450:) are a group of reef fish in which protogyny is common. Wrasses also have an uncommon life history strategy, which is termed diandry (literally, two males). In these species, two male morphs exists: an initial phase male and a terminal phase male. Initial phase males do not look like males and spawn in groups with females. They are not territorial. They are, perhaps, female mimics (which is why they are found swimming in group with females). Terminal phase males are territorial and have a distinctively bright coloration. Individuals are born as males or females, but if they are born males, they are not born as terminal phase males. Females and initial phase males can become terminal phase males. Usually, the most dominant female or initial phase male replaces any terminal phase male when those males die or abandon the group.
1418:
679:
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427:. Generally one anemone contains a 'harem', consisting of a large female, a smaller reproductive male, and even smaller non-reproductive males. If the female is removed, the reproductive male will change sex and the largest of the non-reproductive males will mature and become reproductive. It has been shown that fishing pressure can change when the switch from male to female occurs, since fishermen usually prefer to catch the larger fish. The populations are generally changing sex at a smaller size, due to
503:
1379:
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341:
327:
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51:
624:) are simultaneous hermaphrodites, producing both eggs and sperm and routinely reproducing by self-fertilization. Each individual normally fertilizes itself when an egg and sperm produced by an internal organ unite inside the fish's body. This species is also regarded as the only known vertebrate species that can reproduce by
371:
in which the individual first develops as one sex, but can later change into the opposite sex. (Definitions differ on whether sequential hermaphroditism encompasses serial hermaphroditism; for authors who exclude serial hermaphroditism, a sequential hermaphrodite is also stipulated to only change sex
1557:
and ovarian tissues will both be present in the same individual. Though of all total recorded cases of ovotesticular DSD, in only 8% percent of all cases was SRY present, leaving the rest of cases that could be explained to other or less common causes, with the vast majority simply being currently
568:
are an example of a hermaphroditic gastropod. Mating with a partner is more desirable biologically than self-fertilization, as the genetic material of the resultant offspring is varied, but if mating with a partner is not possible, self-fertilization is practiced. The male sexual organ of an adult
177:
A rough estimate of the number of hermaphroditic animal species is 65,000, about 5% of all animal species, or 33% excluding insects. Insects are almost exclusively gonochoric, and no definitive cases of hermaphroditism have been demonstrated in this group. There are no known hermaphroditic species
475:
in which bidirectional sex change occurs. Once a social hierarchy is established a fish changes sex according to its social status, regardless of the initial sex, based on a simple principle: if the fish expresses subordinate behavior then it changes its sex to female, and if the fish expresses
1621:
under certain circumstances, such as if the fertilization is well organized and the average size of groups is small. Simultaneous hermaphroditism that exclusively reproduces through self-fertilization has evolved many times in plants and animals, but it might not last long evolutionarily.
569:
banana slug is quite large in proportion to its size, as well as compared to the female organ. It is possible for banana slugs, while mating, to become stuck together. If a substantial amount of wiggling fails to separate them, the male organ will be bitten off (using the slug's
734:. Monoecy only occurs in about 7% of flowering plant species. Monoecious plants are often referred to as hermaphroditic because they produce both male and female gametes. However, the individual flowers are not hermaphroditic if they only produce gametes of one sex. 65% of
3651:
1986:
Of note, the otherwise well-studied insects, birds, and mammals are strikingly absent here—with not a single species among these groups showing hermaphroditism (for details on a supposedly hermaphroditic scale insect, however, see
Gardner and Ross,
1529:
In some cases, variations in sex characteristics are caused by unusual levels of sex hormones, which may be the result of an atypical set of sex chromosomes. One common cause of variations in sex characteristics traits is the crossing over of the
647:
were first scientifically observed by explorers, they were thought to be hermaphrodites. Early observations of wild spotted hyenas led researchers to believe that all spotted hyenas, male or female, were born with what looked to be a penis. A
169:
groups. Most hermaphroditic species exhibit some degree of self-fertilization. The distribution of self-fertilization rates among animals is similar to that of plants, suggesting that similar pressures are operating to direct the evolution of
1451:
that the people who bore the sexes of both man and woman were regarded by the
Athenians and the Romans as monsters, and thrown into the sea at Athens and into the Tiber at Rome. Similarly, the 17th-century English jurist and judge
4134:
2048:
The mythological term "hermaphrodite" implies that a person is both fully male and fully female. This is a physiologic impossibility. The words "hermaphrodite" and "pseudo-hermaphrodite" are stigmatizing and misleading
595:
are another example of a simultaneous hermaphrodite. Although they possess ovaries and testes, they have a protective mechanism against self-fertilization. Sexual reproduction occurs when two worms meet and exchange
3039:
3528:
use of the words "hermaphrodite," "pseudohermaphrodite," and "intersex" should be abandoned, as they either are confusing or have a negative social connotation that may be perceived as harmful by some patients and
549:
are perhaps the best-known kinds of simultaneous hermaphrodites, and are the most widespread of terrestrial animals possessing this sexual polymorphism. Sexual material is exchanged between both animals via
1473:, medical authors attempted to ascertain whether or not humans could be hermaphrodites, adopting a precise biological definition to the term. From that period until the early 21st century, individuals with
2758:
Kanamori A, Yamamura A, Koshiba S, Lee JS, Orlando EF, Hori H (October 2006). "Methyltestosterone efficiently induces male development in the self-fertilizing hermaphrodite fish, Kryptolebias marmoratus".
1497:) differed from sex expected from internal gonads. This language has fallen out of favor due to misconceptions and stigma associated with the terms, and also a shift to nomenclature based on genetics.
3936:
1700:
Hermaphroditism in plants may promote self fertilization in pioneer populations. However, plants have evolved multiple different mechanisms to avoid self-fertilization in hermaphrodites, including
1637:
for animals was hermaphroditic and that transitions from hermaphroditism to gonochorism were more numerous than the reverse. Other scientists have criticized this argument; saying it’s based on
562:, both animals will lay fertilized eggs after a period of gestation. The eggs will proceed to hatch after a development period. Snails typically reproduce from early spring through late autumn.
1500:
The term "intersex" described a wide variety of combinations of what are ambiguous biological characteristics. Intersex biology may include, for example, ambiguous-looking external genitalia,
372:
once.) This contrasts with simultaneous hermaphrodites, in which an individual possesses fully functional male and female genitalia. Sequential hermaphroditism is common in fish (particularly
232:
in medical literature, this usage is now considered to be outdated as of 2006 and misleading, as people with ovotesticular syndrome do not have functional sets of both male and female organs.
589:
is hermaphroditic, with both male and female organs active at the same time during copulation. After mating, the external portion of the penis detaches, but is able to regrow within 24 hours.
396:
which states that if an individual of a certain sex could significantly increase its reproductive success after reaching a certain size, it would be to their advantage to switch to that sex.
1441:
states that "Whether an hermaphrodite may witness a testament, depends on which sex prevails" ("Hermafroditus an ad testamentum adhiberi possit, qualitas sexus incalescentis ostendit.").
3700:
1610:
may have contributed to the evolution of simultaneous hermaphroditism and sequential hermaphroditism, it remains unclear if the evolution of anisogamy first led to hermaphroditism or
1928:
3608:
656:, when they may become pregnant. When a female spotted hyena gives birth, she passes the cub through the cervix internally, but then passes it out through the elongated clitoris.
762:(hermaphrodite) flowers and separate fertile male flowers that are sterile as female. Andromonoecy occurs in about 4000 species of flowering plants (2% of flowering plants).
3726:
Margarit E, Coll MD, Oliva R, Gómez D, Soler A, Ballesta F (January 2000). "SRY gene transferred to the long arm of the X chromosome in a Y-positive XX true hermaphrodite".
615:
do not practice self-fertilization, but a pair will mate multiple times over several nights, taking turns between which one acts as the male and which acts as the female.
1050:
1462:
on laws of succession stating, "Every heire is either a male, a female, or an hermaphrodite, that is both male and female. And an hermaphrodite (which is also called
1033:
2121:
was widely applied in such cases, but humans are not hermaphroditic. In a truly hermaphroditic species, individuals have functional sets of male and female organs.
3316:
3609:
3110:
Vallejo-Marín M, Rausher MD (February 2007). "The role of male flowers in andromonoecious species: energetic costs and siring success in
Solanum carolinense L".
3652:
3678:
4270:
3660:
161:, hermaphroditism is a normal condition, enabling a form of sexual reproduction in which either partner can act as the female or male. Hermaphroditism is
3669:"Intersex Awareness Day – Wednesday 26 October. End violence and harmful medical practices on intersex children and adults, UN and regional experts urge"
2401:
Rodgers EW, Early RL, Grober MA (2007). "Social status determines sexual phenotype in the bi-directional sex changing bluebanded goby
Lythrypnus dalli".
3585:
3257:
Schultheiss D, Herrmann TR, Jonas U (March 2006). "Early photo-illustration of a hermaphrodite by the French photographer and artist Nadar in 1860".
6154:
3673:
864:
2100:
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3617:
2812:
2705:
Sakakura Y, Soyano K, Noakes D, Hagiwara A (2006). "Gonadal morphology in the self-fertilizing mangrove killifish, Kryptolebias marmoratus".
2158:
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Transitions
Between Sexual Systems: Understanding the Mechanisms of, and Pathways Between, Dioecy, Hermaphroditism and Other Sexual Systems
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is in fact an enlarged clitoris, which contains an external birth canal. It can be difficult to determine the sex of spotted hyenas until
482:
Dichogamy can have both conservation-related implications for humans, as mentioned above, as well as economic implications. For instance,
458:: Where an organism has female and male reproductive organs, but may act either as a female or as a male during different stages in life.
2615:
Barrière A, Félix MA (July 2005). "High local genetic diversity and low outcrossing rate in
Caenorhabditis elegans natural populations".
3706:
531:(or homogamous hermaphrodites) are individuals in which both male and female sexual organs are present and functional at the same time.
3739:
1355:
3207:
Shaw AJ, Goffinet B (2000). "Chapter 12: Population ecology, population genetics, and microevolution". In Shaw AJ, Goffinet B (eds.).
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species, the plants produce hermaphrodite flowers and separate male-sterile pistillate flowers. One example is the meadow saxifrage,
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2881:
2195:
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4821:
3331:
4605:
4521:
1851:
Jarne P, Auld JR (September 2006). "Animals mix it up too: the distribution of self-fertilization among hermaphroditic animals".
6585:
6132:
2137:
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842:
577:. If a banana slug has lost its male sexual organ, it can still mate as a female, making hermaphroditism a valuable adaptation.
4215:
2855:
2826:
780:. Charles Darwin gave several other examples in his 1877 book "The Different Forms of Flowers on Plants of the Same Species".
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988:
859:
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2438:"Seasonal effects on egg production and level of paternity in a natural population of a simultaneous hermaphrodite snail"
1806:
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stage; however, this is now known to be incorrect, as humans are simply undifferentiated before this stage and possess a
1248:
6098:
3304:
1458:
3668:
3625:
1516:.) This is particularly significant because of the relationship between medical terminology and medical intervention.
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4188:
2004:
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759:
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609:
reproduces primarily by self-fertilization, but infrequent out-crossing events occur at a rate of approximately 1%.
