95:
275:
2598:
49:
2586:
2610:
2574:
701:
Gerardo Lamas (2008) Systematics of butterflies (Lepidoptera: Hesperioidea and
Papilionoidea) in the world: current state and future perspectives (in Spanish). In: Jorge Llorente-Bousquets and AnalĂa Lanteri (eds.)
510:
Of the subfamilies of
Nymphalidae, only the Morphinae and Satyrinae are possibly paraphyletic, but these two subfamilies form a strongly supported clade with the Charaxinae as sister group.
559:
deposits dating roughly between 50 and 25 million years ago – suggest the radiation of the
Papilionoidea into the present-day families took place during that epoch.
555:, yet another prehistoric genus, also seems to belong here, but its relationships are quite obscure, indeed. However, these fossils – all found in
2869:
2813:
551:
2761:
2787:
734:
2792:
514:
2864:
565:, from the end of the Eocene, can be quite robustly assigned to the Nymphalidae, and is quite likely a member of the
2818:
2740:
2753:
2551:
2800:
17:
94:
2673:
2546:
2564:
30:
This article is about a group of butterflies. For the group of flowers that look like butterflies, see
727:
312:
The members of the
Papilionoidea may be distinguished by the following combination of characters:
2664:
2831:
2854:
2626:
413:
360:) are true butterflies that should be included within the Papilionoidea superfamily to reflect
278:
2826:
2123:
382:
2874:
2859:
2722:
993:
985:
976:
720:
8:
2139:
1914:
1906:
679:
654:
630:
605:
89:
606:"Cretaceous origin and repeated tertiary diversification of the redefined butterflies"
573:
fossils of
Papilionoidea are usually assignable to an extant family without problems.
2709:
2678:
2590:
1484:
684:
635:
323:
73:
2465:
1956:
1515:
674:
666:
655:"Phylogenomics provides strong evidence for relationships of butterflies and moths"
625:
617:
80:
576:
Taken together, these fossils place the origin of the
Papilionoidea in the latest
549:, is more firmly assigned to the Lycaenidae, but might belong to the Nymphalidae.
2714:
2034:
1710:
1587:
1463:
1455:
1421:
1395:
1387:
1054:
1025:
1019:
1010:
821:
815:
803:
66:
58:
2805:
2748:
2735:
2614:
2602:
2578:
2255:
2042:
1720:
1645:
1611:
1569:
1494:
1479:
1122:
531:
289:
31:
712:
2848:
2649:
2483:
2408:
2316:
2263:
2242:
2234:
1985:
1977:
1697:
1579:
1574:
1564:
1343:
1335:
1322:
1314:
1285:
1246:
1114:
1059:
1046:
1037:
962:
954:
945:
907:
901:
889:
536:
495:
274:
169:
543:. It is variously placed in the Lycaenidae or Riodinidae. A similar fossil,
2597:
2502:
2470:
2452:
2400:
2332:
2292:
2092:
2084:
2050:
1935:
1927:
1748:
1738:
1725:
1679:
1671:
1658:
1650:
1637:
1595:
1556:
1546:
1523:
1502:
1471:
1434:
1426:
1413:
1293:
1241:
787:
688:
670:
639:
621:
502:
390:
254:
48:
2774:
2688:
2658:
2460:
2418:
2342:
2337:
2308:
2276:
2171:
2112:
1948:
1872:
1864:
1851:
1843:
1814:
1764:
1689:
1603:
1538:
1533:
1528:
1510:
1301:
1236:
1164:
1143:
1135:
1085:
864:
858:
846:
744:
525:
489:
450:
438:
424:
417:
376:
353:
330:
249:
239:
208:
156:
146:
603:
2526:
2434:
2426:
2379:
2363:
2350:
2324:
2300:
2284:
2205:
2192:
2184:
2163:
2155:
2071:
2063:
2006:
1998:
1964:
1822:
1756:
1743:
1715:
1702:
1624:
1251:
1193:
1098:
566:
545:
483:
477:
470:
444:
408:
402:
361:
306:
282:
264:
244:
2585:
2779:
2478:
2384:
2221:
2213:
2131:
1806:
1793:
1777:
1629:
1619:
1405:
1272:
1264:
1220:
1106:
775:
763:
604:
Heikkilä, M.; Kaila, L.; Mutanen, M.; Peña, C.; Wahlberg, N. (2012).
584:, while the extant families emerged around the early Eocene onwards.
