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approximately 20 vessels for 12 hours a day during the months May–September. Researchers discovered that these vessels are in the line of sight for these whales for 98–99.5% of daylight hours. With so many vessels, the air quality around these whales deteriorates and impacts their health. Air pollutants that bind with exhaust fumes are responsible for the activation of the cytochrome P450 1A gene family. Researchers have successfully identified this gene in skin biopsies of live whales and also the lungs of deceased whales. A direct correlation between activation of this gene and the air pollutants can not be made because there are other known factors that will induce the same gene. Vessels can have either wet or dry exhaust systems, with wet exhaust systems leaving more pollutants in the water due to various gas solubility. A modelling study determined that the lowest-observed-adverse-effect-level (LOAEL) of exhaust pollutants was about 12% of the human dose.
1861:. Dialects are composed of specific numbers and types of discrete, repetitive calls. They are complex and stable over time. Call patterns and structure are distinctive within matrilines. Newborns produce calls similar to their mothers, but have a more limited repertoire. Individuals likely learn their dialect through contact with pod members. Family-specific calls have been observed more frequently in the days following a calf's birth, which may help the calf learn them. Dialects are probably an important means of maintaining group identity and cohesiveness. Similarity in dialects likely reflects the degree of relatedness between pods, with variation growing over time. When pods meet, dominant call types decrease and subset call types increase. The use of both call types is called biphonation. The increased subset call types may be the distinguishing factor between pods and inter-pod relations.
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950:. The orca is mostly black but with sharply bordered white areas. The entire lower jaw is white and from here, the colouration stretches across the underside to the genital area; narrowing and expanding some, and extending into lateral flank patches close to the end. The tail fluke (fin) is also white on the underside, while the eyes have white oval-shaped patches behind and above them, and a grey or white "saddle patch" exists behind the dorsal fin and across the back. Males and females also have different patterns of black and white skin in their genital areas. In newborns, the white areas are yellow or orange coloured. Antarctic orcas may have pale grey to nearly white backs. Some Antarctic orcas are brown and yellow due to
2504:. The captain of the vessel reported this was the second attack on a vessel under his command and the orcas focused on the rudders. "Looks like they knew exactly what they are doing. They didn't touch anything else." After an orca repeatedly rammed a vessel off the coast of Norway in 2023, there is a concern the behavior is spreading to other areas. This has led to recommendations that sailors now carry bags of sand. Dropping sand into the water near the rudder is thought to confuse the sonar signal. Experts were divided as to whether the behavior was some sort of revenge or protection response to a previous traumatic incident, or playful or frustrated attempts to get a boat's propeller to emit a stream of high-speed water.
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noises detected with underwater microphones were sonar. The sound originated from a U.S. Navy frigate 12 miles (19 kilometres) distant, Balcomb said." The impact of sonar waves on orcas is potentially life-threatening. Three years prior to
Balcomb's discovery, research in the Bahamas showed 14 beaked whales washed up on the shore. These whales were beached on the day U.S. Navy destroyers were activated into sonar exercise. Of the 14 whales beached, six of them died. These six dead whales were studied, and
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1938:, but have also overcome a variety of techniques designed to stop them, such as the use of unbaited lines as decoys. Once, fishermen placed their boats several miles apart, taking turns retrieving small amounts of their catch, in the hope that the whales would not have enough time to move between boats to steal the catch as it was being retrieved. The tactic worked initially, but the orcas figured it out quickly and split into groups.
939:(3.0 to 3.9 long tons; 3.3 to 4.4 short tons). Orcas may attain larger sizes as males have been recorded at 9.8 m (32 ft) and females at 8.5 m (28 ft). Large males can reach a weight of over 10 t (9.8 long tons; 11 short tons). Calves at birth weigh about 180 kg (400 lb) and are about 2.4 m (7 ft 10 in) long. The skeleton of the orca is typical for an oceanic dolphin, but more robust.
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signs of orca attack via tooth rake marks. Pods of female sperm whales sometimes protect themselves by forming a protective circle around their calves with their flukes facing outwards, using them to repel the attackers. There is also evidence that humpback whales will defend against or mob orcas who are attacking either humpback calves or juveniles as well as members of other species.
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of transient offspring from natal matrilines is common, with juveniles and adults of both sexes participating. Some males become "rovers" and do not form long-term associations, occasionally joining groups that contain reproductive females. As in resident clans, transient community members share an acoustic repertoire, although regional differences in vocalizations have been noted.
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2031:, once thought to occur only in captive orcas, was observed in wild populations by researchers off British Columbia on December 2, 2016. In this incident, an adult male killed the calf of a female within the same pod, with the adult male's mother also joining in the assault. It is theorized that the male killed the young calf in order to mate with its mother (something that
993:. The snout is blunt and lacks the beak of other species. The orca's teeth are very strong, and its jaws exert a powerful grip; the upper teeth fall into the gaps between the lower teeth when the mouth is closed. The firm middle and back teeth hold prey in place, while the front teeth are inclined slightly forward and outward to protect them from powerful jerking movements.
1263:: the orcas force the herring into a tight ball by releasing bursts of bubbles or flashing their white undersides. They then slap the ball with their tail flukes, stunning or killing up to 15 fish at a time, then eating them one by one. Carousel feeding has been documented only in the Norwegian orca population, as well as some oceanic dolphin species.
1838:, orcas depend heavily on underwater sound for orientation, feeding, and communication. They produce three categories of sounds: clicks, whistles, and pulsed calls. Clicks are believed to be used primarily for navigation and discriminating prey and other objects in the surrounding environment, but are also commonly heard during social interactions.
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1218:. Different populations or ecotypes may specialize, and some can have a dramatic impact on prey species. However, whales in tropical areas appear to have more generalized diets due to lower food productivity. Orcas spend most of their time at shallow depths, but occasionally dive several hundred metres depending on their prey.
866:"residents", mammal-eating "transients" and "offshores". Other populations have not been as well studied, although specialized fish and mammal eating orcas have been distinguished elsewhere. Mammal-eating orcas in different regions were long thought likely to be closely related, but genetic testing has refuted this hypothesis.
1358:. While hunting these species, orcas usually have to chase them to exhaustion. For highly social species, orca pods try to separate an individual from its group. Larger groups have a better chance of preventing their prey from escaping, which is killed by being thrown around, rammed and jumped on. Arctic orcas may attack
1318:. Competition between orcas and white sharks is probable in regions where their diets overlap. The arrival of orcas in an area can cause white sharks to flee and forage elsewhere. Orcas appear to target the liver of sharks. In one case a single orca was observed killing and eating a great white shark on its own.
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Bigg's techniques also revealed the
Pacific Northwest population was in the low hundreds rather than the thousands that had been previously assumed. The southern resident community alone had lost 48 of its members to captivity; by 1976, only 80 remained. In the Pacific Northwest, the species that had
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Transient pods are smaller than resident pods, typically consisting of an adult female and one or two of her offspring. Males typically maintain stronger relationships with their mothers than other females. These bonds can extend well into adulthood. Unlike residents, extended or permanent separation
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were standing. The sled dogs' barking is speculated to have sounded enough like seal calls to trigger the orca's hunting curiosity. In the 1970s, a surfer in
California was bitten, but the Orca then retreated, and in 2005, a boy in Alaska who was splashing in a region frequented by harbour seals was
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Worldwide population estimates are uncertain, but recent consensus suggests a minimum of 50,000 (2006). Local estimates include roughly 25,000 in the
Antarctic, 8,500 in the tropical Pacific, 2,250–2,700 off the cooler northeast Pacific and 500–1,500 off Norway. Japan's Fisheries Agency estimated in
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indicates the two
Antarctic groups (types B and C) should be recognized as distinct species, as should the North Pacific transients, leaving the others as subspecies pending additional data. A 2019 study of Type D orcas also found them to be distinct from other populations and possibly even a unique
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populations in the 1990s was controversially attributed by some scientists to orca predation, although with no direct evidence. The decline of sea otters followed a decline in seal populations, which in turn may be substitutes for their original prey, now decimated by industrial whaling. Orcas have
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are not a significant food item. Depletion of specific prey species in an area is, therefore, cause for concern for local populations, despite the high diversity of prey. On average, an orca eats 227 kilograms (500 lb) each day. While salmon are usually hunted by an individual whale or a small
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Orcas are the largest extant members of the dolphin family. Males typically range from 6 to 8 m (20 to 26 ft) long and weigh in excess of 6 t (5.9 long tons; 6.6 short tons). Females are smaller, generally ranging from 5 to 7 m (16 to 23 ft) and weighing about 3 to 4 t
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will be split into a number of different species or at least subspecies over the next few years." Although large variation in the ecological distinctiveness of different orca groups complicate simple differentiation into types, research off the west coast of North
America has identified fish-eating
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testing indicates resident males nearly always mate with females from other pods. Clans, the next level of resident social structure, are composed of pods with similar dialects, and common but older maternal heritage. Clan ranges overlap, mingling pods from different clans. The highest association
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abilities, detecting the location and characteristics of prey and other objects in the water by emitting clicks and listening for echoes, as do other members of the dolphin family. The mean body temperature of the orca is 36 to 38 °C (97 to 100 °F). Like most marine mammals, orcas have a
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region. Eleven members (about half) of one resident pod disappeared in the following year. The spill damaged salmon and other prey populations, which in turn damaged local orcas. By 2009, scientists estimated the AT1 transient population (considered part of a larger population of 346 transients),
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consisting of the eldest female (matriarch) and her sons and daughters, and the descendants of her daughters, etc. The average size of a matriline is 5.5 animals. Because females can reach age 90, as many as four generations travel together. These matrilineal groups are highly stable. Individuals
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has also been reported based on analysis of stomach contents, but this is likely to be the result of scavenging remains dumped by whalers. One orca was also attacked by its companions after being shot. Although resident orcas have never been observed to eat other marine mammals, they occasionally
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indicating the availability of prey to scavenge, and compressed air inflation of whale carcasses causing them to float, thus exposing them to scavenging. However, the devastation of great whale populations by unfettered whaling has possibly reduced their availability for orcas, and caused them to
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As a response to this, in 2017 boats off the
British Columbia coast now have a minimum approach distance of 200 metres compared to the previous 100 metres. This new rule complements Washington State's minimum approach zone of 180 metres that has been in effect since 2011. If a whale approaches a
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temporarily to grab prey before wriggling back to the sea. Beaching, usually fatal to cetaceans, is not an instinctive behaviour, and can require years of practice for the young. Orcas can then release the animal near juvenile whales, allowing the younger whales to practice the difficult capture
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off the south coast of
Western Australia, including an estimated 18–22-meter (59–72 ft) individual. Large whales require much effort and coordination to kill and orcas often target calves. A hunt begins with a chase followed by a violent attack on the exhausted prey. Large whales often show
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An individual orca can often be identified from its dorsal fin and saddle patch. Variations such as nicks, scratches, and tears on the dorsal fin and the pattern of white or grey in the saddle patch are unique. Published directories contain identifying photographs and names for hundreds of North
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continues to increase in popularity, but may have some problematic impacts on orcas. Exposure to exhaust gases from large amounts of vessel traffic is causing concern for the overall health of the 75 remaining southern resident orcas (SRKWs) left as of early 2019. This population is followed by
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near the Puget Sound coastline) noticed uncharacteristic behaviour displayed by the orcas. The whales seemed "agitated and were moving haphazardly, attempting to lift their heads free of the water" to escape the sound of the sonars. "Balcomb confirmed at the time that strange underwater pinging
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suggested no significant difference in survivorship between free-ranging and captive orcas. However, in the wild, females who survive infancy live 46 years on average, and up to 70–80 years in rare cases. Wild males who survive infancy live 31 years on average, and up to 50–60 years. Captivity
2823:. From 1976 to 1997, 55 whales were taken from the wild in Iceland, 19 from Japan, and three from Argentina. These figures exclude animals that died during capture. Live captures fell dramatically in the 1990s, and by 1999, about 40% of the 48 animals on display in the world were captive-born.
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Beginning around 2020, one or more pods of orcas began to attack sailing vessels off the southern tip of Europe, and a few were sunk. At least 15 interactions between orcas and boats off the
Iberian coast were reported in 2020. According to the Atlantic Orca Working Group (GTOA) as many as 500
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and
Washington. They do not breed outside of their community, which was once estimated at around 200 animals and later shrank to around 90. In October 2008, the annual survey revealed seven were missing and presumed dead, reducing the count to 83. This is potentially the largest decline in the
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Dialects also distinguish types. Resident dialects contain seven to 17 (mean = 11) distinctive call types. All members of the North American west coast transient community express the same basic dialect, although minor regional variation in call types is evident. Preliminary research indicates
1993:
was estimated by some researchers to have been as old as 105 years at the time of her death, though a biopsy sample indicated her age as 65 to 80 years. It is thought that orcas held in captivity tend to have shorter lives than those in the wild, although this is subject to scientific debate.
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in captivity and are often described as intelligent, although defining and measuring "intelligence" is difficult in a species whose environment and behavioural strategies are very different from those of humans. Orcas imitate others, and seem to deliberately teach skills to their kin. Off the
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Migration patterns are poorly understood. Each summer, the same individuals appear off the coasts of British Columbia and Washington. Despite decades of research, where these animals go for the rest of the year remains unknown. Transient pods have been sighted from southern Alaska to central
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The public's growing appreciation also led to growing opposition to whale–keeping in aquarium. Only one whale has been taken in North American waters since 1976. In recent years, the extent of the public's interest in orcas has manifested itself in several high-profile efforts surrounding
2249:. Many scientists involved in this lawsuit, including Bill Wareham, a marine scientist with the David Suzuki Foundation, noted increased boat traffic, water toxic wastes, and low salmon population as major threats, putting approximately 87 orcas on the British Columbia Coast in danger.
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varies from 15 to 18 months. Mothers usually calve a single offspring about once every five years. In resident pods, births occur at any time of year, although winter is the most common. Mortality is extremely high during the first seven months of life, when 37–50% of all calves die.
2664:, Washington. She became the first whale to be successfully reintegrated into a wild pod after human intervention, crystallizing decades of research into the vocal behaviour and social structure of the region's orcas. The saving of Springer raised hopes that another young orca named
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2855:. Wild orcas may travel up to 160 kilometres (100 mi) in a day, and critics say the animals are too big and intelligent to be suitable for captivity. Captives occasionally act aggressively towards themselves, their tankmates, or humans, which critics say is a result of
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Lachmuth, Cara L.; Barrett-Lennard, Lance G.; Steyn, D. Q.; Milsom, William K. (April 2011). "Estimation of southern resident killer whale exposure to exhaust emissions from whale-watching vessels and potential adverse health effects and toxicity thresholds".
2595:. So little was known at the time, it was nearly two months before the whale's keepers discovered what food (fish) it was willing to eat. To the surprise of those who saw him, Moby Doll was a docile, non-aggressive whale who made no attempts to attack humans.
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regarded orcas as the most powerful animals in the ocean, and their mythology tells of orcas living in houses and towns under the sea. According to these myths, they took on human form when submerged, and humans who drowned went to live with them. For the
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in place to protect endangered animals along with their habitats) for orcas aside from the laws already in place. In response to this decision, six environmental groups sued the federal government, claiming orcas were facing many threats on the
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Resident orcas in the eastern North Pacific live in particularly complex and stable social groups. Unlike any other known mammal social structure, resident whales live with their mothers for their entire lives. These family groups are based on
2021:, Australia, which would have made him up to 90 years old. Examination of his teeth indicated he died around age 35, but this method of age determination is now believed to be inaccurate for older animals. One male known to researchers in the
2710:, their smaller populations, and the difficulty of taking them. Once the stocks of larger species were depleted, orcas were targeted by commercial whalers in the mid-20th century. Between 1954 and 1997, Japan took 1,178 orcas (although the
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McGowen, Michael R.; Tsagkogeorga, Georgia; Álvarez-Carretero, Sandra; dos Reis, Mario; Struebig, Monika; Deaville, Robert; Jepson, Paul D.; Jarman, Simon; Polanowski, Andrea; Morin, Phillip A.; Rossiter, Stephen J. (October 21, 2019).
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In other anecdotes, researchers describe incidents in which wild orcas playfully tease humans by repeatedly moving objects the humans are trying to reach, or suddenly start to toss around a chunk of ice after a human throws a snowball.
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Pitman, Robert L.; Durban, John W.; Greenfelder, Michael; Guinet, Christophe; Jorgensen, Morton; Olson, Paula A.; Plana, Jordi; Tixier, Paul; Towers, Jared R. (August 7, 2010). "Observations of a distinctive morphotype of killer whale
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Western attitudes towards orcas have changed dramatically in recent decades. In the mid-1960s and early 1970s, orcas came to much greater public and scientific awareness, starting with the live-capture and display of an orca known as
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declined to recognize the two species, citing uncertainty as to whether the types constituted unique species or subspecies. "Pending a more complete global review and revision", the Society provisionally recognized them as subspecies
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in winter, and wolves as orcas in summer. Orcas are believed to assist their hunters in driving walrus. Reverence is expressed in several forms: the boat represents the animal, and a wooden carving hung from the hunter's belt. Small
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1306:. With sharks, orcas may herd them to the surface and strike them with their tail flukes, while bottom-dwelling rays are cornered, pinned to the ground and taken to the surface. In other parts of the world, orcas have preyed on
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The "type C" orca has two-toned grey colouring, including a dark "dorsal cape", in body areas where most orcas have solid black colouring. Research is ongoing into whether one or more orca types are distinct species in need of
2035:), while the male's mother supported the breeding opportunity for her son. The attack ended when the calf's mother struck and injured the attacking male. Such behaviour matches that of many smaller dolphin species, such as the
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AD 70, who wrote, "Orcas (the appearance of which no image can express, other than an enormous mass of savage flesh with teeth) are the enemy of ... they charge and pierce them like warships ramming." (see citation in section
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prey of transients hear whale calls well and thus transients are typically silent. Vocal behaviour in these whales is mainly limited to surfacing activities and milling (slow swimming with no apparent direction) after a kill.
1502:. Often, to avoid injury, orcas disable their prey before killing and eating it. This may involve throwing it in the air, slapping it with their tails, ramming it, or breaching and landing on it. In steeply banked beaches off
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They are sometimes referred to as "blackfish", a name also used for other whale species. "Grampus" is a former name for the species, but is now seldom used. This meaning of "grampus" should not be confused with the genus
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usually bears little resemblance to wild habitat, and captive whales' social groups are foreign to those found in the wild. Critics claim captive life is stressful due to these factors and the requirement to perform
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caused by noise and conflicts with boats are the most significant worldwide threats. In January 2020, the first orca in England and Wales since 2001 was found dead with a large fragment of plastic in its stomach.
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As with residents and transients, the lifestyle of these whales appears to reflect their diet; fish-eating orcas off Norway have resident-like social structures, while mammal-eating orcas in Argentina and the
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family groups (pods). Their sophisticated hunting techniques and vocal behaviors, often specific to a particular group and passed along from generation to generation, are considered to be manifestations of
8019:, 'Old Tom' from Eden, Twofold Bay, Australia, in: W. F. Perrin and A. C. Myrick Jr (eds.): Age determination of toothed whales and sirenians, pp. 143–154 Rep. Int. Whal. Comm. (Special Issue 3), cited in
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Pacific animals. Photographic identification has enabled the local population of orcas to be counted each year rather than estimated, and has enabled great insight into life cycles and social structures.
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2668:, which had become separated from his pod, could be returned to it. However, his case was marked by controversy about whether and how to intervene, and in 2006, Luna was killed by a boat propeller.
2025:(identified as J1) was estimated to have been 59 years old when he died in 2010. Orcas are unique among cetaceans, as their caudal sections elongate with age, making their heads relatively shorter.
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Orcas of the same sex and age group may engage in physical contact and synchronous surfacing. These behaviours do not occur randomly among individuals in a pod, providing evidence of "friendships".
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at marine theme parks. Orcas also feature strongly in the mythologies of indigenous cultures, and their reputation in different cultures ranges from being the souls of humans to merciless killers.
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ranging from 7.6 to 10 cm (3.0 to 3.9 in) thick beneath the skin. The pulse is about 60 heartbeats per minute when the orca is at the surface, dropping to 30 beats/min when submerged.
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2532:, the shooting of orcas was accepted and even encouraged by governments. As an indication of the intensity of shooting that occurred until fairly recently, about 25% of the orcas captured in
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In March 2016, SeaWorld announced that they would be ending their orca breeding program and their theatrical shows. However, as of 2020, theatrical shows featuring orcas are still ongoing.
2734:). Other than commercial hunts, orcas were hunted along Japanese coasts out of public concern for potential conflicts with fisheries. Such cases include a semi-resident male-female pair in
1206:, meaning that they themselves have no natural predators. They are sometimes called "wolves of the sea", because they hunt in groups like wolf packs. Orcas hunt varied prey including fish,
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The killer whale dove >750 m on five occasions while depredating (maximum: 1087 m), but these deep dives were always followed by long periods (3.9–4.6 h) of shallow (<100 m) diving.
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vessel it must be placed in neutral until the whale passes. The World Health Organization has set air quality standards in an effort to control the emissions produced by these vessels.
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2196:, Washington. He was also able to study orcas from "his home porch perched above Puget Sound, where the animals hunt and play in summer months". In May 2003, Balcomb (along with other
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2013:
Males sexually mature at the age of 15, but do not typically reproduce until age 21. Wild males live around 29 years on average, with a maximum of about 60 years. One male, known as
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begins at about 12 months of age, and is complete by two years. According to observations in several regions, all male and female pod members participate in the care of the young.
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Orcas are commonly referred to as "killer whales", despite being a type of dolphin. Since the 1960s, the use of "orca" instead of "killer whale" has steadily grown in common use.
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2427:(God of Sea/Offshore) to bring fortunes (whales) to the coasts, and there had been traditional funerals for stranded or deceased orcas akin to funerals for other animals such as
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Closely related matrilines form loose aggregations called pods, usually consisting of one to four matrilines. Unlike matrilines, pods may separate for weeks or months at a time.
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collapse seen in 60–90% of captive males. Captives have vastly reduced life expectancies, on average only living into their 20s. That said, a 2015 study coauthored by staff at
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Totterdell, J. A.; Wellard, R.; Reeves, I. M.; Elsdon, B.; Markovic, P.; Yoshida, M.; Fairchild, A.; Sharp, G.; Pitman, R. (2022). "The first three records of killer whales (
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for aquariums through 1970 bore bullet scars. The U.S. Navy claimed to have deliberately killed hundreds of orcas in Icelandic waters in 1956 with machine guns, rockets, and
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Towner, A; Micarelli, P; Hurwitz, D; Smale, Mj; Booth, Aj; Stopforth, C; Jacobs, E; Reinero, Fr; Ricci, V; Di Bari, A; Gavazzi, S; Carugno, G; Mahrer, M; Gennari, E (2024).
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to locate them on resting on ice floes, and then swim in groups to create waves that wash over the floe. This washes the prey into the water, where other orcas lie in wait.
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cycling separated by non-cycling periods of three to 16 months. Females can often breed until age 40, followed by a rapid decrease in fertility. Orcas are among the few
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such as breaching (jumping completely out of the water) and tail-slapping. These activities may have a variety of purposes, such as courtship, communication, dislodging
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1133:, their range now extends deep into the northwest Atlantic. Occasionally, orcas swim into freshwater rivers. They have been documented 100 mi (160 km) up the
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Demaster, D.; Trites, A.; Clapham, P.; Mizroch, S.; Wade, P.; Small, R.; Hoef, J. (2006). "The sequential megafaunal collapse hypothesis: Testing with existing data".
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Another conservation concern was made public in September 2008 when the Canadian government decided it was not necessary to enforce further protections (including the
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and live for decades after they have finished breeding. The lifespans of wild females average 50 to 80 years. Some are claimed to have lived substantially longer:
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Although there are examples of killer whales living longer, including several over 30 years old, and two captive orcas (Corky II and Lolita) are in their mid-40s.
773:, which first appeared about 11 million years ago. The orca lineage probably branched off shortly thereafter. Although it has morphological similarities with the
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2766:. Whalers more often considered them a nuisance, however, as orcas would gather to scavenge meat from the whalers' catch. Some populations, such as in Alaska's
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led to the discovery of the species' complex social structure, its use of vocal communication, and its extraordinarily stable mother–offspring bonds. Through
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claims that there had been domestic catches of about 1,600 whales between late 1940s to 1960s) and Norway took 987. Extensive hunting of orcas, including an
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Kaiya, Z.; Leatherwood, S.; Jefferson, A. T. (2002). Perrin F. W.; Reeves R. R.; Dolar L. L. M.; Jefferson A. T.; Marsh H.; Wang Y. J.; Estacion J. (eds.).