228:
in mammals, which is a separate and unrelated phenomenon. While people with the condition were previously called "
6495:
6371:
6361:
6014:
1433:
Historically, the term hermaphrodite was used in law to refer to people whose sex was in doubt. The 12th-century
1190:
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31:
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4136:
Building the Most
Complex Structure on Earth: An Epigenetic Narrative of Development and Evolution of Animals
3823:
3696:
1523:
910:
852:
665:
258:
4299:"True hermaphroditism and mixed gonadal dysgenesis in young children: a clinicopathologic study of 10 cases"
2939:
730:
Flowering plant species with separate, imperfect, male and female flowers on the same individual are called
6658:
6653:
6648:
6435:
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6414:
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6203:
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5669:
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are almost all monoecious. Some plants can change their sex throughout their lifetime, a phenomenon called
528:
3663:, UN Subcommittee on Prevention of Torture and other Cruel, Inhuman or Degrading Treatment or Punishment,
3379:
Greenberg J (1999). "Defining Male and Female: Intersexuality and the
Collision Between Law and Biology".
2008:
722:(female, ovule-producing) parts. The overwhelming majority of flowering plant species are hermaphroditic.
295:
resulting in one individual possessing physical traits of male and female sexes. According to the earlier
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6223:
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5041:
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4590:
4585:
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simultaneous hermaphroditism in animals most likely evolved due to a limited number of mating partners.
6558:
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6366:
5778:
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4945:
4809:
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1258:
1160:
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1045:
743:
321:
3546:"DSD is a Perfectly Fine Term": Reasserting Medical Authority through a Shift in Intersex Terminology"
1417:
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3586:"Information about the standard for gender, sex, and variations of sex characteristics | Stats NZ"
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1466:) shall be heire, either as male or female, according to that kind of sexe which doth prevaile."
1018:
915:
3767:
Why Do Men Have
Nipples?: Hundreds of Questions You'd Only Ask a Doctor After Your Third Martini
3496:
Vilain E, Achermann JC, Eugster EA, Harley VR, Morel Y, Wilson JD, et al. (February 2007).
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4772:
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Royer M (1975). "Hermaphroditism in
Insects. Studies on Icerya purchasi". In Reinboth R (ed.).
1562:
1474:
1444:
1402:
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957:
605:
249:
225:
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were outcrossing hermaphrodites. In flowering plants, hermaphroditism is ancestral to dioecy.
649:
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Looking Queer: Body Image and Identity in Lesbian, Bisexual, Gay, and Transgender Communities
3803:. In Feingold KR, Anawalt B, Blackman MR, Boyce A, Chrousos G, Corpas E, et al. (eds.).
3392:
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Reef and Shore Fishes of the South Pacific: New Caledonia to Tahiti and the Pitcairn Islands
4094:"Phylogenetic perspectives on the evolution of functional hermaphroditism in teleost fishes"
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381:
145:, primarily invertebrates, are hermaphrodites, capable of producing viable gametes of both
3063:"Reproductive biology of the andromonoecious Cucumis melo subsp. agrestis (Cucurbitaceae)"
2548:
1429:. The series may be the earliest medical photographic documentation of an intersex person.
8:
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201:, in both plants and animals, where hermaphrodite individuals coexist with males (called
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3963:
Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences
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species are either hermaphroditic (all flowers produce both male and female gametes) or
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2230:
Animal behavior desk reference: a dictionary of animal behavior, ecology, and evolution
1876:
1833:
1752:
787:, meaning that their gametophytes produce either male or female gametes, but not both.
683:
532:
513:. Each has two penises on the undersides of their heads which they use to inject sperm.
385:
138:
42:
4315:
4298:
3433:"Changing the nomenclature/taxonomy for intersex: a scientific and clinical rationale"
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in some areas by beginning a series of events starting with the upregulation of the
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2418:
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1967:
1899:
1860:
1815:
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1704:, molecular recognition systems and mechanical or morphological mechanisms such as
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1664:
1630:
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1570:
1550:
620:
463:
296:
68:
3139:
2522:
1777:
Hermaphroditism: A Primer on the Biology, Ecology, and Evolution of Dual Sexuality
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are simultaneous hermaphrodites, having both male and female reproductive organs.
472:
304:
281:
214:
190:
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Kouonon LC, Jacquemart AL, Bi AI, Bertin P, Baudoin JP, Dje Y (November 2009).
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that include mixed XX and XY chromosome pairs (46XX/46XY, 46XX/47XXY or 45X/XY
1382:
1312:
874:
269:
241:
218:
54:
4059:
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2726:
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2493:
Leonard JL, Pearse JS, Harper AB (2002). "Comparative reproductive biology of
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5074:
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1742:
1470:
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952:
942:
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612:
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551:
510:
299:, he was born with a physical body combining male and female sexes. The word
197:, where both male and female flowers occur on the same plant. There are also
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4359:
3855:
3838:
3785:
1413:
dedicated to the hermaphrodite deity, Lakapati, who presided over fertility.
1378:
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3131:
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2780:
2654:
2601:
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2479:
2333:
Ghiselin MT (June 1969). "The evolution of hermaphroditism among animals".
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1981:
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dominant or non-dominant superior behavior then it changes its sex to male.
439:: Where the organism develops as a female, and then changes sex to a male.
213:). Sometimes, both male and hermaphrodite flowers occur on the same plant (
3740:
10.1002/(SICI)1096-8628(20000103)90:1<25::AID-AJMG5>3.0.CO;2-5
2354:
1538:. The SRY is then activated in only certain areas, causing development of
406:: Where an organism develops as a male, and then changes sex to a female.
6600:
5955:
5887:
5843:
5808:
5798:
5699:
5436:
5131:
5109:
5009:
4110:
4093:
3761:
3361:
3078:
1737:
1705:
1667:
and gonochorism is also ancestral to hermaphroditic fishes. According to
1618:
1611:
1453:
1406:
1305:
1285:
771:
565:
555:
506:
490:. Since the adults take several years to change from female to male, the
424:
202:
131:
3366:
The First Part of the Institutes of the Laws of England, Institutes 8.a.
2568:"Disposable penis and its replenishment in a simultaneous hermaphrodite"
2064:
Rosenfield KA (2018), "Hermaphrodite", in Vonk J, Shackelford T (eds.),
1930:
Physical and Biological Bases of Life Stability: Man, Biota, Environment
27:
Sexually reproducing organism that produces both male and female gametes
6553:
6347:
6332:
5921:
5904:
5716:
5421:
5338:
4957:
4495:"Born True Hermaphrodite – Pictorial Profile", about Lynn Edward Harris
3170:
2735:
2378:
Biological exuberance : animal homosexuality and natural diversity
2159:
Ovid, Metamorphoses, Book IV: The story of Hermaphroditus and Salmacis.
1561:
Fetuses were previously thought to be phenotypically female before the
1554:
1253:
735:
687:, a hermaphrodite plant with perfect flowers that have both functional
678:
542:
491:
468:
415:
377:
356:
206:
166:
6184:
3896:
3879:
2772:
2629:
2461:
2033:
1972:
1955:
814:
399:
Sequential hermaphrodites can be divided into three broad categories:
340:
326:
6575:
6292:
6043:
5793:
5783:
5731:
5709:
5694:
5632:
5627:
5426:
5406:
5268:
5104:
5031:
4743:
4718:
4708:
4696:
1864:
1727:
1607:
1501:
1494:
1390:
1275:
791:
739:
592:
581:
539:
521:
420:
410:
364:
344:
277:
158:
4407:
Business As Ethical and Business As Usual: Text, Readings, and Cases
2965:
Walas Ł, Mandryk W, Thomas PA, Tyrała-Wierucka Ż, Iszkuło G (2018).
1397:
517:
217:) or both female and hermaphrodite flowers occur on the same plant (
130:. Animal species in which individuals are either male or female are
6107:
5914:
5882:
5803:
5761:
5704:
5684:
5557:
5411:
5401:
5141:
5121:
5021:
4985:
4760:
4661:
4344:. Honolulu, Hawaii: University of Hawaii Press. pp. 346, 387.
4217:
Evolution in Isolation: The Search for an Island Syndrome in Plants
2566:
Sekizawa A, Seki S, Tokuzato M, Shiga S, Nakashima Y (April 2013).
2346:
1732:
1641:
1546:(SOX9), and in other areas not being active (causing the growth of
1373:
805:
625:
486:
are favoured fish for eating in many Asian countries and are often
447:
351:
292:
150:
116:
38:
4490:
The Evolution of Self-Fertile Hermaphroditism: The Fog Is Clearing
4297:
Kim KR, Kwon Y, Joung JY, Kim KS, Ayala AG, Ro JY (October 2002).
4241:
Judd WS, Campbell CS, Kellogg EA, Stevens PF, Donoghue MJ (2002).
3431:
Dreger AD, Chase C, Sousa A, Gruppuso PA, Frader J (August 2005).
2298:
Warner RR (June 1988). "Sex change and the size-advantage model".
1617:
A 2023 study argued that hermaphroditism can evolve directly from
6266:
6038:
5909:
5838:
5721:
5657:
5612:
5602:
5416:
5378:
5114:
4962:
4728:
4701:
2964:
1722:
1547:
1535:
982:
673:
483:
384:). Sequential hermaphroditism can best be understood in terms of
373:
368:
347:
are initially male; the largest fish in a group becomes a female.
210:
194:
171:
154:
4092:
Erisman BE, Petersen CW, Hastings PA, Warner RR (October 2013).
50:
6565:
5773:
5258:
5079:
5000:
4990:
4750:
4723:
4556:
3610:
Androgen Insensitivity Support Syndrome Support Group Australia
3211:. Cambridge, UK: Cambridge University Press. pp. 369–402.
2000:
1648:
level than the species level, and methods exclusively based on
1645:
1539:
1386:
1317:
1265:
784:
719:
715:
711:
704:
692:
688:
597:
570:
443:
273:
179:
142:
127:
124:
4035:"A reconstruction of sexual modes throughout animal evolution"
2944:(Second ed.). Oxford University Press. pp. 138–139.
2287:
from the original on 7 November 2017 – via Google Books.
2176:. Vol. IV. p. 361 – via penelope.uchicago.edu.
74:
5982:
4091:
3410:. Baltimore: Johns Hopkins University Press. pp. 55–81.
1484:
289:
3495:
3194:
The different forms of flowers on plants of the same species
2704:
392:, as described in the size-advantage mode first proposed by
6570:
6076:
5622:
5567:
5283:
4551:
3956:
1270:
546:
285:
263:
183:
120:
101:
92:
4240:
2757:
4530:
4480:
Britannica Online Encyclopedia: hermaphroditism (biology)
4245:(2 ed.). Sunderland MA, US: Sinauer Associates Inc.
3792:
3233:"LAKAPATI: The "Transgender" Tagalog Deity? Not so fast…"
3060:
2565:
2068:, Cham: Springer International Publishing, pp. 1–2,
1814:. Cham: Springer International Publishing. pp. 1–3.
1805:(2020). "Gonochorism". In Vonk J, Shackelford TK (eds.).
