215:
201:
55:
189:
176:
33:
269:
is one of the swarm founding species, meaning that new nests are founded by a large group of workers with a smaller number of inseminated females (egg-laying foundresses), while
2685:
675:
2175:
2025:
1855:
1775:
1325:
705:
1765:
1585:
1385:
1105:
1085:
1005:
365:
2075:
1945:
1935:
1885:
1625:
1255:
965:
945:
875:
735:
725:
345:
315:
2275:
2185:
2045:
2015:
1985:
1835:
1435:
1365:
1055:
975:
845:
785:
655:
2285:
2225:
2215:
2145:
2115:
2095:
2055:
1845:
1555:
1355:
1315:
1275:
1265:
1175:
1165:
1155:
1035:
955:
865:
775:
595:
575:
555:
445:
305:
2255:
2235:
2165:
2085:
2005:
1975:
1965:
1865:
1665:
1635:
1535:
1525:
1515:
1505:
1395:
1285:
1205:
1195:
1125:
1065:
1015:
925:
885:
665:
635:
485:
475:
395:
355:
335:
219:
2265:
2245:
2205:
2065:
1955:
1875:
1815:
1755:
1745:
1715:
1695:
1675:
1605:
1595:
1415:
1405:
1375:
1345:
1295:
1245:
1215:
1185:
1145:
1075:
1045:
995:
895:
815:
795:
765:
755:
715:
685:
645:
625:
565:
545:
535:
465:
425:
405:
193:
2195:
2135:
2035:
1995:
1825:
1795:
1735:
1725:
1655:
1615:
1575:
1565:
1495:
1485:
1465:
1445:
1335:
1235:
1135:
1095:
985:
935:
905:
855:
835:
805:
745:
615:
585:
525:
515:
505:
495:
435:
415:
375:
41:
2295:
2155:
2105:
1895:
1785:
1685:
1645:
1455:
1305:
1225:
1025:
695:
605:
455:
1925:
1545:
1475:
1115:
825:
385:
2608:
2621:
2582:
153:
2626:
294:
segment being petiolate but (in dorsal view) not parallel-sided, and the mesepisternum lacking a scrobal sulcus.
2439:"Keys to the genera of social wasps of South-East Asia (Hymenoptera: Vespidae): Social wasps of South-East Asia"
2680:
2530:
2517:
2535:
2321:
2652:
54:
1905:
271:
2470:
2322:
Vespidae (Hymenoptera) of
Vietnam 3: Synoptic key to Vietnamese species of the polistine genus
2613:
2317:
Guérin-Méneville, 1831 (Hymenoptera, Vespidae) from China, with description of one new species
2639:
2556:
1705:
8:
2508:
1805:
1425:
2438:
2330:
2312:
2125:
915:
49:
2675:
2634:
2543:
2454:
1915:
325:
265:
180:
2548:
2450:
2419:
2371:
275:
is independent-founding, meaning that each nest is founded by a single foundress.
2522:
241:
200:
2493:
2669:
214:
263:
is unusual because it contains both independent and swarm-founding species.
248:
2595:
2502:
2376:
2355:
256:
252:
244:
233:
136:
106:
2587:
2423:
237:
126:
2600:
2569:
2406:
Yosiaki, ItĂ´ (1987). "Social behaviour of the
Australian paper wasp,
188:
86:
66:
2464:
175:
2574:
2487:
2360:, with a note on its developmental process (Hymenoptera: Vespidae)"
291:
287:
116:
290:
having a dorsal carina but lacking a pretegular carina, the first
96:
76:
32:
2561:
2647:
286:
can be distinguished from other genera in the tribe by: the
2437:
Carpenter, James M.; Nguyen, Lien Phuong Thi (2003).
2335:in the Indian subcontinent (Hymenoptera: Vespidae)
2667:
2356:"Nest architecture of the Australian paper wasp
2331:Taxonomic notes on the paper wasps of the genus
2436:
2686:Taxa named by Félix Édouard Guérin-Méneville
2395:. Oxford University Press. pp. 57–60.
31:
2375:
2353:
2326:, with notes on taxonomy and distribution
2410:(de Saussure) (Hymenoptera: Vespidae)".
330:Polašek, Bellingan & van Noort, 2022
213:
199:
187:
174:
2405:
2668:
2313:Pictorial key to species of the genus
2469:
2468:
2354:Yamane, Soichi; Ito, Yosiaki (1994).
