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Pinales

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first complete molecular phylogeny was published, based on 90 species representing all extant genera. This established cycads as the basal group, followed by Ginkgoaceae, as sister to the remaining gymnosperms, and supporting the ‘gnepine’ hypothesis. This analysis favours the five clade hypothesis, the remaining clade following divergence of the Pinidae, are referred to as the conifer II clade, or cupressophytes, in distinction from the conifer I clade (Gnetidae, Pinidae). This clade, in turn, has two lineages. The first consisting of Sciadopityaceae and the Araucariales, the second being the Cupressales. In the Christenhusz scheme, the Sciadopityaceae were considered to be within Cupressales. The term Cupressaceae
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However, the exact phylogeny remained a topic that was 'hotly debated", in particular whether the main lineages were best represented by the four subclasses of Christenhusz and colleagues or the more traditional five clades (cycads, ginkgos, cupressophytes, Pinaceae and gnetophytes). In 2014 the
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Syllabus der Pflanzenfamilien: eine Übersicht über das gesamte Pflanzensystem mit Berücksichtigung der Medicinal- und Nutzpflanzen nebst einer Übersicht über die Florenreiche und Florengebiete der Erde zum Gebrauch bei Vorlesungen und Studien über specielle und medicinisch-pharmaceutische
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An introduction to the natural system of botany: or, A systematic view of the organisation, natural affinities, and geographical distribution, of the whole vegetable kingdom: together with the uses of the most important species in medicine, the arts, and rural or domestic
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Brunsfeld, Steven J.; Soltis, Pamela S.; Soltis, Douglas E.; Gadek, Paul A.; Quinn, Christopher J.; Strenge, Darren D.; Ranker, Tom A. (April 1994). "Phylogenetic Relationships Among the Genera of Taxodiaceae and Cupressaceae: Evidence from rbcL Sequences".
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and the Gymnosperm Database. In this restricted model Pinales (Pinaceae) comprisea 11 genera and about 225 species, all of the other conifers originally included in this order, being included in other orders such as Cupressales.
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A system of two groups was maintained by the most commonly used classification in the twentieth century, the revision of the Engler system by Pilger (1926), who grouped 12 families of the Gymnospermae subdivision into 2 classes;
2530:"LIV. Character and description of Kingia, a new genus of plants found on the South-west coast of New Holland: with observations on the structure of its unimpregnated ovulum; and on the female flower of Cycadeœ and Coniferœ" 1375:
A more comprehensive analysis was undertaken by Ran and colleagues in 2018, as part of a detailed phylogeny of all seed plants. This forms the basis of the Tracheophyte Phylogeny Poster and the Angiosperm Phylogeny Website.
430:, Gymnosperms and Angiosperms. In the gymnosperms (or Gymnospermae) Lindley included two orders, the Cycadeae and the Coniferae. In his final work (1853) he described Gymnogens as a class with four orders; 554:
The treatment of Gymnosperms as two groups, though with varying composition and names, was followed for most of the twentieth century, including the systems of Chamberlain (1935), Benson (1957) and
804:, the number of taxonomic publications increased considerably after 2008, however, these approaches have not been uniform. A taxonomic classification has been complicated by the relationship of 1473:, formerly in Cupressaceae, into a separate family (Sciadopityaceae). Cephalotaxaceae had previously been recognized as a separate family, but was subsequently included in Taxaceae. Similarly 1388:(1926), considered the standard through most of the twentieth century. These families were treated as a single order, in distinction to some earlier systems. His families were; 824:
to conifers (‘gnetifer’ hypothesis) or sister to Pinaceae (‘gnepine’ hypothesis) in which they are classified within the conifers. The extant conifers most likely form a
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Ran, Jin-Hua; Shen, Ting-Ting; Wu, Hui; Gong, Xun; Wang, Xiao-Quan (December 2018). "Phylogeny and evolutionary history of Pinaceae updated by transcriptomic analysis".
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Die natürlichen Pflanzenfamilien nebst ihren Gattungen und wichtigeren Arten, insbesondere den Nutzpflanzen, unter Mitwirkung zahlreicher hervorragender Fachgelehrten
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Historically conifers, in the order Pinales have been considered to consist of six to seven extant families, based on the classification of class Coniferae by
2677: 2372:Лекции по морфологии и систематики архегониальных растений (Lekts. Morf. Sist. Archegon. Lectures on the morphology and taxonomy of archegonial plants) 2987:"Phylogenomics resolves the deep phylogeny of seed plants and indicates partial convergent or homoplastic evolution between Gnetales and angiosperms" 193: 3465: 387:, with studies recovering Gnetales as either a sister group to Pinaceae or being more derived than Pinaceae but sister to the rest of the group. 3504: 2175:
Histoire des végétaux fossiles, ou, Recherches botaniques et géologiques sur les végétaux renfermés dans les diverses couches du globe 2 vols
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and colleagues extended the system of Chase and Reveal to provide a revised classification of gymnosperms in 2011, based on the above four
2384:Морфология и систематика архегониальных растений: лекции профессора И.Н. Горожанкина 1894-95 г. Gymnaspermae (Archispermae). Часть III 917:
The term Pinophyta has also been used to include all conifers, extinct and extant, with Pinales representing all the extant conifers.
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In a later revision, in collaboration with two other taxonomists (1966), Cronquist merged all the gymnosperms into a single division,
403:(1825) first discerned that there were two groups of seed plants, distinguished by the form of seed development, based on whether the 3439: 2638: 2235: 3478: 3138: 3034: 2440: 2357: 2332: 2307: 2279: 2198: 2136: 2104: 3483: 2529: 1480:
Christenhusz and colleagues (2011) included only one family in Pinales, Pinaceae, a practice subsequently followed by the
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An Atlas of the World's Conifers: An Analysis of their Distribution, Biogeography, Diversity and Conservation Status
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De La Torre, Amanda R.; Piot, Anthony; Liu, Bobin; Wilhite, Benjamin; Weiss, Matthew; Porth, Ilga (January 2020).