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stuck in pools enclosed by sea ice, the former are also driven into shallower water where juveniles are grabbed. By contrast, orcas appear to be wary of
985:, with those of males about 1.8 m (5 ft 11 in) high, more than twice the size of the female's, with the male's fin more like an elongated
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Towers, Jared R; Tixier, Paul; Ross, Katherine A; Bennett, John; Arnould, John P Y; Pitman, Robert L; Durban, John W; Northridge, Simon (January 2019).
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Morin, Phillip A.; Archer, Frederick; Foote, Andrew D.; Vilstrup, Julia; Allen, Eric E.; Wade, Paul; Durban, John; Parsons, Kim; Pitman, Robert (2010).
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Of the very few confirmed attacks on humans by wild orcas, none have been fatal. In one instance, orcas tried to tip ice floes on which a dog team and
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With their distinctive pigmentation, adult orcas are seldom confused with any other species. When seen from a distance, juveniles can be confused with
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discovered it could regurgitate fish onto the surface, attracting sea gulls, and then eat the birds. Four others then learned to copy the behaviour.
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Kremers, D.; Lemasson, A.; Almunia, J.; Wanker, R. (2012). "Vocal sharing and individual acoustic distinctiveness within a group of captive orcas (
6681:
3480:
LeDuc, R. G.; Perrin, W. F.; Dizon, A. E. (1999). "Phylogenetic relationships among the delphinid cetaceans based on full cytochrome b sequences".
1125:
whales in the Arctic. However, orcas are merely seasonal visitors to Arctic waters, and do not approach the pack ice in the summer. With the rapid
3243:
2698:
The earliest known records of commercial hunting of orcas date to the 18th century in Japan. During the 19th and early 20th centuries, the global
2500:
vessels have been damaged between 2020 and 2023. In one video, an orca can be seen biting on one of the two rudders ripped from a catamaran near
1911:, mothers push their calves onto the beach, waiting to pull the youngster back if needed. In March 2023, a female orca was spotted with a newborn
12801:
10643:
8748:"Поддержка прав коренных народов Cибири - Spiritual culture subsection of the Eskimos page in the Support for Siberian Indigenous Peoples Rights"
7382:"Higher neuron densities in the cerebral cortex and larger cerebellums may limit dive times of delphinids compared to deep-diving toothed whales"
5683:
Towner, AV; Watson, RGA; Kock, AA; Papastamatiou, Y; Sturup, M; Gennari, E; Baker, K; Booth, T; Dicken, M; Chivell, W; Elwen, S (April 3, 2022).
5325:
3952:
Morin, P. A.; McCarthy, M. L.; Fung, C. W.; Durban, J. W.; Parsons, K. M.; Perrin, W. F.; Taylor, B. L.; Jefferson, T. A.; Archer, F. I. (2024).
1427:, great whales may have been the major food source for orcas. The introduction of modern whaling techniques may have aided orcas by the sound of
907:
10120:
9213:
529:
9621:
6987:
Deeck, V. B.; Ford, J. K. B.; Slater, P. J. B. (2005). "The vocal behaviour of mammal-eating killer whales: communicating with costly calls".
4809:
3423:
Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I
13129:
12892:
12840:
10688:
10676:
10633:
7358:
4150:
2722:
to recommend a ban on commercial hunting of the species pending further research. Today, no country carries out a substantial hunt, although
7482:
5233:
3431:
1934:
demonstrating the whales' curiosity, playfulness, and ability to solve problems. Alaskan orcas have not only learned how to steal fish from
1552:
harass and kill porpoises and seals for no apparent reason. Some dolphins recognize resident orcas as harmless and remain in the same area.
9496:
6942:
4819:, January 28, 2007. Iruka (dolphin) and Kujira (whale) Action Network (IKAN): Iruma, Saitama Prefecture, Japan. Retrieved February 17, 2010
2739:
2588:
1975:
9545:
5945:"Feeding Ecology of Elusive Caribbean Killer Whales Inferred From Bayesian Stable Isotope Mixing Models and Whalers' Ecological Knowledge"
579:
Orcas are not usually a threat to humans, and no fatal attack has ever been documented in their natural habitat. There have been cases of
9087:
6557:
6130:
1062:. Systematic surveys indicate the highest densities of orcas (>0.40 individuals per 100 km) in the northeast Atlantic around the
10031:
8944:
8717:. Original data: Е.С. Рубцова: Материалы по языку и фольклору эскимосов (чаплинский диалект). Академия Наук СССР. Москва-Ленинград, 1954
3914:
Josh D. McInnes; Andrew W. Trites; Chelsea R. Mathieson; Marilyn E. Dahlheim; Jeffrey E. Moore; Paula A. Olson; Kevin M. Lester (2024).
1038:, numbers, and density, relative distribution is difficult to estimate, but they clearly prefer higher latitudes and coastal areas over
996:
Orcas have good eyesight above and below the water, excellent hearing, and a good sense of touch. They have exceptionally sophisticated
7897:
7244:
Simon, M.; McGregor, P. K.; Ugarte, F. (2007). "The relationship between the acoustic behaviour and surface activity of killer whales (
6510:
4915:
969:
Sexual dimorphism between male (top) and female (bottom) killer whales. Pectoral fins, dorsal fin, and flukes are larger in adult males
3216:
2112:
have problems in reproductive and immune functions associated with high levels of PCBs and related contaminants, and a survey off the
1945:
The orca's use of dialects and the passing of other learned behaviours from generation to generation have been described as a form of
8528:
7147:
1794:
1710:
layer is the community, which consists of pods that regularly associate with each other but share no maternal relations or dialects.
9667:
9353:
8170:
Desforges, J-P; Hall, A; Mcconnell, B; et al. (2018). "Predicting global killer whale population collapse from PCB pollution".
7212:
6333:
3647:
3632:
2275:, which may stress the whales and alter their behaviour, particularly if boats approach too closely or block their lines of travel.
580:
12775:
9190:
8047:, Research Document 2005/045, Canadian Science Advisory Secretariat, Fisheries and Oceans Canada. p. 33. Retrieved January 27, 2010
1841:
Northeast Pacific resident groups tend to be much more vocal than transient groups in the same waters. Residents feed primarily on
8110:
7836:
6056:
5459:
4009:
917:, were described during the 1980s by Soviet researchers, but most cetacean researchers are skeptical about their status. Complete
13086:
12814:
10006:
9528:
8734:. Original data: Г. А. Меновщиков: Грамматиκа языка азиатских эскимосов. Часть первая. Академия Наук СССР. Москва-Ленинград, 1962
8432:
6080:"Humpback whales interfering when mammal-eating killer whales attack other species: Mobbing behavior and interspecific altruism?"
3328:
3187:
2648:, David Phillips, led the efforts to return Keiko to the Iceland waters. Keiko however did not adapt to the harsh climate of the
2116:
coast found PCB levels in orcas were higher than levels that had caused health problems in harbour seals. Blubber samples in the
9118:
5255:
Similä, T.; Ugarte, F. (1993). "Surface and underwater observations of cooperatively feeding killer whales in Northern Norway".
3676:
2591:
at the time, including the first scientific studies of an orca's sound production, led to two articles about him in the journal
2189:
13144:
6197:
4594:
4424:
2759:
8773:
5794:
Weller, D. W. (2009). "Predation on marine mammals". In Perrin, William F.; Wursig, Bernd; Thewissen, J. G. M. 'Hans' (eds.).
3297:
12941:
12858:
10583:
10562:
10505:
10439:
10410:
10377:
10305:
10251:
10226:
10073:
8975:
8897:
8647:
6504:
6124:
5803:
4793:
4588:
4397:
4217:
3954:"Revised taxonomy of eastern North Pacific killer whales (Orcinus orca): Bigg's and resident ecotypes deserve species status"
3367:
3031:
2238:
13091:
12819:
3455:
3276:
3179:
13154:
8906:
8550:
3807:
2819:, trainability, striking appearance, playfulness in captivity and sheer size have made it a popular exhibit at aquaria and
785:, a study of cytochrome b gene sequences indicates that its closest extant relatives are the snubfin dolphins of the genus
6243:
Visser, Ingrid N.; Smith, Thomas G.; Bullock, Ian D.; Green, Geoffrey D.; Carlsson, Olle G. L.; Imberti, Santiago (2008).
5025:
2221:
and the federal government did nothing to protect them from these threats. A legal and scientific nonprofit organization,
9960:
7380:
Ridgway, Sam H.; Brownson, Robert H.; Van Alstyne, Kaitlin R.; Hauser, Robert A. (December 16, 2019). Li, Songhai (ed.).
6719:
5569:"Predator–prey and competitive interactions between sharks (order Selachii) and dolphins (suborder Odontoceti): a review"
5501:
3639:. Contract report to the Committee on the Status of Endangered Wildlife in Canada, Ottawa, ON, Canada. Also published as
2711:
10660:
9291:
6646:
3386:
Whales, dolphins, and porpoises of the eastern North Pacific and adjacent Arctic waters: a guide to their identification
989:, whereas the female's is more curved. In the skull, adult males have longer lower jaws than females, as well as larger
13149:
13073:
12762:
8813:
8059:
7661:
7515:
6827:
2461:, orcas were historically feared as dangerous, savage predators. The first written description of an orca was given by
706:
13011:
8303:
8020:
3157:
2834:
campaign against the practice of keeping them in captivity. In captivity, they often develop pathologies, such as the
1524:
12905:
10723:
10465:
10431:
10284:
9750:
9576:
9549:
7748:
4753:
3394:
2383:
754:
488:
with a diverse diet. Individual populations often specialize in particular types of prey. This includes a variety of
10173:
8076:
3013:
13019:
11763:
2719:
1631:
is a behaviour in which a whale holds its head above water to view its surroundings. Resident orcas swim alongside
8343:
5601:
4579:. In Estes, James A.; DeMaster, Douglas P.; Doak, Daniel F.; Williams, Terrie M.; Brownell, Robert L. Jr. (eds.).
4388:
Ford, John K. B. (2009). "Killer Whale". In Perrin, William F.; Wursig, Bernd; Thewissen, J. G. M. 'Hans' (eds.).
4353:
2255:
from shipping, drilling, and other human activities is a significant concern in some key orca habitats, including
1082:. They are considered "common" (0.20–0.40 individuals per 100 km) in the eastern Pacific along the coasts of
869:
A 2024 study supported the elevation of Eastern North American resident and transient orcas as distinct species,
12897:
11745:
9792:"Report of the Second Workshop on The Biology and Conservation of Small Cetaceans and Dugongs of South-East Asia"
8991:
8612:
8414:
5740:"Further insights into killer whales Orcinus orca preying on white sharks Carcharodon carcharias in South Africa"
5044:"The diving behaviour of mammal-eating killer whales : variations with ecological not physiological factors"
3427:
2153:
1981:
Female orcas begin to mature at around the age of 10 and reach peak fertility around 20, experiencing periods of
738:
365:
9021:
8747:
4648:
4522:
Spencer, M. P.; Gornall 3rd, T. A.; Poulter, T. C. (1967). "Respiratory and cardiac activity of killer whales".
3916:"Evidence for an Oceanic Population of Killer Whales (Orcinus orca) in Offshore Waters of California and Oregon"
2371:
consider orca to be people, referring to them as "qwe'lhol'mechen" which means "our relations under the waves".
2294:
numbered only seven individuals and had not reproduced since the spill. This population is expected to die out.
12469:
10524:
10325:
8879:
7288:
5685:"Fear at the top: killer whale predation drives white shark absence at South Africa's largest aggregation site"
2831:
2694:, flanking a whale calf. The boat is being towed by a harpooned whale (not visible here), near Eden, Australia.
2513:
1849:
salmon, which are insensitive to orca calls (inferred from the audiogram of Atlantic salmon). In contrast, the
569:
9727:
9697:
7798:
5043:
4829:
4483:
3067:
13139:
13104:
12879:
11886:
11611:
11602:
9313:
9252:
2939:
2619:
694:
10648:
10625:
8826:
A radio interview with Russian scientists about man and animal, examples taken especially from Asian Eskimos
7871:
6756:
4757:
1872:-eating orcas appear to have different vocalizations for activities like hunting. A population that live in
12415:
11432:
11423:
11355:
7547:
4017:
2145:
region, only 75 whales remain with few births over the last few years. On the west coast of Alaska and the
1776:
977:
are large and rounded, resembling paddles, with those of males significantly larger than those of females.
878:
816:
9887:
7450:
6446:"Causes and consequences of marine mammal population declines in southwest Alaska: a food-web perspective"
6299:
565:
12853:
12687:
12664:
12514:
12442:
11653:
11620:
11382:
9802:
9445:
9418:
8731:
8714:
8286:
2382:
also had great respect for orcas, as evidenced by stone carvings found in a 4,000-year-old burial at the
1961:) appear to have no parallel outside humans and represent an independent evolution of cultural faculties.
1646:, that is, killing that is not designed to be for food. As an example, a BBC film crew witnessed orca in
1343:
42:
12692:
7174:"Vocal behaviour of resident killer whale matrilines with newborn calves: The role of family signatures"
3736:
De Bruyn, P. J. N.; Tosh, C. A.; Terauds, A. (2013). "Killer whale ecotypes: Is there a global model?".
3239:
12547:
12538:
11465:
10873:
10615:
10602:
2962:
2567:
2360:
2121:
1674:. Due to orcas' complex social bonds, many marine experts have concerns about how humane it is to keep
1432:
expand their consumption of smaller marine mammals, thus contributing to the decline of these as well.
1018:
832:
10110:
3913:
2267:
were used to deter seals. Orcas also avoided the surrounding waters. High-intensity sonar used by the
2205:
of two of the whale heads showed hemorrhaging around the brain and the ears, which is consistent with
2141:, wild salmon stocks, a main resident food source, have declined dramatically in recent years. In the
13024:
12920:
12433:
12388:
12356:
12347:
12332:
12323:
12173:
11772:
11629:
10882:
8654:, Newfoundland and Labrador Heritage, Memorial University of Newfoundland. Retrieved January 26, 2010
7022:
Foote, A. D.; Osborne, R. W.; Hoelzel, A. (2008). "Temporal and contextual patterns of killer whale (
6352:"Sequential megafaunal collapse in the North Pacific Ocean: An ongoing legacy of industrial whaling?"
5739:
2193:
2032:
1604:
1383:
1371:
1307:
1035:
391:
5082:
Overall, the whales spent 50% of their time 8 m or shallower and 90% of their time 40 m or shallower
3525:"New insights into the phylogenetic relationships among the oceanic dolphins (Cetacea: Delphinidae)"
1440:
239:
12571:
12562:
12496:
12460:
12424:
12397:
12273:
11946:
11895:
11680:
11265:
9606:
8917:
5588:
3124:
2549:
2188:
has extensively studied orcas since 1976; he is the research biologist responsible for discovering
2105:
1616:
1375:
990:
557:
12871:
10630:
7350:
4189:
3421:
2863:
was involved in the death of three people, and was featured in the critically acclaimed 2013 film
180:
5.4 meter (18 foot) female killer whale compared to 1.8 meter (5 foot 11 inches) human scuba diver
12985:
12282:
12090:
11274:
7896:
Robeck, Todd R.; Willis, Kevin; Scarpuzzi, Michael R.; O'Brien, Justine K. (September 29, 2015).
5229:
2827:
2367:
of southeastern Alaska regarded the orca as custodian of the sea and a benefactor of humans. The
2226:
1495:
10276:
10266:
4504:
N. W. Kasting, S. A. L. Adderly, T. Safford, K. G. Hewlett (1989). "Thermoregulation in Beluga (
2982:
According to Baird, killer whales prefer harbour seals to sea lions and porpoises in some areas.
2134:
found that global orca populations are poised to dramatically decline due such toxic pollution.
12972:
12806:
12642:
12604:
12487:
12451:
12406:
12291:
12099:
11176:
11158:
10493:
6685:
6340:
Ned Rozell, Article #1418, Alaska Science Forum, December 10, 1998. Retrieved February 26, 2010
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2218:
2206:
2157:
2065:
2014:
1624:
1511:
6112:
3384:
1970:
1687:
separate for only a few hours at a time, to mate or forage. With one exception, an orca named
12967:
11713:
11704:
10936:
10716:
10394:
10210:
10038:
9350:
6494:
6198:"Cooperative hunting behavior, prey selectivity and prey handling by pack ice killer whales (
5517:
2653:
2481:
2379:
2356:
2230:
2089:
2028:
1889:
1758:
1675:
1548:
1503:
1275:
812:
344:
31:
17:
12959:
10388:
10295:
10262:
10204:
7939:
6884:"Age and sex influence social interactions, but not associations, within a killer whale pod"
3808:"A review of killer whale interactions with other marine mammals: predation to co-existence"
3021:
13006:
12832:
12736:
12651:
12182:
12108:
11346:
10864:
10334:
9926:
8520:
8179:
7966:
7699:
7393:
7188:
7123:
7035:
6596:
6418:
6363:
6263:
6040:
5993:
5897:
5751:
5696:
5639:
5526:
5430:
5386:
5111:
5042:
Miller, Patrick James O'Malley; Shapiro, Ari Daniel; Deecke, Volker Bernt (November 2010).
5001:
4880:
3965:
3870:
3489:
3208:
2767:
2587:
in 1964. He was the first ever orca to be studied at close quarters alive, not postmortem.
2524:
Competition with fishermen also led to orcas being regarded as pests. In the waters of the
2494:
2286:
2252:
2246:
1588:
1042:
environments. Areas which serve as major study sites for the species include the coasts of
470:
9405:
Schevill, William; Watkins, William (Summer 1966). "Sound Structure and Directionality in
7104:
8:
12523:
12478:
12191:
11919:
11406:
11322:
11313:
11185:
10696:
9661:
8894:
8617:
6587:
Pitman, R. L.; Durban, J. W. (2010). "Killer whale predation on penguins in Antarctica".
5094:
Reisinger, Ryan R.; Keith, Mark; Andrews, Russel D.; de Bruyn, P. J. N. (December 2015).
4916:"Killer whales in Hawaiian waters: information on population identity and feeding habits"
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3629:
2816:
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2213:
2061:
1487:
1457:
1394:
1283:
997:
895:
672:
533:
187:
53:
10338:
9930:
9432:
Newman, Murray; McGeer, Patrick (Summer 1966). "The Capture and Care of a Killer Whale,
9182:
8183:
7970:
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7397:
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7127:
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6600:
6422:
6367:
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6267:
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5901:
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5700:
5643:
5530:
5434:
5390:
5115:
5005:
4884:
3986:
3969:
3953:
3874:
3493:
2610:
Between 1964 and 1976, 50 orcas from the Pacific Northwest were captured for display in
1926:
An orca plays with a ball of ice, soon after a researcher threw a snowball at the whale.
1691:, no permanent separation of an individual from a resident matriline has been recorded.
1047:
12505:
11970:
11961:
11728:
11537:
11528:
11513:
11504:
11364:
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10369:
10115:
9856:
9831:
9387:
9253:"Orcas sank a yacht off Spain — the latest in a slew of such 'attacks' in recent years"
8921:
8689:
8681:
8486:
8213:
8102:
7926:
7898:"Comparisons of life-history parameters between free-ranging and captive killer whale (
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7653:
7601:
7572:
7424:
7381:
7269:
7139:
7004:
6918:
6883:
6612:
6470:
6445:
6297:
Transient Killer Whales – Culprits in the Decline of Sea Otters in Western Alaska?
6052:
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6009:
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399:
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9999:
9525:
6386:
6351:
2682:
2060:(International Union for Conservation of Nature) changed its assessment of the orca's
1865:
offshore orcas have group-specific dialects unlike those of residents and transients.
638:, the species is more closely related to other oceanic dolphins than to other whales.
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12954:
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12656:
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3707:
3567:"Phylogenomic Resolution of the Cetacean Tree of Life Using Target Sequence Capture"
3550:
2301:(CITES), meaning international trade (including in parts/derivatives) is regulated.
1330:. They are recorded to prey on other cetacean species, usually smaller dolphins and
1242:
account for 96% of northeast Pacific residents' diet, including 65% of large, fatty
13065:
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12306:
12228:
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12209:
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3119:
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2348:
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2146:
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2002:
1986:
1935:
1671:
1651:
1647:
1531:
1491:
1447:
1351:
1260:
1083:
1067:
837:
The three to five types of orcas may be distinct enough to be considered different
632:
158:
8644:
7135:
7000:
6643:"Whale uses fish as bait to catch seagulls then shares strategy with fellow orcas"
5763:
5708:
4790:
13051:
12928:
11689:
11671:
11662:
11552:
11167:
11149:
11131:
10999:
10692:
10680:
10667:
10655:
10637:
10619:
10606:
10427:
10237:
10068:
10063:
9532:
9357:
9321:
8913:
8889:
8875:
8651:
8310:
8063:
8044:
7591:
7554:
7406:
6430:
6337:
6303:
5910:
5885:
4892:
4816:
4797:
4207:
3651:
3636:
3283:
2743:
2657:
2603:
2477:
2462:
2458:
2101:
2018:
1903:
1643:
1287:
850:
807:
541:
456:
311:
12998:
12884:
12845:
10420:
Heptner, V. G.; Nasimovich, A. A.; Bannikov, A. G.; Hoffmann, Robert S. (1996).
10358:
Operation Orca: Springer, Luna and the Struggle to Save West Coast Killer Whales
8711:
Materials on the Language and Folklore of the Eskimoes, Vol. I, Chaplino Dialect
8557:
5684:
5164:"Movements and dive behaviour of a toothfish-depredating killer and sperm whale"
5123:
4535:
4445:
4430:
4235:"Orca Behavior and Subsequent Aggression Associated with Oceanarium Confinement"
2706:
and sperm whales, but largely ignored orcas because of their limited amounts of
2445:
1027:
An orca leaps out of the water when swimming—a behaviour known as porpoising—in
930:
13134:
11570:
11337:
11209:
11200:
11047:
11038:
10770:
10402:
10272:
10218:
9058:"Scientists are puzzled by Orcas sinking boats in a string of abnormal attacks"
5844:
5651:
2903:
2860:
2778:
2770:, may have been reduced significantly by whalers shooting them in retaliation.
2763:
2687:
2614:, and public interest in the animals grew. In the 1970s, research pioneered by
2584:
2390:
2364:
2315:
2272:
2197:
2178:
2177:
population in the past 10 years. These deaths can be attributed to declines in
2069:
1946:
1908:
1842:
1719:
1694:
1667:
1564:
1507:
1464:
1428:
1409:
1335:
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1243:
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1142:
1134:
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1118:
1105:
1075:
1071:
1051:
918:
838:
801:
745:
537:
522:
478:
202:
12718:
10685:
10673:
9287:
8077:"'Horrified' scientists first to see killer whale infanticide | CBC News"
7641:
7265:
7103:
Filatova, Olga A.; Fedutin, Ivan D.; Burdin, Alexandr M.; Hoyt, Erich (2007).
6711:
6608:
5961:
5944:
5597:
5542:
5013:
4102:
2758:
Orcas have helped humans hunting other whales. One well-known example was the
1916:
811:. In contrast, a 2019 phylogenetic study found the orca to be the second most
13123:
12827:
12163:
11988:
11391:
11116:
11065:
10832:
10823:
10541:
10533:
10243:
9225:
9161:
9150:"Orca rams into yacht off Shetland in first such incident in northern waters"
8901:
8842:
8664:
Tuck, James A. (1971). "An Archaic Cemetery at Port Au Choix, Newfoundland".
8418:
8279:"Granny's Struggle: When Granny is gone, will her story be the last chapter?"
7917:
7415:
6682:"Videos show killer whales frantically hunting for dolphins off San Clemente"
6642:
5920:
5771:
5716:
5180:
5163:
4324:
4301:
4260:
4206:
Webber, Marc A.; Jefferson, Thomas Allen; Pitman, Robert L. (July 28, 2015).
3815:
3590:
3509:
3417:
3110:
3017:
2931:
2843:
2804:
2795:
2735:
2637:
2414:
2117:
2097:
1990:
1873:
1850:
1515:
1483:
1347:
1327:
1303:
1226:
Fish-eating orcas prey on around 30 species of fish. Some populations in the
1203:
1130:
1099:
759:
734:
615:
513:
501:
485:
452:
361:
207:
154:
102:
10005:. Humane Society International and the Humane Society of the United States.
