61:
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markings. The legs are brown with a yellow like base. Wings are born black but grow yellow-brown when they grow. Males have a bright sky blue or turquoise abdomen marked with a diagnostic black dorsal stripe. However, their blue colour may be faded during cold weather spells. The thorax and head of a male is apple green and their prominent eyes are blue. Females have similar markings but they are mainly a duller green. As the females age, their wings become browner. Less immediately visible features for both sexes are the yellow
459:
115:
433:
409:
90:
42:
1083:
366:, and south-western and central Asia. Since the 1990s, its range has expanded in Europe, both northwards and to higher altitudes. For example, the first Scandinavian record was in 1994 in Denmark; in 2002 it was first recorded in Sweden and in 2004 first in Scotland; today it is regular in all three countries. The species' northward expansion has been tied to
482:
and tadpoles. If their hunt is successful, they eat their smaller prey while flying. The dragonflies breed in a variety of aquatic habitats from large ponds to dikes and slow-moving rivers, but require a plentiful supply of vegetation in the water. They do sometimes breed in brackish water. The
382:
The emperor dragonfly is a large and bulky species. It is 73–82 mm (2.9–3.2 in) long, with average being 78 mm (3.1 in) and males growing larger than females. The average wingspan is 104 mm (4.1 in). When they first emerge, both sexes appear pale green with brown
967:
García-Olivares, Víctor; Muñoz-Barrera, Adrián; Lorenzo-Salazar, José M.; Zaragoza-Trello, Carlos; Rubio-Rodríguez, Luis A.; Díaz-de Usera, Ana; Jáspez, David; Iñigo-Campos, Antonio; González-Montelongo, Rafaela; Flores, Carlos (2021-10-15).
841:
495:
of freshwater ecosystems they arrive in. The larvae are also very large–around 46 millimetres (1.8 in). The adult male is highly territorial, and difficult to approach.
387:
and brown spots on the wings. Emperor dragonflies can also be recognised by their flight patterns: they often fly with their abdomen hanging slightly downwards.
867:
1286:
1325:
1392:
1422:
810:
914:
1260:
324:. It is the largest dragonfly in most of Europe, including the United Kingdom, although exceeded in some areas by other species.
851:
1151:
663:
938:"Biometry of the large dragonfly Anax imperator (Odonata: Aeshnidae): A study of traits from larval development to adults"
17:
1417:
1221:
1019:"Complete mitochondrial genome sequence of Anax parthenope (Odonata: Anisoptera: Aeshnidae) and phylogenetic analysis"
290:
1239:
1330:
970:"A benchmarking of human mitochondrial DNA haplogroup classifiers from whole-genome and whole-exome sequence data"
420:
1252:
444:
257:
1412:
1312:
713:
Herzog, Rebecca; Osigus, Hans−Jürgen; Feindt, Wiebke; Schierwater, Bernd; Hadrys, Heike (29 October 2016).
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60:
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279:
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658:. Vol. 1. With the assistance of Lucien van Beek. Leiden, Boston: Brill. pp. 98–99.
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8:
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408:
394:, which occurs only marginally in the east Mediterranean and in length by females of the
268:
79:
519:
of the emperor dragonfly is the longest of all known dragonfly sequences. It has 16,087
1397:
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1002:
758:
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391:
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109:
1351:
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949:
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730:
581:
196:
1187:
1034:
735:
1338:
528:
1317:
488:
402:. Thus, in most of Europe the emperor is the largest dragonfly species present.
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993:
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They frequently fly high up into the sky in search of prey, which includes
390:
One of the largest species in Europe, the emperor dragonfly is exceeded by
359:
953:
1273:
1131:
484:
70:
1161:
1343:
1265:
786:. Insects of Britain and Ireland: dragonflies and damselflies (Odonata)
399:
843:
Europe's
Dragonflies: A field guide to the damselflies and dragonflies
41:
1278:
1208:
520:
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by the IUCN. The species has widespread and has a stable population.