581:
570:
540:
463:
456:
357:
349:
348:
analyses suggest the traditionally circumscribed
Papilionoidea are a
345:
302:
223:
126:
106:
2766:
2701:
2620:
2696:
2643:
2609:
2371:
2147:
1893:
1885:
1830:
1785:
1442:
1360:
1228:
1093:
1076:
929:
781:
577:
561:
431:
396:
386:
259:
195:
182:
2392:
2024:
2016:
1733:
297:
2442:
769:
757:
704:
Contribuiciones taxonĂłmicas en ordens de insectos hiperdiversos
652:
556:
520:
136:
116:
2573:
2727:
2166:(gossamer-winged butterflies: blues, coppers and relatives)
334:
2387:(daggers, sallows, owlet moths, quakers, cutworms, darts)
515:
2374:(underwing, tiger, tussock, litter, snout, owlet moths)
659:
Proceedings of the Royal
Society B: Biological Sciences
610:
Proceedings of the Royal
Society B: Biological Sciences
372:
The six well-supported families of Papilionoidea are:
326:
are straight and clubbed or hooked as in the skippers.
2562:
1825:(castniid moths: giant butterfly-moths, sun moths)
599:
597:
401:Blues and coppers or gossamer-winged butterflies,
2846:
900:
356:) and Neotropical moth-like butterflies (family
814:
742:
594:
1018:
857:
953:
728:
887:
420:which contain the following 13 subfamilies:
367:
2174:(brush-footed, or four-footed butterflies)
735:
721:
653:Kawahara, A. Y.; Breinholt, J. W. (2014).
340:The pupae are angular rather than rounded.
47:
678:
629:
2353:(hawk moths, sphinx moths and hornworms)
801:
459:(including the owls as tribe Brassolini)
301:, meaning "butterfly") contains all the
273:
2150:(whites, yellows, orangetips, sulphurs)
316:The body is smaller and less moth-like.
14:
2847:
2266:(eggars, snout moths, or lappet moths)
844:
2625:
2624:
927:
926:
886:
843:
800:
716:
535:, which has long been mistaken for a
2870:Taxa named by Pierre André Latreille
2754:d15819d0-778c-445c-9971-44ce6cdd650e
24:
1788:(carpenter millers, or goat moths)
1705:(burnet, forester, or smoky moths)
25:
2886:
1109:(Amazonian primitive ghost moths)
427:(formerly the family Libytheidae)
352:group, and that skippers (family
2608:
2596:
2584:
2572:
333:do not spin cocoons in which to
93:
1125:(African primitive ghost moths)
706:. Mexico City: UNAM. Pp. 57-70.
473:(preponas and leaf butterflies)
466:(formerly the family Satyridae)
1429:(Old World spiny-winged moths)
695:
646:
529:is apparently a Papilionoidea
434:(formerly the family Danaidae)
13:
1:
2327:(Kentish glory and relatives)
1146:(New Zealand primitive moths)
587:
2473:(oriental swallowtail moths)
1751:(planthopper parasite moths)
7:
2547:Taxonomy of the Lepidoptera
2134:(American moth-butterflies)
2095:(Old World butterfly-moths)
2074:(picture-winged leaf moths)
824:(mandibulate archaic moths)
10:
2891:
2045:(tropical fruitworm moths)
1582:(case-bearers, case moths)
1346:(trumpet leaf miner moths)
1101:(swift moths, ghost moths)
512:
501:The tropical longwings or
29:
27:Superfamily of butterflies
2865:Lepidoptera superfamilies
2633:
2542:
2523:
2495:
2451:
2417:
2362:
2295:(American silkworm moths)
2275:
2254:
2233:
2204:
2183:
2126:(swallowtail butterflies)
2104:
2083:
2062:
2033:
2015:
1997:
1976:
1947:
1926:
1905:
1884:
1863:
1842:
1805:
1796:(dudgeon carpenter moths)
1776:
1688:
1670:
1555:
1505:(false diamondback moths)
1493:
1454:
1416:(burrowing webworm moths)
1404:
1386:
1382:
1359:
1334:
1313:
1284:
1263:
1219:
1192:
1163:
1134:
1084:
1075:
1045:
1036:
1009:
984:
975:
944:
940:
936:
922:
896:
882:
853:
839:
810:
796:
752:
498:(especially the adelphas)
423:The snout butterflies or
395:Whites or yellow-whites,
368:Families of Papilionoidea
305:except for the moth-like
236:
231:
90:Scientific classification
88:
55:
46:
41:
2287:(Australian lappet moth)
1296:(pigmy, or midget moths)
455:The morphos and owls or
414:Brush-footed butterflies
1117:(miniature ghost moths)
407:Metalmark butterflies,
2496:Superfamily unassigned
1917:(bristle-legged moths)
1466:(ribbed cocoon makers)
1275:(Andean endemic moths)
1231:(fairy longhorn moths)
671:10.1098/rspb.2014.0970
622:10.1098/rspb.2011.1430
285:
279:Brush-footed butterfly
2827:Paleobiology Database
2403:(prominents, kittens)
1938:(fringe-tufted moths)
994:Acanthopteroctetidae
986:Acanthopteroctetoidea
319:The wings are larger.