2653:B44B87D2-FF98-E668-ECBE-FC0AFA2BFA67
2390:
259:biogeographical regions. The genus
13:
2384:
14:
2697:
2347:
2306:
2455:10.1046/j.1343-8786.2003.00016.x
53:
2364:Psyche: A Journal of Entomology
2430:
2399:
278:
1:
2393:The Evolution of Social Wasps
2340:
7:
247:distributed throughout the
10:
2702:
676:Ropalidia constitutionalis
297:
2477:
1230:Polašek and de Beer, 2023
166:
161:
50:Scientific classification
48:
39:
30:
23:
2408:Ropalidia revolutionalis
2358:Ropalidia romandi cabeti
2176:Ropalidia trichophthalma
2026:Ropalidia semihyalineata
1906:Ropalidia revolutionalis
1856:Ropalidia pseudomalayana
1776:Ropalidia philippinensis
1326:Ropalidia longipetiolata
706:Ropalidia crassipunctata
272:Ropalidia revolutionalis
2391:Hunt, James H. (2007).
1766:Ropalidia phalansterica
1586:Ropalidia nigrofemorata
1386:Ropalidia maculiventris
1106:Ropalidia hongkongensis
1086:Ropalidia guttatipennis
1006:Ropalidia fulvopruinosa
366:Ropalidia aristocratica
2076:Ropalidia socialistica
1946:Ropalidia rufoplagiata
1936:Ropalidia rufocollaris
1886:Ropalidia reactionalis
1626:Ropalidia novaeguinaea
1256:Ropalidia latebalteata
966:Ropalidia flavoviridis
946:Ropalidia flavobrunnea
876:Ropalidia erythrospila
736:Ropalidia cyathiformis
726:Ropalidia curvilineata
346:Ropalidia andamanensis
316:Ropalidia albobalteata
224:
211:
197:
185:
2443:Entomological Science
2276:Ropalidia vitripennis
2186:Ropalidia trimaculata
2046:Ropalidia sexmaculata
2016:Ropalidia sculpturata
1986:Ropalidia schulthessi
1970:Das & Gupta, 1989
1836:Ropalidia principalis
1580:Das & Gupta, 1989
1436:Ropalidia mathematica
1366:Ropalidia mackayensis
1056:Ropalidia grandidieri
976:Ropalidia fluviatilis
846:Ropalidia duchaussoyi
786:Ropalidia democratica
656:Ropalidia conservator
510:Das & Gupta, 1989
350:Das & Gupta, 1989
217:
203:
191:
178:
2286:Ropalidia wollastoni
2226:Ropalidia variabilis
2220:Giordani Soika, 1981
2216:Ropalidia unidentata
2146:Ropalidia thailandia
2116:Ropalidia subclavata
2096:Ropalidia spilostoma
2056:Ropalidia shestakowi
1846:Ropalidia proletaria
1706:Ropalidia ornaticeps
1670:Giordani Soika, 1991
1640:Giordani Soika, 1944
1600:Giordani Soika, 1991
1556:Ropalidia nigrescens
1356:Ropalidia luzonensis
1316:Ropalidia linearecta
1276:Ropalidia laticincta
1266:Ropalidia latetergum
1176:Ropalidia irrequieta
1166:Ropalidia interrupta
1156:Ropalidia interjecta
1036:Ropalidia gracilenta
956:Ropalidia flavopicta
900:Giordani Soika, 1977
866:Ropalidia elegantula
776:Ropalidia deminutiva
710:Giordani Soika, 1981
596:Ropalidia cincinnata
576:Ropalidia celebensis
556:Ropalidia catharinae
446:Ropalidia bicolorata
306:Ropalidia aethiopica
232:is a large genus of
168:approx. 200 species
2412:Journal of Ethology
2290:(Meade-Waldo, 1912)
2256:Ropalidia venustula
2236:Ropalidia variegata
2166:Ropalidia tomentosa
2086:Ropalidia spatulata
2030:(Meade-Waldo, 1912)
2006:Ropalidia scottiana
1976:Ropalidia saussurei
1966:Ropalidia santoshae
1866:Ropalidia pulchella
1806:Ropalidia plebeiana
1666:Ropalidia obscurior
1636:Ropalidia novissima
1536:Ropalidia nigerrima
1526:Ropalidia mysterica
1516:Ropalidia myosolica