475:
321:
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186:
176:
146:
126:
1093:
1213:
1116:
51:
811:"Emperor dragonfly videos, photos and facts — Anax imperator"
479:
450:
371:
166:
362:; it is found throughout Africa and through most of Europe, the
1200:
1146:
1082:
1017:
Wang, Ying; Du, Yimin; Song, Xiang; Huang, Aijun (2021-01-02).
936:
Minot, Marceau; Le Gall, Mickael; Huste, Aurelie (2019-09-10).
156:
136:
557:
555:
553:
551:
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547:
545:
712:
649:
337:
774:
772:
715:"The complete mitochondrial genome of the emperor dragonfly
646:
542:
491:
are very aggressive and are likely to influence the native
769:
868:"Anax immaculifrons Rambur, 1842 – Magnificent Emperor"
1077:
Blue
Emperor - OdonataMAP - Atlas of African Odonata
870:. Dragonflies and Damselflies – in and around Europe
719:
LEACH, 1815 (Odonata: Aeshnidae) via NGS sequencing"
523:. For comparison, the human mitogenome has 16,569
935:
1384:
1016:
839:
358:This dragonfly has a wide distribution through
587:10.2305/IUCN.UK.2016-3.RLTS.T59812A72311295.en
840:Smallshire, Dave; Swash, Andy (2020-07-14).
708:
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59:
40:
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1001:
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734:
697:
585:
561:
483:females lay the eggs into plants such as
510:
14:
1385:
641:
503:Emperor dragonflies are assessed as a
1098:
1097:
805:
803:
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635:
612:
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1253:f1565b76-be8b-4417-a7a2-221ec0b6207f
835:
833:
831:
1393:IUCN Red List least concern species
573:IUCN Red List of Threatened Species
398:, a species with an unusually long
24:
1423:Taxa named by William Elford Leach
798:
601:
527:and the closely related dragonfly
524:
25:
1434:
1070:
828:
377:
346:is the Latin for "emperor", from
1081:
656:Etymological Dictionary of Greek
457:
443:
431:
419:
407:
113:
1010:
960:
929:
907:
882:
498:
353:
342:, "lord"; the specific epithet
327:
316:) is a large species of hawker
942:European Journal of Entomology
860:
846:. Princeton University Press.
672:
13:
1:
1035:10.1080/23802359.2020.1848479
736:10.1080/23802359.2016.1186523
536:
487:, and always lay alone. The
469:
370:, and it is among the first
7:
684:Online Etymology Dictionary
10:
1439:
986:10.1038/s41598-021-99895-5
650:
338:
336:is from the ancient Greek
1418:Insects described in 1815
1106:
894:British Dragonfly Society
890:"Golden-ringed Dragonfly"
622:British Dragonfly Society
250:
243:
225:
218:
110:Scientific classification
108:
86:
77:
67:
58:
48:
39:
34:
1023:Mitochondrial DNA Part B
813:. ARKive. Archived from
723:Mitochondrial DNA Part B
396:golden-ringed dragonfly
1413:Dragonflies of Europe
1365:Paleobiology Database
954:10.14411/eje.2019.031
580:: e.T59812A72311295.
511:Mitochrondrial genome
505:least-concern species
1248:Fauna Europaea (new)
1090:at Wikimedia Commons
919:Lincolnshire Species
27:Species of dragonfly
915:"Emperor Dragonfly"
493:species composition
392:magnificent emperor
80:Conservation status
18:Emperor (dragonfly)
974:Scientific Reports
562:Mitra, A. (2016).
438:Female laying eggs
35:Emperor dragonfly
1403:Odonata of Africa
1380:
1379:
1352:Open Tree of Life
1100:Taxon identifiers
1086:Media related to
853:978-0-691-20497-0
414:Male in side view
364:Arabian Peninsula
332:The generic name
304:emperor dragonfly
300:
299:
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211:A. imperator
103:
16:(Redirected from
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1318:NBNSYS0000005630
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921:. Archived from
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680:"imperator (n.)"
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665:978-900417-418-4
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265:Anax mauricianus
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21:
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1408:Odonata of Asia
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1339:Observation.org
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925:on 25 Oct 2003.
