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of limestone. The entrance to the cave is 6 m wide. The first cave tunnel is 80 m long and has 12 to 150-degree slope. The tunnel evolves into steps of 1 - 1.5 m height. At 125 meters from the entrance there is a 5 meter thick flat plate in a large hall measured 30 meters by 25 meters.
499:
Unlike most other
Paleolithic sites found in Georgia that relied primarily on hunting one species, the people of Satsurblia appeared to have hunted a slightly more diverse range of species. The animal remains found at Satsurblia were dominated primarily by
678:"Satsurblia: new insights of human response and survival across the Last Glacial Maximum in the southern Caucasus...Location of Satsurblia Cave and other key sites with Upper Palaeolithic occupation in western Georgia"
596:
The
Satsurblia individual is genetically closest to an ancient individual, dating to around 9,700 BP, found at the Kotias Klde rock shelter in Georgia. Together, they form a genetically distinct cluster referred to as
1984:
1122:
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470:. The next period of human occupation at Satsurblia took place from around 17,000 to 16,200 BP. The hiatus in human occupation at Satsurblia coincides with the
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The ancient individual from
Satsurblia was male with black hair and brown eyes; however, the individual is one of the earliest found to carry the derived
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of the bone yielded an estimated date of 13,300 BP for the age of the bone. Researchers successfully extracted DNA from the
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613:. Around half of the Yamnaya people's DNA come from the Caucasus hunter-gatherers, with the other half being derived from
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Martkoplishvili, Inga (2015). "Some popular medicinal plants and diseases of the Upper
Palaeolithic in Western Georgia".
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In comparison to modern human populations, the
Satsurblia individual is closest to the modern population in Georgia.
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409:. The karst cave was first excavated in 1976 by A. N. Kalandadze. In the Middle Ages the cave was used as a refuge.
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1030:"Satsurblia: New Insights of Human Response and Survival across the Last Glacial Maximum in the Southern Caucasus"
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allele for blue eyes. The
Satsurblia individual also likely had light skin, as he was found to carry the derived
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allele for light skin. The
Satsurblia individual was also lactose intolerant and did not carry the derived
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4013:
2013:
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1605:
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617:. The Caucasus hunter-gatherers also contributed genetically to modern Central Asians and South Asians.
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The
Caucasus hunter-gatherers contributed significantly to modern European populations by way of the
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fibers, and pottery were discovered at the cave. The lithic artefacts show similarities to eastern
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Some popular medicinal plants and diseases of the Upper
Palaeolithic in Western Georgia
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Barjadze, Sh., Arabuli, T., Mumladze, L., Maghradze, E., Asanidze, Z., Kutalia, T.
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and polished bovid bone were also discovered. The remains of yellow, red and brown
32:
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Cave
Biodiversity of Georgia, Open Access Database, 2019. Institute of Zoology at
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Cave of Altamira and Paleolithic Cave Art of Northern Spain World Heritage Site
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393:) is a paleoanthropological site located 1.2 km from Kumistavi village,
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Prehistoric people first occupied the cave from around 25,500 to 24,400
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421:, formed in the Sataphlia-Tskaltubo karst massif. The cave has many
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977:"Upper Palaeolithic genomes reveal deep roots of modern Eurasians"
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2224:
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Rock art of the Iberian Mediterranean Basin World Heritage Site
2551:
5544:
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3100:
2309:
Caves and Ice Age Art in the Swabian Jura World Heritage Site
562:
493:
434:
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573:
allele commonly found in East Asians and Native Americans.
570:
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1130:
859:. Archaeology News from Past Horizons. November 16, 2015
584:. About 1.7-2.4% of the Satsurblia individual's DNA was
883:
Fu, Qiaomei; Posth, Cosimo; et al. (May 2, 2016).
857:"Finding a fourth strand of ancient European ancestry"
700:"International Archaeological Expedition In Tskaltubo"
799:
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782:
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778:
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546:fragment of an ancient human in the cave. Direct
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388:
364:1976, 1985–88, 1989–1992, 2008–2010, 2012–2013
1652:
1116:
1611:Toba Waterfall and Arsen Okrojanashvili Cave
702:. Georgian National Museum. December 8, 2014
591:
4689:
1091:Europe's fourth ancestral 'tribe' uncovered
576:The Satsurblia individual belongs to mtDNA
1667:Prehistoric cave sites, rock shelters and
1659:
1645:
1123:
1109:
1065:
1055:
1010:
926:
4434:Cradle of Humankind, World Heritage Site
1027:
885:"The genetic history of Ice Age Europe"
882:
803:
761:
488:sites. Perforated pendants made out of
250:
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91:
66:
16:Cave and archaeological site in Georgia
5789:Prehistoric sites in Georgia (country)
5774:Natural monuments of Georgia (country)
5761:
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4688:
4140:
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1702:
1640:
1104:
975:Jones, Eppie R.; et al. (2015).
974:
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736:
734:
719:
717:
534:remains were also found at the site.
480:artefacts, bone artefacts, charcoal,
159:0.01 km (0.0039 sq mi)
1561:Oniore Waterfall and Toba First Cave
1132:Protected areas of Georgia (country)
833:. Ancestral Journeys. Archived from
5784:Protected areas established in 2011
13:
1028:Pinhasi, Ron; et al. (2014).
731:
714:
14:
5815:
2822:Mala Balanica and Velika Balanica
1985:Vézère Valley World Heritage Site
1780:Neolithic flint mines of Spiennes
1084:
632:Tsutskhvati Cave Natural Monument
552:petrous part of the temporal bone
727:საწურბლიას მღვიმის ბუნების ძეგლი
542:In 2013, archaeologists found a
446:The inhabitants of the cave are
381:Satsurblia Cave Natural Monument
249:
242:
221:
214:
197:
184:საწურბლიას მღვიმის ბუნების ძეგლი
90:
83:
65:
58:
41:
22:Satsurblia Cave Natural Monument
5799:Mesolithic Homo sapiens fossils
875:
849:
831:"Palaeolithic DNA from Eurasia"
823:
809:
274:Sataplia-Tskaltubo karst massif
692:
670:
656:
461:
1:
637:
496:were also found at the site.