277:
2749:Fauna Europaea (new)
1915:Schreckensteiniidae
1907:Schreckensteinioidea
1896:(false burnet moths)
1767:(slug, or cup moths)
1325:(Gondwanaland moths)
1304:(white eyecap moths)
1062:(archaic bell moths)
2245:(sack bearer moths)
1967:(false plume moths)
1959:(many-plumed moths)
1817:(little bear moths)
1759:(slug caterpillars)
1614:(long-horned moths)
1598:(grass-miner moths)
996:(archaic sun moths)
616:(1731): 1093–1099.
665:(1788): 20140970.
443:The glasswings or
286:
2842:
2841:
2627:Taxon identifiers
2560:
2559:
2538:
2537:
2534:
2533:
2519:
2518:
2515:
2514:
2511:
2510:
2487:
2474:
2438:
2430:
2404:
2396:
2388:
2375:
2354:
2346:
2328:
2320:
2312:
2304:
2296:
2288:
2267:
2246:
2225:
2217:
2196:
2175:
2167:
2159:
2151:
2143:
2135:
2127:
2117:
2096:
2075:
2054:
2053:(fruitworm moths)
2046:
1989:
1968:
1960:
1939:
1918:
1897:
1876:
1855:
1854:(metalmark moths)
1834:
1833:(clearwing moths)
1826:
1818:
1797:
1789:
1768:
1760:
1752:
1729:
1706:
1662:
1654:
1641:
1640:(concealer moths)
1633:
1615:
1607:
1599:
1591:
1583:
1542:
1519:
1516:Glyphipterigidae
1506:
1485:Roeslerstammiidae
1475:
1467:
1446:
1438:
1430:
1417:
1355:
1354:
1347:
1326:
1305:
1297:
1276:
1255:
1232:
1159:
1158:
1155:
1154:
1147:
1126:
1118:
1110:
1102:
1071:
1070:
1063:
1032:
1031:
1005:
1004:
997:
971:
970:
918:
917:
914:
913:
902:Heterobathmioidea
878:
877:
874:
873:
868:
835:
834:
831:
830:
825:
272:
271:
227:
74:Myscelia cyaniris
16:(Redirected from
2882:
2835:
2834:
2822:
2821:
2809:
2808:
2806:NHMSYS0000503590
2796:
2795:
2783:
2782:
2770:
2769:
2757:
2756:
2744:
2743:
2731:
2730:
2718:
2717:
2705:
2704:
2692:
2691:
2682:
2681:
2669:
2668:
2667:
2654:
2653:
2652:
2622:
2621:
2613:
2612:
2601:
2600:
2589:
2588:
2577:
2576:
2568:
2486:(geometer moths)
2485:
2472:
2466:Pseudobistonidae
2436:
2428:
2402:
2394:
2386:
2373:
2352:
2344:
2326:
2318:
2310:
2302:
2294:
2286:
2265:
2244:
2223:
2215:
2194:
2173:
2165:
2157:
2149:
2141:
2133:
2125:
2116:
2109:
2094:
2073:
2052:
2044:
1987:
1966:
1958:
1937:
1916:
1895:
1874:
1853:
1832:
1824:
1816:
1795:
1787:
1766:
1758:
1750:
1727:
1704:
1660:
1652:
1639:
1631:
1613:
1605:
1597:
1589:
1588:Cosmopterigidae
1581:
1540:
1517:
1504:
1473:
1465:
1464:Bucculatricidae
1444:
1436:
1428:
1415:
1384:
1383:
1380:
1379:
1345:
1324:
1303:
1295:
1274:
1253:
1230:
1217:
1216:
1190:
1189:
1145:
1124:
1116:
1108:
1100:
1082:
1081:
1061:
1043:
1042:
1016:
1015:
995:
982:
981:
951:
950:
942:
941:
938:
937:
924:
923:
908:Heterobathmiidae
898:
897:
884:
883:
866:
855:
854:
841:
840:
823:
822:Micropterigidae
816:Micropterigoidea
812:
811:
798:
797:
737:
730:
723:
714:
713:
707:
699:
693:
692:
682:
650:
644:
643:
633:
601:
295:(from the genus
222:
98:
97:
81:Episcada apuleia
51:
39:
38:
21:
2890:
2889:
2885:
2884:
2883:
2881:
2880:
2879:
2845:
2844:
2843:
2838:
2830:
2825:
2817:
2812:
2804:
2799:
2791:
2786:
2778:
2773:
2765:
2760:
2752:
2747:
2739:
2734:
2726:
2721:
2713:
2708:
2700:
2695:
2687:
2685:
2677:
2672:
2663:
2662:
2657:
2648:
2647:
2642:
2629:
2619:
2607:
2595:
2583:
2571:
2563:
2561:
2556:
2552:Lists by region
2530:
2529:is not a clade.