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The Vegetable Kingdom: or, The structure, classification, and uses of plants, illustrated upon the natural system
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were included in Podocarpaceae. Yews (Taxaceae) have sometimes been treated as a separate order (Taxales).
833: 511: 146: 638:, De-Zhi and colleagues (2004) once again proposed a division of 12 gymnosperm families into two classes; 3377: 3207: 2760: 17: 3093:
Tomlinson, P. B. (2012). "Rescuing Robert Brown—The Origins of Angio-Ovuly in Seed Cones of Conifers".
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of the Spermatophytes. Alternative names and the approximate number of genera and species in each are;
829: 671: 3558: 2502: 2480: 3630: 3519: 3217: 3075: 522:. In his classification, Gymnospermae (alternatively named Archespermae) was a class of the division 451:(1880) included only three orders in the class Gymnospermeae, by including taxads within Coniferae; 63: 3306: 890:(41%), and another 27% are oligotypic (2–5 species). The four subclasses have also been treated as 793: 687: 3274: 2415: 2212: 2173: 2396: 801: 635: 2219: 2156: 3545: 3330: 2855:"Functional and morphological evolution in gymnosperms: A portrait of implicated gene families" 2588: 3658: 3540: 3532: 2756: 474: 3581: 3470: 2810: 2240:(in German). Vol. 21 parts (Abteilungen) (2nd ed.). Berlin: Duncker & Humblot. 3413: 2945: 2719: 2596: 2152: 2092:
Brief History of the Gymnosperms: Classification, Biodiversity, Phytogeography and Ecology
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Lu, Ying; Ran, Jin-Hua; Guo, Dong-Mei; Yang, Zu-Yu; Wang, Xiao-Quan (15 September 2014).
515: 3184:"Tracheophyte Phylogeny Poster - Vascular Plants: Systematics and Characteristics, 2019" 2949: 927:. In this scheme, the Pinidae comprise three orders, including Pinales, and 6 families; 407:
were exposed, receiving pollen directly, or enclosed, which do not. Shortly afterwards,
3612: 3163: 3120: 3112: 3059: 3015: 2986: 2968: 2932:"Phylogeny and Divergence Times of Gymnosperms Inferred from Single-Copy Nuclear Genes" 2931: 2881: 2854: 2841: 2785: 2736: 2702: 2613: 2575: 879: 789: 448: 296: 268: 58: 3527: 3400: 3368: 3183: 3155: 3133: 3051: 3020: 2973: 2886: 2819: 2652: 2633: 2558: 2451: 2436: 2353: 2328: 2303: 2275: 2231: 2194: 2132: 2100: 1506: 891: 202: 3063: 2740: 2706: 2617: 3653: 3617: 3405: 3203: 3167: 3147: 3124: 3104: 3095: 3043: 3010: 3000: 2963: 2953: 2916: 2876: 2866: 2833: 2806: 2777: 2748: 2728: 2714: 2694: 2647: 2605: 2567: 2543: 2349: 2324: 797: 764: 555: 50: 3382: 729:). The gymnosperms are represented by four of these subclasses, placing them in a 2958: 2768: 2752: 2426: 2382: 2343: 2318: 2293: 2269: 2184: 2126: 2115: 2090: 1519: 1474: 1456: 1167: 970: 828:
group. In 2018, the Gymnosperm Phylogeny Group was established, analogous to the
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Ran, Jin-Hua; Shen, Ting-Ting; Wang, Ming-Ming; Wang, Xiao-Quan (27 June 2018).
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Genera plantarum ad exemplaria imprimis in herbariis kewensibus servata definita
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Conifers Around the World: Conifers of the Temperate Zones and Adjacent Regions
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refers to the inclusion of Taxodiaceae. These relationships are shown in this
3647: 2249: 1560: 1441: 1427: 1392: 1385: 1214: 1199: 955: 951: 860: 856: 739: 523: 486: 466: 316:. The distinguishing characteristic is the reproductive structure known as a 228: 183: 168: 2164:(in Latin). Vol. 3(1) 1880. London: L Reeve & Co. pp. 417–447. 867:. Among the seed plants, the gymnosperms are a sister group to the subclass 3571: 3159: 3071: 3055: 3024: 3005: 2977: 2907: 2898: 2890: 2824: 2669: 2665: 2625: 2497: 2411: 2392: 2289: 2227: 2208: 2190: 1397: 1264: 1186: 974: 944: 920: 825: 821: 805: 748: 730: 600: 423: 419: 329: 317: 310: 210: 2089:
Anderson, John Malcolm; Anderson, Heidi M.; Cleal, Christopher J. (2007).
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Subsequent revisions merged the Taxodiaceae and Cupressaceae, and placed
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Gernandt, David S; Willyard, Ann; Syring, John V; Liston, Aaron (2011).
3080: 2845: 2789: 2732: 2634:"A phylogenetic classification of the land plants to accompany APG III" 2609: 2579: 1067: 904: 813: 785: 704: 679: 605: 496: 384: 372:
conifers, however, belong to other distinct orders within the division
154: 101: 2871: 2820:"A New Scheme of Classification of Living Gymnosperms at Family Level" 2425:
Plomion, Christophe; Bousquet, Jean; Kole, Chittaranjan, eds. (2011).
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published his treatise on Gymnosperms (1895), for which he bears the
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to describe the former group. The distinction was then formalized by
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In the latter, Cronquist divided Gymnospermae into two divisions;
504: 365: 246: 219: 3431: 848:(seed bearing plants). In turn, the seed plants together with the 477:
approach, including extinct taxa, with the following six classes;
2403: 2178:. Vol. i 1828. Paris: G. Dufour et Ed. d'Ocagne. p. 22. 1874: 1872: 1409: 1405: 1106: 933: 875: 357: 341: 313: 131: 121: 111: 2678:"A new classification and linear sequence of extant gymnosperms" 2420:(3rd. ed.). London: Bradbury & Evans. pp. 221–234. 1611: 1609: 1607: 1605: 1603: 1601: 1599: 1597: 3301: 3190:(DCPS), Institute of Biology – Botany, Freie Universität Berlin 1761: 1759: 1435: 883: 768: 404: 369: 361: 325: 1869: 1830: 1828: 1732: 3553: 3418: 2797:
Davy, J. Burtt (1937). "The classification of Coniferae II".