1487:
tissue contained both testicular and ovarian tissue, and
1322:
146:
83:
4367:
Chase C (1998). "Affronting Reason". In Atkins D (ed.).
3430:
3256:
1591:
For evolution of sexual systems with hermaphodites, see
1508:). Clinically, medicine currently uses the terminology "
1409:, before becoming a Catholic festival, was initially an
4277:. Busch Entertainment Corporation. 2009. Archived from
1534:(SRY) from the Y chromosome to the X chromosome during
1526:
designed to make bodies more typically male or female.
4499:
3725:
3661:
UN Committee on the Rights of People with Disabilities
3550:
Sociology of Diagnosis (Advances in Medical Sociology)
1522:, and many human rights institutions, have criticized
3329:
3112:
Evolution; International Journal of Organic Evolution
1853:
Evolution; International Journal of Organic Evolution
1425:
of a person displaying ambiguous genitalia, one of a
95:
86:
4186:
3109:
2675:
2234:(2nd ed.). Boca Raton, Fla: CRC Press. p.
783:
About 57% of moss species and 68% of liverworts are
104:
98:
89:
4485:
Current Biology – Gender trading in a hermaphrodite
4293:
Discovery Health Channel, (2007) "I Am My Own Twin"
4190:
Evolutionary Ecology of Plant-Herbivore Interaction
3957:Henshaw JM, Bittlingmaier M, Schärer L (May 2023).
3440:
Journal of Pediatric Endocrinology & Metabolism
2492:
2001:"Getting Rid of "Hermaphroditism" Once and For All"
80:
77:
71:
2400:
1898:. Berlin, Heidelberg: Springer. pp. 135–145.
359:start life as females and later change into males.
209:), or all three exist in the same species (called
3925:. Vol. 2. Academic Press. pp. 212–224.
6640:
4409:. Belmont, CA: Wadsworth Pub. pp. 168–170.
3674:Office of the High Commissioner for Human Rights
3408:Bodies in Doubt: an American History of Intersex
3028:. Cambridge University Press. pp. 111–113.
1800:
600:, copulating on damp nights during warm seasons.
4393:
4296:
3152:
2676:Russell PJ, Hertz PE, McMillan B (2016-01-01).
4944:
4397:(12 March 1993). "How Many Sexes Are There?".
4220:. Cambridge University Press. pp. 21–22.
3798:
3760:
3602:
2380:(2nd ed.). New York: St. Martin's Press.
2034:"Is a person who is intersex a hermaphrodite?"
497:
6092:
4930:
4515:
4431:
4371:. New York: Haworth Press. pp. 205–219.
3702:Human rights and intersex people, Issue Paper
3618:Organisation Intersex International Australia
3153:Stevens D, Richards A (1985). "Gynodioecy in
2614:
2066:Encyclopedia of Animal Cognition and Behavior
1808:Encyclopedia of Animal Cognition and Behavior
1349:
670:Plant reproductive morphology § Bisexual
3950:
3206:
3103:
3017:
2126:
2105:. University of Toronto Press. p. 309.
2059:
2057:
315:
4180:
3830:
3807:. South Dartmouth, Mass.: MDText.com, Inc.
3667:, Pῡras D, et al. (October 24, 2016),
3305:"Decretum Gratiani (Kirchenrechtssammlung)"
2795:
2503:Invertebrate Reproduction & Development
2486:
2435:
1780:. Columbia University Press. pp. 1–7.
1597:Androdioecy § Evolution of androdioecy
1434:
134:, which is the opposite of hermaphroditic.
6099:
6085:
4937:
4923:
4522:
4508:
4243:Plant systematics, a phylogenetic approach
4187:Núñez-Farfán J, Valverde PL (2020-07-30).
4032:
3880:"The varied ways of being male and female"
3185:
2886:. Tata McGraw-Hill Education. p. 87.
2819:
2221:
2219:
2217:
2215:
2213:
2173:The Library of History of Diodorus Siculus
2063:
1956:"The varied ways of being male and female"
1356:
1342:
4314:
4109:
4068:
4058:
3999:
3982:
3895:
3871:
3854:
3540:
3534:
3513:
3378:
3323:
3086:
2967:"Sexual systems in gymnosperms: A review"
2734:
2628:
2591:
2549:"Sea slug's 'disposable penis' surprises"
2543:
2469:
2054:
1971:
1933:. Springer Science & Business Media.
1850:
419:) are colorful reef fish found living in
307:as early as the late fourteenth century.
224:Hermaphrodism is not to be confused with
4026:
3912:
3799:Rey R, Josso N, Racine C (2020-05-27) .
3336:Encyclopedia of Diderot & d'Alembert
3294:Decretum Gratiani, C. 4, q. 2 et 3, c. 3
3013:
3011:
2751:
2698:
2608:
2375:
2332:
2326:
2163:
2094:
2092:
1926:
1416:
1396:
1377:
790:Sequential hermaphroditism is common in
677:
632:
603:The free-living hermaphroditic nematode
516:
501:
350:
339:
325:
240:The term hermaphrodite derives from the
49:
4404:
4339:
4275:SeaWorld/Busch Gardens Animal Infobooks
4085:
4005:
3877:
3657:UN Committee on the Rights of the Child
3645:
3319:from the original on December 20, 2016.
3191:
3146:
3054:
2669:
2429:
2394:
2266:
2260:
2225:
2210:
1953:
1920:
1769:
1767:
797:
14:
6641:
3918:
3884:Molecular Reproduction and Development
3836:
3681:from the original on November 21, 2016
2937:
2931:
2906:
2900:
2297:
2291:
2028:
2026:
2011:from the original on 27 September 2011
1992:
1960:Molecular Reproduction and Development
1947:
1644:, assignments of sexual modes for the
6080:
4918:
4503:
4366:
4213:
4207:
4159:
4153:
4132:
4126:
3959:"Hermaphroditic origins of anisogamy"
3544:(2011). McGann PJ, Hutson DJ (eds.).
3498:"We used to call them hermaphrodites"
3008:
2958:
2848:
2098:
2089:
1893:
1773:
1693:It is widely accepted that the first
650:female spotted hyena's apparent penis
4439:. In Williams RH, Wilson JD (eds.).
4234:
4033:Sasson DA, Ryan JF (December 2017).
3922:Encyclopedia of Evolutionary Biology
3728:American Journal of Medical Genetics
3405:
3360:
3200:
2879:
2873:
2803:"The Painful Realities of Hyena Sex"
2198:from the original on 5 November 2013
2152:
2140:from the original on 6 November 2014
1896:Intersexuality in the Animal Kingdom
1887:
1844:
1794:
1764:
1633:and Priya Iyer argued that the last
1520:Intersex civil society organizations
1393:, origin of the word "hermaphrodite"
494:are extremely valuable individuals.
4618:Heterogametic sex / Homogametic sex
4445:. Philadelphia: Saunders. pp.
4098:Integrative and Comparative Biology
3685:
3614:Intersex Trust Aotearoa New Zealand
2941:Understanding Flowers and Flowering
2537:
2180:
2023:
1998:
1593:Trioecy § Evolution of trioecy
1447:(1461–1523) stated, using the term
1249:Genital modification and mutilation
24:
4442:Williams Textbook of Endocrinology
4437:"Disorders of sex differentiation"
4262:
3562:10.1108/S1057-6290(2011)0000012012
2684:. Cengage Learning. p. 1151.
1459:Institutes of the Lawes of England
813:
25:
6675:
4473:
4316:10.1097/01.MP.0000027623.23885.0D
3839:"Minireview: sex differentiation"
3552:. Advances in Medical Sociology.
3042:from the original on 1 April 2021
2913:Evolution and the Origins of Life
2501:(Gastropoda; Stylommiatophora)".
2300:Trends in Ecology & Evolution
2038:Intersex Society of North America
2005:Intersex Society of North America
1514:variations in sex characteristics
6621:
6620:
6183:
6170:Sex characteristics (legal term)
6059:
4899:
4898:
4692:Evolution of sexual reproduction
4193:. Springer Nature. p. 177.
3770:. New York: Three Rivers Press.
3330:chevalier de Jaucourt L (1765).
3271:10.1111/j.1743-6109.2005.00157.x
3124:10.1111/j.1558-5646.2007.00031.x
2815:from the original on 2012-11-19.
2423:10.1111/j.1095-8649.2007.01427.x
1587:Evolution of sexual reproduction
880:Sex characteristics (legal term)
67:
6015:LGBT-affirming religious groups
4166:. Academic Press. p. 176.
3939:from the original on 2021-05-06
3754:
3719:
3578:
3489:
3424:
3399:
3372:
3354:
3297:
3288:
3250:
3225:
3159:Plant Systematics and Evolution
2915:. Geocities.com. Archived from
2559:
2436:Janssen R, Baur B (July 2015).
2369:
2335:The Quarterly Review of Biology
2267:Pandian TJ (2 September 2011).
2074:10.1007/978-3-319-47829-6_329-1
1820:10.1007/978-3-319-47829-6_305-1
749:
163:also found in some fish species
5179:Sexually transmitted infection
4800:Sexual reproduction in animals
3259:The Journal of Sexual Medicine
2134:"Definition of hermaphroditus"
1493:if their external appearance (
765:
32:Hermaphrodite (disambiguation)
13:
1:
5226:Women who have sex with women
3824:National Institutes of Health
3697:Commissioner for Human Rights
3309:Bayerische StaatsBibliothek (
2907:Molnar S (17 February 2004).
2523:10.1080/07924259.2002.9652738
1758:
1685:Andromonoecy § Evolution
1625:
1385:, the "son" of the Greek god
718:(male, pollen-producing) and
666:Sexual reproduction in plants
6415:International Intersex Forum
6204:Disorders of sex development
6106:
5359:History of erotic depictions
5249:Criminal transmission of HIV
4672:Sex as a biological variable
4655:Simultaneous hermaphroditism
4163:Horticultural Plant Breeding
3653:UN Committee against Torture
3624:, et al. (March 2017),
3515:10.1097/GIM.0b013e31802cffcf
2909:"Plant Reproductive Systems"
2682:Biology: The Dynamic Science
2312:10.1016/0169-5347(88)90176-0
2136:. Numen: The Latin Lexicon.
1904:10.1007/978-3-642-66069-6_14
1689:Gynomonoecy § Evolution
1674:
1580:
1510:disorders of sex development
1370:Disorders of sex development
1106:International Intersex Forum
906:Disorders of sex development
758:species, the plants produce
707:to describe, for example, a
554:, and is then stored in the
264:
235:
7:
6410:Intersex Day of Remembrance
6273:Civil society organizations
5344:Counterculture of the 1960s
4271:"Bony Fishes: Reproduction"
3878:Schärer L (February 2017).
3025:The Biology of Reproduction
2192:Online Etymology Dictionary
1954:Schärer L (February 2017).
1711:
1601:Gynodioecy § Evolution
1101:Intersex Day of Remembrance
989:Civil society organizations
738:species are dioecious, but
529:Simultaneous hermaphrodites
498:Simultaneous hermaphrodites
363:Sequential hermaphrodites (
272:(Ἑρμαφρόδιτος), the son of
149:. In the great majority of
10:
6680:
6367:Sex verification in sports
6066:Human sexuality portal
4810:Penile-vaginal intercourse
4650:Sequential hermaphroditism
4401:. New York. p. Op–Ed.
