198:Lesson 5950:WoRMS 5942:73589 5890:73403 5844:56068 5787:IRMNG 5730:96610 5699:EUNIS 5673:eBird 5665:7B4FR 5649:BOW: 5642:17274 5294:Urile 5152:Genus 5063:Genus 5047:genus 4997:Morus 4909:Genus 4815:Genus 4798:genus 4772:Order 4725:(PDF) 4704:(PDF) 4575:S2CID 4559:JSTOR 4516:S2CID 4508:JSTOR 4429:(PDF) 4414:(PDF) 4298:(PDF) 4287:S2CID 4267:(PDF) 4171:(PDF) 4160:JSTOR 4140:(PDF) 4098:–34. 3886:JSTOR 3839:(PDF) 3824:(PDF) 3763:(PDF) 3752:S2CID 3730:(PDF) 3621:(PDF) 3606:(PDF) 3581:S2CID 3523:(PDF) 3490:(PDF) 3457:(PDF) 3442:(PDF) 3413:(PDF) 3398:(PDF) 3090:S2CID 2960:(PDF) 2949:JSTOR 2937:(PDF) 2906:(PDF) 2891:(PDF) 2772:(PDF) 2757:(PDF) 2748:Morus 2716:(PDF) 2701:(PDF) 2589:S2CID 2354:(PDF) 2343:JSTOR 2323:(PDF) 1808:TaĂ­no 1774:tick 1764:tick 1722:Mugil 1495:tarsi 1172:morph 1106:lores 1101:sulid 1094:chain 991:as a 949:black 914:taxon 410:black 257:booby 5898:OBIS 5839:NCBI 5813:IUCN 5800:ITIS 5779:3788 5751:GBIF 5637:BOLD 4934:Sula 4733:2019 4712:ISBN 4682:ISBN 4648:103B 4643:Ibis 4638:Sula 4618:2020 4600:ISBN 4567:PMID 4476:2020 4458:ISBN 4437:2019 4396:2019 4357:2019 4306:2019 4241:2018 4179:2019 4100:OCLC 3989:2020 3971:ISBN 3947:2019 3916:2019 3847:2019 3785:link 3771:2019 3734:Oryx 3709:2020 3664:2019 3629:2019 3531:2019 3465:2019 3421:2018 3353:2019 3314:2020 3296:ISBN 3275:2020 3257:ISBN 3236:2020 3218:ISBN 3197:2020 3179:ISBN 3158:2019 2968:2019 2914:2018 2862:PMID 2780:2018 2752:Sula 2746:and 2744:Sula 2724:2018 2667:2018 2632:2018 2614:ISBN 2581:PMID 2531:2019 2454:PMID 2403:2018 2362:2018 2306:2018 2263:2019 2225:ISBN 2206:2018 2136:2019 2078:2020 2060:ISBN 2036:2019 1999:2018 1963:2019 1920:2021 1903:2018 1830:The 1806:The 1504:and 1226:and 1128:and 1048:and 971:face 579:and 568:Sula 520:had 470:face 448:blue 402:ater 311:down 274:Sula 259:and 233:The 160:Sula 129:Aves 5717:680 5686:EoL 5660:CoL 5585:AFD 5572:ADW 5561:ABA 4652:doi 4551:doi 4539:225 4500:doi 4496:124 4279:doi 4152:doi 4148:112 4047:hdl 4039:doi 4035:164 3878:doi 3800:hdl 3742:doi 3691:doi 3573:doi 3561:105 3557:Emu 3510:doi 3498:362 3082:doi 3078:157 2945:110 2852:PMC 2844:doi 2811:doi 2573:doi 2446:doi 2335:doi 2327:Auk 2290:doi 2190:doi 1907:doi 1733:). 1711:), 1703:), 1695:), 1687:), 1679:), 1671:), 1663:), 1655:), 1647:), 1639:), 1631:), 1607:), 1599:), 1591:), 1373:Egg 1086:on 963:Ăłps 562:). 462:Ăłps 5989:: 5965:: 5952:: 5939:: 5926:: 5913:: 5900:: 5887:: 5854:: 5841:: 5828:: 5815:: 5802:: 5789:: 5776:: 5753:: 5740:: 5727:: 5714:: 5701:: 5688:: 5675:: 5662:: 5639:: 5626:: 5613:: 5600:: 5587:: 5574:: 5563:: 5548:: 5533:: 4800:: 4774:: 4720:. 4706:. 4680:. 4666:; 4646:. 4608:. 4573:. 4565:. 4557:. 4549:. 4537:. 4514:. 4506:. 4494:. 4466:. 4422:10 4420:. 4416:. 4384:98 4382:. 4378:. 4345:71 4343:. 