1594: 1431: 1423: 812:, and within extinct taxa, and particularly the placement of 700: 674:
came a major realignment of the linear classification of the
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produced by all Pinales. All of the extant conifers, such as
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relationship to angiosperms. Subclasses (number of orders);
2374:(in Russian). Moscow: Русская" типо-литография. p. 88. 1825: 1710: 1708: 1706: 1704: 1677: 1450: 1419: 852: 809: 792:
has been considered controversial, and lacks consensus. As
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when used as a class of ferns, synonymous with Equisetidae
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Cole, Theodor C. H.; Bachelier, Julien B (October 2019).
2852: 1987: 1638: 1628: 1626: 1624: 1584: 1582: 994:, although no formal taxonomic revision was undertaken. 337: 1908: 1701: 620:, with three subdivisions reflecting the main lineages; 2992:
Proceedings of the Royal Society B: Biological Sciences
2717:(October 1960). "The divisions and classes of plants". 2009: 2007: 2005: 1920: 1884: 1840: 1786: 1534:, or seed plant, indicating visible sexual organs, and 1000:
Phylogeny of Gymnosperms, subclasses, orders, families
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Gymnosperms form a group of four subclasses among the
686:(2009). In this system, the land plants form a class, 2799:
Forestry: An International Journal of Forest Research
2088: 2019: 1813: 1803: 1801: 1776: 1774: 422:(1830), dividing what he referred to as the subclass 379:
Multiple molecular studies indicate this order being
3076:"Newly required suprageneric names in Magnoliophyta" 2063: 2061: 2002: 1665: 2031: 1944: 1720: 1689: 3248: 2660: 2147: 1963: 1932: 1896: 1878: 1857: 1798: 1771: 1744: 1738: 1615: 1509:because they are not based on an underlying genus 2058: 863:(embryophytes or land plants), as opposed to the 3645: 2554: 2046: 1765: 1653: 469:(1903) Gymnospermae is listed as a subdivision ( 3249:Nagalingum, Nathalie; Burleigh, Gordon (2018). 2984: 2818:De-Zhi, Fu; Yong, Yang; Guang-Hua, Zhu (2004). 2747: 2168: 1834: 1683: 816:. The latter have been variously classified as 2449: 1647: 836:, with the intention of reaching a consensus. 711:, also known as embryophytes or Embryophyceae 3181: 2496:Eckenwalder, James Emory (3 September 2008). 2459: 2424: 2186:Conifers of the World: The Complete Reference 2097:South African National Biodiversity Institute 1996: 3279:Flora of North America. Volume 1, Chapter 13 2817: 2320:A Handbook of the World's Conifers (2 Vols.) 2295:World Checklist and Bibliography of Conifers 2267: 2259: 2226: 1914: 1714: 259:approximate number of species in parentheses 27:Order of seed plants, also known as conifers 3272: 3134:"Evolution and biogeography of gymnosperms" 3132:Wang, Xiao-Quan; Ran, Jin-Hua (June 2014). 2624: 2586: 2495: 2428:Genetics, Genomics and Breeding of Conifers 2366: 2341: 2182: 1926: 1890: 1851: 1792: 1632: 1588: 2473: 2380: 1981: 40: 3092: 3031: 3014: 3004: 2967: 2957: 2920: 2880: 2870: 2713: 2651: 2587:Chamberlain, Charles Joseph (June 1935). 2025: 1819: 1671: 593:, divided gymnosperms into four classes; 2929: 2639:Botanical Journal of the Linnean Society 2013: 3202: 3131: 2903:"The Kew Review: Conifers of the World" 2811:10.1093/oxfordjournals.forestry.a062697 2410: 2391: 2040: 1957: 1726: 1695: 874:There are about 1000 extant gymnosperm 796:transformed from being based solely on 597:Conopsida (conifers, including Pinales) 14: 3646: 3070: 2897: 2316: 2288: 2268:Debreczy, Zsolt; Rácz, István (2011). 2248: 2207: 2124: 2113: 1969: 1938: 1902: 1863: 1807: 1780: 1750: 1559:This term should not be confused with 725:as subclasses, including Magnoliidae ( 589:Benson,(1957) who introduced the term 3329: 3328: 3239: 3139:Molecular Phylogenetics and Evolution 3035:Molecular Phylogenetics and Evolution 2524: 2067: 1659: 913:Pinidae (Pinophyta, conifers 70, 600) 859:(vascular plants), part of the class 3605:a3d0c65a-3cd5-46c3-b83d-8ee3c5e2b42e 3559:4FBA2131-12D7-0484-5984-AB14415A4B88 3520:633c0581-e6f3-456a-8024-3e3d4586c0e1 2796: 2761:"On the Higher Taxa of Embryobionta" 2381:Горожанкин, Иван Николаевич (1895). 