266:'s first draft featured early versions of the MUTOs where they were established as ancient enemies of Godzilla but never established as to why. When Gareth Edwards came aboard the project, he created a back story to bridge the gap between Godzilla's connection with the MUTOs. Edwards brainstormed the idea that, "When these Godzillas were on Earth, there was another creature that would kill them and lay its eggs inside their dead bodies. Therefore, if these creatures ever came back, part of their life cycle would be the ability to attract Godzillas to the surface to kill them for reproduction."
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species. The female is much larger and walks on eight limbs—the male is much smaller, with one pair of his eight limbs modified into wings for powered flight. Though the MUTOs have an arthropod-like appearance, filmmaker
Guillaume Rocheron likened them more to vertebrates. The angularity of the male
294:
for inspiration and reflected back on what made these monsters and their designs so iconic. From this, the design for the MUTO monsters kept evolving and "mutating", according to
Edwards, into a design he felt was more cohesive. Edwards has credited artist Matt Allsopp for creating the majority of
269:
In an interview, Edwards commented that it took over a year to design the MUTO creatures, stressing that it took that long to create something that was aimed to be new and different for today's day and age. Edwards and the design team looked towards past monster characters from such films as
307:. As a side-effect of the radiation they absorb as food, the MUTOs are capable of causing electromagnetic interference, the male emitting EMP shockwaves from his claws and the female having an EMP field "Sphere of Influence" surrounding her instead.
321:
called the MUTOs "Godzilla's most important co-stars", praising them as threatening antagonists that encourage the audience to root for
Godzilla. Some viewers have been critical of the MUTOs' design, which was perceived to be similar to that of the
295:
the MUTOs designs. Edwards stated, "The DNA of the MUTO is 80 to 90 percent from Matt, he was the main guy". Others contributed to the design of the MUTOs as well such as Weta, Rob Bliss, Steambot, and Legacy, who provided a 3-D model of the MUTOs.
337:
added the MUTOs to their list of "Weakest to
Strongest" MonsterVerse Kaiju, calling the male "nimble and durable" and the female "strong enough to toss Godzilla around" and that they pose a serious threat to Godzilla when partnered.
203:(2014). An early draft of the screenplay differentiated the male and female MUTO as Hokmuto and Femuto. Official merchandise connected with the film refer to the male MUTO as "Winged MUTO" and the female as "Eight-Legged MUTO".
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for monsters. Edwards noted that it took over a year to create a design intended to be new and different and credited a majority of the design to Matt
Allsopp; inspiration was drawn from past American monster films.
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René H.B. Fraaije, Eduard V. Mychko, Leonid S. Barsukov, John W.M. Jagt (2023). "A new mid-Cretaceous hermit crab (Crustacea, Anomura) from
Central Russia sheds new light on paguroid evolution".
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164:. While the term "MUTO" is mainly used to label the two parasitic monsters, it is intended to flag unidentified creatures. Edwards likened the term to
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The term "MUTO" is also used to reference giant monsters who are yet to be named. Such was the case in the official prequel comic,
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stated that "it's basically the UFO of monsters". However, the term is also used as a name for the two parasitic monsters in
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The MUTOs have been positively received by critics and journalists; however, their designs had drawn comparisons with the
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384:(2017). The parent super-species to the MUTOs, dubbed MUTO Prime and Titanus Jinshin-Mushi, appears in the graphic novel
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era that feeds on radiation, is identified as a MUTO before being designated as
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232:(2014), specializes in the hunt for Massive Unidentified Terrestrial Organisms.
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855:"Meet MUTO Prime: The Latest Confirmed Titan in Godzilla: King of the Monsters"
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193:"MUTO" is an acronym for "Massive Unidentified Terrestrial Organism". Director
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The term "MUTO" (Massive
Unidentified Terrestrial Organism) was referenced in
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176:. The characters and their variants had appeared in other media outside of
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917:"The Godzilla Easter Egg Star Wars: Rogue One Snuck In For Gareth Edwards"
886:"Godzilla: King of the Monsters Monster List: All 17 Titan Names Revealed"
262:, Legendary Pictures had planned to feature two new monsters in addition.
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700:"Godzilla" Director Gareth Edwards explains why monsters still matter
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238:(2019) changes the monsters' designation from "MUTOs" to "Titans".