2525:Note: division
2507:
2491:
2447:
2413:
2358:
2319:(Dryandra moth)
2311:(Brahmin moths)
2271:
2250:
2229:
2200:
2179:
2110:
2108:
2100:
2079:
2058:
2043:Copromorphidae
2035:Copromorphoidea
2029:
2011:
1993:
1972:
1943:
1922:
1901:
1880:
1875:(tortrix moths)
1859:
1838:
1801:
1772:
1728:(flannel moths)
1711:Himantopteridae
1684:
1666:
1612:Lecithoceridae
1606:(twirler moths)
1551:
1489:
1474:(Douglas moths)
1456:Gracillarioidea
1450:
1437:(bagworm moths)
1422:Arrhenophanidae
1400:
1388:Simaethistoidea
1374:
1372:
1370:
1368:
1366:
1364:
1362:
1351:
1330:
1309:
1280:
1259:
1211:
1209:
1207:
1205:
1203:
1201:
1199:
1197:
1195:
1184:
1182:
1180:
1178:
1176:
1174:
1172:
1170:
1168:
1166:
1151:
1144:Mnesarchaeidae
1136:Mnesarchaeoidea
1130:
1123:Prototheoridae
1067:
1055:Aenigmatineidae
1028:
1020:Lophocoronoidea
1001:
967:
932:
910:
892:
890:Heterobathmiina
870:
865:Agathiphagidae
859:Agathiphagoidea
849:
827:
806:
792:
748:
741:
711:
710:
700:
696:
651:
647:
602:
595:
590:
517:
430:The danaids or
370:
364:relationships.
221:
92:
77:
70:
67:Calinaga buddha
62:
59:Delias eucharis
35:
28:
23:
22:
15:
12:
11:
5:
2888:
2878:
2877:
2872:
2867:
2862:
2857:
2840:
2839:
2837:
2836:
2823:
2810:
2797:
2784:
2771:
2758:
2745:
2736:Fauna Europaea
2732:
2719:
2706:
2693:
2683:
2670:
2655:
2639:
2637:
2631:
2630:
2618:
2617:
2605:
2593:
2581:
2558:
2557:
2555:
2554:
2549:
2543:
2540:
2539:
2536:
2535:
2532:
2531:
2524:
2521:
2520:
2517:
2516:
2513:
2512:
2509:
2508:
2506:
2505:
2499:
2497:
2493:
2492:
2490:
2489:
2481:
2476:
2468:
2463:
2457:
2455:
2449:
2448:
2446:
2445:
2440:
2432:
2423:
2421:
2415:
2414:
2412:
2411:
2406:
2398:
2390:
2382:
2377:
2368:
2366:
2360:
2359:
2357:
2356:
2348:
2340:
2335:
2330:
2322:
2314:
2306:
2298:
2290:
2281:
2279:
2273:
2272:
2270:
2269:
2264:Lasiocampidae
2260:
2258:
2256:Lasiocampoidea
2252:
2251:
2249:
2248:
2239:
2237:
2231:
2230:
2228:
2227:
2219:
2210:
2208:
2202:
2201:
2199:
2198:
2189:
2187:
2181:
2180:
2178:
2177:
2169:
2161:
2153:
2145:
2137:
2129:
2120:
2118:
2102:
2101:
2099:
2098:
2089:
2087:
2081:
2080:
2078:
2077:
2068:
2066:
2060:
2059:
2057:
2056:
2048:
2039:
2037:
2031:
2030:
2028:
2027:
2021:
2019:
2013:
2012:
2010:
2009:
2003:
2001:
1999:Whalleyanoidea
1995:
1994:
1992:
1991:
1986:Pterophoridae
1982:
1980:
1978:Pterophoroidea
1974:
1973:
1971:
1970:
1962:
1953:
1951:
1945:
1944:
1942:
1941:
1932:
1930:
1924:
1923:
1921:
1920:
1911:
1909:
1903:
1902:
1900:
1899:
1890:
1888:
1882:
1881:
1879:
1878:
1869:
1867:
1861:
1860:
1858:
1857:
1848:
1846:
1840:
1839:
1837:
1836:
1828:
1820:
1811:
1809:
1803:
1802:
1800:
1799:
1791:
1782:
1780:
1774:
1773:
1771:
1770:
1762:
1754:
1746:
1741:
1736:
1731:
1726:Megalopygidae
1723:
1721:Somabrachyidae
1718:
1713:
1708:
1700:
1694:
1692:
1686:
1685:
1683:
1682:
1676:
1674:
1668:
1667:
1665:
1664:
1661:(timber moths)
1656:
1653:(flower moths)
1648:
1646:Pterolonchidae
1643:
1635:
1632:(mompha moths)