2218:(in German) (3rd ed.). Berlin: 2052: 1501:Taxon names beginning with the root 871:(angiosperms or flowering plants). 24: 25: 3670: 3294: 2467: 653:subclass Multinervidae (6 orders) 3312: 3300: 3251:"The Gymnosperm Phylogeny Group" 2653:10.1111/j.1095-8339.2009.01002.x 2244:Die Natürlichen Pflanzenfamilien 780: 62: 3188:Dahlem Centre of Plant Sciences 2672:; Gardner, M. F.; Mill, R. R.; 2474:Delevoryas, T (13 April 2015). 2076: 1553: 1541: 3240:Earle, Christopher J. (2020). 2342:—; Filer, Denis (2013). 2183:Eckenwalder, James E. (2009). 2155:(1862–1883). "Gymnospermeae". 1879:Nagalingum & Burleigh 2018 1739:Bentham & Hooker 1862–1883 1525: 1512: 1495: 1379: 13: 1: 3273:Eckenwalder, James E (2007). 2368:Gorozhankin, Ivan Nikolaevich 1573: 901:(Cycadophyta, cycads 10, 300) 3259:Botanical Society of America 3234:Angiosperm Phylogeny Website 2959:10.1371/journal.pone.0107679 2387:. "Русская" типо-литография. 2125:Benson, Lyman David (1979). 2114:Benson, Lyman David (1957). 1482:Angiosperm Phylogeny Website 1251:(including Cephalotaxaceae) 910:Gnetidae (Gnetophyta 3, 100) 878:, distributed over about 12 839: 834:Pteridophyte Phylogeny Group 670:With the development of the 233:yellow-wood family (170-200) 7: 3275:"Gymnosperm classification" 3175: 3152:10.1016/j.ympev.2014.02.005 3048:10.1016/j.ympev.2018.08.011 2518: 886:. Many of these genera are 526:, divided into subclasses; 390: 10: 3675: 2535:The Philosophical Magazine 2300:Royal Botanic Gardens, Kew 2260:Engler & Prantl (1924) 1522:in 1825, published in 1826 907:(Ginkgophyta, ginkgo 1, 1) 830:Angiosperm Phylogeny Group 672:Angiosperm Phylogeny Group 395: 368:, are included here. Some 3337: 3242:"The Gymnosperm Database" 3218:Missouri Botanical Garden 3109:10.1007/s12229-012-9104-5 2922:10.1007/s12225-018-9738-5 2859:Evolutionary Applications 2699:10.11646/phytotaxa.19.1.3 2548:10.1080/14786442608674070 2211:(1903) . "Gymnospermae". 1997:Cole & Bachelier 2019 1550:indicating enclosed seeds 1262: 1246: 1239: 1212: 1197: 1190: 1180: 1165: 1158: 1134: 1127: 1120: 1104: 1097: 1090: 1065: 1058: 1051: 1044: 1037: 1021: 1014: 274: 267: 180: 175: 160: 153: 59:Scientific classification 57: 48: 39: 34: 2453:The Conifers (Pinophyta) 2402:(1st ed.). London: 2395:(1830). "General view". 2131:(2nd) ed.). Heath. 2082: 1915:Debreczy & Rácz 2011 1538:indicating exposed seeds 1488: 794:taxonomic classification 473:) and adopted more of a 242:umbrella-pine family (1) 215:cypress family (130-140) 2503:Encyclopædia Britannica 2481:Encyclopædia Britannica 2222:Verlag. pp. 70–75. 1927:Farjon & Filer 2013 1852:Chase & Reveal 2009 1616:Christenhusz et al 2011 855:subclasses make up the 802:molecular phylogenetics 636:molecular phylogenetics 565:division Coniferophyta 547:Coniferales (Coniferae) 411:(1828) coined the term 3006:10.1098/rspb.2018.1012 2662:Christenhusz, M. J. M. 2414:(1853) . "Gymnogens". 717:. Class Equisetopsida 3541:Paleobiology Database 1766:Brunsfield et al 1994 746:subclass Ginkgooidae 580:division Cycadophyta 224:pine family (220-250) 188:araucaria family (41) 3309:at Wikimedia Commons 2720:The Botanical Review 2676:(18 February 2011). 2597:The Botanical Review 2460:Plomion et al (2011) 2262:vol. 13 Gymnospermae 2220:Gebrüder Borntraeger 2128:Plant Classification 2117:Plant Classification 1835:Cronquist et al 1966 1684:Brongniart 1828–1837 820:to all gymnosperms, 510:During this period, 434:Cycadeaceae (cycads) 309:, comprises all the 2950:2014PLoSO...9j7679L 2256:. pp. 121–403. 2170:Brongniart, Adolphe 1958:Wang & Ran 2014 1648:Gernandt et al 2011 788:(Acrogymnospermae) 737:subclass Cycadidae 721:is divided into 14 516:botanical authority 437:Pinaceae (conifers) 2999:(1881): 20181012. 