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Godzilla, Mothra and King
Ghidorah: Giant Monsters All-Out Attack
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783:"The 10 Major Monsterverse Kaiju Ranked Weakest To Strongest"
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Godzilla: Awakening, Max and Greg
Borenstein, Chapter 1
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228:) states that the Monarch agency, first introduced in
154:. The characters first appeared as the antagonists in
610:"Good Morning Godzilla — More on New Kaiju Muto"
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663:"Godzilla 2 Has Renamed The MonsterVerse Creatures"
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402:and the creatures from Edwards' directorial debut
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408:(2010), were briefly seen in cave paintings in
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838:: CS1 maint: multiple names: authors list (
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487:"Screenwriter Profile: Dave Callaham"
16:Fictional species in the MonsterVerse
1818:Godzilla: King of the Monsters in 3D
927:from the original on January 6, 2023
781:Wooldridge, Zachary (July 6, 2022).
762:Woerner, Meredith (March 12, 2014).
13:
2447:Film characters introduced in 2014
1123:Ghidorah, the Three-Headed Monster
915:Holmes, Adam (November 30, 2016).
793:from the original on July 20, 2022
608:Barton, Steve (October 28, 2013).
303:MUTOs wing design was inspired by
125:Titanus Jinshin-Mushi (MUTO Prime)
14:
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2045:Godzilla and Godzilla Raids Again
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865:from the original on June 2, 2019
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1839:Always: Sunset on Third Street 2
970:Godzilla: The Art of Destruction
853:Millican, Josh (April 2, 2019).
743:Gettell, Oliver (May 20, 2014).
661:Orquiola, John (July 21, 2018).
298:The film depicts the MUTOs as a
2322:Godzilla x Kong: The New Empire
2248:Godzilla x Kong: The New Empire
1738:Godzilla x Kong: The New Empire
1525:Godzilla x Kong: The New Empire
1438:Godzilla, King of the Monsters!
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884:Whalen, Andrew (May 31, 2019).
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258:Upon acquiring the license for
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2015:Battle Soccer: Field no Hasha
1804:The Beast from 20,000 Fathoms
1257:Godzilla vs. Mechagodzilla II
826:10.1016/j.cretres.2023.105749
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140:) are fictional monsters, or
1825:Godzilla vs. Charles Barkley
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252:Godzilla: Art of Destruction
152:MonsterVerse media franchise
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2467:Fictional species and races
2275:Monarch: Legacy of Monsters
1985:Godzilla in popular culture
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2109:Shin Japan Heroes Universe
1832:Monster Planet of Godzilla
1610:Chibi Godzilla Raids Again
1396:City on the Edge of Battle
1265:Godzilla vs. SpaceGodzilla
1241:Godzilla vs. King Ghidorah
1195:Godzilla vs. Mechagodzilla
1139:Ebirah, Horror of the Deep
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55:The MUTOs as depicted in
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1878:Monsterverse incarnation
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993:. Warner Bros. Pictures.
987:Edwards, Gareth (2014).
951:Callaham, David (2012).
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398:. The MUTOs, along with
224:, Bill Randa (played by
2477:Monsterverse characters
2123:Neon Genesis Evangelion
2039:(Cerasini novel series)
1602:Godzilla Singular Point
1300:Godzilla vs. Megaguirus
1273:Godzilla vs. Destoroyah
1203:Terror of Mechagodzilla
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113:In-universe information
2104:Hibiya Godzilla Square
2031:(Ciencin novel series)
2008:The Movie Monster Game
1388:Planet of the Monsters
1233:Godzilla vs. Biollante
1225:The Return of Godzilla
1107:King Kong vs. Godzilla
998:Nelson, Arvid (2019).
315:Oliver Gettell of the
255:
210:, where a hive-minded
1687:Godzilla (Main Theme)
454:Godzilla: Strike Zone
249:
1811:Bambi Meets Godzilla
1758:" (Blue Ă–yster Cult)
1717:King of the Monsters
1637:Godzilla: The Series
1187:Godzilla vs. Megalon
1171:Godzilla vs. Hedorah
1155:Destroy All Monsters
1099:Godzilla Raids Again
1002:. Legendary Comics.
1000:Godzilla: Aftershock
972:. Insight Editions.