3452:10.1515/JPEM.2005.18.8.729
2994:10.1016/j.baae.2018.05.009
2938:Glover B (February 2014).
1702:sequential hermaphroditism
1678:
1590:
1584:
1456:(Lord Coke), wrote in his
1367:
1046:Sex verification in sports
794:and some vascular plants.
744:sequential hermaphroditism
725:
663:
636:
456:Bidirectional sex changers
322:Sequential hermaphroditism
319:
310:
253:
205:) or with females (called
36:
29:
6616:
6608:Sex differences in humans
6541:
6423:
6380:
6259:
6193:
6181:
6114:
6054:
5935:
5824:
5543:
5495:
5372:
5329:
5234:
5211:Men who have sex with men
5187:
5060:
4956:
4894:
4847:
4680:
4640:Testis-determining factor
4537:
4060:10.1186/s12862-017-1071-3
4006:Leonard JL (2019-05-21).
3837:Hughes IA (August 2001).
2974:Basic and Applied Ecology
2727:10.1007/s10228-006-0362-2
2647:10.1016/j.cub.2005.06.022
2270:Sex Determination in Fish
1927:Gorshkov V (2012-12-06).
1532:testis-determining factor
659:
586:Goniobranchus reticulatus
580:The species of colourful
316:Sequential hermaphrodites
6664:Supernumerary body parts
6128:Compulsory sterilization
4581:Sex-determination system
4039:BMC Evolutionary Biology
3801:"Sexual Differentiation"
1681:Monoecy § Evolution
1665:evolved from gonochorism
838:Compulsory sterilization
618:The mangrove killifish (
137:The individuals of many
37:Not to be confused with
4529:
4405:Harwood S, ed. (1996).
4214:Burns KC (2019-05-16).
4160:Orton TJ (2019-11-21).
4139:. Newnes. p. 101.
4133:Cabej NR (2013-04-01).
3856:10.1210/endo.142.8.8406
3284:(subscription required)
2856:"Hermaphrodite Hyenas?"
2707:Ichthyological Research
1774:Avise JC (2011-03-18).
621:Kryptolebias marmoratus
174:in animals and plants.
165:, but is rare in other
6586:Non-binary recognition
6405:Intersex Awareness Day
6328:Ancient Greek religion
6251:Sexual differentiation
5487:Split attraction model
5477:Sexual objectification
5387:Anarchism and love/sex
4564:Sexual differentiation
3975:10.1098/rstb.2022.0283
3311:Bavarian State Library
3196:. London: John Murray.
2883:Fundamentals of Botany
2880:Soni NK (2010-04-01).
2584:10.1098/rsbl.2012.1150
2495:Ariolimax californicus
2117:In the past, the term
2099:Mills A (2018-01-01).
1563:sexual differentiation
1475:ovotesticular syndrome
1445:Alexander ab Alexandro
1435:
1430:
1414:
1403:Obando Fertility Rites
1394:
1291:Non-binary recognition
1096:Intersex Awareness Day
958:Sexual differentiation
818:
696:
606:Caenorhabditis elegans
525:
514:
360:
348:
337:
336:(common slipper shell)
245:
226:ovotesticular syndrome
199:mixed breeding systems
58:
6278:Fictional characters
6219:Medical interventions
6175:Yogyakarta Principles
6160:Medical interventions
5447:Sex-positive movement
5100:Reproductive medicine
5095:Obesity and sexuality
5042:Sexual response cycle
4606:Temperature-dependent
3764:, Goldberg B (2005).
2678:"Animal Reproduction"
2442:Ecology and Evolution
1524:medical interventions
1420:
1400:
1381:
994:Fictional characters
926:Medical interventions
885:Yogyakarta Principles
870:Medical interventions
817:
681:
639:Pseudohermaphroditism
633:Pseudohermaphroditism
520:
505:
354:
343:
329:
268:, which derives from
53:
5973:Buddhist monasticism
5963:Christian demonology
5789:Urethral intercourse
5588:Erotic sexual denial
5279:Cybersex trafficking
5152:Erectile dysfunction
3627:Darlington Statement
3502:Genetics in Medicine
3315:. February 5, 2009.
2547:(12 February 2013).
1718:Asexual reproduction
1567:paramesonephric duct
1544:transcription factor
1490:pseudohermaphrodites
1281:Anti-gender movement
1051:at the Olympic Games
798:Use regarding humans
558:. After exchange of
382:common slipper shell
288:, he fused with the
230:true hermaphrodites"
114:sexually reproducing
30:For other uses, see
6659:Intersex in history
6654:Reproductive system
6649:Sexual reproduction
6260:Society and culture
6027:Sacred prostitution
5816:Wet T-shirt contest
5680:Non-penetrative sex
5638:Mammary intercourse
5319:Revenge pornography
5090:Masters and Johnson
5052:Vaginal lubrication
4822:Hormonal motivation
4795:Fungal reproduction
4714:Reproductive system
4435:, Conte FA (1998).
4340:Randall JE (2005).
4051:2017BMCEE..17..242S
3620:, Black E, Bond K,
3477:on 20 December 2016
3368:(1st Am. ed.).
3155:Saxifraga granulata
2986:2018BApEc..31....1W
2719:2006IchtR..53..427S
2639:2005CBio...15.1176B
2515:2002InvRD..41...83L
2454:2015EcoEv...5.2916J
2415:2007JFBio..70.1660R
2376:Bagemihl B (1999).
2226:Barrows EM (2001).
1655:Hermaphroditism is
1480:true hermaphrodites
1421:1860 photograph by
977:Society and culture
777:Saxifraga granulata
394:Michael T. Ghiselin
390:life history theory
333:Crepidula fornicata
119:that produces both
6381:History and events
5873:Red-light district
5769:Sexual penetration
5757:Sexual intercourse
5675:Nipple stimulation
5354:Golden Age of Porn
5254:Child sexual abuse
5221:Sexual orientation
5147:Sexual dysfunction
5085:Male contraceptive
5047:Sexual stimulation
4996:Nocturnal emission
4827:Human reproduction
4805:Sexual intercourse
4790:Plant reproduction
4399:The New York Times
4111:10.1093/icb/ict077
3969:(1876): 20220283.
3381:Arizona Law Review
3239:. 29 November 2018
3237:THE ASWANG PROJECT
3192:Charles D (1877).
3171:10.1007/BF02418018
3079:10.1093/aob/mcp196
1753:Self-fertilization
1512:" (also known as
1431:
1415:
1395:
819:
697:
684:Hylocereus undatus
626:self fertilization
533:Self-fertilization
526:
515:
386:behavioral ecology
361:
349:
338:
59:
43:Intersex (biology)
6636:
6635:
6581:Non-binary gender
6549:Children's rights
6424:Rights by country
6303:Intersex and LGBT
6209:Genetic diagnosis
6155:Malta declaration
6150:Legal recognition
6074:
6073:
5983:Hindu monasticism
5968:Clerical celibacy
5834:Adult video games
5545:Sexual activities
5462:Sexual attraction
5452:Sexual abstinence
5364:Sexual revolution
5349:Feminist sex wars
5304:Sexual misconduct
5299:Sexual harassment
5264:Indecent exposure
4971:Clitoral erection
4912:
4911:
4832:Lordosis behavior
4547:Sexual dimorphism
4456:978-0-7216-6152-0
4416:978-0-534-54251-1
4395:Fausto-Sterling A
4378:978-1-56023-931-4
4351:978-0-8248-2698-7
4227:978-1-108-42201-7
4200:978-3-030-46012-9
4173:978-0-12-815570-7
4146:978-0-12-401729-0
4019:978-3-319-94139-4
3932:978-0-12-800426-5
3919:Kliman R (2016).
3897:10.1002/mrd.22775
3777:978-1-4000-8231-5
3693:Council of Europe
3590:www.stats.govt.nz
3571:978-0-85724-575-5
3417:978-0-8018-9155-7
3218:978-0-521-66794-4
3209:Bryophyte biology
3035:978-1-108-49985-9
2951:978-0-19-966159-6
2893:978-0-07-068177-4
2811:. 26 April 2006.
2773:10.1002/dvg.20240
2691:978-1-305-85610-3
2499:A. dolichophallus
2462:10.1002/ece3.1560
2387:978-0-312-25377-6
2280:978-1-4398-7919-1
2245:978-0-8493-2005-7
2112:978-1-4875-9337-7
2083:978-3-319-47829-6
1973:10.1002/mrd.22775
1940:978-3-642-85001-1
1913:978-3-642-66071-9
1829:978-3-319-47829-6
1787:978-0-231-52715-6
1650:maximum parsimony
1606:The evolution of
1437:Decretum Gratiani
1366:
1365:
1244:Children's rights
1119:Rights by country
1009:Intersex and LGBT
911:Genetic diagnosis
865:Malta declaration
860:Legal recognition
471:) are a group of
429:natural selection
388:and evolutionary
262:
16:(Redirected from
6671:
6624:
6623:
6313:Military service
6187:
6101:
6094:
6087:
6078:
6077:
6064:
6063:
5988:Jain monasticism
5752:Sexual fetishism
5670:Mechanics of sex
5553:Conventional sex
5457:Sexual addiction
5392:Extramarital sex
5174:Sexual surrogate
5010:male ejaculation
4939:
4932:
4925:
4916:
4915:
4902:
4901:
4862:Animal sexuality
4785:Sexual selection
4524:
4517:
4510:
4501:
4500:
4468:
4428:
4403:, reprinted in:
4402:
4390:
4363:
4336:
4318:
4303:Modern Pathology
4290:
4288:
4286:
4257:
4256:
4238:
4232:
4231:
4211:
4205:
4204:
4184:
4178:
4177:
4157:
4151:
4150:
4130:
4124:
4123:
4113:
4089:
4083:
4082:
4072:
4062:
4030:
4024:
4023:
4003:
3997:
3996:
3986:
3954:
3948:
3947:
3945:
3944:
3916:
3910:
3909:
3899:
3875:
3869:
3868:
3858:
3849:(8): 3281–3287.
3834:
3828:
3827:
3821:
3819:
3796:
3790:
3789:
3758:
3752:
3751:
3723:
3717:
3716:
3715:
3714:
3705:, archived from
3689:
3683:
3682:
3649:
3643:
3642:
3641:
3639:
3630:, archived from
3606:
3600:
3599:
3597:
3596:
3582:
3576:
3575:
3538:
3532:
3531:
3517:
3493:
3487:
3486:
3484:
3482:
3476:
3470:. Archived from
3437:
3428:
3422:
3421:
3403:
3397:
3396:
3376:
3370:
3369:
3358:
3352:
3351:
3349:
3347:
3327:
3321:
3320:
3301:
3295:
3292:
3286:
3285:
3282:
3254:
3248:
3247:
3245:
3244:
3229:
3223:
3222:
3204:
3198:
3197:
3189:
3183:
3182:
3150:
3144:
3143:
3107:
3101:
3100:
3090:
3067:Annals of Botany
3058:
3052:
3051:
3049:
3047:
3015:
3006:
3005:
2971:
2962:
2956:
2955:
2935:
2929:
2928:
2926:
2924:
2904:
2898:
2897:
2877:
2871:
2870:
2868:
2867:
2858:. Archived from
2852:
2846:
2845:
2843:
2842:
2833:. Archived from
2823:
2817:
2816:
2799:
2793:
2792:
2755:
2749:
2748:
2738:
2702:
2696:
2695:
2673:
2667:
2666:
2632:
2612:
2606:
2605:
2595:
2563:
2557:
2556:
2541:
2535:
2534:
2490:
2484:
2483:
2473:
2433:
2427:
2426:
2409:(6): 1660–1668.