2757:Zimmermann, Walter 2733:10.1007/BF02940572 2610:10.1007/BF02872502 755:subclass Gnetidae 449:Bentham and Hooker 251:yew family (30-50) 3641: 3640: 3528:Open Tree of Life 3331:Taxon identifiers 3305:Media related to 2901:(26 March 2018). 2872:10.1111/eva.12839 2749:Cronquist, Arthur 2715:Cronquist, Arthur 2589:"The Gymnosperms" 2559:Systematic Botany 2442:978-1-57808-719-8 2359:978-90-04-21181-0 2334:978-90-04-17718-5 2309:978-1-84246-025-2 2281:978-963-219-061-7 2200:978-0-88192-974-4 2138:978-0-669-01489-1 2106:978-1-919976-39-6 1715:De-Zhi et al 2004 1373: 1372: 1368: 1367: 1359: 1358: 1350: 1349: 1341: 1340: 1332: 1331: 1323: 1322: 1314: 1313: 1305: 1304: 1296: 1295: 1287: 1286: 1278: 1277: 1225: 1224: 1147: 1146: 1079: 1078: 968: 949: 936: 775: 762:subclass Pinidae 758: 751: 742: 656:subclass Taxidae 521: 440:Taxaceae (taxads) 286: 285: 262: 252: 243: 234: 225: 216: 207: 203:Cheirolepidiaceae 198: 189: 149: 16:(Redirected from 3666: 3634: 3633: 3621: 3620: 3608: 3607: 3595: 3594: 3585: 3584: 3575: 3574: 3562: 3561: 3549: 3548: 3536: 3535: 3523: 3522: 3513: 3512: 3500: 3499: 3497:NHMSYS0020826129 3487: 3486: 3474: 3473: 3461: 3460: 3448: 3447: 3435: 3434: 3422: 3421: 3409: 3408: 3396: 3395: 3386: 3385: 3373: 3372: 3371: 3358: 3357: 3356: 3326: 3325: 3317:Data related to 3316: 3304: 3289: 3287: 3285: 3269: 3267: 3265: 3245: 3228: 3226: 3224: 3199: 3197: 3195: 3171: 3128: 3096:Botanical Review 3089: 3067: 3028: 3018: 3008: 2981: 2971: 2961: 2926: 2924: 2894: 2884: 2874: 2849: 2814: 2793: 2765: 2753:Takhtajan, Armen 2744: 2710: 2682: 2657: 2655: 2621: 2593: 2583: 2551: 2542:(337): 352–361. 2514: 2512: 2510: 2492: 2490: 2488: 2457: 2456:. pp. 1–39. 2446: 2421: 2407: 2388: 2375: 2363: 2338: 2317:— (2010). 2313: 2298:(2nd ed.). 2285: 2257: 2254:Klasse Coniferae 2241: 2234:, eds. (1924) . 2223: 2204: 2179: 2165: 2163: 2142: 2121: 2110: 2071: 2065: 2056: 2050: 2044: 2038: 2029: 2023: 2017: 2011: 2000: 1994: 1985: 1979: 1973: 1967: 1961: 1955: 1942: 1936: 1930: 1924: 1918: 1912: 1906: 1900: 1894: 1891:Eckenwalder 2009 1888: 1882: 1876: 1867: 1861: 1855: 1849: 1838: 1832: 1823: 1817: 1811: 1805: 1796: 1793:Chamberlain 1935 1790: 1784: 1778: 1769: 1763: 1754: 1748: 1742: 1736: 1730: 1724: 1718: 1712: 1699: 1693: 1687: 1681: 1675: 1669: 1663: 1657: 1651: 1645: 1636: 1633:Eckenwalder 2008 1630: 1619: 1613: 1592: 1589:Gorozhankin 1904 1586: 1567: 1557: 1551: 1545: 1539: 1529: 1523: 1518:Read before the 1516: 1510: 1499: 1242: 1241: 1193: 1192: 1183: 1182: 1161: 1160: 1130: 1129: 1123: 1122: 1100: 1099: 1093: 1092: 1061: 1060: 1054: 1053: 1047: 1046: 1040: 1039: 1017: 1016: 1007: 1006: 997: 996: 964: 947: 932: 798:plant morphology 763: 756: 747: 738: 519: 383:with respect to 256: 250: 241: 232: 223: 214: 205: 196: 187: 145: 67: 66: 51:Pinus sylvestris 44: 32: 31: 21: 3674: 3673: 3669: 3668: 3667: 3665: 3664: 3663: 3644: 3643: 3642: 3637: 3629: 3624: 3616: 3611: 3603: 3598: 3590: 3588: 3580: 3578: 3570: 3565: 3557: 3552: 3544: 3539: 3531: 3526: 3518: 3516: 3508: 3503: 3495: 3490: 3482: 3477: 3469: 3464: 3456: 3451: 3443: 3438: 3430: 3425: 3417: 3412: 3404: 3399: 3391: 3389: 3381: 3376: 3367: 3366: 3361: 3352: 3351: 3346: 3333: 3297: 3292: 3283: 3281: 3263: 3261: 3222: 3220: 3193: 3191: 3178: 3074:(August 1995). 3072:Reveal, James L 2838:10.2307/4111081 2782:10.2307/1217531 2763: 2680: 2630:Reveal, James L 2591: 2572:10.2307/2419600 2528:(31 May 1826). 2521: 2508: 2506: 2486: 2484: 2470: 2443: 2406:. p. xxxv. 2360: 2335: 2310: 2282: 2274:. DendroPress. 