466:Godzilla: Aftershock
386:Godzilla: Aftershock
160:(2014), directed by
19:For other uses, see
2437:Godzilla characters
1694:Godzilla: The Album
1249:Godzilla vs. Mothra
1163:All Monsters Attack
1115:Mothra vs. Godzilla
814:Cretaceous Research
208:Godzilla: Awakening
174:Cloverfield monster
121:Winged MUTO (male)
26:Fictional character
2457:Fictional monsters
2301:Kong: Skull Island
2224:Kong: Skull Island
1731:Godzilla Minus One
1492:Legendary Pictures
1450:New World Pictures
1369:Godzilla Minus One
1179:Godzilla vs. Gigan
732:Legendary Pictures
543:. April 15, 2014.
392:, was featured in
381:Kong: Skull Island
300:sexually dimorphic
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148:Legendary Pictures
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637:, p. 27, 56.
612:. Dread Central.
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568:, p. 110.
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468:(comic - 2019)
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2442:Toho monsters
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2344:King Ghidorah
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2022:Godzilla Game
2019:
2017:
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2012:
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1995:
1991:
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1954:SpaceGodzilla
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1939:Mechagodzilla
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1909:King Ghidorah
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1853:Shin Ultraman
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1786:Miscellaneous
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1459:Godzilla 1985
1456:
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1361:Shin Godzilla
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1292:Godzilla 2000
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960:(Screenplay).
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859:Dread Central
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689:, p. 22.
688:
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635:Callaham 2012
631:
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523:, p. 82.
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273:Jurassic Park
267:
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212:superorganism
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136:(acronym for
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30:
22:
2414:
2393:
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2320:
2313:
2306:
2299:
2292:
2273:
2267:Skull Island
2265:
2246:
2238:
2230:
2222:
2214:
2201:Monsterverse
2130:
2122:
2114:
2096:
2091:Monsterverse
2071:Miki Saegusa
2043:
2036:
2028:
2020:
2013:
2006:
1968:
1863:
1851:
1844:
1837:
1830:
1823:
1816:
1809:
1802:
1736:
1729:
1722:
1715:
1708:
1700:
1692:
1643:
1635:
1627:
1608:
1600:
1590:(1997–1998)
1585:
1577:
1571:Zone Fighter
1569:
1523:
1515:
1507:
1499:
1478:
1457:
1436:
1402:
1394:
1386:
1367:
1359:
1343:
1330:
1324:Tokyo S.O.S.
1322:
1314:
1306:
1298:
1290:
1271:
1263:
1255:
1247:
1239:
1231:
1223:
1214:
1201:
1193:
1185:
1177:
1169:
1161:
1153:
1145:
1137:
1129:
1121:
1113:
1105:
1097:
1089:
1080:
1048:
999:
988:
969:
953:
929:. Retrieved
920:
910:
898:. Retrieved
889:
879:
867:. Retrieved
858:
848:
834:cite journal
817:
813:
807:
795:. Retrieved
786:
776:
767:
757:
748:
738:
723:
711:
694:
682:
670:. Retrieved
666:
642:
630:
618:. Retrieved
603:
591:. Retrieved
573:
561:
551:November 15,
549:. Retrieved
540:
499:. Retrieved
491:Fresh Voices
490:
481:
465:
453:
446:
431:
423:
409:
403:
393:
385:
379:
377:
344:Mutotylaspis
342:
340:
332:
326:
324:monster from
316:
314:
297:
289:
283:
277:
271:
268:
257:
251:
233:
229:
226:John Goodman
219:
207:
205:
198:
192:
177:
171:
155:
141:
137:
133:
131:
108:Matt Allsopp
81:
68:
56:
40:MonsterVerse
37:
2396:(franchise)
2388:(franchise)
2286:Soundtracks
2115:Kamen Rider
2056:Rhedosaurus
2002:Video games
1679:Soundtracks
1640:(1998–2000)
1632:(1978–1979)
1582:(1992–1996)
1353:Live-action
921:CinemaBlend
672:October 17,
667:Screen Rant
620:October 16,
566:Nelson 2019
541:SciFi Japan
441:Video games
374:Appearances
352:hermit crab
328:Cloverfield
242:Development
105:Designed by
2431:Categories
2259:Television
1959:Megaguirus
1765:" (Eminem)
1702:Final Wars
1579:Godzilland
1553:Television
1332:Final Wars
931:January 5,
820:: 105749.