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530:Anax parthenope
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1235:Fauna Europaea
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1152:anax-imperator
1143:
1138:Anax imperator
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1108:Anax imperator
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1088:Anax imperator
1079:
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1071:External links
1069:
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1066:
1029:(1): 122–123.
1009:
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784:(Leach, 1815)"
782:Anax imperator
768:
729:(1): 783–786.
717:Anax imperator
696:
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664:
643:Beekes, Robert
634:
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566:Anax imperator
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489:aquatic larvae
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426:Male in flight
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379:
378:Identification
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368:global warming
355:
352:
350:, to command.
329:
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320:of the family
313:Anax imperator
298:
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287:Aeschna lunata
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276:Aeschna azurea
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980:(1): 20510.
977:
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931:
923:the original
918:
909:
897:. Retrieved
884:
872:. Retrieved
862:
842:
819:. Retrieved
815:the original
788:. Retrieved
781:
726:
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687:. Retrieved
683:
674:
655:
637:
625:. Retrieved
591:. Retrieved
577:
571:
565:
533:has 15,366.
529:
514:
502:
499:Conservation
473:
389:
381:
357:
354:Distribution
347:
343:
333:
331:
328:Nomenclature
312:
311:
308:blue emperor
307:
303:
301:
286:
275:
264:
254:Anax formosa
253:
228:
226:
210:
209:
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173:Infraorder:
29:
1274:iNaturalist
1132:Wikispecies
948:: 269–280.
899:25 November
874:25 November
790:25 November
593:19 November
476:butterflies
360:Afroeurasia
280:Charpentier
71:Oxfordshire
1387:Categories
821:2013-08-08
645:(2010) . "
537:References
521:base pairs
517:mitogenome
400:ovipositor
374:to do so.
177:Anisoptera
147:Arthropoda
73:, England
1398:Aeshnidae
1043:2380-2359
994:2045-2322
689:27 August
627:25 August
618:"Emperor"
470:Behaviour
344:imperator
322:Aeshnidae
318:dragonfly
205:Species:
187:Aeshnidae
133:Kingdom:
127:Eukaryota
1292:11246064
1159:BioLib:
1117:Wikidata
1061:33490598
763:89088747
755:33473626
485:pondweed
478:, other
464:Emerging
348:imperare
293:, 1856
291:Kolenati
282:, 1825
271:, 1842
260:, 1823
245:Synonyms
183:Family:
143:Phylum:
137:Animalia
123:Domain:
100:IUCN 3.1
54:, Spain
52:Tenerife
1266:5051775
1123:Q527496
1052:7808743
1003:8519921
746:7799497
480:odonata
451:Exuviae
372:odonata
193:Genus:
167:Odonata
163:Order:
157:Insecta
153:Class:
98: (
1370:194367
1357:485521
1240:214314
1227:241277
1214:ANXXIM
1201:131867
1175:198426
1147:ARKive
1059:
1049:
1041:
1000:
992:
850:
761:
753:
743:
662:
269:Rambur
238:, 1815
68:female
1331:39274
1305:59812
1287:IRMNG
1279:94043
1222:EUNIS
759:S2CID
385:costa
236:Leach
1326:NCBI
1300:IUCN
1261:GBIF
1209:EPPO
1188:DJX8
1170:BOLD
1162:1622
1057:PMID
1039:ISSN
990:ISSN
901:2023
876:2023
848:ISBN
792:2023
751:PMID
691:2021
660:ISBN
651:ἄναξ
647:S.v.
629:2010
595:2021
578:2016
515:The
339:ἄναξ
334:Anax
302:The
198:Anax
49:Male
1344:616
1313:NBN
1196:EoL
1183:CoL
1047:PMC
1031:doi
998:PMC
982:doi
950:doi
946:116
741:PMC
731:doi
654:".
582:doi
306:or
1389::
1367::
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1119::
1055:.
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1037:.
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1021:.
996:.
988:.
978:11
976:.
972:.
944:.
940:.
917:.
892:.
830:^
800:^
771:^
757:.
749:.
739:.
725:.
721:.
699:^
682:.
620:.
603:^
576:.
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