2201: 2161: 2139: 2107: 2085: 2079: 2074: 2066: 2059: 2051: 2047: 2039: 2032: 2024: 2020: 2012: 2003: 1995: 1988: 1982:Delevoryas 2015 1980: 1976: 1968: 1964: 1956: 1945: 1937: 1933: 1925: 1921: 1913: 1909: 1901: 1897: 1889: 1885: 1877: 1870: 1862: 1858: 1850: 1841: 1833: 1826: 1818: 1814: 1806: 1799: 1791: 1787: 1779: 1772: 1764: 1757: 1749: 1745: 1737: 1733: 1725: 1721: 1713: 1702: 1694: 1690: 1682: 1678: 1670: 1666: 1658: 1654: 1646: 1639: 1631: 1622: 1614: 1595: 1587: 1580: 1576: 1571: 1570: 1558: 1554: 1546: 1542: 1530: 1526: 1520:Linnean Society 1517: 1513: 1505:are considered 1500: 1496: 1491: 1475:Phyllocladaceae 1457:Cephalotaxaceae 1382: 1369: 1360: 1351: 1342: 1333: 1324: 1315: 1306: 1297: 1288: 1279: 1226: 1168:Sciadopityaceae 1148: 1080: 971:Sciadopityaceae 842: 783: 645:order Cycadales 573:Chlamydospermae 568:class Coniferae 398: 393: 238:Sciadopityaceae 171: 166: 144: 61: 28: 23: 22: 15: 12: 11: 5: 3672: 3662: 3661: 3656: 3639: 3638: 3636: 3635: 3622: 3618:wfo-9000000404 3609: 3596: 3586: 3576: 3563: 3550: 3537: 3524: 3514: 3501: 3488: 3475: 3462: 3449: 3436: 3423: 3410: 3397: 3387: 3374: 3359: 3343: 3341: 3335: 3334: 3323: 3322: 3321:at Wikispecies 3310: 3296: 3295:External links 3293: 3291: 3290: 3270: 3246: 3237: 3200: 3177: 3174: 3173: 3172: 3129: 3103:(4): 310–334. 3090: 3068: 3029: 2982: 2944:(9): e107679. 2927: 2895: 2865:(1): 210–227. 2850: 2815: 2805:(2): 122–123. 2794: 2776:(4): 129–134. 2745: 2727:(4): 425–482. 2711: 2658: 2646:(2): 122–127. 2622: 2604:(6): 183–209. 2584: 2552: 2520: 2517: 2516: 2515: 2493: 2469: 2468:Encyclopaedias 2466: 2465: 2464: 2463: 2462: 2441: 2422: 2408: 2389: 2364: 2358: 2339: 2333: 2314: 2308: 2286: 2280: 2265: 2264: 2263: 2250:Pilger, Robert 2224: 2205: 2199: 2180: 2166: 2145: 2144: 2143: 2137: 2111: 2105: 2084: 2081: 2080: 2078: 2075: 2073: 2072: 2057: 2045: 2030: 2026:Ran et al 2018 2018: 2001: 1986: 1974: 1962: 1943: 1931: 1919: 1907: 1895: 1883: 1868: 1856: 1839: 1824: 1820:Cronquist 1960 1812: 1797: 1785: 1770: 1755: 1743: 1731: 1719: 1700: 1688: 1676: 1672:Tomlinson 2012 1664: 1652: 1637: 1620: 1593: 1577: 1575: 1572: 1569: 1568: 1552: 1540: 1524: 1511: 1493: 1492: 1490: 1487: 1465: 1464: 1459: 1454: 1444: 1439: 1413: 1395: 1381: 1378: 1371: 1370: 1366: 1365: 1362: 1361: 1357: 1356: 1353: 1352: 1348: 1347: 1344: 1343: 1339: 1338: 1335: 1334: 1330: 1329: 1326: 1325: 1321: 1320: 1317: 1316: 1312: 1311: 1308: 1307: 1303: 1302: 1299: 1298: 1294: 1293: 1290: 1289: 1285: 1284: 1281: 1280: 1276: 1275: 1272: 1271: 1261: 1258: 1257: 1254: 1253: 1245: 1240: 1238: 1232: 1231: 1228: 1227: 1223: 1222: 1219: 1218: 1211: 1208: 1207: 1204: 1203: 1196: 1191: 1189: 1181: 1179: 1176: 1175: 1172: 1171: 1164: 1159: 1157: 1154: 1153: 1150: 1149: 1145: 1144: 1141: 1140: 1133: 1128: 1126: 1121: 1119: 1116: 1115: 1112: 1111: 1103: 1098: 1096: 1091: 1089: 1086: 1085: 1082: 1081: 1077: 1076: 1073: 1072: 1064: 1059: 1057: 1052: 1050: 1045: 1043: 1038: 1036: 1033: 1032: 1029: 1028: 1020: 1015: 1013: 1005: 1002: 1001: 983: 982: 959: 942: 915: 914: 911: 908: 902: 846:spermatophytes 841: 838: 782: 779: 778: 777: 776:(conifers) (1) 760: 753: 744: 688:Equisetopsida 668: 667: 666: 665: 664: 663: 660: 654: 650:Coniferopsida 648: 647: 646: 634:In the era of 632: 631: 628: 625: 614: 613: 608: 603: 598: 587: 586: 585: 584: 578: 577: 576: 569: 552: 551: 548: 540: 539: 533: 508: 507: 502: 499: 494: 489: 484: 471:Unterabteilung 463: 462: 459: 456: 445: 444: 441: 438: 435: 397: 394: 392: 389: 284: 283: 282: 281: 278: 272: 271: 265: 264: 254: 253: 244: 235: 226: 217: 208: 206:(fossils only) 199: 197:(fossils only) 194:Arctopityaceae 190: 178: 177: 173: 172: 167: 158: 157: 151: 150: 139: 135: 134: 129: 125: 124: 119: 115: 114: 109: 105: 104: 99: 92: 91: 86: 79: 78: 73: 69: 68: 55: 54: 46: 45: 37: 36: 26: 9: 6: 4: 3: 2: 3671: 3660: 3657: 3655: 3652: 3651: 3649: 3632: 3627: 3623: 3619: 3614: 3610: 3606: 3601: 3597: 3593: 3587: 3583: 3577: 3573: 3568: 3564: 3560: 3555: 3551: 3547: 3542: 3538: 3534: 3529: 3525: 3521: 3515: 3511: 3506: 3502: 3498: 3493: 3489: 3485: 3480: 3476: 3472: 3467: 3463: 3459: 3454: 3450: 3446: 3441: 3437: 3433: 3428: 3424: 3420: 3415: 3411: 3407: 3402: 3398: 3394: 3388: 3384: 3379: 3375: 3370: 3364: 3360: 3355: 3349: 3345: 3344: 3342: 3340: 3336: 3332: 3327: 3320: 3315: 3311: 3308: 3303: 3299: 3298: 3280: 3276: 3271: 3260: 3256: 3252: 3247: 3243: 3238: 3235: 3232: 3219: 3215: 3214: 3209: 3205: 3204:Stevens, P.F. 3201: 3189: 3185: 3180: 3179: 3169: 3165: 3161: 3157: 