1627:
1622:
1617:
1609:
1601:
1593:
1590:(cosmet moths)
1585:
1580:Coleophoridae
1577:
1572:
1570:Batrachedridae
1567:
1561:
1559:
1553:
1552:
1550:
1549:
1544:
1541:(ermine moths)
1539:Yponomeutidae
1536:
1531:
1526:
1521:
1513:
1508:
1499:
1497:
1495:Yponomeutoidea
1491:
1490:
1488:
1487:
1482:
1480:Gracillariidae
1477:
1469:
1460:
1458:
1452:
1451:
1449:
1448:
1445:(fungus moths)
1440:
1432:
1424:
1419:
1410:
1408:
1402:
1401:
1399:
1398:
1396:Simaethistidae
1392:
1390:
1377:
1357:
1356:
1353:
1352:
1350:
1349:
1340:
1338:
1332:
1331:
1329:
1328:
1323:Palaephatidae
1319:
1317:
1315:Palaephatoidea
1311:
1310:
1308:
1307:
1299:
1290:
1288:
1282:
1281:
1279:
1278:
1269:
1267:
1261:
1260:
1258:
1257:
1249:
1244:
1239:
1234:
1225:
1223:
1214:
1187:
1161:
1160:
1157:
1156:
1153:
1152:
1150:
1149:
1140:
1138:
1132:
1131:
1129:
1128:
1120:
1115:Palaeosetidae
1112:
1104:
1096:
1090:
1088:
1079:
1073:
1072:
1069:
1068:
1066:
1065:
1060:Neopseustidae
1057:
1051:
1049:
1047:Neopseustoidea
1040:
1034:
1033:
1030:
1029:
1026:Lophocoronidae
1024:
1022:
1013:
1007:
1006:
1003:
1002:
1000:
999:
990:
988:
979:
973:
972:
969:
968:
966:
965:
959:
957:
955:Eriocranioidea
948:
934:
933:
920:
919:
916:
915:
912:
911:
906:
904:
894:
893:
880:
879:
876:
875:
872:
871:
863:
861:
851:
850:
837:
836:
833:
832:
829:
828:
820:
818:
808:
807:
794:
793:
791:
790:
784:
778:
772:
766:
760:
753:
750:
749:
740:
739:
732:
725:
717:
709:
708:
694:
645:
592:
591:
589:
586:
532:incertae sedis
508:
507:
506:
505:
499:
492:
488:The nymphs or
486:
480:
474:
467:
462:The browns or
460:
453:
447:
441:
435:
428:
411:
405:
399:
393:
380:
369:
366:
342:
341:
338:
327:
320:
317:
270:
269:
268:
267:
262:
257:
252:
247:
242:
234:
233:
229:
228:
216:
212:
211:
206:
199:
198:
193:
186:
185:
180:
173:
172:
167:
160:
159:
154:
150:
149:
144:
140:
139:
134:
130:
129:
124:
120:
119:
114:
110:
109:
104:
100:
99:
86:
85:
53:
52:
44:
43:
42:Papilionoidea
32:Papilionoideae
26:
9:
6:
4:
3:
2:
2887:
2876:
2873:
2871:
2868:
2866:
2863:
2861:
2858:
2856:
2855:Papilionoidea
2853:
2852:
2850:
2833:
2828:
2824:
2820:
2815:
2811:
2807:
2802:
2798:
2794:
2789:
2785:
2781:
2776:
2772:
2768:
2763:
2759:
2755:
2750:
2746:
2742:
2737:
2733:
2729:
2724:
2720:
2716:
2711:
2707:
2703:
2698:
2694:
2690:
2684:
2680:
2679:Papilionoidea
2675:
2671:
2666:
2665:Papilionoidea
2660:
2656:
2651:
2645:
2641:
2640:
2638:
2636:
2635:Papilionoidea
2632:
2628:
2623:
2616:
2611:
2606:
2604:
2599:
2594:
2592:
2587:
2582:
2580:
2575:
2570:
2569:
2566:
2553:
2550:
2548:
2545:
2544:
2541:
2528:
2522:
2504:
2501:
2500:
2498:
2494:
2488:
2482:
2480:
2477:
2475:
2471:Epicopeiidae
2469:
2467:
2464:
2462:
2459:
2458:
2456:
2454:
2450:
2444:
2441:
2439:
2433:
2431:
2425:
2424:
2422:
2420:
2416:
2410:
2409:Oenosandridae
2407:
2405:
2401:Notodontidae
2399:
2397:
2391:
2389:
2383:
2381:
2378:
2376:
2370:
2369:
2367:
2365:
2361:
2355:
2349:
2347:
2341:
2339:
2336:
2334:
2331:
2329:
2323:
2321:
2315:
2313:
2307:
2305:
2299:
2297:
2293:Apatelodidae
2291:
2289:
2283:
2282:
2280:
2278:
2274:
2268:
2262:
2261:
2259:
2257:
2253:
2247:
2243:Mimallonidae
2241:
2240:
2238:
2236:
2235:Mimallonoidea
2232:
2226:
2220:
2218:
2216:(snout moths)
2212:
2211:
2209:
2207:
2203:
2197:
2191:
2190:
2188:
2186:
2182:
2176:
2170:
2168:
2162:
2160:
2154:
2152:
2146:
2144:
2138:
2136:
2130:
2128:
2124:Papilionidae
2122:
2121:
2119:
2114:
2107:
2106:Papilionoidea
2103:
2097:
2093:Callidulidae
2091:
2090:
2088:
2086:
2085:Calliduloidea
2082:
2076:
2070:
2069:
2067:
2065:
2061:
2055:
2051:Carposinidae
2049:
2047:
2041:
2040:
2038:
2036:
2032:
2026:
2023:
2022:
2020:
2018:
2014:
2008:
2007:Whalleyanidae
2005:
2004:
2002:
2000:
1996:
1990:
1988:(plume moths)
1984:
1983:
1981:
1979:
1975:
1969:
1963:
1961:
1955:
1954:
1952:
1950:
1946:
1940:
1936:Epermeniidae
1934:
1933:
1931:
1929:
1928:Epermenioidea
1925:
1919:
1913:
1912:
1910:
1908:
1904:
1898:
1892:
1891:
1889:
1887:
1883:
1877:
1871:
1870:
1868:
1866:
1862:
1856:
1850:
1849:
1847:
1845:
1841:
1835:
1829:
1827:
1821:
1819:
1813:
1812:
1810:
1808:
1804:
1798:
1794:Dudgeoneidae
1792:
1790:
1784:
1783:
1781:
1779:
1775:
1769:
1763:
1761:
1755:
1753:
1749:Epipyropidae
1747:
1745:
1744:Cyclotornidae
1742:
1740:
1737:
1735:
1732:
1730:
1724:
1722:
1719:
1717:
1714:
1712:
1709:
1707:
1701:
1699:
1698:Heterogynidae
1696:
1695:
1693:
1691:
1687:
1681:
1678:
1677:
1675:
1673:
1672:Galacticoidea
1669:
1663:
1659:Xyloryctidae
1657:
1655:
1651:Scythrididae
1649:
1647:
1644:
1642:
1638:Oecophoridae
1636:
1634:
1628:
1626:
1625:Metachandidae
1623:
1621:
1618:
1616:
1610:
1608:
1602:
1600:
1596:Elachistidae
1594:
1592:
1586:
1584:
1578:
1576:
1575:Blastobasidae
1573:
1571:
1568:
1566:
1565:Autostichidae
1563:
1562:
1560:
1558:
1554:
1548:
1545:
1543:
1537:
1535:
1532:
1530:
1527:
1525:
1522:
1520:
1518:(sedge moths)
1514:
1512:
1509:
1507:
1503:Acrolepiidae
1501:
1500:
1498:
1496:
1492:
1486:
1483:
1481:
1478:
1476:
1472:Douglasiidae
1470:
1468:
1462:
1461:
1459:
1457:
1453:
1447:
1441:
1439:
1433:
1431:
1427:Eriocottidae
1425:
1423:
1420:
1418:
1414:Acrolophidae
1412:
1411:
1409:
1407:
1403:
1397:
1394:
1393:
1391:
1389:
1385:
1381:
1378:
1376:
1358:
1348:
1344:Tischeriidae
1342:
1341:
1339:
1337:
1336:Tischerioidea
1333:
1327:
1321:
1320:
1318:
1316:
1312:
1306:
1300:
1298:
1294:Nepticulidae
1292:
1291:
1289:
1287:
1286:Nepticuloidea
1283:
1277:
1273:Andesianidae
1271:
1270:
1268:
1266:
1265:Andesianoidea
1262:
1256:
1254:(yucca moths)
1250:
1248:
1247:Incurvariidae
1245:
1243:
1240:
1238:
1235:
1233:
1227:
1226:
1224:
1222:
1218:
1215:
1213:
1191:
1188:
1186:
1162:
1148:
1142:
1141:
1139:
1137:
1133:
1127:
1121:
1119:
1113:
1111:
1107:Neotheoridae
1105:
1103:
1097:
1095:
1092:
1091:
1089:
1087:
1083:
1080:
1078:
1074:
1064:
1058:
1056:
1053:
1052:
1050:
1048:
1044:
1041:
1039:
1035:
1027:
1023:
1021:
1017:
1014:
1012:
1011:Lophocoronina
1008:
998:
992:
991:
989:
987:
983:
980:
978:
977:Acanthoctesia
974:
964:
963:Eriocraniidae
961:
960:
958:
956:
952:
949:
947:
943:
939:
935:
931:
925:
921:
909:
905:
903:
899:
895:
891:
885:
881:
869:
867:(kauri moths)
862:
860:
856:
852:
848:
842:
838:
826:
819:
817:
813:
809:
805:
799:
795:
789:
785:
783:
779:
777:
773:
771:
767:
765:
761:
759:
755:
754:
751:
746:
738:
733:
731:
726:
724:
719:
718:
715:
705:
698:
690:
686:
681:
676:
672:
668:
664:
660:
656:
649:
641:
637:
632:
627:
623:
619:
615:
611:
607:
600:
598:
593:
585:
583:
579:
574:
572:
568:
564:
563:
558:
554:
553:
548:
547:
542:
538:
537:geometer moth
534:
533:
528:
527:
522:
516:
511:
504:
500:
497:
496:Limenitidinae
493:
491:
487:
485:
481:
479:
475:
472:
468:
465:
461:
458:
454:
452:
448:
446:
442:
440:
436:
433:
429:
426:
422:
421:
419:
415:
412:
410:
406:
404:
400:
398:
394:
392:
388:
384:
381:
378:
375:
374:
373:
365:
363:
359:
355:
351:
347:
339:
336:
332:
328:
325:
321:
318:
315:
314:
313:
310:
308:
304:
300:
299:
294:
293:Papilionoidea
291:
284:
281:of subfamily
280:
276:
266:
263:
261:
258:
256:
253:
251:
248:
246:
243:
241:
238:
237:
235:
230:
225:
220:
219:Papilionoidea
217:
215:Superfamily:
214:
213:
210:
207:
204:
201:
200:
197:
194:
191:
188:
187:
184:
181:
178:
175:
174:
171:
170:Eulepidoptera
168:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
96:
91:
87:
84:
83:
82:
78:Below right:
76:
75:
69:
68:
61:
60:
54:
50:
45:
40:
37:
33:
19:
2634:
2503:Millieriidae
2484:Geometridae
2453:Geometroidea
2437:(gold moths)
2395:(tuft moths)
2345:(saturniids)
2343:Saturniidae
2333:Eupterotidae
2317:Carthaeidae
2309:Brahmaeidae
2303:(silk moths)
2224:(grass moth)
2195:(teak moths)
2172:Nymphalidae
2158:(metalmarks)
2140:Hesperiidae
2105:
1873:Tortricidae
1865:Tortricoidea
1852:Choreutidae
1844:Choreutoidea
1815:Brachodidae
1765:Limacodidae
1739:Anomoeotidae
1680:Galacticidae
1604:Gelechiidae
1557:Gelechioidea
1547:Ypsolophidae
1524:Heliodinidae
1302:Opostegidae
1242:Heliozelidae
1094:Anomosetidae
1038:Neopseustina
788:Holometabola
786:Superorder:
780:Infraclass:
703:
697:
662:
658:
648:
613:
609:
575:
560:
550:
544:
530:
524:
518:
509:
503:Heliconiinae
391:Papilionidae
383:Swallowtails
371:
350:paraphyletic
346:phylogenetic
343:
331:caterpillars
311:
296:
292:
287:
255:Papilionidae
218:
202:
189:
176:
163:
153:Infraorder:
79:
72:
71:Below left:
65:
63:
57:
36:
18:Papilionidea
2875:Obtectomera
2860:Butterflies
2775:iNaturalist
2659:Wikispecies
2461:Sematuridae
2435:Cimeliidae
2429:(hook-tips)
2427:Drepanidae
2419:Drepanoidea
2351:Sphingidae
2338:Phiditiidae
2325:Endromidae
2301:Bombycidae
2285:Anthelidae
2277:Bombycoidea
2193:Hyblaeidae
2185:Hyblaeoidea
2164:Lycaenidae
2156:Riodinidae
2113:butterflies
2072:Thyrididae
2064:Thyridoidea
1965:Tineodidae
1957:Alucitidae
1949:Alucitoidea
1823:Castniidae
1757:Dalceridae
1703:Zygaenidae
1690:Zygaenoidea
1534:Plutellidae