9961:"Boats to Maintain Greater Distance from Killer Whales | Whales online"
8807:
8411:
8191:
8056:
7625:
7530:
7483:"'Extraordinary' sighting of orca with baby pilot whale astounds scientists"
6376:
6119:. Wursig, Bernd & Thewissen, J. G. M. Academic Press. pp. 357–361.
5310:
3931:
3582:
3563:
3041:
2746:
in 1957, the killing of five whales from a pod of 11 members that swam into
2320:
1922:
13078:
12767:
12627:
12246:
11809:
11800:
11790:
10918:
10798:
10782:
10551:
10361:
9946:
9865:
9847:
8482:
8209:
7935:
7649:
7610:
7577:
7433:
7208:
7172:
Weiß, Brigitte M.; Ladich, Friedrich; Spong, Paul; Symonds, Helena (2006).
7089:
6927:
6899:
6479:
6461:
6395:
5929:
5862:
5669:
5373:
Visser, Ingrid N. (1999). "Benthic foraging on stingrays by killer whales (
4900:
4278:
4070:
3995:
3892:
3769:
3699:
3608:
3149:
2848:
2703:
2649:
2615:
2537:
2368:
2343:
2324:
2290:
2165:
2109:
1572:
1520:
1479:
1413:
1359:
1171:
1138:
1039:
974:
792:
724:
603:
553:
474:
460:
7758:
5500:
Pyle, Peter; Schramm, Mary Jane; Keiper, Carol; Anderson, Scot D. (1999).
4934:
4543:
4209:
Marine Mammals of the World: A Comprehensive Guide to Their Identification
4052:
4033:"Complete mitochondrial genome phylogeographic analysis of killer whales (
12980:
12933:
12866:
12788:
12679:
12636:
12007:
11248:
10847:
10776:
10701:
9723:
9693:
8034:
Life History and Population Dynamics of Northern Resident Killer Whales (
8015:
Mitchell, E. and Baker, A. N. (1980). Age of reputedly old Killer Whale,
6306:" B.C. Cetacean Sightings Network, June 1, 2004. Retrieved March 13, 2010
4837:
2661:
2644:
in 2002. The director of the International Marine Mammal Project for the
2533:
2440:
2423:
2406:
2264:
2260:
2185:
2169:
2142:
2077:
1982:
1912:
1902:
and more cortical neurons than any mammal, including humans. They can be
1899:
1895:
1857:
All members of a resident pod use similar calls, known collectively as a
1846:
1688:
1416:. On three separate occasions in 2019 orcas were recorded to have killed
1401:
1367:
1311:
1255:
1247:
1215:
1178:
965:
782:
770:
635:
517:
291:
77:
12728:
9799:
CMS Technical Series Publication Nº 9 at Convention on Migratory Species
8464:"Behavioural responses of male killer whales to a 'leapfrogging' vessel"
7688:"Quantifying the effects of prey abundance on killer whale reproduction"
6908:
5324:
Visser, Ingrid N.; Berghan, Jo; van Meurs, Rinie; Fertl, Dagmar (2000).
4683:. Northwest Fisheries Science Center. February 14, 2007 . Archived from
3977:
3566:
2626:
unthinkingly been targeted became a cultural icon within a few decades.
1547:
swimming between islands off the northwest coast of North America. Orca
1519:
technique on the now-weakened prey. In the Antarctic, type B orcas hunt
12780:
11023:
11014:
10972:
10813:
10347:
10316:
9497:"Death of Sea World trainer: Do 'killer whales' belong in theme parks?"
9391:
8200:
8141:"Orca 'apocalypse': half of killer whales doomed to die from pollution"
6029:"Killer Whale Predation on Sperm Whales: Observations and Implications"
4943:
4357:
4251:
4177:
3458:(in German). St. Mary's Church, Greifswald. Retrieved February 16, 2010
2835:
2632:
2450:
2428:
2271:
disturbs orcas along with other marine mammals. Orcas are popular with
2222:
2125:
1998:
1877:
1815:
1628:
1563:
species recorded as prey in Antarctic and sub-Antarctic waters include
1417:
1405:
1295:
1207:
1079:
1055:
1028:
978:
959:
842:
144:
122:
87:
9535:
Associated Press. Published December 13, 2003. Retrieved June 7, 2020.
9119:"Orcas filmed destroying ship's rudder in shocking clip: "Surrounded""
8685:
7978:
7200:
6558:"Killer whales eat dolphins. So why are these dolphins tempting fate?"
6450:
Philosophical Transactions of the Royal Society B: Biological Sciences
6096:
6079:
6005:
5443:
5418:
5285:
Visser, Ingrid N. (2005). "First Observations of Feeding on Thresher (
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3760:
3541:
3524:
2925:
2598:
665:
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12749:
12264:
12126:
10990:
10945:
10398:
10214:
10000:"Killer Controversy: Why Orcas Should No Longer Be Kept in Captivity"
8755:
8582:
8580:
8578:
7081:
4515:
2747:
2727:
2723:
2715:
2707:
2691:
2576:
2501:
2399:
2328:
2085:
2017:, was reportedly spotted every winter between the 1840s and 1930 off
1835:
1683:
1580:
1535:
1463:
Orcas swimming in close synchronization to create a wave to wash the
1231:
902:
627:), which referred (among other things) to a whale species, perhaps a
251:
127:
12598:
9383:
8371:"Ottawa Sued over Lack of Legislation to Protect B.C. Killer Whales"
6444:
Estes, J. A.; Doak, D. F.; Springer, A. M.; Williams, T. M. (2009).
5268:
5062:
4169:
3883:
3858:
2072:, recognizing that one or more orca types may actually be separate,
171:
13045:
12754:
12621:
12025:
11998:
11910:
11851:
11644:
11447:
10764:
10752:
9719:
9689:
8677:
6943:"Killer whales form killer friendships, new drone footage suggests"
6561:
5829:) in Icelandic coastal waters and their interspecific interactions"
5628:"Killer whales redistribute white shark foraging pressure on seals"
5472:
American Elasmobranch Society 16th Annual Meeting, June 14–20, 2000
2950:
2839:
2800:
2611:
2418:
2410:
2352:
2202:
1954:
1663:
1632:
1620:
1612:
1475:
1331:
1271:
1117:
and are believed to venture into the denser pack ice, finding open
1114:
787:
505:
271:
117:
112:
97:
92:
82:
70:
9915:
9183:"Sprinkling sand 'deters orcas' according to Cruising Association"
8859:
8857:
8855:
8853:
8575:
8122:
8120:
7686:
Ward, Eric J.; Holmes, Elizabeth E.; Balcomb, Ken C. (June 2009).
4994:
Journal of the Marine Biological Association of the United Kingdom
3026:(3rd ed.). Johns Hopkins University Press. pp. 723–743.
2192:. He studied orcas from the Center for Whale Research, located in
2149:, seal and sea lion populations have also substantially declined.
2047:
1894:
Orcas have the second-heaviest brains among marine mammals (after
1615:, travelling, resting and socializing. Orcas frequently engage in
1238:
and follow that fish's autumnal migration to the Norwegian coast.
1023:
861:
of this genus is clearly in need of review, and it is likely that
13099:
12741:
12071:
11937:
11833:
11824:
11480:
10900:
10733:
10461:
10419:
8809:Животные и отражение их прихода к человеку в самых разных текстах
8032:
Olesiuk, Peter F.; Ellis, Graeme M. and Ford, John K. B. (2005).
7867:
6760:
4981:
Weir, C. R.; Collins, T.; Carvalho, I.; Rosenbaum, H. C. (2010).
4451:
4439:
3784:
3023:
Mammal Species of the World: A Taxonomic and Geographic Reference
2699:
2677:
2641:
2529:
2351:, the orca was regarded as the ruler of the undersea world, with
2007:
1869:
1858:
1636:
1560:
1556:
1389:
1379:
1363:
1235:
1211:
1095:
1054:, New Zealand and parts of the west coast of North America, from
1043:
1002:
951:
846:
729:
677:
676:
fossil, an extinct species of the same genus, Museo Capellini in
628:
509:
497:
465:
321:
301:
132:
107:
12946:
10294:
Ford, John K. B.; Ellis, Graeme M.; Balcomb, Kenneth C. (2000).
9452:
8806:Ковалева, Ирина & Богословская, Людмила (December 3, 2002).
8393:
7895:
7379:
7105:"The structure of the discrete call repertoire of killer whales
4233:
Anderson, Robert; Waayers, Robyn; Knight, Andrew (August 2016).
3286:, Seaworld.org, September 23, 2010, Retrieved September 9, 2010.
11561:
10758:
10746:
9879:
9088:"Gladis the killer whale and her gang of orcas out for revenge"
8939:
8850:
8592:
8255:
8117:
6863:
6739:
6737:
6537:
6296:
2869:. Tilikum lived at SeaWorld from 1992 until his death in 2017.
2160:. This community comprises three pods which live mostly in the
1898:, which have the largest brain of any animal). Orcas have more
1499:
1239:
1227:
1091:
1063:
1059:
955:
281:
261:
162:
10490:
Guardians of the Whales: The Quest to Study Whales in the Wild
8233:"Orcas of the Pacific Northwest Are Starving and Disappearing"
7351:"Big and brilliant: complex whale behavior tied to brain size"
6152:
6150:
6148:
4583:. Berkeley: University of California Press. pp. 145–162.
4406:
3095:
3093:
3091:
3089:
3087:
3085:
2488:
2434:
12993:
12145:
12136:
12035:
5682:
5093:
4987:) in Angolan and Gulf of Guinea waters, tropical West Africa"
4480:"Killer Whales — Adaptations for an Aquatic Environment"
4083:
3258:
2750:
in 1970, and a catch record in southern Taiwan in the 1990s.
2622:
techniques, individuals were named and tracked over decades.
2485:
bumped by an orca that apparently misidentified him as prey.
1930:
People who have interacted closely with orcas offer numerous
1544:
1267:
796:
618:
599:
493:
375:
224:
218:
10599:
9476:
9360:, SeaWorld/Busch Gardens ANIMALS. Retrieved January 11, 2010
7818:
7816:
7464:
7462:
7063:
6734:
6496:
Killer Whales of the World: Natural History and Conservation
6408:
4980:
2512:
Unlike wild orcas, captive orcas have made nearly two dozen
934:
Different angle views of a typical female orca's appearance
12081:
10686:
Watch: Killer Whales Charge Blue Whale (Rare Drone Footage)
9644:
9642:
7953:
Jett, John; Ventre, Jeffrey (2015). "Captive killer whale (
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4457:
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Journal of Zoological Systematics and Evolutionary Research
3082:
2394:
2057:
1584:
854:
791:. However, a more recent (2018) study places the orca as a
489:
8103:"First stranded Orca found in almost 20 years in the Wash"
7785:
7783:
7289:"The smallest killer whale has a large musical repertoire"
5416:
4871:
Morell, Virginia (2011). "Killer Whales Earn Their Name".
2402:
such as tobacco or meat are strewn into the sea for them.
1158:
the 2000s that 2,321 orcas were in the seas around Japan.
1034:
Orcas are found in all oceans and most seas. Due to their
540:
because of the likelihood that two or more orca types are
469:
and is recognizable its black-and-white patterned body. A
11114:
10522:
Hoyt, Erich (August 1984). "The Whales Called "Killer"".
9789:
9748:
9666:(in Japanese). Osaka College of Eco & Animals. 2011.
9345:
Killer Whales Destroyed: VP-7 Accomplishes Special Task,
9257:
8421:, Raincoast Research Society. Retrieved February 18, 2010
8023:, The Dolphin's Encyclopaedia. Retrieved January 27, 2010
7813:
7459:
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7225:
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6661:
6443:
6179:
6177:
5499:
4550:
4521:
2897:
1953:
The complex and stable vocal and behavioural cultures of
1706:
1599:
748:
wrote the first scientific description of an orca in his
733:, and one of many animal species originally described by
10612:
10451:"Conservation Plan for Southern Resident Killer Whales (
10037:. Whale and Dolphin Conservation Society. Archived from
9911:
9909:
9907:
9905:
9751:"Records of Small Cetaceans in Chinese Waters: A Review"
9639:
7902:) populations for application toward species management"
7493:
6162:
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Engelbrecht, T. M.; Kock, A. A.; O'Riain, M. J. (2019).
4719:
4717:
4612:
4331:
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3801:
3799:
3461:
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Males mate with females from other pods, which prevents
10796:
10488:
Obee, Bruce; Ellis, Graeme (1992). Elaine Jones (ed.).
10143:"SeaWorld to End Controversial Orca Shows and Breeding"
9464:
7837:"Oldest Southern Resident killer whale considered dead"
7780:
7729:
Advances in Veterinary Science and Comparative Medicine
6888:
Proceedings of the Royal Society B: Biological Sciences
6242:
5161:
4574:"Worldwide distribution and abundance of killer whales"
3951:
2299:
Convention on International Trade in Endangered Species
1137:
in the United States. They have also been found in the
10622: – Listen live to orcas in Washington State, U.S.
10174:"Even years after Blackfish, SeaWorld still has Orcas"
9979:
9832:"Killer whales and whaling: the scavenging hypothesis"
9720:"Stranding Date Base (1901-2012) for Chiba Prefecture"
8169:
7222:
7171:
6970:
6968:
6853:
6851:
6797:
6795:
6782:
6780:
6778:
6314:
6312:
6174:
4660:
4658:
4427:. American Cetacean Society. Retrieved January 2, 2009
3806:
Jefferson, T. A.; Stacey, P. J.; Baird, R. W. (1991).
3430: ed.). Holmiae. (Laurentii Salvii). p. 824.
2421:
often referred to orcas in their folklore and myth as
473:, they are found in diverse marine environments, from
10644:
Voices in the Sea - Sounds of the Orca (Killer Whale)
10553:
Orca: The Whale Called Killer Camden House Publishing
9902:
9749:
Kaiya, Z.; Leatherwood, S.; Jefferson, A. T. (1995).
9398:
9324:, CNN, February 25, 2010, Retrieved September 9, 2010
9214:"Sailors have a new way to stop killer whale attacks"
8336:"U.S. Navy Sonar May Harm Killer Whales, Expert Says"
8066:, Center for Whale Research. Retrieved March 23, 2012
7573:"Cetaceans Have Complex Brains for Complex Cognition"
6814:
Ford, J. K. B.; Ellis, G. M.; Balcomb, K. C. (1999).
6525:
6493:
Baird, Robert W.; Baird, Robin W. (August 31, 2006).
4714:
4702:
4232:
4205:
3796:
3360:
Whales and Dolphins of the Southern African Subregion
2652:, and died a year into his release after contracting
2507:
2363:, orcas may embody the souls of deceased chiefs. The
2100:, the orca is particularly at risk of poisoning from
1607:, often lifting their entire bodies out of the water.
598:
means "of the kingdom of the dead", or "belonging to
9742:
6822:. University of British Columbia Press. p. 25.
6816:
Killer Whales: The Natural History and Genealogy of
6759:. Keep Whales Wild. January 14, 2011. Archived from
6623:
4624:
4117:
3805:
3729:
3451:
3449:
2879:
2630:
individuals. Following the success of the 1993 film
1662:
Orcas are notable for their complex societies. Only
1559:
but are more likely to kill and leave them uneaten.
1374:
and chase them. Nevertheless, possible predation on
1113:
In the Antarctic, orcas range up to the edge of the
10609: – Orcas in Johnstone Strait, British Columbia
9327:
9148:Hoare, Philip; Hoekendijk, Jeroen (June 21, 2023).
8968:
The Worst Journey in the World: Antarctic 1910–1913
7997:
7985:
7243:
7021:
6965:
6848:
6836:
6792:
6775:
6309:
5041:
4655:
4645:"Canada Finds Killer Whales Drawn to Warmer Arctic"
3735:
2965:– a New Zealand biologist who swims with wild orcas
1326:Orcas are sophisticated and effective predators of
766:that had attracted a great deal of local interest.
10550:
9425:
8057:How Southern Resident Killer Whales are Identified
7548:Scientist Has 'Snowball Fight' With a Killer Whale
5510:) and a possible case of competitive displacement"
5209:
5104:Journal of Experimental Marine Biology and Ecology
4852:
4772:
4077:
3100:Reeves, R.; Pitman, R. L.; Ford, J. K. B. (2017).
10317:"Selective foraging by fish-eating killer whales
10095:Zimmerman, Tim (2011). "The Killer in the Pool".
9690:"Stranding Base (1901–2012) for Hyogo Prefecture"
9288:"ABC News: Killer Whale Attacks SeaWorld Trainer"
7685:
5886:"Prey of killer whales (Orcinus orca) in Iceland"
3446:
2336:indigenous peoples of the Pacific Northwest Coast
2152:In 2005, the United States government listed the
516:; some populations are composed of highly stable
13121:
10570:
10468:(NMFS) Northwest Regional Office. Archived from
10293:
9546:"Orphaned orca's reunion with family celebrated"
8598:
8521:"Unique Killer-Whale Pod Doomed by Exxon Valdez"
8399:
8261:
8126:
7727:Bowden, D. M.; Williams, D. D. (1984). "Aging".
7681:
7679:
7623:
7181:The Journal of the Acoustical Society of America
6869:
6813:
6543:
4412:
3790:
3660:(4) (2001), 676–701. Retrieved January 26, 2010.
3479:
3382:
3264:
3099:
2859:. Between 1991 and 2010, the bull orca known as
2516:since the 1970s, some of which have been fatal.
2263:. In the mid-1990s, loud underwater noises from
2156:community as an endangered population under the
1611:Day-to-day orca behaviour generally consists of
581:captive orcas killing or injuring their handlers
10640:, HowStuffWorks.com, article by Jacob Silverman
10528:. Vol. 166, no. 2. pp. 220–237.
9829:
9538:
9404:
9147:
8730:. Vol. I., pp. 439, 441. Moscow and Leningrad:
8313:, Associated Press. USA Today, October 25, 2008
8298:
8296:
6986:
6486:
6356:Proceedings of the National Academy of Sciences
4148:
1880:have 28 complex burst-pulse and whistle calls.
10355:
9825:
9823:
9458:
8965:
8586:
7791:"Orcas don't do well in captivity. Here's why"
7726:
6940:
5457:
5412:
5410:
5408:
4144:
4142:
4140:
4138:
4136:
4134:
4132:
4010:"List of Marine Mammal Species and Subspecies"
3383:Leatherwood, Stephen; Hobbs, Larry J. (1988).
3125:10.2305/IUCN.UK.2017-3.RLTS.T15421A50368125.en
2851:that are not part of wild orca behaviour, see
530:International Union for Conservation of Nature
10717:
10386:
10300:(Second ed.). Vancouver, BC: UBC Press.
9482:
8927:
8433:"State expert urges Navy to stop sonar tests"
8304:Researchers: 7 Orcas Missing from Puget Sound
7676:
6743:
6156:
4567:
4565:
3670:
3668:
3666:
3327:Liddell, Henry George; Scott, Robert (1940).
2570:'s research team filming orcas in New Zealand
1510:, orcas feed on South American sea lions and
10056:
9431:
9370:"Naval War Declared Against Killer Whales".
9250:
8329:
8327:
8325:
8323:
8321:
8319:
8293:
6673:
6586:
6331:Killer Whales Develop a Taste For Sea Otters
6195:
5818:
5419:"Running scared: when predators become prey"
5280:
5278:
5254:
4474:
4472:
4319:. Vol. 39, no. 1. pp. 54–59.
4315:Stewart, Doug (2001). "Tales of two orcas".
4296:. Vol. 44, no. 6. pp. 22–30.
3326:
3011:
2718:catch of 916 in 1979–80 alone, prompted the
10661:Orcas Preying On Dolphins (Caught On Drone)
10315:Ford, John K. B.; Ellis, Graeme M. (2006).
9820:
9583:. Seattle, Washington: Hearst Corporation.
9282:
9280:
8471:Journal of Cetacean Research and Management
7774:Physiological Basis of Aging and Geriatrics
7561:, February 6, 2009. Retrieved March 7, 2010
7445:
7443:
6249:) hunt seals and a penguin on floating ice"
5405:
4129:
3688:Journal of Cetacean Research and Management
3674:
3389:. Courier Dover Publications. p. 118.
3296:Lewis, Charlton T.; Short, Charles (1879).
2811:, was one of the oldest orcas in captivity.
2543:
2489:Orca attacks on sailboats and small vessels
2435:Attacks by wild orcas on humans and animals
2304:
2120:show higher levels of PCBs, pesticides and
1603:Orcas, like this one near Alaska, commonly
10731:
10724:
10710:
10235:
9014:
8767:
8765:
8138:
7952:
7891:
7889:
7822:
7776:(Fourth ed.). CRC Press. p. 161.
7468:
6712:"Orcas Kill, But Not Just for Food (2:06)"
6667:
6492:
6168:
5468:off Bahia de Los Angeles, Baja California"
5150:maximum dive depths were 767.5 and 499.5 m
4618:
4571:
4562:
4463:
4383:
4381:
4379:
4377:
4375:
4340:
3663:
3467:
3295:
3240:"Orcas: How Science Debunked Superstition"
2753:
2342:, history, spirituality and religion. The
1400:Orcas also prey on larger species such as
750:Piscium & aquatilium animantium natura
374:
196:
170:
143:
12217:
12171:
10571:Kirkevold, B. C.; Lockard, J. S. (1986).
10487:
10346:
10314:
10099:. Houghton Mifflin Harcourt. p. 336.
10094:
9855:
9654:
9648:
9526:Movie-Star Whale Keiko Dies of Pneumonia.
8935:"Orca shares the waves with local surfer"
8863:
8801:
8799:
8742:
8740:
8316:
8199:
8139:Carrington, Damian (September 27, 2018).
7925:
7859:
7711:
7600:
7590:
7502:
7451:Whale Attack Renews Captive Animal Debate
7423:
7405:
7059:
7057:
6941:Lesté-Lasserre, Christa (June 17, 2021).
6917:
6907:
6469:
6385:
6375:
6202:), type B, in Antarctic Peninsula waters"
6095:
5960:
5919:
5909:
5852:
5821:"Occurrence of long-finned pilot whales (
5789:
5787:
5785:
5783:
5781:
5659:
5625:
5587:
5442:
5275:
5189:
5179:
5131:
4942:
4907:
4822:
4800:, sha-chi.jp. Retrieved February 17, 2010
4723:
4678:"Southern Resident Killer Whale Research"
4556:
4469:
4268:
4250:
4060:
3985:
3882:
3759:
3598:
3540:
3201:
3123:
3012:Mead, J. G.; Brownell, R. L. Jr. (2005).
2589:Moby Doll's impact in scientific research
2519:
2355:for slaves and dolphins for warriors. In
2297:Orcas are included in Appendix II of the
1425:Prior to the advent of industrial whaling
815:member of the Delphinidae, with only the
9830:Reeves, Randall; Whitehead, Hal (2005).
9607:"海域自然環境保全基礎調査 - 海棲動物調査報告書, (2)- 19. シャチ
9277:
8835:
8728:Grammar of the language of Asian Eskimos
8551:"Marine Ecology Progress Series 356:269"
8461:
8333:
8272:
8270:
7617:
7456:, March 1, 2010. Retrieved March 7, 2010
7440:
6820:in British Columbia and Washington State
6679:
6349:
5566:
5474:. La Paz, B.C.S., México. Archived from
4292:Olsen, Ken (2006). "Orcas on the edge".
4149:Heyning, J. E.; Dahlheim, M. E. (1988).
3620:
3618:
3416:
3277:Killer Whales. Scientific Classification
3215:. Whale & Dolphin Conservation USA.
2799:
2681:
2656:, at the age of 27. In 2002, the orphan
2640:was returned to the coast of his native
2606:was successfully returned to her family.
2597:
2553:
2444:
2319:
2046:
1969:
1921:
1693:
1598:
1446:Orca beaching to capture sea lion along
1388:
1104:
1022:
964:
929:
614:) for these animals, possibly borrowing
544:. Some local populations are considered
11996:
10171:
9574:
9314:SeaWorld trainer killed by killer whale
8869:
8762:
8430:
8365:
8363:
8361:
8230:
8151:from the original on September 28, 2018
7886:
7771:
6645:. Associated Press. September 7, 2005.
6104:
6071:
6027:Pitman, Robert L.; et al. (2001).
5883:
5495:
5493:
5096:"Movement and diving of killer whales (
4736:
4734:
4732:
4372:
4314:
3856:
3850:
3675:Pitman, Robert L.; Ensor, Paul (2003).
3188:Integrated Taxonomic Information System
3144:
3142:
1523:and other prey by "wave-hunting". They
1274:appear to be important prey, including
1195:Resident orca pursuing a chinook salmon
901:Four types have been documented in the
14:
13122:
12134:
11073:
10356:Francis, Daniel; Hewlett, Gil (2007).