3153: 3149: 3145: 3141: 3140: 3135: 3130: 3126: 3122: 3118: 3114: 3110: 3106: 3102: 3098: 3097: 3091: 3087: 3083: 3082: 3077: 3073: 3069: 3065: 3061: 3057: 3053: 3049: 3045: 3041: 3037: 3036: 3030: 3026: 3022: 3017: 3012: 3007: 3002: 2998: 2994: 2993: 2988: 2983: 2979: 2975: 2970: 2965: 2960: 2955: 2951: 2947: 2943: 2939: 2938: 2933: 2928: 2923: 2918: 2914: 2910: 2909: 2904: 2900: 2899:Farjon, Aljos 2896: 2892: 2888: 2883: 2878: 2873: 2868: 2864: 2860: 2856: 2851: 2847: 2843: 2839: 2835: 2831: 2827: 2826: 2821: 2816: 2812: 2808: 2804: 2800: 2795: 2791: 2787: 2783: 2779: 2775: 2771: 2770: 2762: 2758: 2754: 2750: 2746: 2742: 2738: 2734: 2730: 2726: 2722: 2721: 2716: 2712: 2708: 2704: 2700: 2696: 2692: 2688: 2687: 2679: 2675: 2671: 2667: 2666:Reveal, J. L. 2663: 2659: 2654: 2649: 2645: 2641: 2640: 2635: 2631: 2627: 2626:Chase, Mark W 2623: 2619: 2615: 2611: 2607: 2603: 2599: 2598: 2590: 2585: 2581: 2577: 2573: 2569: 2565: 2561: 2560: 2553: 2549: 2545: 2541: 2537: 2536: 2531: 2527: 2526:Brown, Robert 2523: 2522: 2505: 2504: 2499: 2494: 2483: 2482: 2477: 2472: 2471: 2461: 2455: 2454: 2448: 2447: 2444: 2438: 2434: 2430: 2429: 2423: 2419: 2418: 2413: 2412:Lindley, John 2409: 2405: 2401: 2400: 2394: 2393:Lindley, John 2390: 2386: 2385: 2379: 2373: 2369: 2365: 2361: 2355: 2351: 2347: 2346: 2340: 2336: 2330: 2326: 2322: 2321: 2315: 2311: 2305: 2301: 2297: 2296: 2291: 2290:Farjon, Aljos 2287: 2283: 2277: 2273: 2272: 2266: 2261: 2255: 2251: 2247: 2246: 2245: 2239: 2238: 2233: 2229: 2228:Engler, Adolf 2225: 2221: 2217: 2216: 2210: 2209:Engler, Adolf 2206: 2202: 2196: 2192: 2188: 2187: 2181: 2177: 2176: 2172:(1828–1837). 2171: 2167: 2160: 2159: 2154: 2150: 2146: 2140: 2134: 2130: 2129: 2123: 2122: 2119: 2118: 2112: 2108: 2102: 2098: 2094: 2093: 2087: 2086: 2069: 2064: 2062: 2054: 2049: 2042: 2037: 2035: 2027: 2022: 2015: 2014:Lu et al 2014 2010: 2008: 2006: 1998: 1993: 1991: 1983: 1978: 1971: 1966: 1959: 1954: 1952: 1950: 1948: 1940: 1935: 1928: 1923: 1916: 1911: 1904: 1899: 1892: 1887: 1880: 1875: 1873: 1865: 1860: 1853: 1848: 1846: 1844: 1836: 1831: 1829: 1821: 1816: 1809: 1804: 1802: 1794: 1789: 1782: 1777: 1775: 1767: 1762: 1760: 1752: 1747: 1740: 1735: 1728: 1723: 1716: 1711: 1709: 1707: 1705: 1697: 1692: 1685: 1680: 1673: 1668: 1661: 1656: 1649: 1644: 1642: 1634: 1629: 1627: 1625: 1617: 1612: 1610: 1608: 1606: 1604: 1602: 1600: 1598: 1590: 1585: 1583: 1578: 1565: 1564:sensu stricto 1562: 1561:Equisetopsida 1556: 1549: 1544: 1537: 1533: 1528: 1521: 1515: 1508: 1504: 1498: 1494: 1486: 1483: 1478: 1476: 1472: 1471: 1463: 1460: 1458: 1455: 1452: 1448: 1445: 1443: 1442:Podocarpaceae 1440: 1437: 1433: 1429: 1425: 1421: 1417: 1414: 1411: 1407: 1403: 1399: 1396: 1394: 1393:Araucariaceae 1391: 1390: 1389: 1387: 1377: 1364: 1363: 1355: 1354: 1346: 1345: 1337: 1336: 1328: 1327: 1319: 1318: 1310: 1309: 1301: 1300: 1292: 1291: 1283: 1282: 1274: 1273: 1270: 1269: 1266: 1260: 1259: 1256: 1255: 1252: 1250: 1244: 1243: 1237: 1234: 1233: 1230: 1229: 1221: 1220: 1217: 1216: 1215:Podocarpaceae 1210: 1209: 1206: 1205: 1202: 1201: 1200:Araucariaceae 1195: 1194: 1188: 1185: 1184: 1178: 1177: 1174: 1173: 1170: 1169: 1163: 1162: 1156: 1155: 1152: 1151: 1143: 1142: 1139: 1138: 1132: 1131: 1125: 1124: 1118: 1117: 1114: 1113: 1110: 1108: 1102: 1101: 1095: 1094: 1088: 1087: 1084: 1083: 1075: 1074: 1071: 1070:(Ginkgoales) 1069: 1063: 1062: 1056: 1055: 1049: 1048: 1042: 1041: 1035: 1034: 1031: 1030: 1027: 1025: 1019: 1018: 1012: 1009: 1008: 1004: 1003: 999: 998: 995: 993: 989: 980: 976: 972: 967: 963: 960: 957: 956:Podocarpaceae 953: 952:Araucariaceae 946: 943: 940: 935: 930: 929: 928: 926: 922: 918: 912: 909: 906: 903: 900: 897: 896: 895: 893: 889: 885: 881: 877: 872: 870: 866: 862: 861:Equisetopsida 858: 857:tracheophytes 854: 851: 847: 837: 835: 831: 827: 823: 819: 815: 811: 807: 803: 799: 795: 791: 787: 781:Controversies 774: 770: 766: 761: 754: 750: 745: 741: 736: 735: 734: 732: 728: 724: 720: 716: 715: 710: 706: 703: 702: 697: 695: 691: 