1529:Lyonetiidae
1511:Bedelliidae
1252:Prodoxidae
1237:Cecidosidae
1099:Hepialidae
1086:Hepialoidea
804:Zeugloptera
745:Lepidoptera
526:Lithopsyche
490:Nymphalinae
451:Calinaginae
439:Tellervinae
425:Libytheinae
418:Nymphalidae
377:Hesperiidae
354:Hesperiidae
303:butterflies
290:superfamily
250:Nymphalidae
240:Hesperiidae
209:Obtectomera
196:Apoditrysia
157:Heteroneura
147:Lepidoptera
64:Top right:
2849:Categories
2591:Arthropods
2527:Monotrysia
2385:Noctuidae
2380:Euteliidae
2364:Noctuoidea
2222:Crambidae
2214:Pyralidae
2206:Pyraloidea
2142:(skippers)
2132:Hedylidae
1716:Lacturidae
1630:Momphidae
1435:Psychidae
946:Dacnonypha
774:Subclass:
764:Arthropoda
588:References
567:Nymphalini
552:Riodinella
546:Lithodryas
513:See also:
484:Apaturinae
478:Biblidinae
471:Charaxinae
445:Ithomiinae
409:Riodinidae
403:Lycaenidae
379:(skippers)
307:Hedyloidea
283:Charaxinae
265:Riodinidae
245:Lycaenidae
127:Arthropoda
56:Top left:
2479:Uraniidae
2372:Erebidae
2148:Pieridae
1894:Urodidae
1886:Urodoidea
1831:Sesiidae
1807:Sesioidea
1786:Cossidae
1778:Cossoidea
1620:Lypusidae
1443:Tineidae
1406:Tineoidea
1229:Adelidae
1221:Adeloidea
928:Suborder
888:Suborder
847:Aglossata
845:Suborder
802:Suborder
776:Pterygota
756:Kingdom:
582:Paleogene
580:or early
571:Oligocene
541:Boarmiini
464:Satyrinae
457:Morphinae
387:birdwings
362:cladistic
358:Hedylidae
232:Families
224:Latreille
113:Kingdom:
107:Eukaryota
2697:BugGuide
2686:BioLib:
2644:Wikidata
2393:Nolidae
2017:Immoidea
1077:Exoporia
930:Glossata
782:Neoptera
762:Phylum:
758:Animalia
747:families
689:24966318
640:21920981
578:Mesozoic
562:Prodryas
432:Danainae
397:Pieridae
324:antennae
260:Pieridae
183:Ditrysia
123:Phylum:
117:Animalia
103:Domain:
2650:Q756153
2615:Biology
2603:Animals
2579:Insects
2565:Portals
2025:Immidae
1734:Aididae
770:Insecta
768:Class:
743:Extant
680:4083801
631:3267136
539:of the
344:Recent
298:Papilio
143:Order:
137:Insecta
133:Class:
2832:212998
2793:117235
2443:Doidae
687:
677:
638:
628:
557:Eocene
523:genus
521:fossil
335:pupate
226:, 1802
2819:37572
2780:47224
2689:17297
203:Clade
190:Clade
177:Clade
164:Clade
2814:NCBI
2788:ITIS
2767:1875
2762:GBIF
2741:6944
685:PMID
636:PMID
519:The
494:The
482:The
476:The
469:The
449:The
437:The
385:and
329:The
322:The
288:The
2801:NBN
2728:854
2723:EoL
2715:5G9
2710:CoL
2674:AFD
675:PMC
667:doi
663:281
626:PMC
618:doi
614:279
2851::
2829::
2816::
2803::
2790::
2777::
2764::
2751::
2738::
2725::
2712::
2702:81
2699::
2676::
2661::
2646::
683:.
673:.
661:.
657:.
634:.
624:.
612:.
608:.
596:^
569:.
416:,
389:,
309:.
205::
192::
179::
166::
2567::
2115:)
2111:(
1375:a
1373:i
1371:s
1369:y
1367:r
1365:t
1363:i
1361:D
1212:a
1210:i
1208:s
1206:y
1204:r
1202:t
1200:o
1198:n
1196:o
1194:M
1185:a
1183:r
1181:u
1179:e
1177:n
1175:o
1173:r
1171:e
1169:t
1167:e
1165:H
736:e
729:t
722:v
691:.
669::
642:.
620::
337:.
34:.
20:)
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