2398:
2392:
2391:
2373:
2367:
2366:
2330:
2324:
2323:
2295:
2289:
2288:
2264:
2258:
2257:
2233:
2223:
2208:
2207:
2205:
2203:
2184:
2178:
2177:
2170:"Chapters 1–7".
2167:
2161:
2156:
2150:
2149:
2147:
2145:
2130:
2124:
2123:
2096:
2087:
2086:
2061:
2052:
2051:
2045:
2044:
2030:
2021:
2020:
2018:
2016:
1996:
1990:
1989:
1975:
1951:
1945:
1944:
1924:
1918:
1917:
1891:
1885:
1884:
1865:10.1554/06-246.1
1848:
1842:
1841:
1813:
1798:
1792:
1791:
1771:
1748:Self-pollination
1631:Joan Roughgarden
1575:genital tubercle
1571:mesonephric duct
1440:
1427:nine-part series
1389:and the goddess
1358:
1351:
1344:
802:
801:
464:Lythrypnus dalli
355:Most species of
297:Diodorus Siculus
267:
257:
255:
111:
110:
107:
106:
103:
100:
97:
94:
91:
88:
85:
82:
79:
76:
73:
21:
6679:
6678:
6674:
6673:
6672:
6670:
6669:
6668:
6639:
6638:
6637:
6632:
6612:
6537:
6419:
6376:
6255:
6189:
6188:
6179:
6110:
6105:
6075:
6070:
6058:
6050:
5978:Catholic Church
5938:
5931:
5820:
5539:
5491:
5397:Family planning
5376:
5368:
5325:
5314:Sexual violence
5289:Sex trafficking
5230:
5216:Sexual identity
5206:Gender identity
5183:
5169:Sexual medicine
5065:
5056:
5022:Penile erection
4976:Erogenous zones
4960:
4952:
4950:human sexuality
4943:
4913:
4908:
4890:
4877:Differentiation
4867:Human sexuality
4857:Plant sexuality
4843:
4739:Spermatogenesis
4683:
4676:
4539:
4533:
4528:
4476:
4471:
4457:
4417:
4379:
4352:
4284:
4282:
4281:on 25 July 2011
4269:
4265:
4263:Further reading
4260:
4253:
4239:
4235:
4228:
4212:
4208:
4201:
4185:
4181:
4174:
4158:
4154:
4147:
4131:
4127:
4090:
4086:
4031:
4027:
4020:
4004:
4000:
3955:
3951:
3942:
3940:
3933:
3917:
3913:
3876:
3872:
3835:
3831:
3817:
3815:
3797:
3793:
3778:
3759:
3755:
3724:
3720:
3712:
3710:
3690:
3686:
3650:
3646:
3637:
3635:
3607:
3603:
3594:
3592:
3584:
3583:
3579:
3572:
3539:
3535:
3494:
3490:
3480:
3478:
3474:
3435:
3429:
3425:
3418:
3406:Reis E (2009).
3404:
3400:
3377:
3373:
3359:
3355:
3345:
3343:
3332:"Hermaphrodite"
3328:
3324:
3303:
3302:
3298:
3293:
3289:
3283:
3255:
3251:
3242:
3240:
3231:
3230:
3226:
3219:
3205:
3201:
3190:
3186:
3151:
3147:
3108:
3104:
3059:
3055:
3045:
3043:
3036:
3016:
3009:
2969:
2963:
2959:
2952:
2936:
2932:
2922:
2920:
2905:
2901:
2894:
2878:
2874:
2865:
2863:
2854:
2853:
2849:
2840:
2838:
2827:"Hyena Graphic"
2825:
2824:
2820:
2801:
2800:
2796:
2767:(10): 495–503.
2756:
2752:
2703:
2699:
2692:
2674:
2670:
2623:(13): 1176–84.
2617:Current Biology
2613:
2609:
2578:(2): 20121150.
2572:Biology Letters
2564:
2560:
2542:
2538:
2491:
2487:
2448:(14): 2916–28.
2434:
2430:
2399:
2395:
2388:
2374:
2370:
2331:
2327:
2296:
2292:
2281:
2265:
2261:
2246:
2224:
2211:
2201:
2199:
2188:"Hermaphrodite"
2186:
2185:
2181:
2169:
2168:
2164:
2157:
2153:
2143:
2141:
2132:
2131:
2127:
2113:
2097:
2090:
2084:
2062:
2055:
2042:
2040:
2032:
2031:
2024:
2014:
2012:
1997:
1993:
1952:
1948:
1941:
1925:
1921:
1914:
1892:
1888:
1849:
1845:
1830:
1811:
1799:
1795:
1788:
1772:
1765:
1761:
1714:
1695:vascular plants
1691:
1677:
1635:common ancestor
1628:
1604:
1589:
1583:
1558:unexplainable.
1376:
1368:Main articles:
1362:
1333:
1332:
1328:Sex differences
1234:
1226:
1225:
1121:
1111:
1110:
1074:
1063:
1062:
978:
970:
969:
901:
890:
889:
833:
800:
768:
756:andromonoecious
752:
728:
676:
664:Main articles:
662:
654:sexual maturity
641:
635:
500:
473:coral reef fish
376:fish) and many
324:
318:
313:
305:English lexicon
284:. According to
282:Greek mythology
238:
191:flowering plant
70:
66:
46:
39:Intersex people
35:
28:
23:
22:
15:
12:
11:
5:
6677:
6667:
6666:
6661:
6656:
6651:
6634:
6633:
6631:
6630:
6617:
6614:
6613:
6611:
6610:
6605:
6604:
6603:
6598:
6593:
6588:
6583:
6578:
6568:
6563:
6562:
6561:
6551:
6545:
6543:
6539:
6538:
6536:
6535:
6534:
6533:
6523:
6521:United Kingdom
6518:
6513:
6508:
6503:
6498:
6493:
6488:
6483:
6478:
6473:
6468:
6463:
6458:
6453:
6448:
6443:
6438:
6433:
6427:
6425:
6421:
6420:
6418:
6417:
6412:
6407:
6402:
6401:
6400:
6395:
6384:
6382:
6378:
6377:
6375:
6374:
6369:
6364:
6359:
6358:
6357:
6356:
6355:
6350:
6342:
6341:
6340:
6338:Hermaphroditus
6335:
6320:
6315:
6310:
6305:
6300:
6295:
6290:
6285:
6280:
6275:
6270:
6263:
6261:
6257:
6256:
6254:
6253:
6248:
6243:
6238:
6233:
6228:
6227:
6226:
6216:
6211:
6206:
6200:
6198:
6191:
6190:
6182:
6180:
6178:
6177:
6172:
6167:
6165:Sex assignment
6162:
6157:
6152:
6147:
6146:
6145:
6135:
6133:Discrimination
6130:
6124:
6122:
6112:
6111:
6104:
6103:
6096:
6089:
6081:
6072:
6071:
6069:
6068:
6055:
6052:
6051:
6049:
6048:
6047:
6046:
6036:
6035:
6034:
6032:Ancient Greece
6024:
6019:
6018:
6017:
6007:
6002:
5997:
5992:
5991:
5990:
5985:
5980:
5975:
5965:
5960:
5959:
5958:
5943:
5941:
5933:
5932:
5930:
5929:
5924:
5919:
5918:
5917:
5907:
5902:
5901:
5900:
5895:
5885:
5880:
5875:
5870:
5869:
5868:
5863:
5853:
5852:
5851:
5841:
5836:
5830:
5828:
5822:
5821:
5819:
5818:
5813:
5812:
5811:
5806:
5796:
5791:
5786:
5781:
5776:
5771:
5766:
5765:
5764:
5754:
5749:
5747:Sexual fantasy
5744:
5739:
5734:
5729:
5724:
5719:
5714:
5713:
5712:
5707:
5702:
5697:
5692:
5682:
5677:
5672:
5667:
5662:
5661:
5660:
5655:
5650:
5648:Anal fingering
5640:
5635:
5630:
5625:
5620:
5615:
5610:
5608:Foot fetishism
5605:
5600:
5595:
5590:
5585:
5580:
5575:
5570:
5565:
5560:
5555:
5549:
5547:
5541:
5540:
5538:
5537:
5532:
5527:
5522:
5517:
5512:
5507:
5501:
5499:
5493:
5492:
5490:
5489:
5484:
5479:
5474:
5469:
5467:Sexual capital
5464:
5459:
5454:
5449:
5444:
5439:
5434:
5432:Premarital sex
5429:
5424:
5419:
5414:
5409:
5404:
5399:
5394:
5389:
5383:
5381:
5370:
5369:
5367:
5366:
5361:
5356:
5351:
5346:
5341:
5335:
5333:
5327:
5326:
5324:
5323:
5322:
5321:
5316:
5311:
5309:Sexual slavery
5306:
5301:
5296:
5294:Sexual assault
5291:
5286:
5281:
5271:
5266:
5261:
5256:
5251:
5246:
5244:Age of consent
5240:
5238:
5232:
5231:
5229:
5228:
5223:
5218:
5213:
5208:
5203:
5197:
5195:
5185:
5184:
5182:
5181:
5176:
5171:
5166:
5165:
5164:
5159:
5157:Hypersexuality
5154:
5144:
5139:
5129:
5124:
5119:
5118:
5117:
5112:
5107:
5097:
5092:
5087:
5082:
5077:
5071:
5069:
5058:
5057:
5055:
5054:
5049:
5044:
5039:
5037:Sexual arousal
5034:
5029:
5024:
5019:
5014:
5013:
5012:
4998:
4993:
4988:
4983:
4978:
4973:
4967:
4965:
4954:
4953:
4942:
4941:
4934:
4927:
4919:
4910:
4909:
4907:
4906:
4895:
4892:
4891:
4889:
4888:
4887:
4886:
4885:
4884:
4879:
4874:
4859:
4853:
4851:
4845:
4844:
4842:
4841:
4840:
4839:
4834:
4829:
4824:
4819:
4814:
4813:
4812:
4797:
4792:
4787:
4782:
4781:
4780:
4775:
4765:
4764:
4763:
4758:
4748:
4747:
4746:
4741:
4731:
4726:
4721:
4716:
4711:
4706:
4705:
4704:
4699:
4688:
4686:
4678:
4677:
4675:
4674:
4669:
4664:
4659:
4658:
4657:
4652:
4642:
4637:
4636:
4635:
4630:
4623:Sex chromosome
4620:
4615:
4614:
4613:
4608:
4603:
4598:
4593:
4588:
4578:
4577:
4576:
4571:
4561:
4560:
4559:
4554:
4543:
4541:
4535:
4534:
4527:
4526:
4519:
4512:
4504:
4498:
4497:
4492:
4487:
4482:
4475:
4474:External links
4472:
4470:
4469:
4455:
4429:
4415:
4391:
4377:
4364:
4350:
4337:
4309:(10): 1013–9.