10180:from the original on December 13, 2020
10076:from the original on November 21, 2013
10012:from the original on December 30, 2017
9700:from the original on December 24, 2014
9670:from the original on December 24, 2014
9349:, December 1956, p. 19. Reproduced at
9294:from the original on February 28, 2010
9002:from the original on February 18, 2024
8796:
8737:
8720:
8700:
8610:
8113:from the original on January 16, 2020.
7624:Rendell, Luke; Whitehead, Hal (2001).
7570:
7480:
7348:
7311:
7299:from the original on February 27, 2020
7286:
7054:
6692:from the original on December 10, 2019
6113:"Ecological effects of marine mammals"
6077:
6059:from the original on December 11, 2015
6026:
5942:
5793:
5778:
5626:Jorgensen, S. J.; et al. (2019).
5372:
5330:) Predation on a Shortfin Mako Shark (
5297:) Specialising on Elasmobranch Prey".
5284:
5230:"National Geographic creature feature"
5227:
4870:
4089:), type D, from subantarctic waters".
3522:
3320:
2309:
2225:, led the lawsuit and represented the
1259:group, herring are often caught using
1066:coast, in the north Pacific along the
905:, Types A–D. Two dwarf species, named
12603:
12602:
11113:
10795:
10705:
10387:Heimlich, Sara; Boran, James (2001).
10202:
9890:from the original on October 31, 2019
9556:from the original on November 6, 2015
9488:
9470:
9211:
9116:
9022:"Boy survives bump from killer whale"
8924:, 1855.) Retrieved February 19, 2010.
8645:Religion in Newfoundland and Labrador
8625:from the original on December 5, 2023
8267:
7874:from the original on October 15, 2018
7866:Podt, Annemieke (December 31, 2016).
7828:
7317:
6953:from the original on December 5, 2022
6881:
6629:
6555:
6531:
6318:
6196:Pitman, R. L.; Durban, J. W. (2011).
6183:
6110:
5354:from the original on January 26, 2012
4962:from the original on October 23, 2017
4913:
4664:
4630:
4291:
4123:
3625:
3615:
3473:
3246:from the original on January 31, 2017
3237:
3219:from the original on February 8, 2022
3160:from the original on December 5, 2017
2955:– occupied a similar ecological niche
2239:International Fund for Animal Welfare
1094:, in the North Atlantic Ocean around
769:The orca is one of 35 species in the
13130:IUCN Red List data deficient species
13105:0B8C8927-FB2B-4564-85A7-769DE506635F
12921:8a4a806c-d997-4960-9e7a-82ac875054aa
12033:
10548:
10521:
10448:
10260:
9997:
9985:
9730:from the original on January 9, 2015
9587:from the original on August 11, 2011
9494:
9333:
8992:"Whale Takes Bite From Surfer's Leg"
8943:. September 12, 2008. Archived from
8663:
8611:Gibson, Caitlin (December 5, 2023).
8531:from the original on January 6, 2010
8518:
8443:from the original on October 4, 2012
8358:
8276:
8003:
7991:
7865:
7847:from the original on January 3, 2017
7320:"Killer whales: How smart are they?"
7231:
6974:
6857:
6842:
6801:
6786:
6245:"Antarctic peninsula killer whales (
5798:. Academic Press. pp. 927–930.
5490:
5215:
4974:
4858:
4778:
4729:
4708:
4642:
4581:Whales, whaling and ocean ecosystems
4392:. Academic Press. pp. 550–556.
4387:
4356:. Vancouver Aquarium. Archived from
3357:
3308:from the original on October 7, 2022
3231:
3139:
2730:permit small subsistence hunts (see
2393:people, orcas are said to appear as
1321:
1012:
12079:
10573:Behavioral Biology of Killer Whales
9967:from the original on March 30, 2019
8812:(in Russian). Эхо Москвы. Арсенал.
8709:, p. 156 in Rubcova, E. S. (1954).
8431:McClure, Robert (October 2, 2003).
8083:from the original on March 23, 2018
7508:
7287:Szondy, David (February 26, 2020).
6649:from the original on March 22, 2010
6568:from the original on August 7, 2023
6224:from the original on August 2, 2023
5228:Hughes, Catherine D. (March 2014).
4643:Kwan, Jennifer (January 22, 2007).
4600:from the original on August 7, 2020
4002:
3717:from the original on April 27, 2020
3434:from the original on March 25, 2017
3209:"Facts about orcas (killer whales)"
3111:IUCN Red List of Threatened Species
3005:
2702:industry caught immense numbers of
2449:Male orca depicted in St Mary's in
1657:
37:"Killer whale" redirects here. For
24:
10631:Why are orca called killer whales?
10515:
10123:from the original on June 13, 2012
9771:from the original on July 17, 2015
9627:from the original on July 17, 2015
9575:McClure, Robert (March 11, 2006).
9495:Wood, Daniel (February 24, 2010).
9098:from the original on June 20, 2023
9038:from the original on April 2, 2011
8895:LacusCurtius: Into the Roman World
8500:from the original on March 6, 2014
8381:from the original on June 26, 2012
8243:from the original on July 10, 2018
7868:"Orca Granny: was she really 105?"
7741:10.1016/B978-0-12-039228-5.50015-2
7571:Marino, Lori; et al. (2007).
7361:from the original on July 19, 2021
7153:from the original on July 15, 2011
6882:Weiss, M. N.; et al. (2021).
6680:Connelly, Laylan (July 30, 2019).
6556:Chung, Emily (February 18, 2019).
6513:from the original on July 21, 2011
6111:Estes, James (February 26, 2009).
6053:10.1111/j.1748-7692.2001.tb01000.x
5980:) killing and eating blue whales (
5819:Selbmann, A.; et al. (2022).
5539:10.1111/j.1748-7692.1999.tb00822.x
5399:10.1111/j.1748-7692.1999.tb00793.x
5140:from the original on June 30, 2022
5100:) at a Southern Ocean archipelago"
5072:from the original on July 20, 2018
4914:Baird, R. W.; et al. (2006).
4810:Ten Years after Taiji Orca Capture
3838:from the original on July 22, 2011
3829:10.1111/j.1365-2907.1991.tb00291.x
3502:10.1111/j.1748-7692.1999.tb00833.x
2508:Attacks on humans by captive orcas
2096:Like other animals at the highest
1814:Problems playing these files? See
1747:
1654:to exhaustion, but not eating it.
1579:. Orcas in many areas may prey on
1474:Other marine mammal prey includes
1386:are among Carribean Orcas' prey.
925:
25:
13166:
11746:Indo-Pacific bottlenose dolphin (
10593:
10466:National Marine Fisheries Service
10434:and National Science Foundation.
10432:Smithsonian Institution Libraries
9577:"Luna the orca killed by tugboat"
9550:Canadian Broadcasting Corporation
9507:from the original on May 13, 2012
9265:from the original on July 8, 2023
9232:from the original on July 4, 2023
9193:from the original on July 4, 2023
9129:from the original on July 5, 2023
9068:from the original on July 4, 2023
8771:
7834:
7664:from the original on May 30, 2010
7481:Weston, Phoebe (March 10, 2023).
7349:Dunham, Will (October 16, 2017).
7070:Journal of Comparative Psychology
6722:from the original on June 2, 2023
5744:African Journal of Marine Science
5689:African Journal of Marine Science
5460:"A fatal attack on a whale shark
5236:from the original on June 2, 2023
4791:Ecology of Japanese Coastal Orcas
4754:National Marine Fisheries Service
4752:. Office of Protected Resources,
3859:"Food chains: Killer in the kelp"
3630:Status of Killer Whales in Canada
3339:from the original on May 26, 2024
2773:
2384:Port au Choix Archaeological Site
2243:Raincoast Conservation Foundation
2033:occurs in other carnivore species
1974:Adult female orca with calf near
758:, based on examination of a dead
27:Largest living species of dolphin
10172:Stevens, Blair (June 11, 2020).
10165:
10149:. March 17, 2016. Archived from
10135:
10103:
10088:
10024:
9991:
9953:
9872:
9783:
9712:
9682:
9599:
9568:
9519:
9363:
9339:
9306:
9244:
9205:
9175:
9141:
9110:
9080:
9050:
8998:. MONTEREY. UPI. July 11, 1972.
8984:
8959:
8829:
8816:from the original on May 5, 2008
8657:
8637:
8604:
8543:
8519:Keim, Brandon (March 24, 2009).
8512:
8455:
8424:
8405:
8277:Lyke, M. L. (October 14, 2006).
8224:
8163:
8132:
8095:
8069:
8050:
8026:
8009:
7946:
7797:. March 25, 2019. Archived from
7765:
7720:
7713:10.1111/j.1365-2664.2009.01647.x
7626:"Culture in whales and dolphins"
7564:
7540:
7474:
7373:
7342:
7280:
7237:
7165:
7096:
7048:10.1111/j.1439-0310.2008.01496.x
7015:
6980:
6934:
6875:
6807:
6749:
6704:
6635:
6580:
6549:
6499:. Voyageur Press. pp. 23–.
6437:
6402:
6276:10.1111/j.1748-7692.2007.00163.x
6218:10.1111/j.1748-7692.2010.00453.x
6133:from the original on May 9, 2016
4572:Forney, K. A.; Wade, P. (2007).
3752:10.1111/j.1469-185X.2012.00239.x
3242:. National Wildlife Federation.
2985:
2924:
2910:
2896:
2882:
2720:International Whaling Commission
2389:In the tales and beliefs of the
1799:
1781:
1763:
1728:
1456:
1439:
1188:
1170:
704:
686:
664:
238:
75:
11887:Indo-Pacific finless porpoise (
11764:Tamanend's bottlenose dolphin (
11612:Indian Ocean humpback dolphin (
11603:Indo-Pacific humpback dolphin (
10670:, off the coast of San Clemente
9939:10.1016/j.marpolbul.2011.01.002
9290:. ABC News. November 30, 2006.
9117:Dewan, Pandora (June 9, 2023).
8966:Cherry-Garrard, Apsley (2004).
8732:Academy of Sciences of the USSR
8715:Academy of Sciences of the USSR
6343:
6324:
6289:
6236:
6189:
6115:. In Perrin, William F. (ed.).
6020:
5969:
5936:
5877:
5812:
5731:
5676:
5619:
5560:
5451:
5366:
5317:
5248:
5221:
5155:
5087:
5035:
4864:
4836:. Marinebio.org. Archived from
4803:
4784:
4670:
4636:
4498:
4418:
4346:
4308:
4285:
4226:
4212:. Academic Press. p. 189.
4199:
4024:
3945:
3907:
3677:"Three forms of killer whales (
3557:
3516:
3410:
3376:
3351:
3289:
2976:
2042:
1883:
1738:Multimedia relating to the orca
705:
52:. For 1962 tokusatsu film, see
11433:Southern right whale dolphin (
11424:Northern right whale dolphin (
11356:Atlantic white-sided dolphin (
10649:Orca devours great white shark
10326:Marine Ecology Progress Series
10268:Encyclopedia of Marine Mammals
10196:
10097:The Best American Sampler 2011
8707:The orphan boy with his sister
8599:Ford, Ellis & Balcomb 2000
8400:Ford, Ellis & Balcomb 2000
8262:Ford, Ellis & Balcomb 2000
8127:Ford, Ellis & Balcomb 2000
7318:Spear, Kevin (March 7, 2010).
6870:Ford, Ellis & Balcomb 2000
6544:Ford, Ellis & Balcomb 2000
6117:Encyclopedia of Marine Mammals
5796:Encyclopedia of Marine Mammals
5168:ICES Journal of Marine Science
4482:. Seaworld.org. Archived from
4413:Ford, Ellis & Balcomb 2000
4390:Encyclopedia of Marine Mammals
3791:Ford, Ellis & Balcomb 2000
3362:. Cambridge University Press.
3270:
3265:Ford, Ellis & Balcomb 2000
3172:
3048:
2852:
2832:Whale and Dolphin Conservation
2762:, including the male known as
2190:U.S. Navy sonar may harm orcas
1987:animals that undergo menopause
1370:, which have been recorded to
566:conflicts with human fisheries
455:and the largest member of the
13:
1:
13145:Animals that use echolocation
12470:Deraniyagala's beaked whale (
12443:Ginkgo-toothed beaked whale (
11654:Pantropical spotted dolphin (
11621:Australian humpback dolphin (
11383:Pacific white-sided dolphin (
10613:Salish Sea Hydrophone Network
9501:The Christian Science Monitor
9351:Longevity and Causes of Death
9251:Scott Neuman (May 15, 2024).
9212:Crisp, James (May 27, 2023).
8726:Menovshchikov, G. A. (1962).
8334:Pickrell, John (March 2004).
8231:Robbins, Jim (July 9, 2018).
7630:Behavioral and Brain Sciences
7136:10.1080/09524622.2007.9753581
7001:10.1016/j.anbehav.2004.04.014
5764:10.2989/1814232X.2024.2311272
5709:10.2989/1814232X.2022.2066723
5502:"Predation on a white shark (
4524:Journal of Applied Physiology
4037:) indicates multiple species"
3655:The Canadian Field-Naturalist
3238:Price, Mary (July 22, 2013).
2998:
2940:List of marine mammal species
1965:
1722:behave more like transients.
1152:
1109:Variations in Antarctic orcas
11466:Australian snubfin dolphin (
10874:North Atlantic right whale (
8970:. Globe Pequot. p. 92.
8754:(in Russian). Archived from
7592:10.1371/journal.pbio.0050139
7407:10.1371/journal.pone.0226206
6431:10.1016/j.pocean.2006.02.007
5911:10.1371/journal.pone.0207287
4893:10.1126/science.331.6015.274
4834: – Orca (Killer Whale)"
4018:Society for Marine Mammalogy
2969:
2789:
2285:adversely affected orcas in
2128:. A 2018 study published in
1594:
1344:Pacific white-sided dolphins
1250:are also eaten, but smaller
962:orcas have been documented.
879:Society for Marine Mammalogy
817:Atlantic white-sided dolphin
752:of 1558, part of the larger
574:U.S. Endangered Species list
7:
13155:Taxa named by Carl Linnaeus
12416:Blainville's beaked whale (
12357:Tropical bottlenose whale (
12333:Southern bottlenose whale (
12324:Northern bottlenose whale (
11773:Common bottlenose dolphin (
11630:Atlantic humpback dolphin (
10883:North Pacific right whale (
10423:Mammals of the Soviet Union
9446:New York Zoological Society
9419:New York Zoological Society
8888:September 15, 2020, at the
7553:September 15, 2020, at the
5949:Frontiers in Marine Science
5293:) Sharks by Killer Whales (
5257:Canadian Journal of Zoology
5124:10.1016/j.jembe.2015.08.008
5051:Canadian Journal of Zoology
4536:10.1152/jappl.1967.22.5.974
3456:Zum Wal in der Marienkirche
3426:(in Latin). Vol. v.1 (
3335:. Perseus Digital Library.
3304:. Perseus Digital Library.
2875:
2712:Ministry of the Environment
2636:, the movie's captive star
2470:
2122:brominated flame-retardants
1670:live in comparably complex
795:to the Lissodelphininae, a
655:
10:
13171:
12515:Stejneger's beaked whale (
11896:Yangtze finless porpoise (
11681:Atlantic spotted dolphin (
11266:Short-finned pilot whale (
10691:November 16, 2019, at the
10636:February 19, 2023, at the
10206:Killer Whales of the World
10111:"Corpse Is Found on Whale"
9722:. The Whale Laboratory of
9692:. The Whale Laboratory of
9581:Seattle Post-Intelligencer
9459:Francis & Hewlett 2007
8892:(Latin), in Bill Thayer's
8838:(北海道)「シャチ送り」の遺構か 礼文島の遺跡で出土
8587:Francis & Hewlett 2007
8437:Seattle Post Intelligencer
8283:Seattle Post Intelligencer
8062:November 28, 2009, at the
7772:Timiras, Paola S. (2013).
7692:Journal of Applied Ecology
6078:Pitman, Robert L. (2016).
5845:10.1007/s10211-022-00394-1
5652:10.1038/s41598-019-39356-2
5567:Heithaus, Michael (2001).
5458:O'Sullivan, J. B. (2000).
5377:) in New Zealand waters".
3958:Royal Society Open Science
2963:Ingrid Visser (researcher)
2793:
2675:
2671:
2547:
2492:
2438:
2313:
1887:
1827:
1395:strap-toothed beaked whale
1308:broadnose sevengill sharks
1161:
1019:Orca types and populations
1016:
833:Orca types and populations
830:
619:
36:
29:
13150:Mammals described in 1758
13035:
12611:
12560:
12548:Shepherd's beaked whale (
12536:
12369:
12345:
12304:
12262:
12245:
12208:
12162:
12125:
12091:Araguaian river dolphin (
12070:
12024:
11987:
11959:
11908:
11867:
11850:
11822:
11798:
11789:
11726:
11702:
11642:
11583:
11550:
11526:
11502:
11478:
11445:
11404:
11335:
11311:
11287:
11275:Long-finned pilot whale (
11246:
11222:
11198:
11147:
11130:
11126:
11109:
11064:
11036:
11012:
10916:
10899:
10845:
10821:
10812:
10808:
10791:
10741:
10261:Ford, John K. B. (2002).
10236:Carwardine, Mark (2001).
9919:Marine Pollution Bulletin
9483:Heimlich & Boran 2001
8774:"Siberian Yupik (Eskimo)"
8309:October 28, 2008, at the
7642:10.1017/S0140525X0100396X
7529:: 5. 2005. Archived from
7266:10.1007/s10211-007-0029-7
7109:from Southeast Kamchatka"
7026:) call type production".
6744:Heimlich & Boran 2001
6609:10.1007/s00300-010-0853-5
6336:November 7, 2015, at the
6157:Heimlich & Boran 2001
5962:10.3389/fmars.2021.648421
5598:10.1017/S0952836901000061
5014:10.1017/S002531541000072X
4647:. Reuters. Archived from
4103:10.1007/s00300-010-0871-3
3641:Status of Killer Whales,
3635:November 8, 2011, at the
2338:feature orcas throughout
2106:Polychlorinated biphenyls
2078:Depletion of prey species
1957:groups of killer whales (
1539:been observed preying on
1384:short-finned pilot whales
1382:, and one study suggests
1078:off much of the coast of
586:
405:
398:
382:
373:
350:
343:
235:Scientific classification
233:
216:
194:
185:
178:
169:
151:
142:
63:
12434:Gervais's beaked whale (
12389:Sowerby's beaked whale (
12100:Bolivian river dolphin (
10605:August 15, 2000, at the
10426:. Vol. II, part 3.
10203:Baird, Robin W. (2002).
10147:National Geographic News
9662:
9436:, in British Columbia".
8837:
8808:
8340:National Geographic News
7248:) that feed on herring (
6411:Progress in Oceanography
6350:Springer, A. M. (2003).
6295:Pinell, Nadine, et al. "
5884:Samarra, Filipa (2018).
5334:) in New Zealand Waters"
4796:August 22, 2009, at the
3523:Horreo, Jose L. (2018).
3282:August 15, 2012, at the
3020:; Reeder, D. M. (eds.).
2760:orcas of Eden, Australia
2550:Orcas in popular culture
2544:Modern Western attitudes
2305:Relationship with humans
1868:Norwegian and Icelandic
1795:Vocalizations of an orca
1777:Orca calls at a distance
1496:South American sea lions
1376:long-finned pilot whales
919:mitochondrial sequencing
826:
12572:Cuvier's beaked whale (
12497:Perrin's beaked whale (
12461:Hector's beaked whale (
12425:Ramari's beaked whale (
12398:Andrews' beaked whale (
12274:Arnoux's beaked whale (
11947:Burmeister's porpoise (
11714:Rough-toothed dolphin (
11138:(Oceanic dolphins)
10937:Antarctic minke whale (
10679:April 24, 2020, at the
10666:March 17, 2021, at the
9372:The Science News-Letter
8643:Rollmann, Hans (1999).
8417:September 11, 2012, at
8192:10.1126/science.aat1953
8043:April 19, 2011, at the
6377:10.1073/pnas.1635156100
5943:Kiszka, Jeremy (2021).
5311:10.1578/AM.31.1.2005.83
4014:MarineMammalScience.org
3932:10.1578/am.50.2.2024.93
3333:A Greek-English Lexicon
2828:World Animal Protection
2754:Cooperation with humans
2361:Kwakwaka'wakw mythology
2227:David Suzuki Foundation
1698:A pair of orcas in the
1514:in shallow water, even
1512:southern elephant seals
1221:
857:reported in 2008, "The
723:is the only recognized
648:, whose only member is
570:southern resident orcas
552:due to prey depletion,
459:family. It is the only
428:(Bonnaterre, 1789)
12283:Baird's beaked whale (
12183:Ganges river dolphin (
12109:Amazon river dolphin (
11490:Orca or killer whale (
11414:(Right whale dolphins)
11347:White-beaked dolphin (
10865:Southern right whale (
10654:April 5, 2012, at the
9848:10.1098/rsbl.2005.0348
9356:June 19, 2010, at the
9320:April 3, 2010, at the
8876:Gaius Plinius Secundus
8836:Miyanaga, T. (2014).
8483:10.47536/jcrm.v4i3.844
8462:Williams, Rob (2023).
8289:on September 15, 2020.
7918:10.1093/jmammal/gyv113
6900:10.1098/rspb.2021.0617
6462:10.1098/rstb.2008.0231
6302:June 30, 2017, at the
5504:Carcharodon carcharias
5181:10.1093/icesjms/fsy118
5031:on September 30, 2014.
4815:March 1, 2014, at the
3857:Schrope, Mark (2007).
3700:10.47536/jcrm.v5i2.813
3681:) in Antarctic waters"
3650:July 14, 2010, at the
2826:Organizations such as
2812:
2695:
2646:Earth Island Institute
2607:
2581:southern resident orca
2571:
2520:Human attacks on orcas
2454:
2331:
2219:British Columbia Coast
2207:decompression sickness
2158:Endangered Species Act
2066:conservation dependent
2053:
1978:
1963:
1927:
1752:
1702:
1608:
1397:
1110:
1031:
970:
935:
771:oceanic dolphin family
693:Modern orca skeleton,
12968:Paleobiology Database
12524:Spade-toothed whale (
12488:True's beaked whale (
12479:Strap-toothed whale (
12452:Gray's beaked whale (
12407:Hubbs' beaked whale (
12292:Sato's beaked whale (
12192:Indus river dolphin (
11920:Spectacled porpoise (
11736:(Bottlenose dolphins)
11177:Heaviside's dolphin (
11159:Commerson's dolphin (
10964:Pygmy Bryde's whale (
10674:Orcas vs Sperm Whales
10618:July 9, 2013, at the
9649:Obee & Ellis 1992
9531:June 7, 2020, at the
8912:June 4, 2011, at the
8866:, pp. Chapter 1.
8864:Obee & Ellis 1992
8613:"The call of Tokitae"
8038:) in British Columbia
8021:Know the Killer Whale
7959:Marine Mammal Science
7503:Obee & Ellis 1992
6256:Marine Mammal Science
6206:Marine Mammal Science
6084:Marine Mammal Science
6033:Marine Mammal Science
5986:Marine Mammal Science
5982:Balaenoptera musculus
5825:) and killer whales (
5518:Marine Mammal Science
5506:) by a killer whale (
5379:Marine Mammal Science
5191:10536/DRO/DU:30120094
4935:10.1353/psc.2006.0024
4724:Ford & Ellis 2006
4557:Obee & Ellis 1992
4506:Delphinapterus luecas
4053:10.1101/gr.102954.109
3583:10.1093/sysbio/syz068
3482:Marine Mammal Science
3118:: e.T15421A50368125.
2803:
2685:
2601:
2566:
2482:Terra Nova Expedition
2448:
2439:Further information:
2323:
2231:Environmental Defence
2104:of toxins, including
2050:
1973:
1951:
1925:
1890:Cetacean intelligence
1751:
1697:
1642:Orcas will engage in
1602:
1506:, Argentina, and the
1392:
1378:has been recorded in
1108:
1026:
1017:Further information:
968:
933:
831:Further information:
737:in his landmark 1758
727:species in the genus
463:species in the genus
420:Bonnaterre, 1789
153:Transient orcas near
32:Orca (disambiguation)
13140:Cosmopolitan mammals
12506:Pygmy beaked whale (
12379:(Mesoplodont whales)
11538:False killer whale (
11514:Melon-headed whale (
11234:Pygmy killer whale (
10928:Common minke whale (
10549:Hoyt, Erich (1998),
10321:in British Columbia"
10032:"Orcas in captivity"
9998:Rose, N. A. (2011).