4339:. 4329:) 4293:. 4285:. 4275:24 4273:. 4269:. 4227:. 4223:. 4199:^ 4166:. 4158:. 4146:. 4142:. 4112:^ 4096:33 4045:. 4033:. 4009:^ 3979:. 3955:^ 3933:. 3884:. 3876:. 3864:. 3832:45 3830:. 3826:. 3781:}} 3777:{{ 3758:. 3738:44 3736:. 3732:. 3717:^ 3699:. 3687:24 3685:. 3681:. 3652:38 3650:. 3646:. 3614:57 3612:. 3608:. 3593:^ 3579:. 3571:. 3559:. 3539:^ 3518:. 3508:. 3496:. 3492:. 3473:^ 3450:33 3448:. 3444:. 3429:^ 3406:32 3404:. 3400:. 3380:50 3378:. 3374:. 3341:84 3339:. 3335:. 3304:. 3265:. 3226:. 3187:. 3123:^ 3102:^ 3088:. 3076:. 3052:^ 3037:^ 2976:^ 2955:. 2943:. 2939:. 2922:^ 2899:38 2897:. 2893:. 2874:^ 2860:. 2850:. 2838:. 2834:. 2807:27 2805:. 2801:. 2765:35 2763:. 2759:. 2732:^ 2709:18 2707:. 2703:. 2653:. 2649:. 2622:. 2587:. 2579:. 2571:. 2561:14 2559:. 2555:. 2553:)" 2539:^ 2517:. 2514:)" 2487:^ 2479:24 2452:. 2444:. 2434:58 2432:. 2389:. 2370:^ 2349:. 2341:. 2331:37 2329:. 2325:. 2296:. 2286:18 2284:. 2280:. 2196:. 2186:14 2184:. 2180:. 2122:. 2099:. 2068:. 2044:^ 1951:46 1949:. 1945:. 1901:. 1895:. 1879:^ 1242:. 1124:, 1120:, 957:ᜄψ 778:) 758:) 722:) 702:) 671:) 644:) 475:. 456:ᜄψ 42:, 5318:† 5137:( 5052:) 5041:( 4894:( 4804:) 4792:( 4764:e 4757:t 4750:v 4735:. 4697:" 4690:. 4658:. 4654:: 4620:. 4581:. 4553:: 4545:: 4522:. 4502:: 4478:. 4439:. 4398:. 4359:. 4325:( 4321:" 4308:. 4281:: 4264:" 4243:. 4229:8 4181:. 4154:: 4106:. 4053:. 4049:: 4041:: 3991:. 3949:. 3918:. 3892:. 3880:: 3872:: 3866:9 3849:. 3808:. 3802:: 3787:) 3773:. 3744:: 3711:. 3693:: 3666:. 3631:. 3587:. 3575:: 3567:: 3533:. 3512:: 3504:: 3467:. 3423:. 3355:. 3316:. 3277:. 3238:. 3199:. 3160:. 3096:. 3084:: 2970:. 2916:. 2868:. 2846:: 2840:6 2819:. 2813:: 2782:. 2726:. 2669:. 2655:1 2634:. 2595:. 2575:: 2567:: 2533:. 2481:. 2460:. 2448:: 2440:: 2405:. 2364:. 2337:: 2308:. 2292:: 2265:. 2233:. 2208:. 2192:: 2138:. 2105:I 2097:" 2093:" 2080:. 2038:. 2001:. 1965:. 1922:. 1909:: 1893:" 1889:" 1842:( 1794:( 1752:( 1744:( 1729:( 1715:( 1707:( 1699:( 1691:( 1683:( 1675:( 1667:( 1659:( 1651:( 1643:( 1635:( 1627:( 1619:( 1611:( 1603:( 1595:( 1587:( 1564:( 1556:( 1210:( 1194:( 1186:( 1178:( 1052:) 976:. 974:' 968:' 960:( 952:' 946:' 938:( 774:( 754:( 718:( 667:( 640:( 603:( 595:( 587:( 558:( 500:' 494:' 473:' 467:' 459:( 451:' 445:' 437:( 413:' 407:' 397:' 391:' 383:( 285:( 237:( 74:)

Index

white seabird with long pale bill standing on a beach and angled part toward camera
Midway Atoll
Conservation status
Least Concern
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Aves
Suliformes
Sulidae
Sula
Binomial name
Lesson

seabird
booby
gannet
Sulidae
René-PrimevÚre Lesson
Sula
Atlantic
Nazca booby
subspecies
colonies
atolls
agonistic displays
down

Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.

↑