412:
1057:10.1371/journal.pone.0111271
947:Journal of Ethnopharmacology
7:
5365:
3010:Other caves with decoration
2092:Other caves with decoration
2014:Les Eyzies-de-Tayac-Sireuil
1690:Caves containing pictograms
620:
537:
33:natural monument or feature
10:
5820:
5779:Caves of Georgia (country)
4141:
2994:Other World Heritage Sites
2081:Other World Heritage Sites
1703:
1680:Paleoanthropological sites
1486:Goderdzi Petrified Forest
1431:Alazani Floodplain Forests
663:Satsurblia Cave in Georgia
319:360 m (1,181 ft)
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1977:
1961:
1920:
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1827:
1806:
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1506:Keterisi Mineral Vaucluse
1416:
1400:
1292:
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960:10.1016/j.jep.2015.03.003
592:Caucasus hunter-gatherers
389:
368:
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344:
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331:
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315:
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258:Satsurblia Cave (Imereti)
208:
196:
179:
175:Agency of Protected Areas
171:
163:
155:
118:
108:
52:
40:
26:
21:
5658:Northern Mariana Islands
3295:
615:Eastern Hunter Gatherers
600:Caucasus Hunter-Gatherer
441:
554:and managed to recover
300:42.388111°N 42.606167°E
140:42.388111°N 42.606167°E
4182:Manyana Rock Paintings
2233:La Chapelle-aux-Saints
1212:Strict Nature Reserves
526:were also found. Some
407:meters above sea level
395:Tsqaltubo Municipality
47:Inside Satsurblia Cave
4745:Actun Tunichil Muknal
1501:Jvari Pass Travertine
1436:Artsivi (Eagle) Gorge
982:Nature Communications
746:Ilia State University
417:Satsurblia Cave is a
361:Excavation dates
5804:Geography of Imereti
4319:Ifri N'Amr Ou Moussa
3259:Red Lady of Paviland
2151:Mayrières supérieure
1546:Ochkhomuri Waterfall
1456:Bodorna Rock Columns
1401:Protected Landscapes
651:Martkoplishvili 2015
582:Haplogroup J1-Y6313*
472:Last Glacial Maximum
305:42.388111; 42.606167
145:42.388111; 42.606167
5134:Colorado Millennial
3772:Shiraho Saonetabaru
2576:Bukuroshja e Fjetur
1048:2014PLoSO...9k1271P
995:2015NatCo...6.8912J
911:10.1038/nature17993
903:2016Natur.534..200F
804:Fu & Posth 2016
627:Prehistoric Georgia
296: /
261:Show map of Imereti
233:Show map of Georgia
230:Location in Georgia
193:
172:Governing body
136: /
99:Show map of Imereti
74:Show map of Georgia
4952:Dominican Republic
4713:Cueva de las Manos
3609:Huto and Kamarband
3019:Barranc del Migdia
2905:Caves in Cantabria
2329:Hohlenstein-Stadel
2136:Grotte de Gabillou
2121:La Chaire a Calvin
1621:Tskaltsitela Gorge
1551:Kinchkha Waterfall
1426:Abano Mineral Lake
1153:Borjomi-Kharagauli
1003:10.1038/ncomms9912
837:on October 3, 2016
680:. VolcanoDiscovery
372:A. N. Kalandadze,
191:
5794:Neanderthal sites
5769:IUCN Category III
5756:
5755:
5726:
5725:
5722:
5721:
5356:
5355:
5352:
5351:
4869:Piedras del Tunjo
4801:Toca da Tira Peia
4680:
4679:
4676:
4675:
4132:
4131:
4128:
4127:
3286:
3285:
3282:
3281:
2922:Hornos de la Peña
1634:
1633:
1616:Truso Travertines
1586:Samshvilde Canyon
1526:Mukhura Waterfall
1418:Natural Monuments
1229:Bichvinta-Miuseri
897:(7606): 200–205.
666:Protected Planet
530:, wolf, fox, and
508:; the remains of
390:საწურბლიას მღვიმე
378:
377:
189:
188:
109:Nearest city
5811:
5675:Papua New Guinea
5375:
5374:
5362:
5361:
5314:Stanfield-Worley
5289:Rockhouse Cliffs
4703:
4702:
4686:
4685:
4554:Howieson's Poort
4251:Enkapune Ya Muto
4151:
4150:
4138:
4137:
3604:Eshkaft-e Siahoo
3305:
3304:
3292:
3291:
3059:Peñas de Cabrera
2359:Kleine Feldhofer
2319:Geissenklösterle
2171:Roc-aux-Sorciers
1713:
1712:
1700:
1699:
1661:
1654:
1647:
1638:
1637:
1626:Tsutskhvati Cave
1571:Abasha Waterfall
1536:Navenakhevi Cave
1481:Gachedili Canyon
1461:Dashbashi Canyon
1362:Ktsia-Tabatskuri
1294:Managed Reserves
1188:Pshav-Khevsureti
1125:
1118:
1111:
1102:
1101:
1079:
1069:
1059:
1024:
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971:
940:
930:
869:
868:
866:
864:
853:
847:
846:
844:
842:
827:
821:
820:
813:
807:
801:
790:
784:
765:
759:
748:
738:
729:
725:
721:
712:
711:
709:
707:
696:
690:
689:
687:
685:
674:
668:
660:
654:
648:
392:
391:
311:
310:
308:
307:
306:
301:
297:
294:
293:
292:
289:
262:
253:
252:
246:
234:
225:
224:
218:
201:
194:
190:
151:
150:
148:
147:
146:
141:
137:
134:
133:
132:
129:
100:
94:
93:
87:
75:
69:
68:
62:
45:
19:
18:
5819:
5818:
5814:
5813:
5812:
5810:
5809:
5808:
5759:
5758:
5757:
5752:
5718:
5702:
5686:
5670:
5652:
5631:
5613:
5597:
5569:
5485:Mackintosh 90/1
5405:Beginner's Luck
5369:
5348:
5164:Dutchess Quarry
5104:Arnold Research
5092:
5076:
5035:
4979:
4963:
4947:
4929:
4888:
4847:
4826:
4805:
4759:
4755:Midnight Terror
4733:
4717:
4697:
4672:
4656:
4635:
4619:
4593:
4422:
4396:
4380:
4359:
4343:
4334:Kelif el Boroud
4297:
4276:
4260:
4239:
4218:
4202:
4186:
4165:
4145:
4124:
4103:
4082:
4056:
4044:
4018:
3987:
3951:
3933:
3917:
3901:
3885:
3869:
3823:
3802:
3786:
3735:
3654:
3633:
3577:
3546:
3500:
3484:
3458:
3397:
3381:
3335:
3319:
3299:
3278:
3177:
3161:
3140:
3039:los Murciélagos
2987:Roca dels Moros
2883:
2852:
2826:
2795:
2766:Chertovy Vorota
2749:
2718:
2667:
2646:
2630:
2618:North Macedonia
2612:
2596:
2580:
2564:
2546:
2483:
2462:
2401:
2373:
2297:
2060:Abri du Poisson
2025:Les Combarelles
1973:
1957:
1916:
1900:
1859:
1823:
1802:
1798:Trou de l'Abîme
1747:
1707:
1694:
1671:
1665:
1635:
1630:
1591:Satsurblia Cave
1581:Sakhizari Cliff
1566:Prometheus Cave
1541:Nazodelavo Cave
1412:
1396:
1288:
1207:
1134:
1129:
1087:
1082:
1042:(10): e111271.