1506:Ropalidia mutabilis
1470:(Meade-Waldo, 1912)
1426:Ropalidia marginata
1396:Ropalidia magnanima
1286:Ropalidia lefebvrei
1206:Ropalidia jaculator
1196:Ropalidia jacobsoni
1126:Ropalidia impetuosa
1066:Ropalidia granulata
1016:Ropalidia galimatia
980:(Meade-Waldo, 1912)
926:Ropalidia favulorum
886:Ropalidia eurostoma
666:Ropalidia conspicua
636:Ropalidia cocoscola
486:Ropalidia bispinosa
476:Ropalidia bipartita
396:Ropalidia australis
356:Ropalidia antennata
336:Ropalidia anarchica
220:Ropalidia variegata
2681:Hymenoptera genera
2424:10.1007/bf02349943
2377:10.1155/1994/92839
2270:Gusenleitner, 1996
2266:Ropalidia vietnama
2246:Ropalidia velutina
2206:Ropalidia unicolor
2126:Ropalidia sumatrae
2066:Ropalidia socialis
2020:Gusenleitner, 2001
1956:Ropalidia sakalava
1876:Ropalidia ranavali
1816:Ropalidia politica
1756:Ropalidia petulans
1746:Ropalidia perplexa
1716:Ropalidia palawana
1696:Ropalidia opulenta
1676:Ropalidia ochracea
1660:Gusenleitner, 1966
1630:(Schulthess, 1913)
1606:Ropalidia nitidula
1596:Ropalidia nilssoni
1430:(Lepeletier, 1836)
1416:Ropalidia malayana
1406:Ropalidia malaisei
1376:Ropalidia maculata
1346:Ropalidia lugubris
1296:Ropalidia leopoldi
1246:Ropalidia kurandae
1216:Ropalidia javanica
1186:Ropalidia irritata
1146:Ropalidia inquieta
1076:Ropalidia gregaria
1046:Ropalidia gracilis
996:Ropalidia fraterna
916:Ropalidia fasciata
896:Ropalidia excavata
816:Ropalidia distigma
796:Ropalidia dichroma
766:Ropalidia decorata
756:Ropalidia deceptor
716:Ropalidia cristata
686:Ropalidia copiaria
646:Ropalidia colorata
626:Ropalidia clypeata
610:(Lepeletier, 1836)
566:Ropalidia cauponae
546:Ropalidia carinata
536:Ropalidia capensis
466:Ropalidia binghami
426:Ropalidia bicincta
406:Ropalidia bambusae
225:
212:
198:
194:Ropalidia distigma
186:
2663:
2662:
2635:Open Tree of Life
2471:Taxon identifiers
2196:Ropalidia turneri
2150:Gusenleitner 1994
2136:Ropalidia taiwana
2036:Ropalidia sericea
1996:Ropalidia scitula
1916:Ropalidia romandi
1826:Ropalidia prasina
1796:Ropalidia plebeja
1736:Ropalidia pendula
1726:Ropalidia papuana
1656:Ropalidia obscura
1616:Ropalidia nobilis
1576:Ropalidia nigrita
1566:Ropalidia nigrior
1496:Ropalidia montana
1486:Ropalidia modesta
1466:Ropalidia mimikae
1446:Ropalidia melania
1336:Ropalidia loriana
1236:Ropalidia kojimai
1136:Ropalidia incurva
1096:Ropalidia hirsuta
986:Ropalidia formosa
936:Ropalidia festina
906:Ropalidia extrema
856:Ropalidia eboraca
836:Ropalidia ducalis
806:Ropalidia dispila
746:Ropalidia darwini
740:(Fabricius, 1804)
616:Ropalidia clavata
586:Ropalidia chromis
526:Ropalidia canaria
516:Ropalidia brunnea
506:Ropalidia brevita
496:Ropalidia brazzai
436:Ropalidia bicolor
416:Ropalidia bensoni
376:Ropalidia artifex
326:Ropalidia amabala
266:Ropalidia romandi
181:Ropalidia amabala
173:
172:
157:
42:Ropalidia nobilis
2693:
2656:
2655:
2643:
2642:
2630:
2629:
2617:
2616:
2604:
2603:
2591:
2590:
2578:
2577:
2565:
2564:
2552:
2551:
2539:
2538:
2526:
2525:
2513:
2512:
2511:
2498:
2497:
2496:
2466:
2465:
2459:
2458:
2434:
2428:
2427:
2403:
2397:
2396:
2388:
2382:
2381:
2379:
2370:(3–4): 145–158.