501:August 14,
473:References
460:Literature
356:Cretaceous
91:Created by
2394:King Kong
2349:King Kong
2133:franchise
2125:franchise
2117:franchise
2099:franchise
2097:King Kong
1944:Biollante
1904:King Kong
593:August 1,
411:Rogue One
366:(Central
341:In 2023,
311:Reception
291:King Kong
214:from the
44:character
2415:Category
2386:Godzilla
2339:Godzilla
2332:Monsters
2294:Godzilla
2216:Godzilla
2159:Category
2131:Ultraman
2037:Godzilla
2029:Godzilla
1884:Anguirus
1874:Godzilla
1763:Godzilla
1756:Godzilla
1709:Godzilla
1629:Godzilla
1621:American
1594:episodes
1563:Japanese
1501:Godzilla
1480:Godzilla
1423:American
1380:Animated
1091:Godzilla
1073:Japanese
1050:Godzilla
990:Godzilla
968:(2014).
954:Godzilla
925:Archived
894:Archived
890:Newsweek
863:Archived
797:July 20,
791:Archived
787:Collider
749:LA Times
730:, 2014,
728:Godzilla
614:Archived
587:Archived
583:Facebook
545:Archived
495:Archived
425:Godzilla
405:Monsters
400:Godzilla
349:paguroid
334:Collider
260:Godzilla
230:Godzilla
200:Godzilla
184:Overview
180:(2014).
178:Godzilla
157:Godzilla
70:Godzilla
57:Godzilla
2378:Related
2076:Super X
2066:Gotengo
1934:Megalon
1924:Hedorah
1919:Minilla
1914:Baragon
1748:Related
944:Sources
900:June 1,
869:June 1,
705:YouTube
216:Permian
2359:Mothra
2278:(2023)
2270:(2023)
2251:(2024)
2243:(2021)
2235:(2019)
2227:(2017)
2219:(2014)
2061:Jirahs
2051:G-Fest
1997:Comics
1990:-zilla
1899:Mothra
1712:(2014)
1697:(1998)
1648:(2023)
1613:(2023)
1605:(2021)
1574:(1973)
1528:(2024)
1520:(2021)
1512:(2019)
1504:(2014)
1483:(1998)
1462:(1985)
1441:(1956)
1407:(2018)
1399:(2018)
1391:(2017)
1372:(2023)
1364:(2016)
1335:(2004)
1327:(2003)
1319:(2002)
1311:(2001)
1303:(2000)
1295:(1999)
1276:(1995)
1268:(1994)
1260:(1993)
1252:(1992)
1244:(1991)
1236:(1989)
1228:(1984)
1215:Heisei
1206:(1975)
1198:(1974)
1190:(1973)
1182:(1972)
1174:(1971)
1166:(1969)
1158:(1968)
1150:(1967)
1142:(1966)
1134:(1965)
1126:(1964)
1118:(1964)
1110:(1962)
1102:(1955)
1094:(1954)
1006:
976:
436:(2019)
428:(2014)
368:Russia
360:Albian
86:(2019)
73:(2014)
59:(2014)
2472:Kaiju
2369:Rodan
2208:Films
1978:Other
1964:Zilla
1929:Gigan
1894:Varan
1889:Rodan
1865:Kaiju
1796:Films
1669:Music
1344:Reiwa
1081:ShĹŤwa
1063:Films
958:(PDF)
418:Films
279:Alien
146:, in
143:kaiju
134:MUTOs
118:Alias
2364:MUTO
2081:Toho
1969:MUTO
1004:ISBN
974:ISBN
933:2022
902:2019
871:2019
840:link
799:2022
674:2018
622:2022
595:2014
553:2020
503:2015
347:, a
288:and
189:Name
132:The
32:MUTO
21:MutĹŤ
1346:era
1217:era
1083:era
822:doi
818:154
768:io9
703:on
166:UFO
2433::
923:.
919:.
892:.
888:.
861:.
857:.
836:}}
832:{{
816:.
789:.
785:.
766:.
747:.
665:.
651:^
585:.
581:.
539:.
528:^
511:^
493:.
489:.
331:.
282:,
276:,
150:'
2193:e
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2179:v
1880:)
1876:(
1761:"
1754:"
1689:"
1685:"
1041:e
1034:t
1027:v
1012:.
982:.
935:.
904:.
873:.
842:)
828:.
824::
801:.
770:.
751:.
676:.
624:.
597:.
555:.
505:.
358:(
254:.
23:.
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