878:
873:
872:
862:
860:
855:
854:
850:
840:
838:
829:
828:
824:
817:"J-Y6313 YTree"
815:
814:
810:
802:
793:
785:
768:
760:
751:
742:Satsurblia Cave
739:
732:
723:
722:
715:
705:
703:
698:
697:
693:
683:
681:
676:
675:
671:
661:
657:
649:
645:
640:
623:
594:
540:
532:Eurasian beaver
519:Capra caucasica
464:
444:
415:
373:
304:
302:
298:
295:
290:
287:
285:
283:
282:
266:
265:
264:
263:
260:
259:
256:
255:
254:
237:
236:
235:
232:
231:
228:
227:
226:
204:
203:Satsurblia Cave
192:Satsurblia Cave
144:
142:
138:
135:
130:
127:
125:
123:
122:
104:
103:
102:
101:
98:
97:
96:
95:
78:
77:
76:
73:
72:
71:
70:
48:
36:
17:
12:
11:
5:
5817:
5807:
5806:
5801:
5796:
5791:
5786:
5781:
5776:
5771:
5754:
5753:
5751:
5750:
5745:
5740:
5735:
5731:
5728:
5727:
5724:
5723:
5720:
5719:
5717:
5716:
5710:
5708:
5704:
5703:
5701:
5700:
5694:
5692:
5688:
5687:
5685:
5684:
5678:
5676:
5672:
5671:
5669:
5668:
5662:
5660:
5654:
5653:
5651:
5650:
5645:
5639:
5637:
5633:
5632:
5630:
5629:
5623:
5621:
5615:
5614:
5612:
5611:
5605:
5603:
5599:
5598:
5596:
5595:
5590:
5585:
5579:
5577:
5571:
5570:
5568:
5567:
5562:
5557:
5552:
5547:
5542:
5537:
5532:
5527:
5522:
5517:
5512:
5507:
5502:
5497:
5492:
5487:
5482:
5477:
5472:
5467:
5462:
5457:
5452:
5447:
5442:
5437:
5432:
5427:
5422:
5417:
5412:
5407:
5402:
5400:Beeton Shelter
5397:
5392:
5387:
5381:
5379:
5371:
5370:
5358:
5357:
5354:
5353:
5350:
5349:
5347:
5346:
5341:
5336:
5331:
5326:
5321:
5316:
5311:
5306:
5301:
5296:
5291:
5286:
5281:
5276:
5271:
5266:
5261:
5256:
5251:
5246:
5241:
5236:
5231:
5226:
5221:
5216:
5211:
5206:
5201:
5196:
5191:
5186:
5181:
5176:
5171:
5166:
5161:
5156:
5151:
5146:
5141:
5136:
5131:
5126:
5121:
5116:
5111:
5106:
5100:
5098:
5094:
5093:
5091:
5090:
5084:
5082:
5078:
5077:
5075:
5074:
5072:T'uqu T'uquyuq
5069:
5064:
5059:
5054:
5049:
5043:
5041:
5037:
5036:
5034:
5033:
5028:
5023:
5018:
5013:
5008:
5003:
4998:
4993:
4987:
4985:
4981:
4980:
4978:
4977:
4971:
4969:
4965:
4964:
4962:
4961:
4955:
4953:
4949:
4948:
4946:
4945:
4939:
4937:
4931:
4930:
4928:
4927:
4922:
4917:
4912:
4907:
4902:
4896:
4894:
4890:
4889:
4887:
4886:
4881:
4876:
4871:
4866:
4861:
4855:
4853:
4849:
4848:
4846:
4845:
4840:
4834:
4832:
4828:
4827:
4825:
4824:
4819:
4813:
4811:
4807:
4806:
4804:
4803:
4798:
4793:
4788:
4783:
4778:
4773:
4767:
4765:
4761:
4760:
4758:
4757:
4752:
4747:
4741:
4739:
4735:
4734:
4732:
4731:
4725:
4723:
4719:
4718:
4716:
4715:
4709:
4707:
4699:
4698:
4682:
4681:
4678:
4677:
4674:
4673:
4671:
4670:
4664:
4662:
4658:
4657:
4655:
4654:
4649:
4643:
4641:
4637:
4636:
4634:
4633:
4627:
4625:
4621:
4620:
4618:
4617:
4612:
4607:
4601:
4599:
4595:
4594:
4592:
4591:
4586:
4581:
4576:
4574:Pinnacle Point
4571:
4566:
4561:
4556:
4551:
4546:
4541:
4536:
4531:
4526:
4521:
4516:
4512:
4511:
4506:
4501:
4496:
4491:
4486:
4481:
4476:
4471:
4466:
4461:
4456:
4451:
4446:
4441:
4436:
4430:
4428:
4424:
4423:
4421:
4420:
4415:
4410:
4404:
4402:
4398:
4397:
4395:
4394:
4388:
4386:
4382:
4381:
4379:
4378:
4376:The White Lady
4373:
4367:
4365:
4361:
4360:
4358:
4357:
4351:
4349:
4345:
4344:
4342:
4341:
4336:
4331:
4326:
4321:
4316:
4311:
4305:
4303:
4299:
4298:
4296:
4295:
4290:
4284:
4282:
4278:
4277:
4275:
4274:
4268:
4266:
4262:
4261:
4259:
4258:
4253:
4247:
4245:
4241:
4240:
4238:
4237:
4232:
4226:
4224:
4220:
4219:
4217:
4216:
4210:
4208:
4204:
4203:
4201:
4200:
4194:
4192:
4188:
4187:
4185:
4184:
4179:
4173:
4171:
4167:
4166:
4164:
4163:
4157:
4155:
4147:
4146:
4134:
4133:
4130:
4129:
4126:
4125:
4123:
4122:
4117:
4111:
4109:
4105:
4104:
4102:
4101:
4096:
4090:
4088:
4084:
4083:
4081:
4080:
4075:
4070:
4064:
4062:
4058:
4057:
4052:
4050:
4046:
4045:
4043:
4042:
4037:
4032:
4026:
4024:
4020:
4019:
4017:
4016:
4011:
4006:
4001:
3995:
3993:
3989:
3988:
3986:
3985:
3980:
3975:
3970:
3965:
3959:
3957:
3953:
3952:
3950:
3949:
3943:
3941:
3935:
3934:
3932:
3931:
3925:
3923:
3919:
3918:
3916:
3915:
3909:
3907:
3903:
3902:
3900:
3899:
3897:Khoit Tsenkher
3893:
3891:
3887:
3886:
3884:
3883:
3877:
3875:
3871:
3870:
3868:
3867:
3862:
3857:
3852:
3847:
3842:
3837:
3831:
3829:
3825:
3824:
3822:
3821:
3816:
3810:
3808:
3804:
3803:
3801:
3800:
3794:
3792:
3788:
3787:
3785:
3784:
3779:
3774:
3769:
3764:
3759:
3754:
3749:
3743:
3741:
3737:
3736:
3734:
3733:
3728:
3723:
3718:
3713:
3708:
3703:
3698:
3693:
3688:
3683:
3678:
3673:
3668:
3662:
3660:
3656:
3655:
3653:
3652:
3647:
3641:
3639:
3635:
3634:
3632:
3631:
3626:
3621:
3616:
3611:
3606:
3601:
3596:
3591:
3585:
3583:
3579:
3578:
3576:
3575:
3570:
3565:
3560:
3554:
3552:
3548:
3547:
3545:
3544:
3539:
3534:
3529:
3524:
3519:
3514:
3508:
3506:
3502:
3501:
3499:
3498:
3492:
3490:
3486:
3485:
3483:
3482:
3477:
3472:
3466:
3464:
3460:
3459:
3457:
3456:
3451:
3446:
3441:
3436:
3431:
3426:
3421:
3416:
3411:
3405:
3403:
3399:
3398:
3396:
3395:
3389:
3387:
3383:
3382:
3380:
3379:
3374:
3369:
3364:
3359:
3354:
3349:
3343:
3341:
3337:
3336:
3334:
3333:
3327:
3325:
3321:
3320:
3318:
3317:
3311:
3309:
3301:
3300:
3288:
3287:
3284:
3283:
3280:
3279:
3277:
3276:
3271:
3266:
3261:
3256:
3251:
3246:
3241:
3236:
3231:
3226:
3221:
3216:
3211:
3209:Creswell Crags
3206:
3201:
3196:
3191:
3189:Aveline's Hole
3185:
3183:
3182:United Kingdom
3179:
3178:
3176:
3175:
3169:
3167:
3163:
3162:
3160:
3159:
3154:
3148:
3146:
3142:
3141:
3139:
3138:
3133:
3131:Santa Catalina
3128:
3123:
3118:
3113:
3108:
3103:
3098:
3093:
3088:
3083:
3077:
3076:
3071:
3066:
3061:
3056:
3051:
3046:
3041:
3036:
3031:
3026:
3021:
3016:
3011:
3007:
3006:
3001:
2996:
2990:
2989:
2984:
2979:
2972:
2971:
2966:
2961:
2956:
2951:
2950:
2949:
2946:
2945:
2944:
2939:
2936:
2933:
2926:Monte Castillo
2923:
2920:
2915:
2912:
2902:
2897:
2891:
2889:
2885:
2884:
2882:
2881:
2876:
2871:
2866:
2860:
2858:
2854:
2853:
2851:
2850:
2845:
2840:
2834:
2832:
2828:
2827:
2825:
2824:
2819:
2814:
2809:
2807:Hadži-Prodan's
2803:
2801:
2797:
2796:
2794:
2793:
2788:
2783:
2778:
2773:
2768:
2763:
2757:
2755:
2751:
2750:
2748:
2747:
2742:
2737:
2732:
2726:
2724:
2720:
2719:
2717:
2716:
2711:
2706:
2701:
2696:
2691:
2686:
2681:
2675:
2673:
2669:
2668:
2666:
2665:
2660:
2654:
2652:
2648:
2647:
2645:
2644:
2638:
2636:
2632:
2631:
2629:
2628:
2622:
2620:
2614:
2613:
2611:
2610:
2604:
2602:
2598:
2597:
2595:
2594:
2588:
2586:
2582:
2581:
2579:
2578:
2572:
2570:
2566:
2565:
2563:
2562:
2556:
2554:
2548:
2547:
2545:
2544:
2539:
2534:
2529:
2522:
2517:
2512:
2507:
2502:
2497:
2491:
2489:
2485:
2484:
2482:
2481:
2476:
2470:
2468:
2464:
2463:
2461:
2460:
2455:
2450:
2445:
2440:
2435:
2430:
2425:
2420:
2415:
2409:
2407:
2403:
2402:
2400:
2399:
2394:
2389:
2383:
2381:
2375:
2374:
2372:
2371:
2366:
2361:
2356:
2351:
2346:
2342:
2341:
2336:
2331:
2326:
2321:
2316:
2311:
2305:
2303:
2299:
2298:
2296:
2295:
2290:
2285:
2280:
2275:
2270:
2265:
2260:
2255:
2250:
2245:
2240:
2235:
2230:
2227:
2222:
2217:
2214:
2211:
2208:
2205:
2200:
2195:
2189:
2188:
2183:
2178:
2173:
2168:
2163:
2158:
2153:
2148:
2143:
2138:
2133:
2128:
2123:
2118:
2113:
2108:
2103:
2098:
2093:
2089:
2088:
2083:
2077:
2076:
2071:
2066:
2061:
2058:
2053:
2050:
2045:
2044:Laugerie-Haute
2042:
2040:Laugerie-Basse
2037:
2032:
2027:
2022:
2021:Abri Chadourne
2019:
2016:
2011:
2010:
2009:
2004:
1998:
1993:
1990:
1987:
1981:
1979:
1975:
1974:
1972:
1971:
1965:
1963:
1959:
1958:
1956:
1955:
1950:
1945:
1940:
1935:
1930:
1924:
1922:
1921:Czech Republic
1918:
1917:
1915:
1914:
1908:
1906:
1902:
1901:
1899:
1898:
1893:
1891:Romuald's Cave
1888:
1883:
1878:
1873:
1867:
1865:
1861:
1860:
1858:
1857:
1852:
1847:
1842:
1837:
1831:
1829:
1825:
1824:
1822:
1821:
1816:
1810:
1808:
1804:
1803:
1801:
1800:
1795:
1790:
1785:
1782:
1777:
1772:
1767:
1762:
1759:
1755:
1753:
1749:
1748:
1746:
1745:
1740:
1735:
1730:
1725:
1719:
1717:
1709:
1708:
1696:
1695:
1693:
1692:
1687:
1685:Cave paintings
1682:
1676:
1673:
1672:
1669:cave paintings
1664:
1663:
1656:
1649:
1641:
1632:
1631:
1629:
1628:
1623:
1618:
1613:
1608:
1603:
1598:
1593:
1588:
1583:
1578:
1573:
1568:
1563:
1558:
1553:
1548:
1543:
1538:
1533:
1528:
1523:
1518:
1513:
1508:
1503:
1498:
1493:
1488:
1483:
1478:
1473:
1468:
1463:
1458:
1453:
1448:
1443:
1438:
1433:
1428:
1422:
1420:
1414:
1413:
1411:
1410:
1404:
1402:
1398:
1397:
1395:
1394:
1389:
1384:
1379:
1374:
1369:
1364:
1359:
1354:
1349:
1344:
1339:
1334:
1329:
1324:
1319:
1314:
1309:
1304:
1298:
1296:
1290:
1289:
1287:
1286:
1281:
1276:
1271:
1266:
1261:
1256:
1251:
1246:
1241:
1236:
1231:
1226:
1221:
1215:
1213:
1209:
1208:
1206:
1205:
1200:
1195:
1190:
1185:
1180:
1175:
1170:
1165:
1160:
1155:
1150:
1144:
1142:
1140:National parks
1136:
1135:
1128:
1127:
1120:
1113:
1105:
1099:
1098:
1093:
1086:
1085:External links
1083:
1081:
1080:
1025:
972:
941:
919:10211.3/198594
879:
877:
874:
871:
870:
848:
822:
808:
791:
766:
749:
730:
713:
691:
669:
655:
642:
641:
639:
636:
635:
634:
629:
622:
619:
611:Yamnaya people
593:
590:
539:
536:
504:, followed by
463:
460:
443:
440:
414:
411:
376:
375:
374:T. Meshveliani
370:
369:Archaeologists
366:
365:
362:
358:
357:
353:
352:
346:
342:
341:
337:
336:
333:
329:
328:
325:
321:
320:
317:
313:
312:
280:
276:
275:
272:
268:
267:
257:
248:
247:
241:
240:
239:
238:
229:
220:
219:
213:
212:
211:
210:
209:
206:
205:
202:
187:
186:
181:
177:
176:
173:
169:
168:
165:
161:
160:
157:
153:
152:
120:
116:
115:
110:
106:
105:
89:
88:
82:
81:
80:
79:
64:
63:
57:
56:
55:
54:
53:
50:
49:
46:
38:
37:
27:
24:
23:
15:
9:
6:
4:
3:
2:
5816:
5805:
5802:
5800:
5797:
5795:
5792:
5790:
5787:
5785:
5782:
5780:
5777:
5775:
5772:
5770:
5767:
5766:
5764:
5749:
5746:
5744:
5743:Cave painting
5741:
5739:
5738:List of caves
5736:
5733:
5732:
5729:
5715:
5712:
5711:
5709:
5705:
5699:
5696:
5695:
5693:
5689:
5683:
5680:
5679:
5677:
5673:
5667:
5664:
5663:
5661:
5659:
5655:
5649:
5646:
5644:
5641:
5640:
5638:
5634:
5628:
5625:
5624:
5622:
5620:
5619:New Caledonia
5616:
5610:
5607:
5606:
5604:
5600:
5594:
5591:
5589:
5586:
5584:
5581:
5580:
5578:
5576:
5572:
5566:
5563:
5561:
5558:
5556:
5553:
5551:
5548:
5546:
5543:
5541:
5538:
5536:
5533:
5531:
5528:
5526:
5525:New Guinea II
5523:
5521:
5518:
5516:
5513:
5511:
5508:
5506:
5503:
5501:
5500:Mannalargenna
5498:
5496:
5493:
5491:
5488:
5486:
5483:
5481:
5478:
5476:
5473:
5471:
5468:
5466:
5463:
5461:
5458:
5456:
5453:
5451:
5448:
5446:
5443:
5441:
5438:
5436:
5433:
5431:
5428:
5426:
5423:
5421:
5418:
5416:
5413:
5411:
5408:
5406:
5403:
5401:
5398:
5396:
5393:
5391:
5388:
5386:
5383:
5382:
5380:
5376:
5372:
5368:
5363:
5359:
5345:
5342:
5340:
5337:
5335:
5332:
5330:
5327:
5325:
5322:
5320:
5317:
5315:
5312:
5310:
5307:
5305:
5302:
5300:
5297:
5295:
5292:
5290:
5287:
5285:
5282:
5280:
5277:
5275:
5272:
5270:
5269:On Your Knees
5267:
5265:
5262:
5260:
5257:
5255:
5252:
5250:
5247:
5245:
5242:
5240:
5237:
5235:
5232:
5230:
5227:
5225:
5222:
5220:
5217:
5215:
5212:
5210:
5207:
5205:
5202:
5200:
5197:
5195:
5194:Hidden Valley
5192:
5190:
5187:
5185:
5182:
5180:
5177:
5175:
5172:
5170:
5167:
5165:
5162:
5160:
5157:
5155:
5152:
5150:
5147:
5145:
5142:
5140:
5137:
5135:
5132:
5130:
5127:
5125:
5122:
5120:
5117:
5115:
5112:
5110:
5107:
5105:
5102:
5101:
5099:
5097:United States
5095:
5089:
5086:
5085:
5083:
5079:
5073:
5070:
5068:
5065:
5063:
5060:
5058:
5055:
5053:
5050:
5048:
5045:
5044:
5042:
5038:
5032:
5029:
5027:
5024:
5022:
5019:
5017:
5016:Guilá Naquitz
5014:
5012:
5009:
5007:
5004:
5002:
5001:Dzibilchaltún
4999:
4997:
4994:
4992:
4989:
4988:
4986:
4982:
4976:
4973:
4972:
4970:
4966:
4960:
4957:
4956:
4954:
4950:
4944:
4941:
4940:
4938:
4936:
4932:
4926:
4923:
4921:
4918:
4916:
4913:
4911:
4908:
4906:
4903:
4901:
4898:
4897:
4895:
4891:
4885:
4882:
4880:
4877:
4875:
4872:
4870:
4867:
4865:
4862:
4860:
4857:
4856:
4854:
4850:
4844:
4841:
4839:
4836:
4835:
4833:
4829:
4823:
4820:
4818:
4815:
4814:
4812:
4808:
4802:
4799:
4797:
4794:
4792:
4789:
4787:
4784:
4782:
4781:Pedra Pintada
4779:
4777:
4774:
4772:
4769:
4768:
4766:
4762:
4756:
4753:
4751:
4748:
4746:
4743:
4742:
4740:
4736:
4730:
4727:
4726:
4724:
4720:
4714:
4711:
4710:
4708:
4704:
4700:
4696:
4695:South America
4692:
4687:
4683:
4669:
4666:
4665:
4663:
4659:
4653:
4650:
4648:
4645:
4644:
4642:
4638:
4632:
4629:
4628:
4626:
4622:
4616:
4613:
4611:
4608:
4606:
4603:
4602:
4600:
4596:
4590:
4587:
4585:
4582:
4580:
4577:
4575:
4572:
4570:
4567:
4565:
4562:
4560:
4559:Klasies River
4557:
4555:
4552:
4550:
4547:
4545:
4542:
4540:
4537:
4535:
4534:Byneskranskop
4532:
4530:
4527:
4525:
4522:
4520:
4517:
4514:
4513:
4510:
4507:
4505:
4502:
4500:
4497:
4495:
4492:
4490:
4487:
4485:
4482:
4480:
4477:
4475:
4472:
4470:
4467:
4465:
4462:
4460:
4457:
4455:
4452:
4450:
4447:
4445:
4442:
4440:
4437:
4435:
4432:
4431:
4429:
4425:
4419:
4416:
4414:
4411:
4409:
4406:
4405:
4403:
4399:
4393:
4390:
4389:
4387:
4383:
4377:
4374:
4372:
4369:
4368:
4366:
4362:
4356:
4353:
4352:
4350:
4346:
4340:
4337:
4335:
4332:
4330:
4327:
4325:
4324:Ifri Oudadane
4322:
4320:
4317:
4315:
4312:
4310:
4307:
4306:
4304:
4300:
4294:
4293:Uan Muhuggiag
4291:
4289:
4286:
4285:
4283:
4279:
4273:
4270:
4269:
4267:
4263:
4257:
4254:
4252:
4249:
4248:
4246:
4242:
4236:
4233:
4231:
4228:
4227:
4225:
4221:
4215:
4212:
4211:
4209:
4205:
4199:
4196:
4195:
4193:
4189:
4183:
4180:
4178:
4175:
4174:
4172:
4168:
4162:
4159:
4158:
4156:
4152:
4148:
4144:
4139:
4135:
4121:
4118:
4116:
4113:
4112:
4110:
4106:
4100:
4097:
4095:
4092:
4091:
4089:
4085:
4079:
4076:
4074:
4071:
4069:
4066:
4065:
4063:
4059:
4055:
4051:
4047:
4041:
4038:
4036:
4033:
4031:
4028:
4027:
4025:
4021:
4015:
4012:
4010:
4007:
4005:
4002:
4000:
3999:Batadombalena
3997:
3996:
3994:
3990:
3984:
3981:
3979:
3976:
3974:
3971:
3969:
3966:
3964:
3961:
3960:
3958:
3954:
3948:
3945:
3944:
3942:
3940:
3936:
3930:
3927:
3926:
3924:
3920:
3914:
3911:
3910:
3908:
3904:
3898:
3895:
3894:
3892:
3888:
3882:
3879:
3878:
3876:
3872:
3866:
3863:
3861:
3858:
3856:
3855:Ras Baalbek I
3853:
3851:
3848:
3846:
3843:
3841:
3838:
3836:
3833:
3832:
3830:
3826:
3820:
3817:
3815:
3812:
3811:
3809:
3805:
3799:
3796:
3795:
3793:
3789:
3783:
3780:
3778:
3775:
3773:
3770:
3768:
3765:
3763:
3760:
3758:
3755:
3753:
3750:
3748:
3745:
3744:
3742:
3738:
3732:
3729:
3727:
3724:
3722:
3719:
3717:
3714:
3712:
3709:
3707:
3704:
3702:
3699:
3697:
3694:
3692:
3691:Nahal Me'arot
3689:
3687:
3684:
3682:
3679:
3677:
3674:
3672:
3669:
3667:
3664:
3663:
3661:
3657:
3651:
3648:
3646:
3643:
3642:
3640:
3636:
3630:
3627:
3625:
3622:
3620:
3617:
3615:
3612:
3610:
3607:
3605:
3602:
3600:
3597:
3595:
3592:
3590:
3587:
3586:
3584:
3580:
3574:
3571:
3569:
3566:
3564:
3561:
3559:
3556:
3555:
3553:
3549:
3543:
3540:
3538:
3535:
3533:
3530:
3528:
3525:
3523:
3520:
3518:
3515:
3513:
3510:
3509:
3507:
3503:
3497:
3494:
3493:
3491:
3487:
3481:
3478:
3476:
3473:
3471:
3468:
3467:
3465:
3461:
3455:
3452:
3450:
3447:
3445:
3442:
3440:
3437:
3435:
3432:
3430:
3427:
3425:
3422:
3420:
3417:
3415:
3412:
3410:
3407:
3406:
3404:
3400:
3394:
3391:
3390:
3388:
3384:
3378:
3375:
3373:
3370:
3368:
3365:
3363:
3360:
3358:
3355:
3353:
3350:
3348:
3345:
3344:
3342:
3338:
3332:
3329:
3328:
3326:
3322:
3316:
3313:
3312:
3310:
3306:
3302:
3298:
3293:
3289:
3275:
3274:Windmill Hill
3272:
3270:
3267:
3265:
3262:
3260:
3257:
3255:
3252:
3250:
3247:
3245:
3242:
3240:
3237:
3235:
3232:
3230:
3227:
3225:
3222:
3220:
3219:Heathery Burn
3217:
3215:
3212:
3210:
3207:
3205:
3202:
3200:
3197:
3195:
3192:
3190:
3187:
3186:
3184:
3180:
3174:
3171:
3170:
3168:
3164:
3158:
3155:
3153:
3150:
3149:
3147:
3143:
3137:
3134:
3132:
3129:
3127:
3124:
3122:
3119:
3117:
3114:
3112:
3109:
3107:
3104:
3102:
3099:
3097:
3094:
3092:
3089:
3087:
3084:
3082:
3079:
3078:
3075:
3072:
3070:
3067:
3065:
3062:
3060:
3057:
3055:
3052:
3050:
3047:
3045:
3042:
3040:
3037:
3035:
3032:
3030:
3027:
3025:
3022:
3020:
3017:
3015:
3012:
3009:
3008:
3005:
3002:
3000:
2997:
2995:
2992:
2991:
2988:
2985:
2983:
2980:
2977:
2974:
2973:
2970:
2967:
2965:
2962:
2960:
2957:
2955:
2954:Tito Bustillo
2952:
2947:
2943:
2940:
2937:
2935:Las Chimeneas
2934:
2932:
2929:
2928:
2927:
2924:
2921:
2919:
2916:
2913:
2911:
2908:
2907:
2906:
2903:
2901:
2898:
2896:
2893:
2892:
2890:
2886:
2880:
2877:
2875:
2872:
2870:
2867:
2865:
2862:
2861:
2859:
2855:
2849:
2846:
2844:
2841:
2839:
2836:
2835:
2833:
2829:
2823:
2820:
2818:
2815:
2813:
2810:
2808:
2805:
2804:
2802:
2798:
2792:
2789:
2787:
2784:
2782:
2779:
2777:
2774:
2772:
2769:
2767:
2764:
2762:
2761:Akhshtyrskaya
2759:
2758:
2756:
2752:
2746:
2743:
2741:
2738:
2736:
2733:
2731:
2728:
2727:
2725:
2721:
2715:
2712:
2710:
2709:Pego do Diabo
2707:
2705:
2702:
2700:
2697:
2695:
2692:
2690:
2687:
2685:
2682:
2680:
2677:
2676:
2674:
2670:
2664:
2661:
2659:
2656:
2655:
2653:
2649:
2643:
2640:
2639:
2637:
2633:
2627:
2624:
2623:
2621:
2619:
2615:
2609:
2606:
2605:
2603:
2599:
2593:
2590:
2589:
2587:
2583:
2577:
2574:
2573:
2571:
2567:
2561:
2558:
2557:
2555:
2553:
2549:
2543:
2540:
2538:
2535:
2533:
2530:
2528:
2527:
2523:
2521:
2518:
2516:
2513:
2511:
2508:
2506:
2503:
2501:
2500:Arene Candide
2498:
2496:
2493:
2492:
2490:
2486:
2480:
2477:
2475:
2472:
2471:
2469:
2465:
2459:
2456:
2454:
2451:
2449:
2446:
2444:
2441:
2439:
2436:
2434:
2431:
2429:
2426:
2424:
2421:
2419:
2416:
2414:
2411:
2410:
2408:
2404:
2398:
2395:
2393:
2390:
2388:
2387:Devil's Tower
2385:
2384:
2382:
2380:
2376:
2370:
2367:
2365:
2362:
2360:
2357:
2355:
2352:
2350:
2347:
2344:
2343:
2340:
2337:
2335:
2332:
2330:
2327:
2325:
2322:
2320:
2317:
2315:
2312:
2310:
2307:
2306:
2304:
2300:
2294:
2291:
2289:
2286:
2284:
2281:
2279:
2276:
2274:
2271:
2269:
2266:
2264:
2261:
2259:
2256:
2254:
2251:
2249:
2246:
2244:
2241:
2239:
2236:
2234:
2231:
2228:
2226:
2223:
2221:
2218:
2215:
2212:
2209:
2206:
2204:
2201:
2199:
2196:
2194:
2191:
2190:
2187:
2184:
2182:
2179:
2177:
2174:
2172:
2169:
2167:
2164:
2162:
2161:Pair-non-Pair
2159:
2157:
2154:
2152:
2149:
2147:
2146:Le Mas-d'Azil
2144:
2142:
2139:
2137:
2134:
2132:
2129:
2127:
2124:
2122:
2119:
2117:
2114:
2112:
2109:
2107:
2104:
2102:
2099:
2097:
2096:Arcy-sur-Cure
2094:
2091:
2090:
2087:
2084:
2082:
2079:
2078:
2075:
2072:
2070:
2067:
2065:
2062:
2059:
2057:
2054:
2051:
2049:
2046:
2043:
2041:
2038:
2036:
2035:Font-de-Gaume
2033:
2031:
2028:
2026:
2023:
2020:
2017:
2015:
2012:
2008:
2005:
2003:Abri Castanet
2002:
2001:
2000:Castel Merle
1999:
1997:
1994:
1991:
1988:
1986:
1983:
1982:
1980:
1976:
1970:
1967:
1966:
1964:
1960:
1954:
1951:
1949:
1946:
1944:
1941:
1939:
1936:
1934:
1931:
1929:
1926:
1925:
1923:
1919:
1913:
1910:
1909:
1907:
1903:
1897:
1894:
1892:
1889:
1887:
1884:
1882:
1879:
1877:
1874:
1872:
1869:
1868:
1866:
1862:
1856:
1853:
1851:
1848:
1846:
1843:
1841:
1838:
1836:
1833:
1832:
1830:
1826:
1820:
1817:
1815:
1812:
1811:
1809:
1805:
1799:
1796:
1794:
1791:
1789:
1786:
1783:
1781:
1778:
1776:
1773:
1771:
1768:
1766:
1763:
1760:
1757:
1756:
1754:
1750:
1744:
1741:
1739:
1736:
1734:
1731:
1729:
1726:
1724:
1721:
1720:
1718:
1714:
1710:
1706:
1701:
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1582:
1579:
1577:
1574:
1572:
1569:
1567:
1564:
1562:
1559:
1557:
1556:Okatse Canyon
1554:
1552:
1549:
1547:
1544:
1542:
1539:
1537:
1534:
1532:
1531:Nagarevi Cave
1529:
1527:
1524:
1522:
1519:
1517:
1514:
1512:
1509:
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1499:
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1487:
1484:
1482:
1479:
1477:
1474:
1472:
1471:Gabzaruli Tba
1469:
1467:
1466:Didghele Cave
1464:
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1457:
1454:
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1291:
1285:
1282:
1280:
1277:
1275:
1272:
1270:
1267:
1265:
1264:Pskhu-Gumista
1262:
1260:
1257:
1255:
1252:
1250:
1247:
1245:
1242:
1240:
1237:
1235:
1232:
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1227:
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1214:
1210:
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1199:
1196:
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1107:
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1103:
1097:
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984:
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934:
929:
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920:
916:
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904:
900:
896:
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891:
886:
881:
880:
863:September 25,
858:
852:
841:September 25,
836:
832:
826:
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812:
805:
800:
798:
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788:
783:
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777:
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771:
763:
758:
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754:
747:
743:
737:
735:
728:
724:(in Georgian)
720:
718:
706:September 25,
701:
695:
684:September 25,
679:
673:
667:
664:
659:
652:
647:
643:
633:
630:
628:
625:
624:
618:
616:
612:
607:
604:
602:
601:
589:
587:
583:
579:
578:Haplogroup K3
574:
572:
568:
564:
559:
557:
553:
549:
545:
544:temporal bone
535:
533:
529:
525:
521:
520:
515:
511:
507:
503:
497:
495:
491:
487:
486:Epigravettian
483:
479:
475:
473:
469:
459:
457:
456:
451:
450:
439:
436:
432:
428:
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245:
217:
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158:
154:
149:
121:
117:
114:
111:
107:
86:
61:
51:
44:
39:
34:
30:
29:IUCN category
25:
20:
5550:Wargata Mina
5510:Murrawijinie
5445:Eagles Reach
5440:Devil's Lair
5344:Wilson Butte
5129:Cherry Creek
5119:Bull Thistle
4864:Chiribiquete
4822:Charlie Lake
4750:Barton Creek
4564:Melkhoutboom
4504:Sterkfontein
4494:Plovers Lake
4427:South Africa
4329:Jebel Irhoud
4314:Ifri N'Ammar
4049:Turkmenistan
4014:Hunugalagala
3798:Iraq ed-Dubb
3563:Jeriji Saléh
3495:
3224:High Pasture
2781:Shulgan-Tash
2704:Pedra Furada
2525:
2364:Lichtenstein
2354:Brillenhöhle
2283:Le Regourdou
2243:La Ferrassie
2238:Combe Grenal
2181:Trois Frères
2116:Fontéchevade
2069:La Madeleine
1723:Drachenhöhle
1601:Takhti-Tepha
1596:Solkota Cave
1590:
1576:Sakajia Cave
1516:Melouri Cave
1511:Khomuli Cave
1496:Jortsku Cave
1476:Ghliana Cave
1441:Balda Canyon
1039:
1033:
986:
980:
951:
945:
894:
888:
876:Bibliography
861:. Retrieved
851:
839:. Retrieved
835:the original
825:
811:
762:Pinhasi 2014
704:. Retrieved
694:
682:. Retrieved
672:
665:
658:
646:
608:
605:
598:
595:
575:
560:
556:low coverage
541:
517:
514:steppe bison
498:
476:
465:
453:
447:
445:
416:
380:
379:
291:42°36′22.2″E
288:42°23′17.2″N
131:42°36′22.2″E
128:42°23′17.2″N
5636:New Zealand
5460:Gwion Gwion
5329:Trail Creek
5254:Meadowcroft
5214:Last Supper
5144:Daugherty's
5006:Las Flechas
4791:Rei do Mato
4515:Other caves
4499:Rising Star
4474:Makapansgat
4439:Bolt's Farm
4256:Njoro River
4099:Teshik-Tash
4094:Obi-Rakhmat
4078:Yarımburgaz
3978:Lapuz Lapuz
3956:Philippines
3865:Nachcharini
3860:Ras El Kelb
3819:Tam Pa Ling
3814:Tham An Mah
3696:Nahal Hemar
3444:Zhoukoudian
3419:Guanyindong
3393:Laang Spean
3315:Darra-e Kur
3308:Afghanistan
3254:Portbraddon
3244:Marble Arch
3157:Wildkirchli
3145:Switzerland
3081:Other caves
3054:Ojo Guareña
3034:Maltravieso
3029:Los Casares
3004:Siega Verde
2969:Los Aviones
2964:Santimamiñe
2938:Las Monedas
2931:El Castillo
2838:Čertova pec
2786:Mezmaiskaya
2694:Lagar Velho
2626:Cocev Kamen
2505:Balzi Rossi
2345:Other caves
2334:Sirgenstein
2263:Le Moustier
2253:Fontbrégoua
2213:Bonne-Femme
2193:Other caves
1912:Aetokremnos
1855:Utroba Cave
1761:Claminforge
1758:Belle-Roche
1521:Motena Cave
1491:Iazoni Cave
1446:Bgheri Cave
1259:Mariamjvari
588:in origin.
586:Neanderthal
462:Archaeology
431:travertines
427:stalagmites
423:stalactites
350:Paleolithic
348:Late Upper
303: /
279:Coordinates
164:Established
143: /
119:Coordinates
5763:Categories
5748:Speleology
5565:Wellington
5520:Naracoorte
5505:Mudgegonga
5490:Madjedbebe
5455:Gabarnmung
5395:Ballawinne
5385:Abrakurrie
5284:Pictograph
5062:Qillqatani
5057:Pikimachay
5052:Guitarrero
5047:Chivateros
5026:Oxtotitlán
4879:Tequendama
4589:Wonderwerk
4569:Nelson Bay
4549:Elands Bay
4509:Swartkrans
4454:Gladysvale
4401:Somaliland
4348:Mozambique
4288:Haua Fteah
4087:Uzbekistan
3752:Minatogawa
3747:Kosegasawa
3701:Nahal Oren
3645:Hazar Merd
3614:Qaleh Bozi
3599:Do-Ashkaft
3496:Satsurblia
3463:East Timor
3340:Azerbaijan
3229:Kendrick's
3194:Bontnewydd
3116:Don Gaspar
3069:Praileaitz
3024:Las Caldas
2942:La Pasiega
2869:Divje Babe
2791:Okladnikov
2776:Ignatievka
2699:Pala Pinta
2608:Għar Dalam
2585:Luxembourg
2560:St Brelade
2542:Villabruna
2453:Stravomyti
2443:Megalakkos
2428:Eileithyia
2413:Alepotrypa
2324:Hohle Fels
2210:Balauzière
2166:Pech Merle
2074:Rouffignac
2048:La Micoque
2030:Cro-Magnon
1989:Bara Bahau
1933:Býčí skála
1881:Vela Spila
1840:Devetashka
1835:Bacho Kiro
1606:Tetra Cave
1312:Bugdasheni
1284:Vashlovani
1203:Vashlovani
1178:Machakhela
787:Jones 2015
638:References
580:and Y-DNA
548:AMS dating
528:brown bear
490:stalagmite
449:Plutomurus
433:and large
419:karst cave
413:Morphology
401:region of
356:Site notes
327:karst cave
5714:Nanumanga
5698:Falemauga
5609:Makauwahi
5378:Australia
5334:Trinchera
5209:La Grange
5179:Gatecliff
5174:Franktown
5169:Fort Rock
5149:Dry Creek
5067:Toquepala
5031:Sac Actun
5021:Ox Bel Ha
5011:Frightful
4996:Coxcatlan
4975:Long Mile
4729:Quadiriki
4706:Argentina
4544:Diepkloof
4529:Boomplaas
4484:Minnaar's
4469:Kromdraai
4418:Laas Geel
4413:Dhaymoole
4408:Dhambalin
4371:Apollo 11
4272:Liphofung
4198:Shum Laka
4161:Gueldaman
4115:Con Moong
3992:Sri Lanka
3939:Palestine
3913:Padah-Lin
3850:Ksar Akil
3782:Yamashita
3767:Pinza-Abu
3573:Pettakere
3568:Liang Bua
3551:Indonesia
3537:Pahargarh
3522:Bhimbetka
3480:Lene Hara
3470:Jerimalai
3449:Zengpiyan
3439:Yuchanyan
3239:Long Hole
3136:del Valle
3111:dels Bous
3064:la Pileta
2999:Atapuerca
2914:Covalanas
2812:Pešturina
2740:Muierilor
2730:Coliboaia
2642:Svarthola
2458:Theopetra
2448:Petralona
2433:Franchthi
2423:Asphendou
2379:Gibraltar
2349:Baumann's
2339:Vogelherd
2314:Bockstein
2293:Vallonnet
2288:Rochereil
2278:Raymonden
2268:Noisetier
2220:Bruniquel
2141:Marsoulas
2131:Lombrives
2126:La Marche
2052:La Mouthe
2018:Abri Audi
1996:Cap Blanc
1938:Koněprusy
1928:Amatérská
1896:Veternica
1845:Kozarnika
1819:Ledenjača
1743:Tischofer
1733:Lurgrotte
1367:Lagodekhi
1347:Khanchali
1342:Katsoburi
1337:Kartsakhi
1322:Gardabani
1249:Lagodekhi
1239:Kintrishi
1219:Babaneuri
1168:Kintrishi
1158:Javakheti
954:: 42–52.
558:genomes.