9963:. November 8, 2017.
9758:Asian Marine Biology
9611:(Limaeus,1758)マイルカ科"
9187:Practical Boat Owner
8650:May 4, 2009, at the
8346:on September 9, 2011
8109:. January 14, 2020.
7906:Journal of Mammalogy
7536:on October 11, 2007.
7516:"Killer whale games"
7330:on September 8, 2015
6763:on December 16, 2010
5478:on February 28, 2010
4486:on September 4, 2013
4195:on January 18, 2012.
3358:Best, P. B. (2007).
3150:"Appendices | CITES"
3070:on December 12, 2021
2768:Prince William Sound
2742:being killed in the
2620:photo-identification
2602:In 2002, the orphan
2495:Iberian orca attacks
2287:Prince William Sound
2247:Wilderness Committee
1650:playing with a male
1589:Marineland of Canada
1587:. A captive orca at
1488:California sea lions
1284:short-tail stingrays
1001:layer of insulating
823:) being more basal.
713:3D model of skeleton
568:. In late 2005, the
532:assesses the orca's
471:cosmopolitan species
30:For other uses, see
12314:(Bottlenose whales)
12054:Dwarf sperm whale (
12045:Pygmy sperm whale (
11877:(Finless porpoises)
11593:(Humpback dolphins)
11457:Irrawaddy dolphin (
11374:Hourglass dolphin (
11085:Pygmy right whale (
10697:National Geographic
10525:National Geographic
10339:2006MEPS..316..185F
9931:2011MarPB..62..792L
9347:Naval Aviation News
9034:. August 18, 2005.
8907:English translation
8784:on October 28, 2016
8758:on August 30, 2007.
8618:The Washington Post
8589:, pp. 115–120.
8377:. October 9, 2008.
8184:2018Sci...361.1373D
8178:(6409): 1373–1376.
7971:2015MMamS..31.1362J
7704:2009JApEc..46..632W
7398:2019PLoSO..1426206R
7193:2006ASAJ..119..627W
7128:2007Bioac..16..261F
7040:2008Ethol.114..599F
6601:2010PoBio..33.1589P
6456:(1524): 1647–1658.
6423:2006PrOce..68..329D
6368:2003PNAS..10012223S
6362:(21): 12223–12228.
6268:2008MMamS..24..225V
6045:2001MMamS..17..494P
5998:2022MMamS..38.1286T
5902:2018PLoSO..1307287S
5756:2024AfJMS..46....1T
5701:2022AfJMS..44..139T
5644:2019NatSR...9.6153J
5607:on January 15, 2016
5531:1999MMamS..15..563P
5464:, by killer whales
5435:2019Ecosp..10E2531E
5391:1999MMamS..15..220V
5116:2015JEMBE.473...90R
5006:2010JMBUK..90.1601W
4885:2011Sci...331..274M
4651:on August 17, 2017.
4452:Heptner et al. 1996
4440:Heptner et al. 1996
4425:Orca (Killer whale)
3978:10.1098/rsos.231368
3970:2024RSOS...1131368M
3875:2007Natur.445..703S
3494:1999MMamS..15..619L
2821:aquatic theme parks
2310:Indigenous cultures
2214:Species at Risk Act
2090:habitat disturbance
2062:conservation status
1577:rockhopper penguins
1541:terrestrial mammals
1340:bottlenose dolphins
954:in the water. Both
944:false killer whales
896:nominate subspecies
845:, or possibly even
821:Leucopleurus acutus
673:Orcinus citoniensis
572:were placed on the
562:marine mammal parks
534:conservation status
510:dolphins and whales
417:Delphinus gladiator
412:Linnaeus, 1758
188:Conservation status
54:Killer Whale (film)
12229:La Plata dolphin (
11929:Harbour porpoise (
11898:N. asiaeorientalis
11755:Burrunan dolphin (
11323:Fraser's dolphin (
11186:Hector's dolphin (
10500:: Whitecap Books.
10370:Harbour Publishing
10348:10.3354/meps316185
10116:The New York Times
8922:Henry Thomas Riley
8881:Historia Naturalis
8778:East Asian Studies
8666:American Antiquity
8237:The New York Times
7942:on April 23, 2019.
7546:Pitman, Robert L.
7449:Associated Press.
6757:"Keep Whales Wild"
6718:. April 19, 2022.
5823:Globicephala melas
5632:Scientific Reports
5576:Journal of Zoology
5289:) and Hammerhead (
4252:10.3390/ani6080049
3739:Biological Reviews
3571:Systematic Biology
3302:A Latin Dictionary
2918:Marine life portal
2813:
2732:Aboriginal whaling
2696:
2690:swims alongside a
2660:was discovered in
2608:
2572:
2455:
2332:
2074:endangered species
2054:
2037:bottlenose dolphin
1979:
1928:
1753:
1703:
1609:
1569:chinstrap penguins
1429:exploding harpoons
1398:
1316:great white sharks
1292:smooth hammerheads
1234:sea specialize in
1141:in Canada and the
1111:
1050:of Argentina, the
1032:
987:isosceles triangle
971:
936:
877:respectively. The
779:pygmy killer whale
775:false killer whale
755:Historia animalium
512:. They are highly
481:to tropical seas.
13115:
13114:
12955:Open Tree of Life
12605:Taxon identifiers
12596:
12595:
12592:
12591:
12588:
12587:
12584:
12583:
12382:
12317:
12256:
12241:
12240:
12204:
12203:
12158:
12157:
12121:
12120:
12066:
12065:
12020:
12019:
11983:
11982:
11971:Dall's porpoise (
11880:
11861:
11846:
11845:
11785:
11784:
11739:
11690:Spinner dolphin (
11672:Striped dolphin (
11663:Clymene dolphin (
11596:
11417:
11365:Peale's dolphin (
11299:Risso's dolphin (
11259:
11168:Chilean dolphin (
11141:
11120:
11105:
11104:
11101:
11100:
11097:
11096:
11060:
11059:
10910:
10895:
10894:
10858:
10802:
10585:978-0-8451-3100-8
10577:Alan R. Liss Inc.
10564:978-0-920656-25-9
10507:978-1-55110-034-0
10441:978-1-886106-81-9
10412:978-0-89658-545-4
10379:978-1-55017-426-7
10307:978-0-7748-0800-2
10253:978-0-7894-8266-2
10228:978-0-7603-2654-1
10153:on April 20, 2016
9988:, pp. 43–44.
9552:. July 13, 2007.
9473:, pp. 73–80.
9461:, pp. 58–59.
9409:(killer whale)".
9378:(24): 374. 1956.
9189:. June 19, 2023.
9027:The Seattle Times
8996:Los Angeles Times
8977:978-1-59228-212-8
8900:May 24, 2012, at
8772:Vajda, Edward J.
8412:Research on Orcas
7979:10.1111/mms.12225
7523:Blackfish Sounder
7234:, pp. 15–16.
7201:10.1121/1.2130934
6595:(11): 1589–1594.
6506:978-0-7603-2654-1
6186:, pp. 61–62.
6126:978-0-08-091993-5
6097:10.1111/mms.12343
6006:10.1111/mms.12906
5921:20.500.11815/1095
5805:978-0-08-091993-5
5548:on March 22, 2012
5444:10.1002/ecs2.2531
5332:Isurus oxyrinchus
5057:(11): 1103–1112.
4879:(6015): 274–276.
4711:, pp. 24–29.
4590:978-0-520-24884-7
4466:, pp. 30–32.
4399:978-0-08-091993-5
4317:National Wildlife
4294:National Wildlife
4219:978-0-12-409592-2
4162:Mammalian Species
3869:(7129): 703–705.
3793:, pp. 16–21.
3542:10.1111/jzs.12255
3369:978-0-521-89710-5
3184:(Linnaeus, 1758)"
3033:978-0-8018-8221-0
2945:List of cetaceans
2564:
2526:Pacific Northwest
2514:attacks on humans
2235:Greenpeace Canada
2154:southern resident
2139:Pacific Northwest
2108:(PCBs). European
2023:Pacific Northwest
1825:
1824:
1804:
1786:
1768:
1700:Pacific Northwest
1676:them in captivity
1672:social structures
1617:surface behaviour
1555:Orcas do consume
1492:Steller sea lions
1393:Orca attacking a
1352:harbour porpoises
1322:Mammals and birds
1177:Tail-slapping in
1013:Range and habitat
983:sexual dimorphism
914:Orcinus glacialis
888:O. o. rectipinnus
884:Orcinus orca ater
764:Bay of Greifswald
631:. As part of the
479:Antarctic regions
435:
434:
228:
211:
16:(Redirected from
13162:
13108:
13107:
13095:
13094:
13082:
13081:
13069:
13068:
13056:
13055:
13054:
13028:
13027:
13015:
13014:
13002:
13001:
12989:
12988:
12976:
12975:
12963:
12962:
12950:
12949:
12937:
12936:
12924:
12923:
12914:
12913:
12901:
12900:
12888:
12887:
12885:NBNSYS0000005173
12875:
12874:
12862:
12861:
12849:
12848:
12836:
12835:
12823:
12822:
12810:
12809:
12797:
12796:
12784:
12783:
12771:
12770:
12758:
12757:
12745:
12744:
12732:
12731:
12722:
12721:
12709:
12708:
12696:
12695:
12683:
12682:
12673:
12672:
12660:
12659:
12647:
12646:
12645:
12632:
12631:
12630:
12600:
12599:
12381:
12380:
12376:
12316:
12315:
12311:
12260:
12259:
12255:
12254:
12250:
12215:
12214:
12169:
12168:
12132:
12131:
12093:I. araguaiaensis
12077:
12076:
12031:
12030:
12010:P. macrocephalus
11994:
11993:
11879:
11878:
11874:
11865:
11864:
11860:
11859:
11855:
11796:
11795:
11738:
11737:
11733:
11595:
11594:
11590:
11571:Guiana dolphin (
11416:
11415:
11411:
11268:G. macrorhynchus
11258:
11257:
11253:
11210:Common dolphin (
11145:
11144:
11140:
11139:
11135:
11128:
11127:
11119:(Toothed whales)
11118:
11111:
11110:
11071:
11070:
11048:Humpback whale (
10930:B. acutorostrata
10914:
10913:
10909:
10908:
10904:
10857:
10856:
10852:
10819:
10818:
10810:
10809:
10800:
10793:
10792:
10726:
10719:
10712:
10703:
10702:
10626:Keep Whales Wild
10589:
10567:
10557:, Camden House,
10556:
10545:
10511:
10498:British Columbia
10484:
10482:
10480:
10475:on June 26, 2008
10474:
10459:
10445:
10416:
10383:
10352:
10350:
10311:
10290:
10257:
10232:
10190:
10189:
10187:
10185:
10169:
10163:
10162:
10160:
10158:
10139:
10133:
10132:
10130:
10128:
10119:. July 7, 1999.
10107:
10101:
10100:
10092:
10086:
10085:
10083:
10081:
10060:
10054:
10053:
10051:
10049:
10043:
10036:
10028:
10022:
10021:
10019:
10017:
10011:
10004:
9995:
9989:
9983:
9977:
9976:
9974:
9972:
9957:
9951:
9950:
9913:
9900:
9899:
9897:
9895:
9876:
9870:
9869:
9859:
9827:
9818:
9817:
9815:
9813:
9808:on March 4, 2016
9807:
9801:. Archived from
9796:
9787:
9781:
9780:
9778:
9776:
9770:
9755:
9746:
9740:
9739:
9737:
9735:
9716:
9710:
9709:
9707:
9705:
9686:
9680:
9679:
9677:
9675:
9658:
9652:
9646:
9637:
9636:
9634:
9632:
9626:
9615:
9603:
9597:
9596:
9594:
9592:
9572:
9566:
9565:
9563:
9561:
9542:
9536:
9523:
9517:
9516:
9514:
9512:
9492:
9486:
9480:
9474:
9468:
9462:
9456:
9450:
9449:
9429:
9423:
9422:
9402:
9396:
9395:
9367:
9361:
9343:
9337:
9331:
9325:
9310:
9304:
9303:
9301:
9299:
9284:
9275:
9274:
9272:
9270:
9248:
9242:
9241:
9239:
9237:
9209:
9203:
9202:
9200:
9198:
9179:
9173:
9172:
9170:
9168:
9145:
9139:
9138:
9136:
9134:
9114:
9108:
9107:
9105:
9103:
9094:. May 29, 2023.
9084:
9078:
9077:
9075:
9073:
9064:. May 23, 2023.
9054:
9048:
9047:
9045:
9043:
9032:Associated Press
9018:
9012:
9011:
9009:
9007:
8988:
8982:
8981:
8963:
8957:
8956:
8954:
8952:
8947:on July 24, 2011
8931:
8925:
8873:
8867:
8861:
8848:
8847:
8833:
8827:
8825:
8823:
8821:
8803:
8794:
8793:
8791:
8789:
8780:. Archived from
8769:
8760:
8759:
8744:
8735:
8724:
8718:
8704:
8698:
8697:
8661:
8655:
8641:
8635:
8634:
8632:
8630:
8608:
8602:
8596:
8590:
8584:
8573:
8572:
8570:
8568:
8563:on July 28, 2011
8562:
8556:. Archived from
8555:
8547:
8541:
8540:
8538:
8536:
8516:
8510:
8509:
8507:
8505:
8499:
8468:
8459:
8453:
8452:
8450:
8448:
8428:
8422:
8409:
8403:
8397:
8391:
8390:
8388:
8386:
8367:
8356:
8355:
8353:
8351:
8342:. Archived from
8331:
8314:
8300:
8291:
8290:
8285:. Archived from
8274:
8265:
8259:
8253:
8252:
8250:
8248:
8228:
8222:
8221:
8203:
8167:
8161:
8160:
8158:
8156:
8136:
8130:
8124:
8115:
8114:
8099:
8093:
8092:
8090:
8088:
8073:
8067:
8054:
8048:
8030:
8024:
8013:
8007:
8001:
7995:
7989:
7983:
7982:
7965:(4): 1362–1377.
7950:
7944:
7943:
7938:. Archived from
7929:
7912:(5): 1055–1070.
7893:
7884:
7883:
7881:
7879:
7863:
7857:
7856:
7854:
7852:
7832:
7826:
7820:
7811:
7810:
7808:
7806:
7787:
7778:
7777:
7769:
7763:
7762:
7724:
7718:
7717:
7715:
7683:
7674:
7673:
7671:
7669:
7621:
7615:
7614:
7604:
7594:
7568:
7562:
7544:
7538:
7537:
7535:
7520:
7512:
7506:
7500:
7491:
7490:
7478:
7472:
7466:
7457:
7447:
7438:
7437:
7427:
7409:
7392:(12): e0226206.
7377:
7371:
7370:
7368:
7366:
7346:
7340:
7339:
7337:
7335:
7326:. Archived from
7324:Orlando Sentinel
7315:
7309:
7308:
7306:
7304:
7284:
7278:
7277:
7241:
7235:
7229:
7220:
7219:
7218:on May 27, 2011.
7217:
7211:. Archived from
7178:
7169:
7163:
7162:
7160:
7158:
7152:
7113:
7100:
7094:
7093:
7082:10.1037/a0028858
7061:
7052:
7051:
7019:
7013:
7012:
6989:Animal Behaviour
6984:
6978:
6972:
6963:
6962:
6960:
6958:
6938:
6932:
6931:
6921:
6911:
6879:
6873:
6867:
6861:
6855:
6846:
6840:
6834:
6833:
6811:
6805:
6799:
6790:
6784:
6773:
6772:
6770:
6768:
6753:
6747:
6741:
6732:
6731:
6729:
6727:
6708:
6702:
6701:
6699:
6697:
6677:
6671:
6665:
6659:
6658:
6656:
6654:
6639:
6633:
6627:
6621:
6620:
6584:
6578:
6577:
6575:
6573:
6553:
6547:
6541:
6535:
6529:
6523:
6522:
6520:
6518:
6490:
6484:
6483:
6473:
6441:
6435:
6434:
6417:(2–4): 329–342.
6406:
6400:
6399:
6389:
6379:
6347:
6341:
6328:
6322:
6316:
6307:
6293:
6287:
6286:
6285:on May 31, 2011.
6284:
6278:. Archived from
6253:
6240:
6234:
6233:
6231:
6229:
6193:
6187:
6181:
6172:
6166:
6160:
6154:
6143:
6142:
6140:
6138:
6108:
6102:
6101:
6099:
6075:
6069:
6068:
6066:
6064:
6024:
6018:
6017:
5992:(3): 1286–1301.
5973:
5967:
5966:
5964:
5940:
5934:
5933:
5923:
5913:
5881:
5875:
5874:
5856:
5816:
5810:
5809:
5791:
5776:
5775:
5735:
5729:
5728:
5680:
5674:
5673:
5663:
5623:
5617:
5616:
5614:
5612:
5606:
5600:. Archived from
5591:
5573:
5564:
5558:
5557:
5555:
5553:
5547:
5541:. Archived from
5514:
5497:
5488:
5487:
5485:
5483:
5455:
5449:
5448:
5446:
5414:
5403:
5402:
5370:
5364:
5363:
5361:
5359:
5353:
5338:
5321:
5315:
5314:
5287:Alopias vulpinus
5282:
5273:
5272:
5263:(8): 1494–1499.
5252:
5246:
5245:
5243:
5241:
5225:
5219:
5213:
5207:
5206:
5193:
5183:
5159:
5153:
5152:
5147:
5145:
5135:
5091:
5085:
5084:
5079:
5077:
5071:
5048:
5039:
5033:
5032:
5030:
5024:. Archived from
5000:(8): 1601–1611.
4991:
4983:"Killer whales (
4978:
4972:
4971:
4969:
4967:
4961:
4946:
4920:
4911:
4905:
4904:
4868:
4862:
4856:
4850:
4849:
4847:
4845:
4840:on July 13, 2012
4826:
4820:
4807:
4801:
4788:
4782:
4776:
4770:
4769:
4767:
4765:
4756:. Archived from
4738:
4727:
4721:
4712:
4706:
4700:
4699:
4697:
4695:
4690:on March 6, 2014
4689:
4682:
4674:
4668:
4662:
4653:
4652:
4640:
4634:
4628:
4622:
4616:
4610:
4609:
4607:
4605:
4599:
4578:
4569:
4560:
4559:, pp. 1–27.
4554:
4548:
4547:
4519:
4513:
4502:
4496:
4495:
4493:
4491:
4476:
4467:
4461:
4455:
4449:
4443:
4437:
4428:
4422:
4416:
4410:
4404:
4403:
4385:
4370:
4369:
4367:
4365:
4360:on April 5, 2012
4350:
4344:
4338:
4329:
4328:
4312:
4306:
4305:
4289:
4283:
4282:
4272:
4254:
4230:
4224:
4223:
4203:
4197:
4196:
4194:
4188:. Archived from
4159:
4146:
4127:
4121:
4115:
4114:
4081:
4075:
4074:
4064:
4028:
4022:
4021:
4006:
4000:
3999:
3989:
3949:
3943:
3942:
3940:
3938:
3911:
3905:
3904:
3886:
3854:
3848:
3847:
3845:
3843:
3837:
3812:
3803:
3794:
3788:
3782:
3781:
3763:
3733:
3727:
3726:
3724:
3722:
3716:
3685:
3672:
3661:
3622:
3613:
3612:
3602:
3561:
3555:
3554:
3544:
3520:
3514:
3513:
3477:
3471:
3465:
3459:
3453:
3444:
3443:
3441:
3439:
3414:
3408:
3407:
3405:
3403:
3380:
3374:
3373:
3355:
3349:
3348:
3346:
3344:
3324:
3318:
3317:
3315:
3313:
3293:
3287:
3274:
3268:
3262:
3256:
3255:
3253:
3251:
3235:
3229:
3228:
3226:
3224:
3205:
3199:
3198:
3196:
3194:
3176:
3170:
3169:
3167:
3165:
3146:
3137:
3136:
3134:
3132:
3127:
3097:
3080:
3079:
3077:
3075:
3066:. Archived from
3052:
3046:
3045:
3009:
2992:
2989:
2983:
2980:
2934:
2929:
2928:
2920:
2915:
2914:
2913:
2906:
2901:
2900:
2892:
2890:Cetaceans portal
2887:
2886:
2885:
2809:Miami Seaquarium
2565:
2459:Western cultures
2376:Maritime Archaic
2257:Johnstone Strait
2253:Underwater noise
2174:British Columbia
2147:Aleutian Islands
2118:Norwegian Arctic
1806:
1805:
1788:
1787:
1770:
1769:
1750:
1734:
1733:
1658:Social structure
1652:Steller sea lion
1648:British Columbia
1532:Aleutian Islands
1504:Península Valdés
1478:species such as
1460:
1448:Valdes Peninsula
1443:
1356:Dall's porpoises
1288:common threshers
1266:In New Zealand,
1261:carousel feeding
1192:
1174:
1084:British Columbia
1068:Aleutian Islands
1048:Valdés Peninsula
991:occipital crests
948:Risso's dolphins
739:10th edition of
710:
709:
690:
668:
622:
621:
606:originally used
556:, pollution (by
542:separate species
429:
421:
413:
394:
386:
378:
356:
243:
242:
222:
205:
200:
199:
174:
159:Aleutian Islands
147:
137:
69:Temporal range:
61:
60:
46:
21:
13170:
13169:
13165:
13164:
13163:
13161:
13160:
13159:
13120:
13119:
13116:
13111:
13103:
13098:
13090:
13085:
13077:
13072:
13064:
13059:
13050:
13049:
13044:
13031:
13023:
13018:
13010:
13005:
12997:
12992:
12984:
12979:
12971:
12966:
12958:
12953:
12945:
12940:
12932:
12929:Observation.org
12927:
12919:
12917:
12909:
12904:
12896:
12891:
12883:
12878:
12870:
12865:
12857:
12852:
12844:
12839:
12831:
12826:
12818:
12813:
12805:
12800:
12792:
12787:
12779:
12774:
12766:
12761:
12753:
12748:
12740:
12735:
12727:
12725:
12717:
12712:
12704:
12699:
12691:
12686:
12678:
12676:
12668:
12663:
12655:
12650:
12641:
12640:
12635:
12626:
12625:
12620:
12607:
12597:
12580:
12556:
12532:
12418:M. densirostris
12378:
12377:
12375:
12365:
12341:
12313:
12312:
12310:
12300:
12253:(Beaked whales)
12252:
12251:
12249:
12237:
12200:
12154:
12117:
12062:
12016:
11979:
11955:
11904:
11889:N. phocaenoides
11876:
11875:
11873:
11857:
11856:
11854:
11842:
11818:
11781:
11735:
11734:
11732:
11722:
11698:
11692:S. longirostris
11674:S. coeruleoalba
11638:
11592:
11591:
11589:
11579:
11546:
11522:
11498:
11474:
11459:O. brevirostris
11441:
11413:
11412:
11410:
11400:
11392:Dusky dolphin (
11331:
11307:
11283:
11255:
11254:
11252:
11242:
11218:
11194:
11150:Cephalorhynchus
11137:
11136:
11134:
11122:
11093:
11056:
11050:M. novaeangliae
11032:
11008:
10982:Omura's whale (
10955:Bryde's whale (
10906:
10905:
10903:
10901:Balaenopteridae
10891:
10854:
10853:
10851:
10841:
10833:Bowhead whale (
10804:
10801:(Baleen whales)
10787:
10737:
10730:
10693:Wayback Machine
10681:Wayback Machine
10668:Wayback Machine
10656:Wayback Machine
10638:Wayback Machine
10620:Wayback Machine
10607:Wayback Machine
10596:
10586:
10565:
10518:
10516:Further reading
10508:
10494:North Vancouver
10478:
10476:
10472:
10457:
10442:
10428:Washington D.C.