4294:
4291:
4266:
4264:
4261:
4259:
4258:
4251:
4233:
4226:
4206:
4199:
4179:
4172:
4152:
4145:
4125:
4084:
4025:
4018:
3998:
3949:
3931:
3911:
3870:
3829:
3791:
3776:
3753:
3718:
3699:(April 2015),
3684:
3644:
3601:
3577:
3570:
3533:
3488:
3446:(8): 729–733.
3423:
3416:
3398:
3371:
3353:
3322:
3296:
3287:
3249:
3224:
3217:
3199:
3184:
3145:
3102:
3073:(6): 1129–39.
3053:
3034:
3022:(2019-10-10).
3007:
2957:
2950:
2930:
2899:
2892:
2872:
2847:
2818:
2794:
2750:
2713:(4): 427–430.
2697:
2690:
2668:
2607:
2558:
2536:
2509:(1–3): 83–93.
2485:
2428:
2393:
2386:
2368:
2347:10.1086/406066
2341:(2): 189–208.
2325:
2290:
2279:
2259:
2244:
2209:
2179:
2162:
2151:
2125:
2111:
2102:Biology of Sex
2088:
2082:
2053:
2022:
1991:
1946:
1939:
1919:
1912:
1886:
1859:(9): 1816–24.
1843:
1828:
1803:Shackelford TK
1793:
1786:
1762:
1760:
1757:
1756:
1755:
1750:
1745:
1740:
1735:
1730:
1725:
1720:
1713:
1710:
1676:
1673:
1627:
1624:
1585:Main article:
1582:
1579:
1449:hermaphrodite,
1411:Anitist ritual
1383:Hermaphroditus
1364:
1363:
1361:
1360:
1353:
1346:
1338:
1335:
1334:
1331:
1330:
1325:
1320:
1315:
1313:Medical ethics
1310:
1309:
1308:
1303:
1298:
1293:
1288:
1283:
1278:
1268:
1263:
1262:
1261:
1251:
1246:
1241:
1235:
1232:
1231:
1228:
1227:
1224:
1223:
1218:
1216:United Kingdom
1213:
1208:
1203:
1198:
1193:
1188:
1183:
1178:
1173:
1168:
1163:
1158:
1153:
1148:
1143:
1138:
1133:
1128:
1122:
1117:
1116:
1113:
1112:
1109:
1108:
1103:
1098:
1093:
1092:
1091:
1086:
1075:
1069:
1068:
1065:
1064:
1061:
1060:
1055:
1054:
1053:
1043:
1038:
1037:
1036:
1026:
1021:
1016:
1011:
1006:
1001:
996:
991:
986:
979:
976:
975:
972:
971:
968:
967:
960:
955:
950:
945:
940:
935:
934:
933:
923:
918:
913:
908:
902:
896:
895:
892:
891:
888:
887:
882:
877:
875:Sex assignment
872:
867:
862:
857:
856:
855:
845:
843:Discrimination
840:
834:
825:
824:
821:
820:
810:
809:
799:
796:
772:gynomonoecious
767:
764:
751:
748:
727:
724:
714:that has both
661:
658:
645:spotted hyenas
637:Main article:
634:
631:
630:
629:
616:
610:
601:
590:
578:
563:
552:spermatophores
535:often occurs.
499:
496:
480:
479:
478:
477:
453:
452:
451:
434:
433:
432:
320:Main article:
317:
314:
312:
309:
270:Hermaphroditus
265:hermaphroditos
246:hermaphroditus
237:
234:
26:
18:Hermaphroditic
9:
6:
4:
3:
2:
6676:
6665:
6662:
6660:
6657:
6655:
6652:
6650:
6647:
6646:
6644:
6629:
6628:
6619:
6618:
6615:
6609:
6606:
6602:
6599:
6597:
6594:
6592:
6589:
6587:
6584:
6582:
6579:
6577:
6574:
6573:
6572:
6569:
6567:
6564:
6560:
6557:
6556:
6555:
6552:
6550:
6547:
6546:
6544:
6540:
6532:
6529:
6528:
6527:
6526:United States
6524:
6522:
6519:
6517:
6514:
6512:
6509:
6507:
6504:
6502:
6499:
6497:
6494:
6492:
6489:
6487:
6484:
6482:
6479:
6477:
6474:
6472:
6469:
6467:
6464:
6462:
6459:
6457:
6454:
6452:
6449:
6447:
6444:
6442:
6439:
6437:
6434:
6432:
6429:
6428:
6426:
6422:
6416:
6413:
6411:
6408:
6406:
6403:
6399:
6396:
6394:
6391:
6390:
6389:
6386:
6385:
6383:
6379:
6373:
6370:
6368:
6365:
6363:
6360:
6354:
6351:
6349:
6346:
6345:
6343:
6339:
6336:
6334:
6331:
6330:
6329:
6326:
6325:
6324:
6321:
6319:
6316:
6314:
6311:
6309:
6306:
6304:
6301:
6299:
6296:
6294:
6291:
6289:
6288:Gender system
6286:
6284:
6281:
6279:
6276:
6274:
6271:
6268:
6265:
6264:
6262:
6258:
6252:
6249:
6247:
6246:Quigley scale
6244:
6242:
6239:
6237:
6236:Phall-O-Meter
6234:
6232:
6229:
6225:
6222:
6221:
6220:
6217:
6215:
6214:Hermaphrodite
6212:
6210:
6207:
6205:
6202:
6201:
6199:
6196:
6192:
6186:
6176:
6173:
6171:
6168:
6166:
6163:
6161:
6158:
6156:
6153:
6151:
6148:
6144:
6141:
6140:
6139:
6136:
6134:
6131:
6129:
6126:
6125:
6123:
6121:
6117:
6113:
6109:
6102:
6097:
6095:
6090:
6088:
6083:
6082:
6079:
6067:
6062:
6057:
6056:
6053:
6045:
6042:
6041:
6040:
6037:
6033:
6030:
6029:
6028:
6025:
6023:
6020:
6016:
6013:
6012:
6011:
6008:
6006:
6003:
6001:
5998:
5996:
5993:
5989:
5986:
5984:
5981:
5979:
5976:
5974:
5971:
5970:
5969:
5966:
5964:
5961:
5957:
5954:
5953:
5952:
5948:
5945:
5944:
5942:
5940:
5934:
5928:
5925:
5923:
5920:
5916:
5913:
5912:
5911:
5908:
5906:
5903:
5899:
5896:
5894:
5891:
5890:
5889:
5886:
5884:
5881:
5879:
5876:
5874:
5871:
5867:
5864:
5862:
5859:
5858:
5857:
5854:
5850:
5847:
5846:
5845:
5842:
5840:
5837:
5835:
5832:
5831:
5829:
5827:
5823:
5817:
5814:
5810:
5807:
5805:
5802:
5801:
5800:
5797:
5795:
5792:
5790:
5787:
5785:
5782:
5780:
5777:
5775:
5772:
5770:
5767:
5763:
5760:
5759:
5758:
5755:
5753:
5750:
5748:
5745:
5743:
5742:Sex positions
5740:
5738:
5735:
5733:
5730:
5728:
5725:
5723:
5720:
5718:
5715:
5711:
5708:
5706:
5703:
5701:
5698:
5696:
5693:
5691:
5688:
5687:
5686:
5683:
5681:
5678:
5676:
5673:
5671:
5668:
5666:
5663:
5659:
5656:
5654:
5651:
5649:
5646:
5645:
5644:
5641:
5639:
5636:
5634:
5631:
5629:
5626:
5624:
5621:
5619:
5618:Forced orgasm
5616:
5614:
5611:
5609:
5606:
5604:
5601:
5599:
5596:
5594:
5591:
5589:
5586:
5584:
5581:
5579:
5576:
5574:
5571:
5569:
5566:
5564:
5561:
5559:
5556:
5554:
5551:
5550:
5548:
5546:
5542:
5536:
5535:United States
5533:
5531:
5528:
5526:
5523:
5521:
5518:
5516:
5513:
5511:
5508:
5506:
5503:
5502:
5500:
5498:
5494:
5488:
5485:
5483:
5480:
5478:
5475:
5473:
5472:Sexual ethics
5470:
5468:
5465:
5463:
5460:
5458:
5455:
5453:
5450:
5448:
5445:
5443:
5440:
5438:
5435:
5433:
5430:
5428:
5425:
5423:
5420:
5418:
5415:
5413:
5410:
5408:
5405:
5403:
5400:
5398:
5395:
5393:
5390:
5388:
5385:
5384:
5382:
5380:
5375:
5374:Relationships
5371:
5365:
5362:
5360:
5357:
5355:
5352:
5350:
5347:
5345:
5342:
5340:
5337:
5336:
5334:
5332:
5328:
5320:
5317:
5315:
5312:
5310:
5307:
5305:
5302:
5300:
5297:
5295:
5292:
5290:
5287:
5285:
5282:
5280:
5277:
5276:
5275:
5272:
5270:
5267:
5265:
5262:
5260:
5257:
5255:
5252:
5250:
5247:
5245:
5242:
5241:
5239:
5237:
5233:
5227:
5224:
5222:
5219:
5217:
5214:
5212:
5209:
5207:
5204:
5202:
5201:Gender binary
5199:
5198:
5196:
5194:
5190:
5186:
5180:
5177:
5175:
5172:
5170:
5167:
5163:
5162:Hyposexuality
5160:
5158:
5155:
5153:
5150:
5149:
5148:
5145:
5143:
5140:
5137:
5136:PLISSIT model
5133:
5130:
5128:
5127:Sex education
5125:
5123:
5120:
5116:
5113:
5111:
5108:
5106:
5103:
5102:
5101:
5098:
5096:
5093:
5091:
5088:
5086:
5083:
5081:
5078:
5076:
5075:Birth control
5073:
5072:
5070:
5068:
5063:
5059:
5053:
5050:
5048:
5045:
5043:
5040:
5038:
5035:
5033:
5030:
5028:
5027:Pre-ejaculate
5025:
5023:
5020:
5018:
5017:Pelvic thrust
5015:
5011:
5007:
5004:
5003:
5002:
4999:
4997:
4994:
4992:
4989:
4987:
4984:
4982:
4979:
4977:
4974:
4972:
4969:
4968:
4966:
4964:
4959:
4955:
4951:
4947:
4940:
4935:
4933:
4928:
4926:
4921:
4920:
4917:
4905:
4897:
4896:
4893:
4883:
4880:
4878:
4875:
4873:
4870:
4869:
4868:
4865:
4864:
4863:
4860:
4858:
4855:
4854:
4852:
4850:
4846:
4838:
4837:Pelvic thrust
4835:
4833:
4830:
4828:
4825:
4823:
4820:
4818:
4815:
4811:
4808:
4807:
4806:
4803:
4802:
4801:
4798:
4796:
4793:
4791:
4788:
4786:
4783:
4779:
4776:
4774:
4771:
4770:
4769:
4768:Fertilization
4766:
4762:
4759:
4757:
4754:
4753:
4752:
4749:
4745:
4742:
4740:
4737:
4736:
4735:
4734:Gametogenesis
4732:
4730:
4727:
4725:
4722:
4720:
4717:
4715:
4712:
4710:
4707:
4703:
4700:
4698:
4695:
4694:
4693:
4690:
4689:
4687:
4685:
4679:
4673:
4670:
4668:
4667:parasexuality
4665:
4663:
4660:
4656:
4653:
4651:
4648:
4647:
4646:
4645:Hermaphrodite
4643:
4641:
4638:
4634:
4631:
4629:
4626:
4625:
4624:
4621:
4619:
4616:
4612:
4611:Haplodiploidy
4609:
4607:
4604:
4602:
4599:
4597:
4594:
4592:
4589:
4587:
4584:
4583:
4582:
4579:
4575:
4572:
4570:
4567:
4566:
4565:
4562:
4558:
4555:
4553:
4550:
4549:
4548:
4545:
4544:
4542:
4536:
4532:
4525:
4520:
4518:
4513:
4511:
4506:
4505:
4502:
4496:
4493:
4491:
4488:
4486:
4483:
4481:
4478:
4477:
4466:
4462:
4458:
4452:
4448:
4444:
4443:
4438:
4434:
4430:
4426:
4422:
4418:
4412:
4408:
4400:
4396:
4392:
4388:
4384:
4380:
4374:
4370:
4365:
4361:
4357:
4353:
4347:
4343:
4338:
4334:
4330:
4326:
4322:
4317:
4312:
4308:
4304:
4300:
4295:
4292:
4280:
4276:
4272:
4268:
4267:
4254:
4252:0-87893-403-0
4248:
4244:
4237:
4229:
4223:
4219:
4218:
4210:
4202:
4196:
4192:
4191:
4183:
4175:
4169:
4165:
4164:
4156:
4148:
4142:
4138:
4137:
4129:
4121:
4117:
4112:
4107:
4104:(4): 736–54.