502:wild boar
455:Oxychilus
397:, in the
113:Tsqaltubo
5555:Warratyi
5540:Tarragal
5530:Nunamira
5515:Murujuga
5480:Kutikina
5475:Koonalda
5470:Koongine
5430:Cliefden
5425:Cave Bay
5324:Tomaquag
5309:Sisyphus
5239:Mantle's
5229:Lovelock
5224:LoDaisKa
5204:Humboldt
5088:Werehpai
5081:Suriname
4991:Chan Hol
4910:Centella
4900:Ambrosio
4852:Colombia
4817:Bluefish
4661:Zimbabwe
4598:Tanzania
4584:Stadsaal
4489:Motsetsi
4459:Gondolin
4449:Drimolen
4444:Cooper's
4339:Taforalt
4309:Hercules
4235:Swimmers
4207:DR Congo
4191:Cameroon
4170:Botswana
4040:Tham Lod
4030:Pha Taem
4023:Thailand
4004:Belilena
3922:Pakistan
3890:Mongolia
3874:Malaysia
3835:Antelias
3777:Tobayama
3731:Zuttiyeh
3650:Shanidar
3512:Ambadevi
3429:Tianyuan
3409:Baishiya
3386:Cambodia
3126:El Mirón
3121:Guanches
3096:Armintxe
3014:Bacinete
2959:Altxerri
2948:El Pendo
2918:La Garma
2900:Altamira
2857:Slovenia
2848:Jasovská
2831:Slovakia
2817:Risovača
2771:Denisova
2735:Cuciulat
2689:Furninha
2684:Escoural
2672:Portugal
2663:Paradise
2658:Obłazowa
2592:Loschbur
2532:Paglicci
2515:Corbeddu
2397:Vanguard
2392:Gorham's
2273:La Quina
2216:Bouillon
2203:Aurignac
2007:Reverdit
1992:Bernifal
1871:Grapčeva
1828:Bulgaria
1788:Scladina
1775:Naulette
1738:Salzofen
1451:Birtvisi
1382:Sataplia
1372:Madatapa
1352:Kobuleti
1317:Chachuna
1274:Sataplia
1244:Kobuleti
1173:Kolkheti
1076:25354048
1035:PLOS ONE
1021:26567969
989:: 8912.
968:25769538
937:27135931
621:See also
538:Genetics
524:roe deer
506:red deer
385:Georgian
316:Altitude
5666:Chugai'
5648:Ruakuri
5583:Gadao's
5560:Warreen
5495:Mammoth
5465:Jenolan
5410:Blanche
5390:Acheron
5367:Oceania
5339:Ventana
5319:Tainter
5294:Russell
5279:Pendejo
5274:Paisley
5234:Mammoth
5154:Durango
5114:Bonfire
4968:Jamaica
4935:Curaçao
4874:Sáchica
4859:El Abra
4843:Milodón
4786:Peruaçu
4776:Maquiné
4668:Bambata
4647:Kalemba
4519:Blombos
4464:Haasgat
4385:Nigeria
4364:Namibia
4302:Morocco
4265:Lesotho
4177:Tsodilo
4154:Algeria
4108:Vietnam
4068:Belbaşı
4054:Dzhebel
4009:Fa Hien
3968:Kalanay
3929:Sanghao
3906:Myanmar
3845:Kaukaba
3828:Lebanon
3716:Raqefet
3686:Misliya
3671:HaYonim
3619:Warwasi
3594:Darband
3589:Bisitun
3558:Harimau
3532:Gudiyam
3527:Edakkal
3489:Georgia
3434:Xianren
3362:Damjili
3357:Buzeyir
3331:Areni-1
3324:Armenia
3249:Oldbury
3214:Gough's
3199:Cathole
3173:Verteba
3166:Ukraine
2723:Romania
2714:Salemas
2679:Aroeira
2510:Cavallo
2495:Addaura
2474:Baradla
2467:Hungary
2418:Apidima
2302:Germany
2258:Lazaret
2186:Villars
2106:Cosquer
2086:Chauvet
2064:Lascaux
1962:Finland
1886:Vindija
1876:Krapina
1864:Croatia
1784:Ramioul
1752:Belgium
1728:Gudenus
1716:Austria
1408:Tusheti
1392:Tetrobi
1302:Ajameti
1279:Tusheti
1254:Liakhvi
1234:Borjomi
1224:Batsara
1198:Tusheti
1193:Tbilisi
1183:Mtirala
1163:Kazbegi
1067:4213019
1044:Bibcode
1012:4660371
991:Bibcode
928:4943878
899:Bibcode
603:(CHG).
567:SLC24A5
510:aurochs
403:Georgia
399:Imereti
345:Periods
340:History
180:Website
5707:Tuvalu
5643:Moncks
5627:Pindai
5602:Hawaii
5593:Talagi
5588:Mahlac
5450:Fossil
5435:Cloggs
5420:Burrup
5299:Sandia
5244:Marmes
5189:Hidden
5184:Graham
5139:Danger
5124:Burnet
4984:Mexico
4959:Pomier
4920:Patana
4905:Calero
4884:Tibitó
4810:Canada
4771:Gentio
4764:Brazil
4738:Belize
4652:Mumbwa
4640:Zambia
4624:Uganda
4605:Kondoa
4579:Sibudu
4524:Border
4479:Malapa
4355:Ngalue
4230:Beasts
4214:Matupi
4143:Africa
4073:Karain
4061:Turkey
4035:Spirit
3973:Minori
3963:Callao
3947:Shuqba
3791:Jordan
3762:Ōzakai
3757:Muroya
3706:Qafzeh
3676:Kebara
3659:Israel
3629:Yafteh
3624:Wezmeh
3454:Zhiren
3367:Tağlar
3269:Thor's
3204:Coygan
3152:Bichon
3106:Bedmar
3074:Sidrón
2910:Chufín
2843:Domica
2800:Serbia
2754:Russia
2651:Poland
2635:Norway
2569:Kosovo
2552:Jersey
2537:Romito
2479:Szelim
2438:Kleidi
2406:Greece
2111:Cussac
2101:Gargas
2056:Pataud
1978:France
1948:Mladeč
1905:Cyprus
1850:Magura
1814:Badanj
1807:Bosnia
1705:Europe
1377:Nedzvi
1357:Korugi
1148:Algeti
1074:
1064:
1019:
1009:
966:
935:
925:
890:Nature
522:, and
478:Lithic
435:gourds
405:, 287
387::
332:Length
271:Region
5691:Samoa
5545:Ubirr
5535:ORS 7
5304:Shoup
5264:Mummy
5259:Modoc
5249:Martz
5199:Hogup
5109:Baker
4925:Pluma
4831:Chile
4796:Santo
4722:Aruba
4691:North
4631:Nyero
4615:Mumba
4539:Cango
4281:Libya
4244:Kenya
4223:Egypt
4120:Ngườm
3983:Tabon
3840:Jeita
3740:Japan
3726:Tabun
3721:Skhul
3711:Qesem
3681:Manot
3542:Tikla
3517:Belum
3505:India
3475:Laili
3424:Luobi
3414:Fuyan
3402:China
3377:Gazma
3352:Azykh
3347:Allar
3234:Kents
3101:Axlor
3091:Antón
3086:Ángel
3044:Nerja
2982:Araña
2888:Spain
2879:Potok
2874:Pekel
2864:Betal
2601:Malta
2526:Felci
2488:Italy
2369:Ofnet
2229:Cauna
2225:Calès
2198:Arago
2176:Renne
2156:Niaux
1953:Šipka
1943:Kůlna
1770:Goyet
1765:Engis
1387:Sulda
1269:Ritsa
563:HERC2
494:ochre
442:Fauna
335:125 m
31:III (
5734:Cave
5682:Kilu
5575:Guam
5415:Bone
5219:Levi
5159:Dust
5040:Peru
4943:Hato
4915:Cura
4893:Cuba
4838:Fell
4693:and
4610:Bahi
3881:Niah
3807:Laos
3666:Amud
3638:Iraq
3582:Iran
3297:Asia
3264:Sand
3049:Niño
2745:Oase
2520:Deer
2248:Fées
1969:Wolf
1332:Iori
1327:Ilto
1072:PMID
1017:PMID
964:PMID
933:PMID
865:2016
843:2016
708:2016
686:2016
571:EDAR
482:flax
452:and
324:Type
167:2011
156:Area
4392:Rop
3372:Zar
2207:Azé
1793:Spy
1307:Asa
1062:PMC
1052:doi
1007:PMC
999:doi
956:doi
952:166
923:PMC
915:hdl
907:doi
895:534
5765::
1070:.
1060:.
1050:.
1038:.
1032:.
1015:.
1005:.
997:.
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