2351:
2301:
2296:Ropalidia zonata
2291:
2281:
2280:(Saussure, 1890)
2271:
2261:
2260:(Saussure, 1900)
2251:
2250:(Saussure, 1890)
2241:
2231:
2230:(Saussure, 1890)
2221:
2211:
2201:
2191:
2181:
2171:
2161:
2156:Ropalidia timida
2151:
2141:
2131:
2121:
2120:(Saussure, 1890)
2111:
2106:Ropalidia stigma
2101:
2091:
2081:
2080:(Saussure, 1854)
2071:
2070:(Saussure, 1862)
2061:
2060:Schulthess, 1931
2051:
2041:
2031:
2021:
2011:
2010:(Saussure, 1890)
2001:
1991:
1990:(Saussure, 1890)
1981:
1971:
1961:
1960:(Saussure, 1900)
1951:
1941:
1931:
1921:
1911:
1910:(Saussure, 1854)
1901:
1900:(Saussure, 1900)
1896:Ropalidia regina
1891:
1890:(Saussure, 1854)
1881:
1880:(Saussure, 1890)
1871:
1870:(Saussure, 1900)
1861:
1851:
1841:
1831:
1830:(Saussure, 1900)
1821:
1820:(Saussure, 1854)
1811:
1801:
1791:
1786:Ropalidia pilosa
1781:
1780:(Saussure, 1854)
1771:
1770:(Saussure, 1853)
1761:
1751:
1741:
1731:
1721:
1711:
1701:
1691:
1686:Ropalidia opifex
1681:
1671:
1661:
1651:
1646:Ropalidia nursei
1641:
1631:
1621:
1611:
1610:(Saussure, 1890)
1601:
1591:
1581:
1571:
1561:
1551:
1541:
1531:
1521:
1511:
1501:
1491:
1481:
1471:
1461:
1456:Ropalidia merina
1451:
1441:
1431:
1421:
1411:
1401:
1391:
1381:
1371:
1361:
1351:
1341:
1331:
1321:
1311:
1306:Ropalidia lepida
1301:
1291:
1281:
1271:
1261:
1251:
1241:
1231:
1226:Ropalidia jemmae
1221:
1211:
1201:
1191:
1181:
1171:
1161:
1160:(Saussure, 1900)
1151:
1141:
1131:
1121:
1111:
1110:(Saussure, 1854)
1101:
1091:
1090:(Saussure, 1853)
1081:
1080:(Saussure, 1854)
1071:
1061:
1060:(Saussure, 1890)
1051:
1041:
1031:
1026:Ropalidia gemmea
1021:
1020:(Saussure, 1853)
1011:
1001:
1000:(Saussure, 1900)
991:
990:(Saussure, 1854)
981:
971:
961:
951:
941:
931:
921:
920:(Fabricius 1804)
911:
901:
891:
881:
871:
861:
851:
841:
840:(Saussure, 1900)
831:
830:(Saussure, 1853)
821:
811:
801:
791:
790:(Saussure, 1853)
781:
771:
761:
751:
741:
731:
721:
711:
701:
696:Ropalidia crassa
691:
690:(Saussure, 1862)
681:
680:(Saussure, 1853)
671:
661:
651:
641:
631:
621:
620:(Saussure, 1853)
611:
606:Ropalidia cincta
601:
591:
581:
571:
561:
551:
550:(Saussure, 1890)
541:
540:(Saussure, 1862)
531:
521:
511:
501:
491:
481:
471:
461:
456:Ropalidia bidens
451:
441:
431:
430:(Saussure, 1890)
421:
411:
401:
400:(Saussure, 1854)
391:
390:(Saussure, 1890)
381:
380:(Saussure, 1853)
371:
370:(Saussure, 1853)
361:
360:(Saussure, 1890)
351:
341:
340:(Saussure, 1853)
331:
321:
311:
154:Guérin-Méneville
152:
58:
57:
35:
21:
20:
2701:
2700:
2696:
2695:
2694:
2692:
2691:
2690:
2666:
2665:
2664:
2659:
2651:
2646:
2638:
2633:
2625:
2620:
2612:
2607:
2599:
2594:
2586:
2581:
2573:
2568:
2560:
2555:
2547:
2542:
2534:
2529:
2521:
2516:
2507:
2506:
2501:
2492:
2491:
2486:
2473:
2463:
2462:
2435:
2431:
2404:
2400:
2389:
2385:
2352:
2348:
2343:
2309:
2304:
2299:
2289:
2279:
2269:
2259:
2249:
2239:
2229:
2219:
2209:
2199:
2189:
2179:
2169:
2159:
2149:
2139:
2129:
2119:
2109:
2100:(Cameron, 1906)
2099:
2089:
2079:
2069:
2059:
2049:
2039:
2029:
2019:
2009:
2000:(Bingham, 1897)