4103:
4099:
4095:
4088:
4080:
4076:
4071:
4066:
4061:
4056:
4052:
4048:
4044:
4040:
4036:
4029:
4021:
4015:
4011:
4010:
4002:
3994:
3990:
3985:
3980:
3976:
3972:
3968:
3964:
3960:
3953:
3938:
3934:
3928:
3924:
3923:
3915:
3907:
3903:
3898:
3893:
3890:(2): 94–104.
3889:
3885:
3881:
3874:
3866:
3862:
3857:
3852:
3848:
3844:
3843:Endocrinology
3840:
3833:
3825:
3814:
3810:
3806:
3802:
3795:
3787:
3783:
3779:
3773:
3769:
3768:
3763:
3757:
3749:
3745:
3741:
3737:
3733:
3729:
3722:
3709:on 2016-01-06
3708:
3704:
3703:
3698:
3694:
3688:
3680:
3676:
3675:
3670:
3666:
3662:
3658:
3654:
3648:
3634:on 2017-03-22
3633:
3629:
3628:
3623:
3619:
3615:
3611:
3605:
3591:
3587:
3581:
3573:
3567:
3563:
3559:
3555:
3551:
3547:
3543:
3537:
3530:
3525:
3521:
3516:
3511:
3507:
3503:
3499:
3492:
3473:
3469:
3465:
3461:
3457:
3453:
3449:
3445:
3441:
3434:
3427:
3419:
3413:
3409:
3402:
3394:
3390:
3386:
3382:
3375:
3367:
3363:
3357:
3341:
3337:
3333:
3326:
3318:
3314:
3312:
3306:
3300:
3291:
3280:
3276:
3272:
3268:
3265:(2): 355–60.
3264:
3260:
3253:
3238:
3234:
3228:
3220:
3214:
3210:
3203:
3195:
3188:
3180:
3176:
3172:
3168:
3164:
3160:
3156:
3149:
3141:
3137:
3133:
3129:
3125:
3121:
3118:(2): 404–12.
3117:
3113:
3106:
3098:
3094:
3089:
3084:
3080:
3076:
3072:
3068:
3064:
3057:
3041:
3037:
3031:
3027:
3026:
3021:
3014:
3012:
3003:
2999:
2995:
2991:
2987:
2983:
2979:
2975:
2968:
2961:
2953:
2947:
2943:
2942:
2934:
2919:on 2009-10-22
2918:
2914:
2910:
2903:
2895:
2889:
2885:
2884:
2876:
2862:on 2010-11-23
2861:
2857:
2851:
2837:on 2011-03-10
2836:
2832:
2828:
2822:
2814:
2810:
2809:
2804:
2798:
2790:
2786:
2782:
2778:
2774:
2770:
2766:
2762:
2754:
2746:
2742:
2737:
2732:
2728:
2724:
2720:
2716:
2712:
2708:
2701:
2693:
2687:
2683:
2679:
2672:
2664:
2660:
2656:
2652:
2648:
2644:
2640:
2636:
2631:
2630:q-bio/0508003
2626:
2622:
2618:
2611:
2603:
2599:
2594:
2589:
2585:
2581:
2577:
2573:
2569:
2562:
2554:
2550:
2546:
2540:
2532:
2528:
2524:
2520:
2516:
2512:
2508:
2504:
2500:
2496:
2489:
2481:
2477:
2472:
2467:
2463:
2459:
2455:
2451:
2447:
2443:
2439:
2432:
2424:
2420:
2416:
2412:
2408:
2404:
2397:
2389:
2383:
2379:
2372:
2364:
2360:
2356:
2352:
2348:
2344:
2340:
2336:
2329:
2321:
2317:
2313:
2309:
2305:
2301:
2294:
2286:
2282:
2276:
2273:. CRC Press.
2272:
2271:
2263:
2255:
2251:
2247:
2241:
2237:
2232:
2231:
2222:
2220:
2218:
2216:
2214:
2197:
2193:
2189:
2183:
2175:
2174:
2166:
2160:
2155:
2139:
2135:
2129:
2122:
2120:
2119:hermaphrodite
2114:
2108:
2104:
2103:
2095:
2093:
2085:
2079:
2075:
2071:
2067:
2060:
2058:
2050:
2039:
2035:
2029:
2027:
2010:
2006:
2002:
1995:
1988:
1983:
1979:
1974:
1969:
1966:(2): 94–104.
1965:
1961:
1957:
1950:
1942:
1936:
1932:
1931:
1923:
1915:
1909:
1905:
1901:
1897:
1890:
1882:
1878:
1874:
1870:
1866:
1862:
1858:
1854:
1847:
1839:
1835:
1831:
1825:
1821:
1817:
1810:
1809:
1804:
1797:
1789:
1783:
1779:
1778:
1770:
1768:
1763:
1754:
1751:
1749:
1746:
1744:
1743:Gynandromorph
1741:
1739:
1736:
1734:
1731:
1729:
1726:
1724:
1721:
1719:
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1715:
1709:
1707:
1703:
1698:
1696:
1690:
1686:
1682:
1672:
1670:
1666:
1662:
1661:invertebrates
1658:
1653:
1651:
1647:
1643:
1640:
1636:
1632:
1623:
1620:
1615:
1613:
1609:
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1594:
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1572:
1568:
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1517:
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1492:
1491:
1486:
1482:
1481:
1476:
1472:
1471:Victorian era
1467:
1465:
1461:
1460:
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1439:
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1428:
1424:
1419:
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1388:
1384:
1380:
1375:
1371:
1359:
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1352:
1347:
1345:
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1324:
1321:
1319:
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1264:
1260:
1257:
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1255:
1252:
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1239:Hermaphrodite
1237:
1236:
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1229:
1222:
1221:United States
1219:
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959:
956:
954:
953:Quigley scale
951:
949:
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943:Phall-O-Meter
941:
939:
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929:
928:
927:
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721:
717:
713:
710:
706:
702:
701:hermaphrodite
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690:
686:
685:
680:
675:
671:
667:
657:
655:
651:
646:
640:
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623:
622:
617:
614:
611:
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607:
602:
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594:
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576:
575:apophallation
572:
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564:
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548:
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541:
538:
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536:
534:
530:
523:
519:
512:
511:penis fencing
508:
507:Turbellarians
504:
495:
493:
489:
485:
474:
470:
466:
465:
460:
459:
457:
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445:
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440:
438:
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426:
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417:
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409:Example: The
408:
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402:
401:
400:
397:
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391:
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383:
380:(such as the
379:
375:
370:
366:
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346:
342:
335:
334:
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323:
308:
306:
302:
301:hermaphrodite
298:
294:
291:
287:
283:
279:
275:
271:
266:
260:
251:
250:Ancient Greek
247:
243:
233:
231:
227:
222:
220:
216:
212:
208:
204:
200:
196:
192:
189:About 94% of
187:
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173:
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160:
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144:
140:
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126:
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118:
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64:
63:hermaphrodite
56:
55:Garden snails
52:
48:
44:
40:
33:
19:
6625:
6596:Third gender
6591:Queer theory
6496:South Africa
6241:Prader scale
6231:Orchidometer
6213:
6138:Human rights
6120:legal issues
6116:Human rights
5937:Religion and
5927:Webcam model
5866:Survival sex
5856:Prostitution
5826:Sex industry
5737:Sex in space
5665:Masturbation
5505:Ancient Rome
5482:Sexual slang
5274:Sexual abuse
4981:Insemination
4756:spermatozoon
4684:reproduction
4644:
4633:Y chromosome
4628:X chromosome
4574:Virilization
4569:Feminization
4441:
4406:
4398:
4368:
4341:
4306:
4302:
4285:12 September
4283:. Retrieved
4279:the original
4274:
4242:
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4189:
4182:
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4038:
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4012:. Springer.
4008:
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3952:
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3921:
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3873:
3846:
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3832:
3822:– via
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3804:
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3727:
3721:
3711:, retrieved
3707:the original
3701:
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3647:
3636:, retrieved
3632:the original
3626:
3604:
3593:. Retrieved
3589:
3580:
3553:
3549:
3536:
3527:
3508:(2): 65–66.
3505:
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3472:the original
3443:
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3339:
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3236:
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3024:
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2973:
2960:
2940:
2933:
2923:12 September
2921:. Retrieved
2917:the original
2912:
2902:
2882:
2875:
2864:. Retrieved
2860:the original
2850:
2839:. Retrieved
2835:the original
2830:
2821:
2808:Live Science
2806:
2797:
2764:
2760:
2753:
2710:
2706:
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2681:
2671:
2620:
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2431:
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2396:
2377:
2371:
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2334:
2328:
2306:(6): 133–6.