10413:
10380:
10308:
10287:
10254:
10229:
10199:
10194:
10193:
10183:
10181:
10170:
10166:
10156:
10154:
10141:
10140:
10136:
10126:
10124:
10109:
10108:
10104:
10093:
10089:
10079:
10077:
10069:Rotten Tomatoes
10062:
10061:
10057:
10047:
10045:
10044:on July 6, 2010
10041:
10034:
10030:
10029:
10025:
10015:
10013:
10009:
10002:
9996:
9992:
9984:
9980:
9970:
9968:
9959:
9958:
9954:
9914:
9903:
9893:
9891:
9878:
9877:
9873:
9836:Biology Letters
9828:
9821:
9811:
9809:
9805:
9794:
9788:
9784:
9774:
9772:
9768:
9753:
9747:
9743:
9733:
9731:
9718:
9717:
9713:
9703:
9701:
9688:
9687:
9683:
9673:
9671:
9664:
9660:
9659:
9655:
9647:
9640:
9630:
9628:
9624:
9613:
9605:
9604:
9600:
9590:
9588:
9573:
9569:
9559:
9557:
9544:
9543:
9539:
9533:Wayback Machine
9524:
9520:
9510:
9508:
9493:
9489:
9481:
9477:
9469:
9465:
9457:
9453:
9430:
9426:
9403:
9399:
9384:10.2307/3936617
9369:
9368:
9364:
9358:Wayback Machine
9344:
9340:
9332:
9328:
9322:Wayback Machine
9311:
9307:
9297:
9295:
9286:
9285:
9278:
9268:
9266:
9249:
9245:
9235:
9233:
9210:
9206:
9196:
9194:
9181:
9180:
9176:
9166:
9164:
9146:
9142:
9132:
9130:
9115:
9111:
9101:
9099:
9092:The Independent
9086:
9085:
9081:
9071:
9069:
9056:
9055:
9051:
9041:
9039:
9020:
9019:
9015:
9005:
9003:
8990:
8989:
8985:
8978:
8964:
8960:
8950:
8948:
8933:
8932:
8928:
8914:Wayback Machine
8905:. (See also an
8890:Wayback Machine
8874:
8870:
8862:
8851:
8839:
8834:
8830:
8819:
8817:
8810:
8805:
8804:
8797:
8787:
8785:
8770:
8763:
8746:
8745:
8738:
8725:
8721:
8705:
8701:
8662:
8658:
8652:Wayback Machine
8642:
8638:
8628:
8626:
8609:
8605:
8597:
8593:
8585:
8576:
8566:
8564:
8560:
8553:
8549:
8548:
8544:
8534:
8532:
8517:
8513:
8503:
8501:
8497:
8466:
8460:
8456:
8446:
8444:
8429:
8425:
8410:
8406:
8398:
8394:
8384:
8382:
8369:
8368:
8359:
8349:
8347:
8332:
8317:
8311:Wayback Machine
8301:
8294:
8275:
8268:
8260:
8256:
8246:
8244:
8229:
8225:
8168:
8164:
8154:
8152:
8137:
8133:
8125:
8118:
8101:
8100:
8096:
8086:
8084:
8075:
8074:
8070:
8064:Wayback Machine
8055:
8051:
8045:Wayback Machine
8031:
8027:
8014:
8010:
8002:
7998:
7990:
7986:
7951:
7947:
7894:
7887:
7877:
7875:
7864:
7860:
7850:
7848:
7833:
7829:
7823:Carwardine 2001
7821:
7814:
7804:
7802:
7801:on June 6, 2020
7789:
7788:
7781:
7770:
7766:
7751:
7725:
7721:
7684:
7677:
7667:
7665:
7622:
7618:
7569:
7565:
7555:Wayback Machine
7545:
7541:
7533:
7518:
7514:
7513:
7509:
7501:
7494:
7479:
7475:
7469:Carwardine 2001
7467:
7460:
7448:
7441:
7378:
7374:
7364:
7362:
7347:
7343:
7333:
7331:
7316:
7312:
7302:
7300:
7285:
7281:
7254:Acta Ethologica
7250:Clupea harengus
7242:
7238:
7230:
7223:
7215:
7176:
7170:
7166:
7156:
7154:
7150:
7111:
7101:
7097:
7062:
7055:
7020:
7016:
6985:
6981:
6973:
6966:
6956:
6954:
6939:
6935:
6880:
6876:
6868:
6864:
6856:
6849:
6841:
6837:
6830:
6812:
6808:
6800:
6793:
6785:
6776:
6766:
6764:
6755:
6754:
6750:
6742:
6735:
6725:
6723:
6710:
6709:
6705:
6695:
6693:
6678:
6674:
6668:Carwardine 2001
6666:
6662:
6652:
6650:
6641:
6640:
6636:
6628:
6624:
6585:
6581:
6571:
6569:
6554:
6550:
6542:
6538:
6530:
6526:
6516:
6514:
6507:
6491:
6487:
6442:
6438:
6407:
6403:
6348:
6344:
6338:Wayback Machine
6329:
6325:
6317:
6310:
6304:Wayback Machine
6294:
6290:
6282:
6251:
6241:
6237:
6227:
6225:
6194:
6190:
6182:
6175:
6169:Carwardine 2001
6167:
6163:
6155:
6146:
6136:
6134:
6127:
6109:
6105:
6076:
6072:
6062:
6060:
6025:
6021:
5974:
5970:
5941:
5937:
5882:
5878:
5817:
5813:
5806:
5792:
5779:
5736:
5732:
5681:
5677:
5624:
5620:
5610:
5608:
5604:
5571:
5565:
5561:
5551:
5549:
5545:
5512:
5498:
5491:
5481:
5479:
5462:Rhincodon typus
5456:
5452:
5415:
5406:
5371:
5367:
5357:
5355:
5351:
5341:Aquatic Mammals
5336:
5326:"Killer Whale (
5322:
5318:
5299:Aquatic Mammals
5291:Sphyrna zygaena
5283:
5276:
5269:10.1139/z93-210
5253:
5249:
5239:
5237:
5226:
5222:
5214:
5210:
5160:
5156:
5143:
5141:
5092:
5088:
5075:
5073:
5069:
5063:10.1139/Z10-080
5046:
5040:
5036:
5028:
4989:
4979:
4975:
4965:
4963:
4959:
4923:Pacific Science
4918:
4912:
4908:
4869:
4865:
4857:
4853:
4843:
4841:
4828:
4827:
4823:
4817:Wayback Machine
4808:
4804:
4798:Wayback Machine
4789:
4785:
4777:
4773:
4763:
4761:
4760:on July 9, 2017
4742:"Killer whale (
4740:
4739:
4730:
4722:
4715:
4707:
4703:
4693:
4691:
4687:
4680:
4676:
4675:
4671:
4663:
4656:
4641:
4637:
4629:
4625:
4619:Carwardine 2001
4617:
4613:
4603:
4601:
4597:
4591:
4576:
4570:
4563:
4555:
4551:
4520:
4516:
4503:
4499:
4489:
4487:
4478:
4477:
4470:
4464:Carwardine 2001
4462:
4458:
4450:
4446:
4438:
4431:
4423:
4419:
4411:
4407:
4400:
4386:
4373:
4363:
4361:
4352:
4351:
4347:
4341:Carwardine 2001
4339:
4332:
4313:
4309:
4290:
4286:
4231:
4227:
4220:
4204:
4200:
4192:
4170:10.2307/3504225
4157:
4147:
4130:
4122:
4118:
4082:
4078:
4041:Genome Research
4029:
4025:
4008:
4007:
4003:
3950:
3946:
3936:
3934:
3926:(2 }): 93–106.
3920:Aquatic Mammals
3912:
3908:
3884:10.1038/445703a
3855:
3851:
3841:
3839:
3835:
3810:
3804:
3797:
3789:
3785:
3734:
3730:
3720:
3718:
3714:
3683:
3673:
3664:
3652:Wayback Machine
3637:Wayback Machine
3623:
3616:
3562:
3558:
3521:
3517:
3478:
3474:
3468:Carwardine 2001
3466:
3462:
3454:
3447:
3437:
3435:
3415:
3411:
3401:
3399:
3397:
3381:
3377:
3370:
3356:
3352:
3342:
3340:
3325:
3321:
3311:
3309:
3294:
3290:
3284:Wayback Machine
3275:
3271:
3263:
3259:
3249:
3247:
3236:
3232:
3222:
3220:
3207:
3206:
3202:
3192:
3190:
3178:
3177:
3173:
3163:
3161:
3148:
3147:
3140:
3130:
3128:
3098:
3083:
3073:
3071:
3054:
3053:
3049:
3034:
3014:"Order Cetacea"
3010:
3006:
3001:
2996:
2995:
2990:
2986:
2981:
2977:
2972:
2930:
2923:
2916:
2911:
2909:
2902:
2895:
2888:
2883:
2881:
2878:
2798:
2792:
2776:
2756:
2744:Seto Inland Sea
2708:recoverable oil
2686:The orca named
2680:
2674:
2554:
2552:
2546:
2522:
2510:
2497:
2491:
2463:Pliny the Elder
2453:, Germany, 1545
2443:
2437:
2417:, and southern
2318:
2312:
2307:
2102:bioaccumulation
2045:
2019:New South Wales
1968:
1892:
1886:
1832:
1826:
1821:
1820:
1812:
1810:
1809:
1808:
1807:
1800:
1797:
1791:
1790:
1789:
1782:
1779:
1773:
1772:
1771:
1764:
1761:
1754:
1748:
1731:
1668:higher primates
1660:
1644:surplus killing
1597:
1565:gentoo penguins
1534:, a decline in
1472:
1471:
1470:
1469:
1468:
1461:
1452:
1451:
1450:
1444:
1410:humpback whales
1336:common dolphins
1324:
1224:
1200:
1199:
1198:
1197:
1196:
1193:
1184:
1183:
1182:
1175:
1164:
1155:
1129:decline in the
1021:
1015:
928:
926:Characteristics
851:Species problem
835:
829:
808:Cephalorhynchus
741:Systema Naturae
718:
717:
716:
715:
714:
711:
700:
699:
698:
691:
682:
681:
680:
669:
658:
650:Risso's dolphin
594:The genus name
589:
560:), capture for
457:oceanic dolphin
427:
419:
411:
384:
383:
369:
358:
352:
339:
237:
229:
212:
201:
197:
190:
181:
165:
138:
136:
135:
130:
125:
120:
115:
110:
105:
100:
95:
90:
85:
80:
74:
67:
65:
57:
44:
35:
28:
23:
22:
15:
12:
11:
5:
13168:
13158:
13157:
13152:
13147:
13142:
13137:
13132:
13113:
13112:
13110:
13109:
13096:
13083:
13070:
13057:
13041:
13039:
13037:Delphinus orca
13033:
13032:
13030:
13029:
13016:
13003:
12990:
12977:
12964:
12951:
12938:
12925:
12915:
12902:
12889:
12876:
12863:
12850:
12837:
12824:
12811:
12798:
12785:
12772:
12759:
12746:
12733:
12723:
12710:
12697:
12684:
12674:
12661:
12648:
12633:
12617:
12615:
12609:
12608:
12594:
12593:
12590:
12589:
12586:
12585:
12582:
12581:
12579:
12578:
12574:Z. cavirostris
12568:
12566:
12558:
12557:
12555:
12554:
12544:
12542:
12534:
12533:
12531:
12530:
12521:
12512:
12503:
12494:
12485:
12476:
12467:
12458:
12449:
12440:
12431:
12422:
12413:
12404:
12395:
12385:
12383:
12367:
12366:
12364:
12363:
12353:
12351:
12343:
12342:
12340:
12339:
12330:
12320:
12318:
12302:
12301:
12299:
12298:
12289:
12280:
12270:
12268:
12257:
12243:
12242:
12239:
12238:
12236:
12235:
12231:P. blainvillei
12225:
12223:
12212:
12206:
12205:
12202:
12201:
12199:
12198:
12189:
12179:
12177:
12166:
12160:
12159:
12156:
12155:
12153:
12152:
12142:
12140:
12129:
12123:
12122:
12119:
12118:
12116:
12115:
12111:I. geoffrensis
12106:
12102:I. boliviensis
12097:
12087:
12085:
12074:
12068:
12067:
12064:
12063:
12061:
12060:
12051:
12041:
12039:
12028:
12022:
12021:
12018:
12017:
12015:
12014:
12004:
12002:
11991:
11985:
11984:
11981:
11980:
11978:
11977:
11967:
11965:
11957:
11956:
11954:
11953:
11949:P. spinipinnis
11944:
11935:
11926:
11916:
11914:
11906:
11905:
11903:
11902:
11893:
11883:
11881:
11862:
11848:
11847:
11844:
11843:
11841:
11840:
11830:
11828:
11820:
11819:
11817:
11816:
11810:Beluga whale (
11806:
11804:
11801:Delphinapterus
11793:
11787:
11786:
11783:
11782:
11780:
11779:
11770:
11761:
11752:
11742:
11740:
11724:
11723:
11721:
11720:
11716:S. bredanensis
11710:
11708:
11700:
11699:
11697:
11696:
11687:
11678:
11669:
11660:
11650:
11648:
11640:
11639:
11637:
11636:
11627:
11618:
11609:
11599:
11597:
11581:
11580:
11578:
11577:
11568:
11564:S. fluviatilis
11558:
11556:
11548:
11547:
11545:
11544:
11534:
11532:
11524:
11523:
11521:
11520:
11510:
11508:
11500:
11499:
11497:
11496:
11486:
11484:
11476:
11475:
11473:
11472:
11463:
11453:
11451:
11443:
11442:
11440:
11439:
11430:
11420:
11418:
11402:
11401:
11399:
11398:
11389:
11385:L. obliquidens
11380:
11371:
11362:
11353:
11349:L. albirostris
11343:
11341:
11338:Lagenorhynchus
11333:
11332:
11330:
11329:
11319:
11317:
11309:
11308:
11306:
11305:
11295:
11293:
11285:
11284:
11282:
11281:
11272:
11262:
11260:
11256:(Pilot whales)
11244:
11243:
11241:
11240:
11230:
11228:
11220:
11219:
11217:
11216:
11206:
11204:
11196:
11195:
11193:
11192:
11183:
11174:
11165:
11161:C. commersonii
11155:
11153:
11142:
11124:
11123:
11107:
11106:
11103:
11102:
11099:
11098:
11095:
11094:
11092:
11091:
11081:
11079:
11068:
11062:
11061:
11058:
11057:
11055:
11054:
11044:
11042:
11034:
11033:
11031:
11030:
11020:
11018:
11010:
11009:
11007:
11006:
11000:Rice's whale (
10997:
10988:
10979:
10970:
10961:
10952:
10943:
10939:B. bonaerensis
10934:
10924:
10922:
10911:
10897:
10896:
10893:
10892:
10890:
10889:
10880:
10871:
10861:
10859:
10855:(Right whales)
10843:
10842:
10840:
10839:
10829:
10827:
10816:
10806:
10805:
10789:
10788:
10786:
10785:
10779:
10773:
10771:Laurasiatheria
10767:
10761:
10755:
10749:
10742:
10739:
10738:
10729:
10728:
10721:
10714:
10706:
10700:
10699:
10683:
10671:
10658:
10646:
10641:
10628:
10623:
10610:
10595:
10594:External links
10592:
10591:
10590:
10584:
10568:
10563:
10546:
10517:
10514:
10513:
10512:
10506:
10485:
10446:
10440:
10417:
10411:
10403:Voyageur Press
10384:
10378:
10353:
10312:
10306:
10291:
10285:
10273:Academic Press
10263:"Killer Whale"
10258:
10252:
10233:
10227:
10219:Voyageur Press
10198:
10195:
10192:
10191:
10164:
10134:
10102:
10087:
10055:
10023:
9990:
9978:
9952:
9925:(4): 792–805.
9901:
9871:
9842:(4): 415–418.
9819:
9782:
9741:
9711:
9681:
9653:
9638:
9598:
9567:
9537:
9518:
9487:
9475:
9463:
9451:
9424:
9397:
9362:
9338:
9326:
9305:
9276:
9243:
9204:
9174:
9140:
9109:
9079:
9049:
9013:
8983:
8976:
8958:
8926:
8868:
8849:
8846:(in Japanese).
8828:
8795:
8761:
8736:
8719:
8699:
8678:10.2307/277719
8672:(3): 343–358.
8656:
8636:
8603:
8591:
8574:
8542:
8511:
8477:(3): 305–310.
8454:
8423:
8404:
8402:, p. 100.
8392:
8357:
8315:
8292:
8266:
8254:
8223:
8162:
8131:
8116:
8094:
8068:
8049:
8025:
8008:
7996:
7984:
7945:
7885:
7858:
7827:
7812:
7779:
7764:
7749:
7719:
7698:(3): 632–640.
7675:
7636:(2): 309–324.
7616:
7585:(e139): e139.
7563:
7539:
7507:
7492:
7473:
7458:
7439:
7372:
7341:
7310:
7279:
7236:
7221:
7187:(1): 627–635.
7164:
7122:(3): 261–280.
7095:
7076:(4): 433–445.
7053:
7034:(6): 599–606.
7014:
6995:(2): 395–405.
6979:
6964:
6933:
6874:
6862:
6847:
6835:
6829:978-0774804691
6828:
6806:
6791:
6774:
6748:
6733:
6703:
6672:
6660:
6634:
6622:
6579:
6548:
6536:
6534:, p. 124.
6524:
6505:
6485:
6436:
6401:
6342:
6323:
6308:
6288:
6262:(1): 225–234.
6235:
6188:
6173:
6161:
6144:
6125:
6103:
6070:
6039:(3): 494–507.
6019:
5968:
5935:
5876:
5839:(3): 141–154.
5811:
5804:
5777:
5730:
5695:(2): 139–152.
5675:
5618:
5589:10.1.1.404.130
5559:
5525:(2): 563–568.
5489:
5450:
5404:
5385:(1): 220–227.
5365:
5347:(3): 229–231.
5316:
5274:
5247:
5220:
5208:
5174:(1): 298–311.
5154:
5086:
5034:
4973:
4929:(4): 523–530.
4906:
4863:
4851:
4821:
4802:
4783:
4771:
4750:NOAA Fisheries
4728:
4713:
4701:
4669:
4654:
4635:
4633:, p. 128.
4623:
4611:
4589:
4561:
4549:
4530:(5): 974–981.
4514:
4508:) and Killer (
4497:
4468:
4456:
4454:, p. 683.
4444:
4442:, p. 681.
4429:
4417:
4405:
4398:
4371:
4345:
4330:
4307:
4284:
4225:
4218:
4198:
4128:
4126:, p. 129.
4116:
4097:(2): 303–306.
4076:
4047:(7): 908–916.
4023:
4001:
3944:
3906:
3849:
3823:(4): 151–180.
3795:
3783:
3728:
3694:(2): 131–139.
3662:
3614:
3577:(3): 479–501.
3556:
3535:(2): 476–480.
3515:
3488:(3): 619–648.
3472:
3460:
3445:
3409:
3395:
3375:
3368:
3350:
3319:
3288:
3269:
3257:
3230:
3200:
3171:
3138:
3081:
3060:Linnaeus 1758"
3047:
3032:
3003:
3002:
3000:
2997:
2994:
2993:
2984:
2974:
2973:
2971:
2968:
2967:
2966:
2960:
2959:
2958:
2957:
2956:
2936:
2935:
2921:
2907:
2904:Mammals portal
2893:
2877:
2874:
2794:Main article:
2791:
2788:
2779:Whale watching
2775:
2774:Whale watching
2772:
2755:
2752:
2676:Main article:
2673:
2670:
2585:Saturna Island
2583:harpooned off
2545:
2542:
2521:
2518:
2509:
2506:
2493:Main article:
2490:
2487:
2436:
2433:
2391:Siberian Yupik
2357:Nuu-chah-nulth
2316:Animal worship
2311:
2308:
2306:
2303:
2273:whale watchers
2198:whale watchers
2179:Chinook salmon
2098:trophic levels
2084:, large-scale
2070:data deficient
2044:
2041:
1967:
1964:
1947:animal culture
1909:Crozet Islands
1888:Main article:
1885:
1882:
1823:
1822:
1811:
1798:
1793:
1792:
1780:
1775:
1774:
1762:
1757:
1756:
1755:
1746:
1745:
1744:
1741:
1740:
1732:
1730:
1727:
1720:Crozet Islands
1659:
1656:
1596:
1593:
1508:Crozet Islands
1484:elephant seals
1465:crabeater seal
1462:
1455:
1454:
1453:
1445:
1438:
1437:
1436:
1435:
1434:
1348:dusky dolphins
1328:marine mammals
1323:
1320:
1304:shortfin makos
1300:basking sharks
1223:
1220:
1204:apex predators
1194:
1187:
1186:
1185:
1176:
1169:
1168:
1167:
1166:
1165:
1163:
1160:
1154:
1151:
1143:Horikawa River
1135:Columbia River
1127:Arctic sea ice
1076:Southern Ocean
1072:Gulf of Alaska
1052:Crozet Islands
1046:, Norway, the
1036:enormous range
1014:
1011:
927:
924:
875:O. rectipinnus
828:
825:
802:Lagenorhynchus
799:that includes
762:animal in the
746:Konrad Gessner
712:
703:
702:
701:
692:
685:
684:
683:
670:
663:
662:
661:
660:
659:
657:
654:
604:Ancient Romans
588:
585:
538:data deficient
523:animal culture
502:marine mammals
486:apex predators
433:
432:
431:
430:
425:Orca gladiator
422:
414:
409:Delphinus orca
403:
402:
396:
395:
380:
379:
371:
370:
359:
348:
347:
341:
340:
333:
331:
327:
326:
319:
315:
314:
309:
305:
304:
299:
295:
294:
289:
285:
284:
279:
275:
274:
269:
265:
264:
259:
255:
254:
249:
245:
244:
231:
230:
217:
214:
213:
203:Data Deficient
195:
192:
191:
186:
183:
182:
179:
176:
175:
167:
166:
152:
149:
148:
140:
139:
131:
126:
121:
116:
111:
106:
101:
96:
91:
86:
81:
76:
68:
26:
9:
6:
4:
3:
2:
13167:
13156:
13153:
13151:
13148:
13146:
13143:
13141:
13138:
13136:
13133:
13131:
13128:
13127:
13125:
13118:
13106:
13101:
13097:
13093:
13088:
13084:
13080:
13075:
13071:
13067:
13062:
13058:
13053:
13047:
13043:
13042:
13040:
13038:
13034:
13026:
13021:
13017:
13013:
13008:
13004:
13000:
12995:
12991:
12987:
12982:
12978:
12974:
12969:
12965:
12961:
12956:
12952:
12948:
12943:
12939:
12935:
12930:
12926:
12922:
12916:
12912:
12907:
12903:
12899:
12894:
12890:
12886:
12881:
12877:
12873:
12868:
12864:
12860:
12855:
12851:
12847:
12842:
12838:
12834:
12829:
12825:
12821:
12816:
12812:
12808:
12803:
12799:
12795:
12790:
12786:
12782:
12777:
12773:
12769:
12764:
12760:
12756:
12751:
12747:
12743:
12738:
12734:
12730:
12724:
12720:
12715:
12711:
12707:
12702:
12698:
12694:
12689:
12685:
12681:
12675:
12671:
12666:
12662:
12658:
12653:
12649:
12644:
12638:
12634:
12629:
12623:
12619:
12618:
12616:
12614:
12610:
12606:
12601:
12577:
12575:
12570:
12569:
12567:
12565:
12564:
12559:
12553:
12551:
12546:
12545:
12543:
12541:
12540:
12535:
12529:
12527:
12522:
12520:
12518:
12517:M. stejnegeri
12513:
12511:
12509:
12508:M. peruvianus
12504:
12502:
12500:
12495:
12493:
12491:
12486:
12484:
12482:
12477:
12475:
12473:
12468:
12466:
12464:
12459:
12457:
12455:
12450:
12448:
12446:
12445:M. ginkgodens
12441:
12439:
12437:
12432:
12430:
12428:
12423:
12421:
12419:
12414:
12412:
12410:
12409:M. carlhubbsi
12405:
12403:
12401:
12396:
12394:
12392:
12387:
12386:
12384:
12374:
12373:
12368:
12362:
12360:
12355:
12354:
12352:
12350:
12349:
12344:
12338:
12336:
12335:H. planifrons
12331:
12329:
12327:
12326:H. ampullatus
12322:
12321:
12319:
12309:
12308:
12303:
12297:
12295:
12290:
12288:
12286:
12281:
12279:
12277:
12272:
12271:
12269:
12267:
12266:
12261:
12258:
12248:
12244:
12234:
12232:
12227:
12226:
12224:
12222:
12221:
12216:
12213:
12211:
12210:Pontoporiidae
12207:
12197:
12195:
12190:
12188:
12186:
12181:
12180:
12178:
12176:
12175:
12170:
12167:
12165:
12164:Platanistidae
12161:
12151:
12149:
12148:L. vexillifer
12144:
12143:
12141:
12139:
12138:
12133:
12130:
12128:
12124:
12114:
12112:
12107:
12105:
12103:
12098:
12096:
12094:
12089:
12088:
12086:
12084:
12083:
12078:
12075:
12073:
12069:
12059:
12057:
12052:
12050:
12048:
12043:
12042:
12040:
12038:
12037:
12032:
12029:
12027:
12023:
12013:
12011:
12008:Sperm whale (
12006:
12005:
12003:
12001:
12000:
11995:
11992:
11990:
11986:
11976:
11974:
11969:
11968:
11966:
11964:
11963:
11958:
11952:
11950:
11945:
11943:
11941:
11936:
11934:
11932:
11927:
11925:
11923:
11918:
11917:
11915:
11913:
11912:
11907:
11901:
11899:
11894:
11892:
11890:
11885:
11884:
11882:
11872:
11871:
11866:
11863:
11853:
11849:
11839:
11837:
11832:
11831:
11829:
11827:
11826:
11821:
11815:
11813:
11808:
11807:
11805:
11803:
11802:
11797:
11794:
11792:
11788:
11778:
11776:
11771:
11769:
11767:
11762:
11760:
11758:
11753:
11751:
11749:
11744:
11743:
11741:
11731:
11730:
11725:
11719:
11717:
11712:
11711:
11709:
11707:
11706:
11701:
11695:
11693:
11688:
11686:
11684:
11679:
11677:
11675:
11670:
11668:
11666:
11661:
11659:
11657:
11652:
11651:
11649:
11647:
11646:
11641:
11635:
11633:
11628:
11626:
11624:
11623:S. sahulensis
11619:
11617:
11615:
11610:
11608:
11606:
11601:
11600:
11598:
11588:
11587:
11582:
11576:
11574:
11573:S. guianensis
11569:
11567:
11565:
11560:
11559:
11557:
11555:
11554:
11549:
11543:
11541:
11540:P. crassidens
11536:
11535:
11533:
11531:
11530:
11525:
11519:
11517:
11512:
11511:
11509:
11507:
11506:
11505:Peponocephala
11501:
11495:
11493:
11488:
11487:
11485:
11483:
11482:
11477:
11471:
11469:
11464:
11462:
11460:
11455:
11454:
11452:
11450:
11449:
11444:
11438:
11436:
11431:
11429:
11427:
11422:
11421:
11419:
11409:
11408:
11403:
11397:
11395:
11390:
11388:
11386:
11381:
11379:
11377:
11372:
11370:
11368:
11363:
11361:
11359:
11354:
11352:
11350:
11345:
11344:
11342:
11340:
11339:
11334:
11328:
11326:
11321:
11320:
11318:
11316:
11315:
11314:Lagenodelphis
11310:
11304:
11302:
11297:
11296:
11294:
11292:
11291:
11286:
11280:
11278:
11273:
11271:
11269:
11264:
11263:
11261:
11251:
11250:
11245:
11239:
11237:
11232:
11231:
11229:
11227:
11226:
11221:
11215:
11213:
11208:
11207:
11205:
11203:
11202:
11197:
11191:
11189:
11184:
11182:
11180:
11179:C. heavisidii
11175:
11173:
11171:
11166:
11164:
11162:
11157:
11156:
11154:
11152:
11151:
11146:
11143:
11133:
11129:
11125:
11121:
11112:
11108:
11090:
11088:
11083:
11082:
11080:
11078:
11077:
11072:
11069:
11067:
11066:Cetotheriidae
11063:
11053:
11051:
11046:
11045:
11043:
11041:
11040:
11035:
11029:
11027:
11022:
11021:
11019:
11017:
11016:
11011:
11005:
11003:
10998:
10996:
10994:
10989:
10987:
10985:
10980:
10978:
10976:
10971:
10969:
10967:
10962:
10960:
10958:
10953:
10951:
10949:
10944:
10942:
10940:
10935:
10933:
10931:
10926:
10925:
10923:
10921:
10920:
10915:
10912:
10902:
10898:
10888:
10886:
10881:
10879:
10877:
10872:
10870:
10868:
10863:
10862:
10860:
10850:
10849:
10844:
10838:
10836:
10835:B. mysticetus
10831:
10830:
10828:
10826:
10825:
10820:
10817:
10815:
10811:
10807:
10803:
10794:
10790:
10784:
10780:
10778:
10774:
10772:
10768:
10766:
10762:
10760:
10756:
10754:
10750:
10748:
10744:
10743:
10740:
10735:
10727:
10722:
10720:
10715:
10713:
10708:
10707:
10704:
10698:
10694:
10690:
10687:
10684:
10682:
10678:
10675:
10672:
10669:
10665:
10662:
10659:
10657:
10653:
10650:
10647:
10645:
10642:
10639:
10635:
10632:
10629:
10627:
10624:
10621:
10617:
10614:
10611:
10608:
10604:
10601:
10598:
10597:
10587:
10581:
10578:
10574:
10569:
10566:
10560:
10555:
10554:
10547:
10543:
10539:
10535:
10531:
10527:
10526:
10520:
10519:
10509:
10503:
10499:
10495:
10491:
10486:
10471:
10467:
10463:
10456:
10454:
10449:NMFS (2005).