1999:
1989:
1979:
1969:
1959:
1949:
1940:(Cameron, 1900)
1939:
1929:
1926:Ropalidia rosae
1919:
1909:
1899:
1889:
1879:
1869:
1859:
1849:
1839:
1829:
1819:
1809:
1800:(Saussure 1862)
1799:
1789:
1779:
1769:
1759:
1749:
1739:
1729:
1719:
1710:(Cameron, 1900)
1709:
1699:
1689:
1679:
1669:
1659:
1649:
1639:
1629:
1619:
1609:
1599:
1590:(Cameron, 1910)
1589:
1579:
1569:
1559:
1549:
1546:Ropalidia nigra
1539:
1529:
1519:
1509:
1499:
1489:
1479:
1476:Ropalidia minor
1469:
1459:
1449:
1439:
1429:
1419:
1409:
1399:
1389:
1379:
1369:
1359:
1349:
1340:(Buysson, 1910)
1339:
1330:(Cameron, 1911)
1329:
1319:
1309:
1299:
1289:
1279:
1269:
1260:(Cameron, 1902)
1259:
1249:
1239:
1229:
1219:
1209:
1200:(Buysson, 1908)
1199:
1189:
1179:
1169:
1159:
1149:
1139:
1129:
1119:
1116:Ropalidia horni
1109:
1099:
1089:
1079:
1069:
1059:
1049:
1039:
1029:
1019:
1010:(Cameron, 1906)
1009:
999:
989:
979:
969:
959:
949:
939:
929:
919:
909:
899:
889:
880:(Cameron, 1908)
879:
869:
859:
850:(Gribodo, 1896)
849:
839:
829:
826:Ropalidia dubia
819:
809:
799:
789:
779:
769:
759:
749:
739:
729:
719:
709:
699:
689:
679:
669:
659:
649:
639:
629:
619:
609:
599:
589:
579:
569:
560:(Cameron, 1913)
559:
549:
539:
529:
519:
509:
499:
489:
479:
469:
459:
449:
439:
429:
419:
409:
399:
389:
379:
369:
359:
349:
339:
329:
320:(Cameron, 1906)
319:
310:(Buysson, 1906)
309:
300:
281:
151:
52:
17:
12:
11:
5:
2699:
2689:
2688:
2683:
2678:
2661:
2660:
2658:
2657:
2644:
2631:
2618:
2605:
2592:
2579:
2566:
2553:
2540:
2527:
2514:
2499:
2483:
2481:
2475:
2474:
2461:
2460:
2449:(3): 183–192.
2429:
2418:(2): 115–124.
2398:
2383:
2345:
2344:
2342:
2339:
2338:
2337:
2328:
2319:
2308:
2307:Identification
2305:
2303:
2302:
2292:
2282:
2272:
2262:
2252:
2242:
2232:
2222:
2212:
2202:
2200:Richards, 1978
2192:
2182:
2180:Richards, 1978
2172:
2170:(Gerst., 1857)
2162:
2152:
2142:
2132:
2122:
2112:
2102:
2092:
2082:
2072:
2062:
2052:
2042:
2032:
2022:
2012:
2002:
1992:
1982:
1972:
1962:
1952:
1950:(Cameron 1905)
1942:
1932:
1930:Blommers, 2012
1922:
1920:(Guill., 1841)
1912:
1902:
1892:
1882:
1872:
1862:
1852:
1842:
1832:
1822:
1812:
1810:Richards, 1978
1802:
1792:
1782:
1772:
1762:
1752:
1750:Blommers, 2012
1742:
1732:
1722:
1712:
1702:
1692:
1682:
1672:
1662:
1652:
1642:
1632:
1622:
1620:(Gerst., 1857)
1612:
1602:
1592:
1582:
1572:
1570:Richards, 1978
1562:
1552:
1542:
1532:
1530:Blommers, 2012
1522:
1512:
1510:Richards, 1978
1502:
1492:
1482:
1480:Saussure, 1900
1472:
1462:
1460:Blommers, 2012
1452:
1450:Richards, 1978
1442:
1432:
1422:
1420:(Cameron 1903)
1412:
1402:
1392:
1382:
1372:
1370:Richards, 1978
1362:
1352:
1342:
1332:
1322:
1320:Blommers, 2012
1312:
1302:
1292:
1282:
1272:
1270:Richards, 1978
1262:
1252:
1250:Richards, 1978
1242:
1240:Blommers, 2012
1232:
1222:
1212:
1202:
1192:
1182:
1172:
1162:
1152:
1142:
1132:
1122:
1112:
1102:
1092:
1082:
1072:
1062:
1052:
1042:
1040:Richards, 1978
1032:
1022:
1012:
1002:
992:
982:
972:
962:
952:
942:
932:
930:Blommers, 2012
922:
912:
902:
892:
882:
872:
870:Richards, 1978
862:
860:Richards, 1978
852:
842:
832:
822:
820:(Gerst., 1857)
812:
810:(Cameron 1913)
802:
792:
782:
772:
762:
752:
750:Richards, 1978
742:
732:
730:(Cameron 1908)
722:
712:
702:
692:
682:
672:
662:
652:
642:
640:Blommers, 2012
632:
622:
612:
602:
600:Blommers, 2012
592:
582:
572:
570:Blommers, 2012
562:
552:
542:
532:
522:
512:
502:
500:(Buysson 1906)
492:
482:
472:
462:
452:
442:
432:
422:
420:Richards, 1978
412:
410:Richards, 1978
402:
392:
386:Ropalidia atra
382:
372:
362:
352:
342:
332:
322:
312:
301:
299:
296:
280:
277:
171:
170:
164:
163:
159:
158:
144:
140:
139:
134:
130:
129:
124:
120:
119:
114:
110:
109:
104:
100:
99:
94:
90:
89:
84:
80:
79:
74:
70:
69:
64:
60:
59:
46:
45:
37:
36:
28:
27:
16:Genus of wasps
15:
9:
6:
4:
3:
2:
2698:
2687:
2684:
2682:
2679:
2677:
2674:
2673:
2671:
2654:
2649:
2645:
2641:
2636:
2632:
2628:
2623:
2619:
2615:
2610:
2606:
2602:
2597:
2593:
2589:
2584:
2580:
2576:
2571:
2567:
2563:
2558:
2554:
2550:
2545:
2541:
2537:
2532:
2528:
2524:
2519:
2515:
2510:
2504:
2500:
2495:
2489:
2485:
2484:
2482:
2480:
2476:
2472:
2467:
2456:
2452:
2448:
2444:
2440:
2433:
2425:
2421:
2417:
2413:
2409:
2402:
2394:
2387:
2378:
2373:
2369:
2365:
2361:
2359:
2350:
2346:
2336:
2334:
2329:
2327:
2325:
2320:
2318:
2316:
2311:
2310:
2298:
2297:
2293:
2288:
2287:
2283:
2278:
2277:
2273:
2268:
2267:
2263:
2258:
2257:
2253:
2248:
2247:
2243:
2240:(Smith, 1852)
2238:
2237:
2233:
2228:
2227:
2223:
2218:
2217:
2213:
2210:(Smith, 1859)
2208:
2207:
2203:
2198:
2197:
2193:
2188:
2187:
2183:
2178:
2177:
2173:
2168:
2167:
2163:
2158:
2157:
2153:
2148:
2147:
2143:
2138:
2137:
2133:
2130:(Weber, 1801)
2128:
2127:
2123:
2118:
2117:
2113:
2110:(Smith, 1858)
2108:
2107:
2103:
2098:
2097:
2093:
2088:
2087:
2083:
2078:
2077:
2073:
2068:
2067:
2063:
2058:
2057:
2053:
2048:
2047:
2043:
2040:Cameron, 1905
2038:
2037:
2033:
2028:
2027:
2023:
2018:
2017:
2013:
2008:
2007:
2003:
1998:
1997:
1993:
1988:
1987:
1983:
1978:
1977:
1973:
1968:
1967:
1963:
1958:
1957:
1953:
1948:
1947:
1943:
1938:
1937:
1933:
1928:
1927:
1923:
1918:
1917:
1913:
1908:
1907:
1903:
1898:
1897:
1893:
1888:
1887:
1883:
1878:
1877:
1873:
1868:
1867:
1863:
1858:
1857:
1853:
1850:Richards 1978
1848:
1847:
1843:
1838:
1837:
1833:
1828:
1827:
1823:
1818:
1817:
1813:
1808:
1807:
1803:
1798:
1797:
1793:
1790:(Smith, 1859)
1788:
1787:
1783:
1778:
1777:
1773:
1768:
1767:
1763:
1760:Cheesm., 1952
1758:
1757:
1753:
1748:
1747:
1743:
1740:(Smith, 1857)
1738:
1737:
1733:
1728:
1727:
1723:
1718:
1717:
1713:
1708:
1707:
1703:
1700:(Smith, 1857)
1698:
1697:
1693:
1688:
1687:
1683:
1678:
1677:
1673:
1668:
1667:
1663:
1658:
1657:
1653:
1648:
1647:
1643:
1638:
1637:
1633:
1628:
1627:
1623:
1618:
1617:
1613:
1608:
1607:
1603:
1598:
1597:
1593:
1588:
1587:
1583:
1578:
1577:
1573:
1568:
1567:
1563:
1558:
1557:
1553:
1548:
1547:
1543:
1538:
1537:
1533:
1528:
1527:
1523:
1520:(Dalla Torre)
1518:
1517:
1513:
1508:
1507:
1503:
1498:
1497:
1493:
1490:(Smith, 1858)
1488:
1487:
1483:
1478:
1477:
1473:
1468:
1467:
1463:
1458:
1457:
1453:
1448:
1447:
1443:
1440:(Smith, 1861)
1438:
1437:
1433:
1428:
1427:
1423:
1418:
1417:
1413:
1408:
1407:
1403:
1398:
1397:
1393:
1388:
1387:
1383:
1378:
1377:
1373:
1368:
1367:
1363:
1358:
1357:
1353:
1350:(Smith, 1858)
1348:
1347:
1343:
1338:
1337:
1333:
1328:
1327:
1323:
1318:
1317:
1313:
1308:
1307:
1303:
1300:Bequard, 1932
1298:
1297:
1293:
1288:
1287:
1283:
1278:
1277:
1273:
1268:
1267:
1263:
1258:
1257:
1253:
1248:
1247:
1243:
1238:
1237:
1233:
1228:
1227:
1223:
1218:
1217:
1213:
1208:
1207:
1203:
1198:
1197:
1193:
1190:(Smith, 1864)
1188:
1187:
1183:
1178:
1177:
1173:
1168:
1167:
1163:
1158:
1157:
1153:
1150:Cheesm., 1952
1148:
1147:
1143:
1140:Cheesm., 1952
1138:
1137:
1133:
1130:(Smith, 1861)
1128:
1127:
1123:
1118:
1117:
1113:
1108:
1107:
1103:
1100:Richards 1978
1098:
1097:
1093:
1088:
1087:
1083:
1078:
1077:
1073:
1068:
1067:
1063:
1058:
1057:
1053:
1050:(Smith, 1859)
1048:
1047:
1043:
1038:
1037:
1033:
1030:Cheesm., 1952
1028:
1027:
1023:
1018:
1017:
1013:
1008:
1007:
1003:
998:
997:
993:
988:
987:
983:
978:
977:
973:
968:
967:
963:
960:(Smith, 1857)
958:
957:
953:
948:
947:
943:
940:(Smith, 1864)
938:
937:
933:
928:
927:
923:
918:
917:
913:
908:
907:
903:
898:
897:
893:
890:Richards 1978
888:
887:
883:
878:
877:
873:
868:
867:
863:
858:
857:
853:
848:
847:
843:
838:
837:
833:
828:
827:
823:
818:
817:
813:
808:
807:
803:
798:
797:
793:
788:
787:
783:
780:Cheesm., 1952
778:
777:
773:
768:
767:
763:
760:(Smith, 1863)
758:
757:
753:
748:
747:
743:
738:
737:
733:
728:
727:
723:
718:
717:
713:
708:
707:
703:
698:
697:
693:
688:
687:
683:
678:
677:
673:
668:
667:
663:
660:(Smith, 1860)
658:
657:
653:
648:
647:
643:
638:
637:
633:
628:
627:
623:
618:
617:
613:
608:
607:
603:
598:
597:
593:
590:Polašek, 2024
588:
587:
583:
578:
577:
573:
568:
567:
563:
558:
557:
553:
548:
547:
543:
538:
537:
533:
530:Cheesm., 1952
528:
527:
523:
518:
517:
513:
508:
507:
503:
498:
497:
493:
490:(Meade-Waldo)
488:
487:
483:
478:
477:
473:
468:
467:
463:
458:
457:
453:
448:
447:
443:
440:(Smith, 1865)
438:
437:
433:
428:
427:
423:
418:
417:
413:
408:
407:
403:
398:
397:
393:
388:
387:
383:
378:
377:
373:
368:
367:
363:
358:
357:
353:
348:
347:
343:
338:
337:
333:
328:
327:
323:
318:
317:
313:
308:
307:
303:
302:
295:
293:
289:
285:
276:
274:
273:
268:
267:
262:
258:
254:
250:
246:
243:
239:
236:paper wasps (
235:
231:
230:
222:
221:
216:
210:
206:
202:
196:
195:
190:
183:
182:
177:
169:
165:
160:
155:
150:
149:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
98:
95:
92:
91:
88:
85:
82:
81:
78:
75:
72:
71:
68:
65:
62:
61:
56:
51:
47:
44:
43:
38:
34:
29:
26:
22:
19:
2478:
2446:
2442:
2432:
2415:
2411:
2407:
2401:
2392:
2386:
2367:
2363:
2357:
2349:
2332:
2323:
2314:
2294:
2284:
2274:
2264:
2254:
2244:
2234:
2224:
2214:
2204:
2194:
2184:
2174:
2164:
2154:
2144:
2134:
2124:
2114:
2104:
2094:
2084:
2074:
2064:
2054:
2044:
2034:
2024:
2014:
2004:
1994:
1984:
1980:Kojima, 1998
1974:
1964:
1954:
1944:
1934:
1924:
1914:
1904:
1894:
1884:
1874:
1864:
1860:Kojima, 1996
1854:
1844:
1840:Kojima, 2001
1834:
1824:
1814:
1804:
1794:
1784:
1774:
1764:
1754:
1744:
1734:
1724:
1720:Kojima, 1985
1714:
1704:
1694:
1684:
1674:
1664:
1654:
1644:
1634:
1624:
1614:
1604:
1594:
1584:
1574:
1564:
1554:
1550:(Smith 1859)
1544:
1534:
1524:
1514:
1504:
1494:
1484:
1474:
1464:
1454:
1444:
1434:
1424:
1414:
1404:
1394:
1390:Guerin, 1831
1384:
1380:(Rad., 1881)
1374:
1364:
1360:Kojima, 1996
1354:
1344:
1334:
1324:
1314:
1304:
1294:
1290:Guill., 1841
1284:
1274:
1264:
1254:
1244:
1234:
1224:
1214:
1210:(Smith 1871)
1204:
1194:
1184:
1180:(Kohl, 1906)
1174:
1164:
1154:
1144:
1134:
1124:
1114:
1104:
1094:
1084:
1074:
1064:
1054:
1044:
1034:
1024:
1014:
1004:
994:
984:
974:
970:Kojima, 1998
964:
954:
944:
934:
924:
914:
904:
894:
884:
874:
864:
854:
844:
834:
824:
814:
804:
794:
784:
774:
770:(Smith 1858)
764:
754:
744:
734:
724:
720:Kojima, 1989
714:
704:
694:
684:
674:
664:
654:
650:Vecht, 1941
644:
634:
624:
614:
604:
594:
584:
574:
564:
554:
544:
534:
524:
520:(Smith 1859)
514:
504:
494:
484:
474:
470:Vecht, 1941
464:
454:
450:Vecht, 1962
444:
434:
424:
414:
404:
394:
384:
374:
364:
354:
344:
334:
324:
314:
304:
283:
282:
270:
264:
260:
257:Australasian
249:Afrotropical
228:
227:
226:
218:
208:
204:
192:
179:
167:
147:
146:
40:
24:
18:
2596:iNaturalist
2503:Wikispecies
2190:Vecht, 1962
2160:Vecht, 1962
2140:Sonan 1935
2090:Vecht, 1962
1690:Vecht, 1962
1680:Vecht, 1962
1650:Vecht, 1941
1560:Vecht, 1962
1540:Vecht, 1962
1410:Vecht, 1962
1400:Vecht, 1941
1310:Vecht, 1962
1280:Vecht, 1962
1220:Vecht, 1962
1170:Vecht, 1941
1120:Sonan, 1938
1070:Vecht, 1941
950:Vecht, 1962
910:Vecht, 1962
800:Vecht, 1941
700:Vecht, 1941
670:Smith, 1863
630:Kojima 1996
580:Vecht, 1941
480:Vecht, 1962
460:Vecht, 1996
279:Description
253:Indomalayan
245:Ropalidiini
137:Ropalidiini
123:Subfamily:
107:Hymenoptera
2670:Categories
2341:References
261:Ropalidia
238:Polistinae
209:ornaticeps
127:Polistinae
87:Arthropoda
2523:Ropalidia
2509:Ropalidia
2479:Ropalidia
2333:Ropalidia
2324:Ropalidia
2315:Ropalidia
2300:(Cameron)
2050:(Cameron)
1730:(Cameron)
1500:Carl 1934
292:metasomal
284:Ropalidia
240:) in the
229:Ropalidia
205:Ropalidia
148:Ropalidia
73:Kingdom:
67:Eukaryota
25:Ropalidia
2676:Vespidae
2494:Q1956829
2488:Wikidata
288:pronotum
234:eusocial
162:Species
117:Vespidae
113:Family:
83:Phylum:
77:Animalia
63:Domain:
2640:1036418
2614:1302531
2588:1311012
298:Species
184:on nest
143:Genus:
133:Tribe:
103:Order:
97:Insecta
93:Class:
2601:120832
2575:1ROPAG
223:female
2648:Plazi
2627:91393
2609:IRMNG
2562:31290
2549:63RV3
242:tribe
207:c.f.
2622:NCBI
2583:GBIF
2570:EPPO
2536:5364
2531:BOLD
255:and
156:1831
2557:EoL
2544:CoL
2518:AFD
2451:doi
2420:doi
2372:doi
2368:101
2672::
2650::
2637::
2624::
2611::
2598::
2585::
2572::
2559::
2546::
2533::
2520::
2505::
2490::
2445:.
2441:.
2414:.
2366:.
2362:.
251:,
2457:.
2453::
2447:6
2426:.
2422::
2416:5
2380:.
2374::
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