2303:
2299:
2293:
2269:
2262:
2229:
2200:. Retrieved
2191:
2182:
2172:
2165:
2154:
2142:. Retrieved
2128:
2118:
2116:
2101:
2065:
2047:
2041:. Retrieved
2037:
2013:. Retrieved
1994:
1985:
1963:
1959:
1949:
1929:
1922:
1895:
1889:
1856:
1852:
1846:
1807:
1796:
1776:
1699:
1692:
1669:Nelson Çabej
1657:polyphyletic
1654:
1639:paraphyletic
1629:
1619:mating types
1616:
1605:
1560:
1528:
1518:
1499:
1489:
1479:
1477:were termed
1468:
1463:
1457:
1448:
1443:
1432:
1301:Third gender
1296:Queer theory
1238:
1191:South Africa
962:
948:Prader scale
938:Orchidometer
848:Human rights
831:legal issues
827:Human rights
789:
782:
775:
769:
753:
750:Andromonoecy
729:
700:
698:
682:
642:
619:
604:
584:
566:Banana slugs
527:
488:aquacultured
481:
462:
455:
436:
425:sea anemones
414:
403:
398:
362:
331:
303:entered the
300:
254:ἑρμαφρόδιτος
239:
229:
223:
215:andromonoecy
188:
176:
136:
62:
60:
47:
6601:Transgender
6506:Switzerland
6491:New Zealand
6362:Researchers
6197:and biology
6010:LGBT people
5956:Tantric sex
5888:Sex tourism
5844:Pornography
5809:Erotic talk
5799:Virtual sex
5700:Cunnilingus
5530:South Korea
5525:Philippines
5437:Promiscuity
5132:Sex therapy
5110:Gynaecology
4433:Grumbach MM
3734:(1): 25–8.
3556:: 155–182.
3542:Georgiann D
3387:: 277–278.
2831:EurekAlert!
2736:10069/35713
2403:J Fish Biol
1999:Herndon A.
1738:Gonochorism
1706:heterostyly
1612:gonochorism
1469:During the
1454:Edward Coke
1407:Philippines
1306:Transgender
1201:Switzerland
1186:New Zealand
1041:Researchers
916:Definitions
900:and biology
766:Gynomonoecy
703:is used in
560:spermatozoa
556:spermatheca
543:land snails
367:) occur in
219:gynomonoecy
203:androdioecy
6643:Categories
6554:Disability
6393:of surgery
6372:Television
6348:Androgynos
6333:Aphroditus
6308:Literature
5922:Strip club
5905:Sex worker
5878:Sex museum
5849:Film actor
5717:Paraphilia
5643:Manual sex
5497:By country
5422:Orgasm gap
5339:Blue Movie
4958:Physiology
4817:Copulation
4538:Biological
4045:(1): 242.
3943:2021-09-29
3713:2018-06-10
3595:2024-03-12
3243:2019-07-03
2866:2011-03-19
2841:2011-03-20
2043:2024-01-20
1801:Holub AM,
1759:References
1679:See also:
1626:In animals
1555:testicular
1502:karyotypes
1464:Androgynus
1286:Non-binary
1254:Disability
1084:of surgery
1073:and events
1058:Television
1014:Literature
921:Healthcare
792:bryophytes
736:gymnosperm
732:monoecious
720:carpellate
593:Earthworms
522:Earthworms
509:mating by
492:broodstock
469:Lythrypnus
416:Amphiprion
378:gastropods
357:parrotfish
330:Shells of
207:gynodioecy
195:monoecious
167:vertebrate
159:earthworms
141:groups of
132:gonochoric
6576:Androgyny
6436:Australia
6431:Argentina
6293:Guevedoce
6269:(antonym)
6044:Sex magic
6022:Mormonism
5939:sexuality
5794:Urolagnia
5784:Tribadism
5732:Sexercise
5710:Irrumatio
5695:Anilingus
5653:Fingering
5633:Group sex
5628:Gang bang
5598:Fetishism
5573:Child sex
5427:Polyamory
5407:Free love
5269:Obscenity
5193:diversity
5105:Andrology
5067:education
5032:Pregnancy
4872:Mechanics
4849:Sexuality
4744:Oogenesis
4719:Sex organ
4709:Germ cell
4697:Anisogamy
4447:1303–1425
4425:141382073
4325:357415945
3805:Endotext
3638:March 21,
3342:: 165–167
3165:: 43–54.
3020:Minelli A
3018:Fusco G,
2545:Morelle R
2254:299866547
2015:2 October
1838:240938739
1728:Androgyny
1675:In plants
1663:where it
1608:anisogamy
1581:Evolution
1553:). Thus,
1495:phenotype
1483:if their
1391:Aphrodite
1276:Androgyny
1131:Australia
1126:Argentina
985:(antonym)
785:unisexual
716:staminate
699:The term
582:sea slugs
545:and land
540:Pulmonate
461:Example:
442:Example:
437:Protogyny
421:symbiosis
411:clownfish
404:Protandry
365:dichogamy
345:Clownfish
278:Aphrodite
259:romanized
236:Etymology
151:tunicates
139:taxonomic
6627:Category
6542:See also
6531:military
6456:Colombia
6398:timeline
6344:Judaism
6323:Religion
6195:Medicine
6108:Intersex
5951:Hinduism
5947:Buddhism
5883:Sex shop
5804:Cybersex
5779:Swinging
5762:Foreplay
5705:Fellatio
5685:Oral sex
5578:Creampie
5563:Bareback
5558:Anal sex
5412:Marriage
5402:Flirting
5189:Identity
5142:Sexology
5122:Safe sex
4986:Intersex
4904:Category
4882:Activity
4778:Internal
4773:External
4662:Intersex
4465:35364729
4387:38519315
4360:52152732
4333:12379746
4120:23817661
4079:29207942
3993:36934747
3984:10024982
3937:Archived
3906:28032683
3865:11459768
3813:25905232
3786:57722472
3762:Leyner M
3748:10602113
3679:archived
3665:Méndez J
3622:Briffa T
3529:parents.
3524:17304046
3468:39459050
3460:16200837
3364:(1812).
3346:13 April
3317:Archived
3279:16490032
3179:21632274
3132:17348949
3097:19671577
3046:29 March
3040:Archived
3002:90740232
2813:Archived
2789:13639701
2781:17029221
2655:16005289
2602:23407499
2553:BBC News
2531:83829239
2480:26306176
2363:38139187
2320:21227182
2285:Archived
2196:Archived
2138:Archived
2009:Archived
1982:28032683
1881:23849389
1873:17089966
1733:Futanari
1712:See also
1642:Spiralia
1573:, and a
1374:Intersex
1233:See also
1151:Colombia
1089:timeline
1029:Military
1024:Religion
898:Medicine
806:Intersex
740:conifers
484:groupers
467:(Family
448:Labridae
446:(Family
293:Salmacis
155:mollusks
117:organism
6466:Germany
6388:History
6267:Endosex
6224:history
6143:reports
6039:Thelema
6005:Judaism
5910:Sex toy
5839:Erotica
5727:Quickie
5722:Pompoir
5658:Handjob
5613:Footjob
5603:Fisting
5442:Romance
5417:Modesty
5379:society
5331:History
5115:Urology
4963:biology
4946:Outline
4729:Meiosis
4702:Isogamy
4070:5717846
4047:Bibcode
3481:27 July
3088:2766191
2982:Bibcode
2980:: 1–9.
2761:Genesis
2745:9474211
2715:Bibcode
2663:2229622
2635:Bibcode
2593:3639767
2511:Bibcode
2471:4541995
2450:Bibcode
2411:Bibcode
2355:4901396
2144:19 July
1723:Trioecy
1551:tissues
1548:ovarian
1536:meiosis
1485:gonadal
1405:in the
1161:Germany
1079:History
1071:History
983:Endosex
964:more...
931:history
853:reports
760:perfect
726:Monoecy
709:perfect
693:stamens
689:carpels
674:Monoicy
613:Hamlets
598:gametes
573:), see
444:Wrasses
413:(genus
374:teleost
369:species
311:Animals
261::
248:, from
211:trioecy
180:mammals
172:selfing
143:animals
128:gametes
112:) is a
6566:Gender
6559:rights
6516:Uganda
6511:Taiwan
6481:Mexico
6461:France
6441:Canada
6353:Tumtum
6318:People
5995:Daoism
5898:Female
5861:Sacred
5774:Sumata
5593:Facial
5583:Edging
5259:Incest
5080:Condom
5062:Health
5006:Female
5001:Orgasm
4991:Libido
4751:Gamete
4724:Mating
4682:Sexual
4557:Female
4463:
4453:
4423:
4413:
4385:
4375:
4358:
4348:
4331:
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4249:
4224:
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4170:
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4077:
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3991:
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3929:
3904:
3863:
3818:28 Mar
3811:
3784:
3774:
3746:
3568:
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3466:
3458:
3414:
3393:896307
3391:
3362:Coke E
3277:
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3177:
3140:781464
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3130:
3095:
3085:
3032:
3000:
2948:
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2743:
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2661:
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2529:
2478:
2468:
2384:
2361:
2353:
2318:
2277:
2252:
2242:
2202:3 June
2109:
2080:
2049:words.
1987:2011).
1980:
1937:
1910:
1879:
1871:
1836:
1826:
1784:
1687:, and
1646:phylum
1599:, and
1540:testes
1506:mosaic
1387:Hermes
1318:Rights
1266:Gender
1259:rights
1211:Uganda
1206:Taiwan
1176:Mexico
1156:France
1136:Canada
1019:People
808:topics
712:flower
705:botany
672:, and
660:Plants
571:radula
274:Hermes
178:among
157:, and
125:female
57:mating
6501:Spain
6486:Nepal
6476:Malta
6471:Kenya
6451:China
6446:Chile
6283:Films
6000:Islam
5893:Child
5520:Japan
5515:India
5510:China
4540:terms
3475:(PDF)
3464:S2CID
3436:(PDF)
3175:S2CID
3157:L.".
3136:S2CID
2998:S2CID
2970:(PDF)
2785:S2CID
2741:S2CID
2659:S2CID
2625:arXiv
2527:S2CID
2359:S2CID
1877:S2CID
1834:S2CID
1812:(PDF)
1423:Nadar
1196:Spain
1181:Nepal
1171:Malta
1166:Kenya
1146:China
1141:Chile
999:Films
643:When
547:slugs
423:with
290:nymph
242:Latin
184:birds
147:sexes
6571:LGBT
6298:Flag
6118:and
5949:and
5915:Doll
5623:Frot
5568:BDSM
5377:and
5284:Rape
5191:and
5008:and
4961:and
4761:ovum
4552:Male
4461:OCLC
4451:ISBN
4421:OCLC
4411:ISBN
4383:OCLC
4373:ISBN
4356:OCLC
4346:ISBN
4329:PMID
4321:OCLC
4287:2009
4247:ISBN
4222:ISBN
4195:ISBN
4168:ISBN
4141:ISBN
4116:PMID
4075:PMID
4014:ISBN
3989:PMID
3927:ISBN
3902:PMID
3861:PMID
3820:2023
3809:PMID
3782:OCLC
3772:ISBN
3744:PMID
3640:2017
3566:ISBN
3520:PMID
3483:2016
3456:PMID
3412:ISBN
3389:SSRN
3348:2023
3275:PMID
3213:ISBN
3128:PMID
3093:PMID
3048:2021
3030:ISBN
2946:ISBN
2925:2009
2888:ISBN
2777:PMID
2686:ISBN
2651:PMID
2598:PMID
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