10447:
10443:
10437:
10433:
10429:
10425:
10424:
10418:
10414:
10408:
10404:
10400:
10396:
10392:
10391:
10390:Killer Whales
10385:
10381:
10375:
10371:
10367:
10363:
10359:
10354:
10349:
10344:
10340:
10336:
10332:
10328:
10327:
10322:
10320:
10313:
10309:
10303:
10299:
10298:
10297:Killer Whales
10292:
10288:
10286:0-12-551340-2
10282:
10278:
10274:
10270:
10269:
10264:
10259:
10255:
10249:
10245:
10244:BBC Worldwide
10241:
10240:
10239:Killer Whales
10234:
10230:
10224:
10220:
10216:
10212:
10208:
10207:
10201:
10200:
10179:
10175:
10168:
10157:September 27,
10152:
10148:
10144:
10138:
10127:September 11,
10122:
10118:
10117:
10112:
10106:
10098:
10091:
10075:
10071:
10070:
10065:
10059:
10040:
10033:
10027:
10008:
10001:
9994:
9987:
9982:
9966:
9962:
9956:
9948:
9944:
9940:
9936:
9932:
9928:
9924:
9920:
9912:
9910:
9908:
9906:
9889:
9885:
9884:Be Whale Wise
9881:
9875:
9867:
9863:
9858:
9853:
9849:
9845:
9841:
9837:
9833:
9826:
9824:
9804:
9800:
9793:
9786:
9767:
9763:
9759:
9752:
9745:
9729:
9725:
9721:
9715:
9699:
9695:
9691:
9685:
9669:
9665:
9657:
9651:, p. 34.
9650:
9645:
9643:
9623:
9619:
9612:
9610:
9602:
9586:
9582:
9578:
9571:
9555:
9551:
9547:
9541:
9534:
9530:
9527:
9522:
9506:
9502:
9498:
9491:
9485:, p. 11.
9484:
9479:
9472:
9467:
9460:
9455:
9447:
9443:
9439:
9435:
9428:
9420:
9416:
9412:
9408:
9401:
9393:
9389:
9385:
9381:
9377:
9373:
9366:
9359:
9355:
9352:
9348:
9342:
9336:, p. 41.
9335:
9330:
9323:
9319:
9315:
9309:
9293:
9289:
9283:
9281:
9264:
9260:
9259:
9254:
9247:
9231:
9227:
9223:
9219:
9218:The Telegraph
9215:
9208:
9192:
9188:
9184:
9178:
9163:
9159:
9155:
9151:
9144:
9128:
9124:
9120:
9113:
9097:
9093:
9089:
9083:
9067:
9063:
9059:
9053:
9037:
9033:
9029:
9028:
9023:
9017:
9001:
8997:
8993:
8987:
8979:
8973:
8969:
8962:
8946:
8942:
8941:
8936:
8930:
8923:
8919:
8915:
8911:
8908:
8904:
8903:
8902:archive.today
8899:
8896:
8891:
8887:
8884:
8882:
8877:
8872:
8865:
8860:
8858:
8856:
8854:
8845:
8844:
8843:Asahi Shimbun
8832:
8815:
8811:
8802:
8800:
8783:
8779:
8775:
8768:
8766:
8757:
8753:
8749:
8743:
8741:
8733:
8729:
8723:
8716:
8713:. Leningrad:
8712:
8708:
8703:
8695:
8691:
8687:
8683:
8679:
8675:
8671:
8667:
8660:
8653:
8649:
8646:
8640:
8624:
8620:
8619:
8614:
8607:
8601:, p. 11.
8600:
8595:
8588:
8583:
8581:
8579:
8559:
8552:
8546:
8530:
8526:
8522:
8515:
8496:
8492:
8488:
8484:
8480:
8476:
8472:
8465:
8458:
8442:
8438:
8434:
8427:
8420:
8419:archive.today
8416:
8413:
8408:
8401:
8396:
8380:
8376:
8372:
8366:
8364:
8362:
8345:
8341:
8337:
8330:
8328:
8326:
8324:
8322:
8320:
8312:
8308:
8305:
8299:
8297:
8288:
8284:
8280:
8273:
8271:
8264:, p. 98.
8263:
8258:
8242:
8238:
8234:
8227:
8219:
8215:
8211:
8207:
8202:
8197:
8193:
8189:
8185:
8181:
8177:
8173:
8166:
8155:September 28,
8150:
8146:
8142:
8135:
8129:, p. 99.
8128:
8123:
8121:
8112:
8108:
8104:
8098:
8082:
8078:
8072:
8065:
8061:
8058:
8053:
8046:
8042:
8039:
8037:
8029:
8022:
8018:
8012:
8006:, p. 35.
8005:
8000:
7994:, p. 33.
7993:
7988:
7980:
7976:
7972:
7968:
7964:
7960:
7957:) survival".
7956:
7949:
7941:
7937:
7933:
7928:
7923:
7919:
7915:
7911:
7907:
7903:
7901:
7892:
7890:
7878:September 11,
7873:
7869:
7862:
7846:
7842:
7838:
7831:
7825:, p. 26.
7824:
7819:
7817:
7800:
7796:
7792:
7786:
7784:
7775:
7768:
7760:
7756:
7752:
7750:9780120392285
7746:
7742:
7738:
7734:
7730:
7723:
7714:
7709:
7705:
7701:
7697:
7693:
7689:
7682:
7680:
7663:
7659:
7655:
7651:
7647:
7643:
7639:
7635:
7631:
7627:
7620:
7612:
7608:
7603:
7598:
7593:
7588:
7584:
7580:
7579:
7574:
7567:
7560:
7556:
7552:
7549:
7543:
7532:
7528:
7524:
7517:
7511:
7505:, p. 42.
7504:
7499:
7497:
7488:
7484:
7477:
7471:, p. 67.
7470:
7465:
7463:
7455:
7452:
7446:
7444:
7435:
7431:
7426:
7421:
7417:
7413:
7408:
7403:
7399:
7395:
7391:
7387:
7383:
7376:
7360:
7356:
7352:
7345:
7329:
7325:
7321:
7314:
7298:
7294:
7290:
7283:
7275:
7271:
7267:
7263:
7259:
7255:
7251:
7247:
7240:
7233:
7228:
7226:
7214:
7210:
7206:
7202:
7198:
7194:
7190:
7186:
7182:
7175:
7168:
7149:
7145:
7141:
7137:
7133:
7129:
7125:
7121:
7117:
7110:
7108:
7099:
7091:
7087:
7083:
7079:
7075:
7071:
7067:
7060:
7058:
7049:
7045:
7041:
7037:
7033:
7029:
7025:
7018:
7010:
7006:
7002:
6998:
6994:
6990:
6983:
6977:, p. 20.
6976:
6971:
6969:
6952:
6948:
6944:
6937:
6929:
6925:
6920:
6915:
6910:
6905:
6901:
6897:
6893:
6889:
6885:
6878:
6872:, p. 27.
6871:
6866:
6860:, p. 14.
6859:
6854:
6852:
6845:, p. 13.
6844:
6839:
6831:
6825:
6821:
6817:
6810:
6804:, p. 39.
6803:
6798:
6796:
6789:, p. 12.
6788:
6783:
6781:
6779:
6762:
6758:
6752:
6746:, p. 35.
6745:
6740:
6738:
6721:
6717:
6713:
6707:
6691:
6687:
6683:
6676:
6670:, p. 64.
6669:
6664:
6648:
6644:
6638:
6632:, p. 14.
6631:
6626:
6618:
6614:
6610:
6606:
6602:
6598:
6594:
6590:
6589:Polar Biology
6583:
6567:
6563:
6559:
6552:
6546:, p. 19.
6545:
6540:
6533:
6528:
6512:
6508:
6502:
6498:
6497:
6489:
6481:
6477:
6472:
6467:
6463:
6459:
6455:
6451:
6447:
6440:
6432:
6428:
6424:
6420:
6416:
6412:
6405:
6397:
6393:
6388:
6383:
6378:
6373:
6369:
6365:
6361:
6357:
6353:
6346:
6339:
6335:
6332:
6327:
6321:, p. 23.
6320:
6315:
6313:
6305:
6301:
6298:
6292:
6281:
6277:
6273:
6269:
6265:
6261:
6257:
6250:
6248:
6239:
6223:
6219:
6215:
6211:
6207:
6203:
6201:
6192:
6185:
6180:
6178:
6171:, p. 29.
6170:
6165:
6159:, p. 45.
6158:
6153:
6151:
6149:
6132:
6128:
6122:
6118:
6114:
6107:
6098:
6093:
6089:
6085:
6081:
6074:
6058:
6054:
6050:
6046:
6042:
6038:
6034:
6030:
6023:
6015:
6011:
6007:
6003:
5999:
5995:
5991:
5987:
5983:
5979:
5972:
5963:
5958:
5954:
5950:
5946:
5939:
5931:
5927:
5922:
5917:
5912:
5907:
5903:
5899:
5895:
5891:
5887:
5880:
5872:
5868:
5864:
5860:
5855:
5850:
5846:
5842:
5838:
5834:
5830:
5828:
5824:
5815:
5807:
5801:
5797:
5790:
5788:
5786:
5784:
5782:
5773:
5769:
5765:
5761:
5757:
5753:
5749:
5745:
5741:
5734:
5726:
5722:
5718:
5714:
5710:
5706:
5702:
5698:
5694:
5690:
5686:
5679:
5671:
5667:
5662:
5657:
5653:
5649:
5645:
5641:
5637:
5633:
5629:
5622:
5603:
5599:
5595:
5590:
5585:
5581:
5577:
5570:
5563:
5544:
5540:
5536:
5532:
5528:
5524:
5520:
5519:
5511:
5509:
5505:
5496:
5494:
5477:
5473:
5469:
5467:
5463:
5454:
5445:
5440:
5436:
5432:
5429:(1): e02531.
5428:
5424:
5420:
5413:
5411:
5409:
5400:
5396:
5392:
5388:
5384:
5380:
5376:
5369:
5350:
5346:
5342:
5335:
5333:
5329:
5320:
5312:
5308:
5304:
5300:
5296:
5292:
5288:
5281:
5279:
5270:
5266:
5262:
5258:
5251:
5235:
5231:
5224:
5218:, p. 18.
5217:
5212:
5205:
5201:
5197:
5192:
5187:
5182:
5177:
5173:
5169:
5165:
5158:
5151:
5139:
5134:
5129:
5125:
5121:
5117:
5113:
5109:
5105:
5101:
5099:
5090:
5083:
5068:
5064:
5060:
5056:
5052:
5045:
5038:
5027:
5023:
5019:
5015:
5011:
5007:
5003:
4999:
4995:
4988:
4986:
4977:
4958:
4954:
4950:
4945:
4940:
4936:
4932:
4928:
4924:
4917:
4910:
4902:
4898:
4894:
4890:
4886:
4882:
4878:
4874:
4867:
4861:, p. 17.
4860:
4855:
4839:
4835:
4833:
4825:
4818:
4814:
4811:
4806:
4799:
4795:
4792:
4787:
4781:, p. 46.
4780:
4775:
4759:
4755:
4751:
4747:
4745:
4737:
4735:
4733:
4725:
4720:
4718:
4710:
4705:
4686:
4679:
4673:
4667:, p. 10.
4666:
4661:
4659:
4650:
4646:
4639:
4632:
4627:
4621:, p. 21.
4620:
4615:
4596:
4592:
4586:
4582:
4575:
4568:
4566:
4558:
4553:
4545:
4541:
4537:
4533:
4529:
4525:
4518:
4511:
4507:
4501:
4490:September 14,
4485:
4481:
4475:
4473:
4465:
4460:
4453:
4448:
4441:
4436:
4434:
4426:
4421:
4415:, p. 45.
4414:
4409:
4401:
4395:
4391:
4384:
4382:
4380:
4378:
4376:
4359:
4355:
4354:"Wild Whales"
4349:
4343:, p. 20.
4342:
4337:
4335:
4326:
4322:
4318:
4311:
4303:
4299:
4295:
4288:
4280:
4276:
4271:
4266:
4262:
4258:
4253:
4248:
4244:
4240:
4236:
4229:
4221:
4215:
4211:
4210:
4202:
4191:
4187:
4183:
4179:
4175:
4171:
4167:
4163:
4156:
4154:
4145:
4143:
4141:
4139:
4137:
4135:
4133:
4125:
4120:
4112:
4108:
4104:
4100:
4096:
4092:
4091:Polar Biology
4088:
4080:
4072:
4068:
4063:
4058:
4054:
4050:
4046:
4042:
4038:
4036:
4027:
4019:
4015:
4011:
4005:
3997:
3993:
3988:
3983:
3979:
3975:
3971:
3967:
3963:
3959:
3955:
3948:
3933:
3929:
3925:
3921:
3917:
3910:
3902:
3898:
3894:
3890:
3885:
3880:
3876:
3872:
3868:
3864:
3860:
3853:
3834:
3830:
3826:
3822:
3818:
3817:
3816:Mammal Review
3809:
3802:
3800:
3792:
3787:
3779:
3775:
3771:
3767:
3762:
3757:
3753:
3749:
3745:
3741:
3740:
3732:
3713:
3709:
3705:
3701:
3697:
3693:
3689:
3682:
3680:
3671:
3669:
3667:
3659:
3656:
3653:
3649:
3646:
3644:
3638:
3634:
3631:
3627:
3621:
3619:
3610:
3606:
3601:
3596:
3592:
3588:
3584:
3580:
3576:
3572:
3568:
3560:
3552:
3548:
3543:
3538:
3534:
3530:
3526:
3519:
3511:
3507:
3503:
3499:
3495:
3491:
3487:
3483:
3476:
3470:, p. 19.
3469:
3464:
3457:
3452:
3450:
3433:
3429:
3425:
3424:
3419:
3413:
3398:
3396:0-486-25651-0
3392:
3388:
3387:
3379:
3371:
3365:
3361:
3354:
3338:
3334:
3330:
3323:
3307:
3303:
3299:
3292:
3285:
3281:
3278:
3273:
3267:, p. 69.
3266:
3261:
3245:
3241:
3234:
3218:
3214:
3210:
3204:
3189:
3185:
3183:
3175:
3159:
3155:
3151:
3145:
3143:
3126:
3121:
3117:
3113:
3112:
3107:
3105:
3096:
3094:
3092:
3090:
3088:
3086:
3069:
3065:
3061:
3059:
3051:
3043:
3039:
3035:
3029:
3025:
3024:
3019:
3018:Wilson, D. E.
3015:
3008:
3004:
2988:
2979:
2975:
2964:
2961:
2954:
2952:
2948:
2947:
2946:
2943:
2942:
2941:
2938:
2937:
2933:
2932:Oceans portal
2927:
2922:
2919:
2908:
2905:
2899:
2894:
2891:
2880:
2873:
2870:
2868:
2867:
2862:
2858:
2854:
2850:
2849:circus tricks
2845:
2844:Minnesota Zoo
2841:
2837:
2833:
2829:
2824:
2822:
2818:
2810:
2806:
2802:
2797:
2796:Captive orcas
2787:
2783:
2780:
2771:
2769:
2765:
2761:
2751:
2749:
2745:
2741:
2737:
2736:Akashi Strait
2733:
2729:
2725:
2721:
2717:
2713:
2709:
2705:
2701:
2693:
2689:
2684:
2679:
2669:
2667:
2663:
2659:
2655:
2651:
2647:
2643:
2639:
2635:
2634:
2627:
2623:
2621:
2617:
2613:
2605:
2600:
2596:
2594:
2590:
2586:
2582:
2578:
2569:
2568:Ingrid Visser
2551:
2541:
2539:
2538:depth charges
2535:
2531:
2527:
2517:
2515:
2505:
2503:
2496:
2486:
2483:
2479:
2474:
2472:
2467:
2464:
2460:
2452:
2447:
2442:
2432:
2430:
2426:
2425:
2420:
2416:
2415:Kuril Islands
2412:
2408:
2403:
2401:
2396:
2392:
2387:
2385:
2381:
2377:
2372:
2370:
2366:
2362:
2358:
2354:
2350:
2349:Kwakwaka'wakw
2345:
2341:
2337:
2330:
2327:sculpture by
2326:
2322:
2317:
2302:
2300:
2295:
2292:
2289:and Alaska's
2288:
2284:
2282:
2276:
2274:
2270:
2266:
2262:
2258:
2254:
2250:
2248:
2244:
2240:
2236:
2232:
2228:
2224:
2220:
2215:
2210:
2208:
2204:
2199:
2195:
2194:Friday Harbor
2191:
2187:
2182:
2180:
2175:
2171:
2167:
2163:
2159:
2155:
2150:
2148:
2144:
2140:
2135:
2133:
2132:
2127:
2123:
2119:
2115:
2111:
2110:harbour seals
2107:
2103:
2099:
2094:
2091:
2087:
2083:
2079:
2075:
2071:
2067:
2063:
2059:
2056:In 2008, the
2049:
2040:
2038:
2034:
2030:
2026:
2024:
2020:
2016:
2011:
2009:
2004:
2000:
1995:
1992:
1988:
1984:
1977:
1976:South Georgia
1972:
1962:
1960:
1956:
1950:
1948:
1943:
1939:
1937:
1933:
1924:
1920:
1918:
1914:
1910:
1905:
1901:
1897:
1891:
1881:
1879:
1875:
1874:McMurdo Sound
1871:
1866:
1862:
1860:
1855:
1852:
1851:marine mammal
1848:
1844:
1839:
1837:
1831:
1819:
1817:
1796:
1778:
1760:
1743:
1742:
1739:
1736:
1735:
1729:Vocalizations
1726:
1723:
1721:
1715:
1711:
1708:
1701:
1696:
1692:
1690:
1685:
1679:
1677:
1673:
1669:
1665:
1655:
1653:
1649:
1645:
1640:
1638:
1634:
1630:
1626:
1622:
1618:
1614:
1606:
1601:
1592:
1590:
1586:
1582:
1578:
1574:
1573:king penguins
1570:
1566:
1562:
1558:
1553:
1550:
1546:
1542:
1537:
1533:
1528:
1526:
1522:
1521:Weddell seals
1517:
1513:
1509:
1505:
1501:
1497:
1493:
1489:
1485:
1481:
1480:harbour seals
1477:
1466:
1459:
1449:
1442:
1433:
1430:
1426:
1422:
1419:
1415:
1411:
1407:
1403:
1396:
1391:
1387:
1385:
1381:
1377:
1373:
1369:
1365:
1361:
1360:beluga whales
1357:
1353:
1349:
1345:
1341:
1337:
1333:
1329:
1319:
1317:
1313:
1309:
1305:
1301:
1297:
1293:
1289:
1285:
1281:
1277:
1273:
1269:
1264:
1262:
1257:
1253:
1249:
1245:
1241:
1237:
1233:
1229:
1219:
1217:
1213:
1209:
1205:
1191:
1180:
1173:
1159:
1150:
1146:
1144:
1140:
1136:
1132:
1131:Hudson Strait
1128:
1124:
1120:
1116:
1107:
1103:
1101:
1100:Faroe Islands
1097:
1093:
1089:
1085:
1081:
1077:
1073:
1069:
1065:
1061:
1057:
1053:
1049:
1045:
1041:
1037:
1030:
1025:
1020:
1010:
1006:
1004:
999:
994:
992:
988:
984:
981:also exhibit
980:
976:
975:pectoral fins
967:
963:
961:
957:
953:
949:
945:
940:
932:
923:
920:
916:
915:
910:
909:
908:Orcinus nanus
904:
899:
897:
893:
889:
885:
880:
876:
872:
867:
864:
860:
856:
852:
848:
844:
840:
834:
824:
822:
818:
814:
810:
809:
804:
803:
798:
794:
790:
789:
784:
780:
776:
772:
767:
765:
761:
757:
756:
751:
747:
743:
742:
736:
735:Carl Linnaeus
732:
731:
726:
722:
708:
696:
689:
679:
675:
674:
667:
653:
651:
647:
646:
639:
637:
634:
630:
626:
617:
616:Ancient Greek
613:
609:
605:
601:
597:
592:
584:
582:
577:
575:
571:
567:
563:
559:
555:
551:
547:
543:
539:
535:
531:
526:
524:
519:
515:
511:
507:
503:
499:
495:
491:
487:
482:
480:
476:
472:
468:
467:
462:
458:
454:
453:toothed whale
450:
446:
445:
440:
426:
423:
418:
415:
410:
407:
406:
404:
401:
397:
393:
390:
381:
377:
372:
367:
363:
357:
355:
349:
346:
345:Binomial name
342:
338:
337:
332:
329:
328:
325:
324:
320:
317:
316:
313:
310:
307:
306:
303:
300:
297:
296:
293:
290:
287:
286:
283:
280:
277:
276:
273:
270:
267:
266:
263:
260:
257:
256:
253:
250:
247:
246:
241:
236:
232:
226:
220:
215:
209:
204:
193:
189:
184:
177:
173:
168:
164:
160:
156:
155:Unimak Island
150:
146:
141:
134:
129:
124:
119:
114:
109:
104:
99:
94:
89:
84:
79:
72:
62:
59:
55:
51:
49:
43:Killer Whale
41:episode, see
40:
33:
19:
13117:
13036:
12911:killer-whale
12719:orcinus-orca
12670:Orcinus_orca
12657:Orcinus_orca
12643:Orcinus orca
12613:Orcinus orca
12612:
12573:
12561:
12550:T. shepherdi
12549:
12537:
12526:M. traversii
12525:
12516:
12507:
12498:
12489:
12480:
12471:
12462:
12453:
12444:
12436:M. europaeus
12435:
12426:
12417:
12408:
12399:
12390:
12370:
12359:I. pacificus
12358:
12346:
12334:
12325:
12305:
12293:
12284:
12275:
12263:
12230:
12218:
12193:
12185:P. gangetica
12184:
12172:
12147:
12135:
12110:
12101:
12092:
12080:
12055:
12047:K. breviceps
12046:
12034:
12009:
11997:
11989:Physeteridae
11972:
11962:Phocoenoides
11960:
11948:
11939:
11930:
11922:P. dioptrica
11921:
11909:
11897:
11888:
11868:
11836:M. monoceros
11835:
11823:
11811:
11799:
11791:Monodontidae
11775:T. truncatus
11774:
11766:T. erebennus
11765:
11757:T. australis
11756:
11747:
11727:
11715:
11703:
11691:
11683:S. frontalis
11682:
11673:
11664:
11656:S. attenuata
11655:
11643:
11631:
11622:
11613:
11605:S. chinensis
11604:
11584:
11572:
11563:
11551:
11539:
11527:
11515:
11503:
11491:
11489:
11479:
11468:O. heinsohni
11467:
11458:
11446:
11434:
11425:
11407:Lissodelphis
11405:
11393:
11384:
11375:
11367:L. australis
11366:
11357:
11348:
11336:
11324:
11312:
11300:
11288:
11276:
11267:
11249:Globicephala
11247:
11236:F. attenuata
11235:
11223:
11211:
11199:
11187:
11178:
11169:
11160:
11148:
11087:C. marginata
11086:
11074:
11049:
11037:
11025:
11024:Gray whale (
11015:Eschrichtius
11013:
11001:
10992:
10983:
10974:
10973:Blue whale (
10965:
10956:
10947:
10938:
10929:
10919:Balaenoptera
10917:
10884:
10876:E. glacialis
10875:
10867:E. australis
10866:
10846:
10834:
10822:
10783:Whippomorpha
10777:Artiodactyla
10572:
10552:
10523:
10489:
10477:. Retrieved
10470:the original
10453:Orcinus orca
10452:
10422:
10389:
10362:Madeira Park
10357:
10330:
10324:
10319:Orcinus orca
10318:
10296:
10267:
10238:
10205:
10184:December 23,
10182:. Retrieved
10167:
10155:. Retrieved
10151:the original
10146:
10137:
10125:. Retrieved
10114:
10105:
10096:
10090:
10080:November 23,
10078:. Retrieved
10067:
10058:
10046:. Retrieved
10039:the original
10026:
10016:December 21,
10014:. Retrieved
9993:
9981:
9969:. Retrieved
9955:
9922:
9918:
9892:. Retrieved
9883:
9874:
9839:
9835:
9812:December 24,
9810:. Retrieved
9803:the original
9798:
9785:
9775:December 24,
9773:. Retrieved
9761:
9757:
9744:
9732:. Retrieved
9714:
9704:December 24,
9702:. Retrieved
9684:
9674:December 24,
9672:. Retrieved
9656:
9629:. Retrieved
9620:: 54. 1998.
9617:
9609:Orcinus orca
9608:
9601:
9589:. Retrieved
9580:
9570:
9558:. Retrieved
9540:
9521:
9509:. Retrieved
9500:
9490:
9478:
9466:
9454:
9441:
9437:
9434:Orcinus orca
9433:
9427:
9414:
9410:
9406:
9400:
9375:
9371:
9365:
9346:
9341:
9329:
9308:
9296:. Retrieved
9267:. Retrieved
9256:
9246:
9234:. Retrieved
9217:
9207:
9195:. Retrieved
9186:
9177:
9165:. Retrieved
9154:The Guardian
9153:
9143:
9131:. Retrieved
9122:
9112:
9100:. Retrieved
9091:
9082:
9070:. Retrieved
9061:
9052:
9040:. Retrieved
9025:
9016:
9004:. Retrieved
8995:
8986:
8967:
8961:
8949:. Retrieved
8945:the original
8938:
8929:
8918:John Bostock
8893:
8880:
8871:
8841:
8831:
8818:. Retrieved
8786:. Retrieved
8782:the original
8777:
8756:the original
8751:
8727:
8722:
8710:
8706:
8702:
8669:
8665:
8659:
8639:
8629:December 10,
8627:. Retrieved
8616:
8606:
8594:
8567:December 31,
8565:. Retrieved
8558:the original
8545:
8535:December 31,
8533:. Retrieved
8524:
8514:
8504:February 25,
8502:. Retrieved
8474:
8470:
8457:
8445:. Retrieved
8436:
8426:
8407:
8395:
8383:. Retrieved
8374:
8348:. Retrieved
8344:the original
8339:
8287:the original
8282:
8257:
8245:. Retrieved
8236:
8226:
8175:
8171:
8165:
8153:. Retrieved
8145:The Guardian
8144:
8134:
8106:
8097:
8085:. Retrieved
8071:
8052:
8036:Orcinus orca
8035:
8028:
8017:Orcinus orca
8016:
8011:
7999:
7987:
7962:
7958:
7955:Orcinus orca
7954:
7948:
7940:the original
7909:
7905:
7900:Orcinus orca
7899:
7876:. Retrieved
7861:
7849:. Retrieved
7840:
7830:
7803:. Retrieved
7799:the original
7794:
7773:
7767:
7732:
7728:
7722:
7695:
7691:
7666:. Retrieved
7633:
7629:
7619:
7582:
7578:PLOS Biology
7576:
7566:
7559:Live Science
7558:
7542:
7531:the original
7526:
7522:
7510:
7487:The Guardian
7486:
7476:
7453:
7389:
7385:
7375:
7365:December 23,
7363:. Retrieved
7354:
7344:
7332:. Retrieved
7328:the original
7323:
7313:
7303:February 27,
7301:. Retrieved
7292:
7282:
7260:(2): 47–53.
7257:
7253:
7249:
7246:Orcinus orca
7245:
7239:
7213:the original
7184:
7180:
7167:
7157:February 23,
7155:. Retrieved
7119:
7116:Bioacoustics
7115:
7107:Orcinus orca
7106:
7098:
7073:
7069:
7066:Orcinus orca
7065:
7031:
7027:
7024:Orcinus orca
7023:
7017:
6992:
6988:
6982:
6955:. Retrieved
6946:
6936:
6909:10871/125706
6891:
6887:
6877:
6865:
6838:
6819:
6818:Orcinus orca
6815:
6809:
6767:February 16,
6765:. Retrieved
6761:the original
6751:
6724:. Retrieved
6715:
6706:
6696:November 24,
6694:. Retrieved
6675:
6663:
6653:February 18,
6651:. Retrieved
6637:
6625:
6592:
6588:
6582:
6570:. Retrieved
6551:
6539:
6527:
6515:. Retrieved
6495:
6488:
6453:
6449:
6439:
6414:
6410:
6404:
6359:
6355:
6345:
6326:
6291:
6280:the original
6259:
6255:
6247:Orcinus orca
6246:
6238:
6226:. Retrieved
6212:(1): 16–36.
6209:
6205:
6200:Orcinus orca
6199:
6191:
6164:
6137:December 30,
6135:. Retrieved
6116:
6106:
6087:
6083:
6073:
6063:February 25,
6061:. Retrieved
6036:
6032:
6022:
5989:
5985:
5981:
5978:Orcinus orca
5977:
5971:
5952:
5948:
5938:
5893:
5889:
5879:
5836:
5832:
5827:Orcinus orca
5826:
5822:
5814:
5795:
5747:
5743:
5733:
5692:
5688:
5678:
5635:
5631:
5621:
5609:. Retrieved
5602:the original
5582:(1): 53–68.
5579:
5575:
5562:
5550:. Retrieved
5543:the original
5522:
5516:
5508:Orcinus orca
5507:
5503:
5482:February 18,
5480:. Retrieved
5476:the original
5471:
5466:Orcinus orca
5465:
5461:
5453:
5426:
5422:
5382:
5378:
5375:Orcinus orca
5374:
5368:
5356:. Retrieved
5344:
5340:
5331:
5328:Orcinus orca
5327:
5319:
5305:(1): 83–88.
5302:
5298:
5295:Orcinus orca
5294:
5290:
5286:
5260:
5256:
5250:
5238:. Retrieved
5223:
5211:
5203:
5171:
5167:
5157:
5149:
5142:. Retrieved
5107:
5103:
5098:Orcinus orca
5097:
5089:
5081:
5074:. Retrieved
5054:
5050:
5037:
5026:the original
4997:
4993:
4985:Orcinus orca
4984:
4976:
4964:. Retrieved
4926:
4922:
4909:
4876:
4872:
4866:
4854:
4842:. Retrieved
4838:the original
4832:Orcinus orca
4831:
4824:
4805:
4786:
4774:
4762:. Retrieved
4758:the original
4749:
4744:Orcinus orca
4743:
4704:
4692:. Retrieved
4685:the original
4672:
4649:the original
4638:
4626:
4614:
4604:February 25,
4602:. Retrieved
4580:
4552:
4527:
4523:
4517:
4510:Orcinus orca
4509:
4505:
4500:
4488:. Retrieved
4484:the original
4459:
4447:
4420:
4408:
4389:
4362:. Retrieved
4358:the original
4348:
4316:
4310:
4293:
4287:
4242:
4238:
4228:
4208:
4201:
4190:the original
4164:(304): 1–9.
4161:
4153:Orcinus orca
4152:
4119:
4094:
4090:
4087:Orcinus orca
4086:
4079:
4044:
4040:
4035:Orcinus orca
4034:
4026:
4020:. June 2024.
4013:
4004:
3961:
3957:
3947:
3935:. Retrieved
3923:
3919:
3909:
3866:
3862:
3852:
3842:February 23,
3840:. Retrieved
3820:
3814:
3786:
3746:(1): 62–80.
3743:
3737:
3731:
3719:. Retrieved
3691:
3687:
3679:Orcinus orca
3678:
3657:
3654:
3643:Orcinus orca
3642:
3574:
3570:
3559:
3532:
3528:
3518:
3485:
3481:
3475:
3463:
3436:. Retrieved
3422:
3418:Linnaeus, C.
3412:
3400:. Retrieved
3385:
3378:
3359:
3353:
3341:. Retrieved
3332:
3322:
3310:. Retrieved
3301:
3291:
3272:
3260:
3248:. Retrieved
3233:
3221:. Retrieved
3212:
3203:
3191:. Retrieved
3182:Orcinus orca
3181:
3174:
3162:. Retrieved
3153:
3131:November 19,
3129:. Retrieved
3115:
3109:
3104:Orcinus orca
3103:
3074:December 17,
3072:. Retrieved
3068:the original
3063:
3058:Orcinus orca
3057:
3050:
3022:
3007:
2987:
2978:
2949:
2871:
2865:
2825:
2817:intelligence
2814:
2784:
2777:
2757:
2697:
2650:Arctic Ocean
2631:
2628:
2624:
2616:Michael Bigg
2609:
2592:
2573:
2523:
2511:
2498:
2478:photographer
2475:
2465:
2456:
2422:
2404:
2388:
2380:Newfoundland
2373:
2333:
2296:
2291:Kenai Fjords
2281:Exxon Valdez
2280:
2277:
2265:salmon farms
2251:
2211:
2183:
2166:Haro Straits
2151:
2136:
2129:
2095:
2055:
2043:Conservation
2027:
2012:
1996:
1980:
1959:Orcinus orca
1958:
1952:
1944:
1940:
1929:
1896:sperm whales
1893:
1884:Intelligence
1867:
1863:
1856:
1840:
1833:
1813:
1737:
1724:
1716:
1712:
1704:
1680:
1661:
1641:
1610:
1554:
1529:
1473:
1467:off the floe
1423:
1414:minke whales
1402:sperm whales
1399:
1368:pilot whales
1325:
1312:whale sharks
1265:
1225:
1201:
1156:
1149:California.
1147:
1139:Fraser River
1112:
1033:
1007:
998:echolocation
995:
972:
941:
937:
912:
906:
900:
891:
887:
883:
874:
870:
868:
862:
836:
820:
806:
800:
793:sister taxon
786:
783:pilot whales
768:
753:
749:
740:
728:
721:Orcinus orca
720:
719:
671:
643:
640:
624:
611:
607:
595:
593:
590:
578:
554:habitat loss
527:
483:
464:
449:killer whale
448:
444:Orcinus orca
443:
442:
438:
436:
424:
416:
408:
389:Orcinus orca
388:
354:Orcinus orca
353:
351:
336:O. orca
335:
334:
322:
298:Infraorder:
292:Artiodactyla
66:Killer whale
58:
48:The Avengers
47:
39:The Avengers
38:
12981:SeaLifeBase
12867:NatureServe
12789:iNaturalist
12637:Wikispecies
12481:M. layardii
12400:M. bowdoini
12348:Indopacetus
11931:P. phocoena
11870:Neophocoena
11858:(Porpoises)
11852:Phocoenidae
11426:L. borealis
11394:L. obscurus
11376:L. cruciger
11170:C. eutropia
11132:Delphinidae
11117:Odontoceti
11026:E. robustus
10993:B. physalus
10991:Fin whale (
10975:B. musculus
10948:B. borealis
10946:Sei whale (
10885:E. japonica
10769:Superorder
10763:Infraclass
10333:: 185–199.
10275:. pp.
10197:Works cited
10064:"Blackfish"
10048:January 26,
9894:October 31,
9764:: 119–139.
9724:Shimonoseki
9694:Shimonoseki
9631:January 14,
9560:November 6,
8951:October 13,
8302:Le Phuong.
8201:10023/16189
7735:: 305–341.
6517:February 2,
5638:(1): 6153.
5611:January 18,
5144:December 6,
4966:October 23,
4944:10125/22585
4694:January 26,
3645:, in Canada
3438:January 14,
3402:January 28,
3223:February 8,
3164:January 14,
3064:Fossilworks
2815:The orca's
2662:Puget Sound
2534:Puget Sound
2441:Orca attack
2429:brown bears
2424:Repun Kamuy
2407:Ainu people
2261:Haro Strait
2186:Ken Balcomb
2170:Puget Sound
2143:Puget Sound
2126:polar bears
2052:protection.
2029:Infanticide
1991:Granny (J2)
1983:polyestrous
1917:Snæfellsnes
1913:pilot whale
1900:gray matter
1830:Whale sound
1549:cannibalism
1418:blue whales
1406:grey whales
1296:blue sharks
1256:pink salmon
1248:Chum salmon
1216:sea turtles
1210:, mammals,
1208:cephalopods
1179:Vestfjorden
1074:and in the
979:Dorsal fins
636:Delphinidae
518:matrilineal
312:Delphinidae
221:Appendix II
13124:Categories
12539:Tasmacetus
12499:M. perrini
12472:M. hotaula
12463:M. hectori
12372:Mesoplodon
12307:Hyperoodon
12294:B. minimus
12285:B. bairdii
12276:B. arnuxii
12220:Pontoporia
12174:Platanista
11748:T. aduncus
11665:S. clymene
11632:S. teuszii
11614:S. plumbea
11516:P. electra
11435:L. peronii
11301:G. griseus
11212:D. delphis
11188:C. hectori
11115:Parvorder
10907:(Rorquals)
10814:Balaenidae
10799:Mysticeti
10797:Parvorder
10479:January 2,
10395:Stillwater
10242:. London:
10211:Stillwater
9734:January 9,
9618:自然環境保全基礎調査
9471:Baird 2002
9298:January 3,
9042:January 3,
7851:January 3,
6630:Baird 2002
6532:Baird 2002
6319:Baird 2002
6184:Baird 2002
5833:Acta Ethol
5750:(1): 1–5.
5133:2263/49986
5110:: 90–102.
5076:August 23,
4764:August 15,
4665:Baird 2002
4631:Baird 2002
4124:Baird 2002
3761:2263/21531
3721:January 9,
3626:Baird 2002
3343:October 9,
3312:October 7,
3213:Whales.org
2999:References
2836:dorsal fin
2740:Harimanada
2633:Free Willy
2548:See also:
2473:, above).
2451:Greifswald
2400:sacrifices
2378:people of
2314:See also:
2245:, and the
2223:Ecojustice
2184:Scientist
2114:Washington
2086:oil spills
1999:inbreeding
1966:Life cycle
1878:Antarctica
1828:See also:
1816:media help
1759:Orca calls
1684:matrilines
1635:and other
1629:Spyhopping
1581:cormorants
1543:, such as
1276:eagle rays
1202:Orcas are
1153:Population
1145:in Japan.
1121:much like
1088:Washington
1080:Antarctica
1056:California
1029:Hood Canal
960:melanistic
892:O. o. orca
843:subspecies
550:endangered
546:threatened
508:and other
484:Orcas are
157:, eastern
13052:Q67206427
12391:M. bidens
12265:Berardius
12247:Ziphiidae
12127:Lipotidae
11938:Vaquita (
11834:Narwhal (
11812:D. leucas
11529:Pseudorca
11358:L. acutus
11201:Delphinus
11039:Megaptera
10984:B. omurai
10957:B. brydei
10848:Eubalaena
10781:Suborder
10600:Orca-Live
10542:643483454
10534:0027-9358
9986:NMFS 2005
9971:March 14,
9511:March 19,
9438:Zoologica
9411:Zoologica
9334:NMFS 2005
9226:0307-1235
9162:0261-3077
9006:March 23,
8820:April 29,
8788:August 7,
8694:163391715
8385:March 19,
8350:March 19,
8087:March 22,
8004:NMFS 2005
7992:NMFS 2005
7416:1932-6203
7293:New Atlas
7232:NMFS 2005
6975:NMFS 2005
6858:NMFS 2005
6843:NMFS 2005
6802:NMFS 2005
6787:NMFS 2005
6572:August 7,
6014:246167673
5896:(12): 6.
5871:249487897
5772:1814-232X
5725:250118179
5717:1814-232X
5584:CiteSeerX
5552:April 28,
5423:Ecosphere
5216:NMFS 2005
4859:NMFS 2005
4779:NMFS 2005
4709:NMFS 2005
4512:) Whales"
4364:March 23,
4325:0028-0402
4302:0028-0402
4261:2076-2615
4245:(8): 49.
4186:253914153
3937:March 22,
3591:1063-5157
3510:0824-0469
3154:cites.org
2970:Footnotes
2953:melvillei
2866:Blackfish
2807:, at the
2790:Captivity
2748:Tokyo Bay
2728:Greenland
2724:Indonesia
2716:Antarctic
2692:whaleboat
2654:pneumonia
2593:Zoologica
2577:Moby Doll
2502:Gibraltar
2353:sea lions
2340:their art
2329:Bill Reid
2283:oil spill
2203:CAT scans
2082:pollution
2003:Gestation
1955:sympatric
1936:longlines
1932:anecdotes
1836:cetaceans
1834:Like all
1664:elephants
1633:porpoises
1621:parasites
1595:Behaviour
1536:sea otter
1525:"spy-hop"
1332:porpoises
1314:and even
1280:long-tail
1232:Greenland
1228:Norwegian
1064:Norwegian
922:species.
903:Antarctic
695:Naturalis
330:Species:
258:Kingdom:
252:Eukaryota
73:to recent
13046:Wikidata
12994:Species+
12872:2.105428
12859:14300074
12807:10193882
12742:46559273
12677:BioLib:
12622:Wikidata
12563:Ziphius
12490:M. mirus
12454:M. grayi
12194:P. minor
12056:K. simus
12026:Kogiidae
11999:Physeter
11973:P. dalli
11940:P. sinus
11911:Phocoena
11729:Tursiops
11645:Stenella
11562:Tucuxi (
11448:Orcaella
11325:L. hosei
11277:G. melas
11002:B. ricei
10966:B. edeni
10765:Eutheria
10759:Mammalia
10753:Chordata
10747:Animalia
10745:Kingdom
10689:Archived
10677:Archived
10664:Archived
10652:Archived
10634:Archived
10616:Archived
10603:Archived
10464:, U.S.:
10178:Archived
10121:Archived
10074:Archived
10007:Archived
9965:Archived
9947:21276987
9888:Archived
9866:17148221
9766:Archived
9728:Archived
9726:. 2014.
9698:Archived
9696:. 2014.
9668:Archived
9622:Archived
9591:April 8,
9585:Archived
9554:Archived
9529:Archived
9505:Archived
9448:: 59–70.
9421:: 71–76.
9354:Archived
9318:Archived
9292:Archived
9263:Archived
9230:Archived
9191:Archived
9127:Archived
9123:Newsweek
9096:Archived
9066:Archived
9062:NBC News
9036:Archived
9000:Archived
8910:Archived
8898:Archived
8886:Archived
8814:Archived
8648:Archived
8623:Archived
8529:Archived
8495:Archived
8491:55958971
8447:June 25,
8441:Archived
8415:Archived
8379:Archived
8375:CBC News
8307:Archived
8241:Archived
8218:52876312
8210:30262502
8149:Archived
8111:Archived
8107:BBC News
8081:Archived
8060:Archived
8041:Archived
7936:26937049
7872:Archived
7845:Archived
7668:March 7,
7662:Archived
7658:24052064
7650:11530544
7611:17503965
7551:Archived
7454:CBS News
7434:31841529
7386:PLOS ONE
7359:Archived
7334:March 7,
7297:Archived
7274:29828311
7209:16454316
7148:Archived
7144:56304541
7090:22866769
7028:Ethology
7009:16899659
6957:June 18,
6951:Archived
6928:34130498
6894:(1953).
6720:Archived
6690:Archived
6647:Archived
6617:44055219
6566:Archived
6562:CBC News
6511:Archived
6480:19451116
6396:14526101
6334:Archived
6300:Archived
6228:July 27,
6222:Archived
6131:Archived
6090:: 7–58.
6057:Archived
5930:30540762
5890:PLOS ONE
5863:35694552
5670:30992478
5349:Archived
5240:July 25,
5234:Archived
5200:91256980
5138:Archived
5067:Archived
5022:84721171
4957:Archived
4953:16788148
4901:21252323
4844:June 26,
4813:Archived
4794:Archived
4595:Archived
4279:27548232
4111:20734772
4071:20413674
3996:38545612
3987:10966402
3893:17301765
3833:Archived
3770:22882545
3712:Archived
3708:52257732
3648:Archived
3633:Archived
3609:31633766
3551:91933816
3432:Archived
3420:(1758).
3337:Archived
3306:Archived
3280:Archived
3250:July 30,
3244:Archived
3217:Archived
3193:March 9,
3158:Archived
3042:62265494
2951:Livyatan
2876:See also
2842:and the
2840:SeaWorld
2830:and the
2658:Springer
2604:Springer
2471:"Naming"
2419:Sakhalin
2411:Hokkaido
2124:than in
1637:dolphins
1613:foraging
1557:seabirds
1516:beaching
1500:walruses
1364:narwhals
1334:such as
1310:, small
1212:seabirds
1181:, Norway
1115:pack ice
1098:and the
859:taxonomy
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