685: 681: 677: 673: 662:order Pinales 661: 659:order Taxales 658: 657: 655: 652: 651: 649: 644: 643: 641: 640: 639: 637: 629: 626: 623: 622: 621: 619: 612: 609: 607: 604: 602: 599: 596: 595: 594: 592: 583:class Cycadae 582: 581: 579: 574: 570: 567: 566: 564: 563: 562: 559: 557: 549: 546: 545: 544: 537: 534: 532: 529: 528: 527: 525: 524:Archegoniatae 518:for Pinales, 517: 513: 506: 503: 500: 498: 495: 493: 490: 488: 487:Bennettitales 485: 483: 480: 479: 478: 476: 472: 468: 467:Engler system 460: 457: 454: 453: 452: 450: 447:In contrast, 442: 439: 436: 433: 432: 431: 429: 425: 421: 417: 415: 410: 406: 402: 388: 386: 382: 377: 375: 371: 367: 363: 359: 355: 351: 347: 343: 339: 335: 331: 327: 323: 319: 315: 312: 308: 305: 301: 298: 294: 291: 280:Coniferophyta 279: 276: 275: 273: 270: 266: 263: 260: 248: 245: 239: 236: 230: 229:Podocarpaceae 227: 221: 218: 212: 209: 204: 200: 195: 191: 185: 184:Araucariaceae 182: 181: 179: 174: 170: 165: 164: 159: 156: 152: 148: 143: 140: 137: 136: 133: 130: 127: 126: 123: 120: 117: 116: 113: 110: 107: 106: 103: 100: 97: 94: 93: 90: 89:Tracheophytes 87: 84: 81: 80: 77: 74: 71: 70: 65: 60: 56: 53: 52: 47: 43: 38: 33: 30: 19: 3659:Plant orders 3338: 3282:. Retrieved 3278: 3262:. Retrieved 3257:(Abstract). 3254: 3230: 3221:. Retrieved 3213:AP Web v. 14 3211: 3192:. Retrieved 3143: 3137: 3100: 3094: 3085: 3079: 3039: 3033: 2996: 2990: 2941: 2935: 2912: 2908:Kew Bulletin 2906: 2862: 2858: 2829: 2825:Kew Bulletin 2823: 2802: 2798: 2773: 2767: 2724: 2718: 2693:(1): 55–70. 2690: 2684: 2674:Chase, M. W. 2643: 2637: 2601: 2595: 2563: 2557: 2539: 2533: 2507:. Retrieved 2501: 2485:. Retrieved 2479: 2476:"Gymnosperm" 2452: 2427: 2416: 2397: 2383: 2377: 2371: 2344: 2319: 2294: 2270: 2253: 2236: 2232:Prantl, Karl 2213: 2191:Timber Press 2185: 2174: 2157: 2153:Hooker, J.D. 2127: 2116: 2091: 2077:Bibliography 2048: 2041:Stevens 2020 2021: 1977: 1965: 1934: 1922: 1910: 1898: 1886: 1859: 1815: 1788: 1746: 1734: 1727:Lindley 1853 1722: 1696:Lindley 1830 1691: 1679: 1667: 1655: 1563: 1555: 1547: 1543: 1535: 1531: 1527: 1514: 1507:illegitimate 1502: 1497: 1479: 1468: 1466: 1398:Cupressaceae 1383: 1374: 1267: 1265:Cupressaceae 1263: 1247: 1213: 1198: 1187:Araucariales 1166: 1135: 1105: 1066: 1026:(Cycadales) 1022: 1010: 987: 984: 975:Cupressaceae 945:Araucariales 921:Christenhusz 919: 916: 873: 843: 826:monophyletic 822:sister group 784: 731:sister group 718: 712: 699: 693: 689: 669: 642:Cycadopsida 633: 617: 615: 601:Ephedropsida 590: 588: 560: 553: 541: 509: 470: 464: 446: 424:Dicotyledons 414:Phanérogames 412: 399: 381:paraphyletic 378: 292: 287: 258: 255: 211:Cupressaceae 161: 141: 102:Gymnospermae 95: 82: 49: 29: 3453:iNaturalist 3363:Wikispecies 3255:Botany 2018 3088:(2): 68–76. 3042:: 106–116. 2242:, see also 2149:Bentham, G. 1970:Reveal 1995 1939:Farjon 2018 1903:Farjon 2010 1864:Farjon 2001 1808:Benson 1957 1781:Pilger 1926 1751:Engler 1903 1470:Sciadopitys 1462:Taxodiaceae 1380:Subdivision 1236:Cupressales 1011:Gymnosperms 962:Cupressales 869:Magnoliidae 865:green algae 850:monilophyte 808:to extinct 727:angiosperms 714:nom. illeg. 676:land plants 611:Cycadopsida 538:(Coniferae) 531:Cycadoideae 512:Gorozhankin 492:Cordaitales 416:gymnosperms 330:celery-pine 277:Coniferales 18:Coniferales 3648:Categories 3081:Phytologia 2832:(1): 111. 2670:Farjon, A. 2566:(2): 253. 2068:Earle 2020 1660:Brown 1826 1574:References 1548:Angiosperm 1536:gymnosperm 1532:Phanerogam 1109:(Pinales) 1068:Ginkgoidae 905:Ginkgoidae 814:Gnetophyta 786:Gymnosperm 694:sensu lato 606:Gnetopsida 575:(Gnetales) 497:Ginkgoales 461:Cycadaceae 409:Brongniart 155:Type genus 128:Subclass: 108:Division: 3284:11 August 3208:"Pinales" 3206:(2020) . 3194:24 August 3146:: 24–40. 2686:Phytotaxa 2498:"Conifer" 2433:CRC Press 2292:(2001) . 2162:(3 vols.) 2053:Davy 1937 1402:cypresses 1024:Cycadidae 992:cladogram 925:subclades 899:Cycadidae 892:divisions 888:monotypic 840:Phylogeny 765:Cronquist 723:subclades 624:Cycadicae 618:Pinophyta 556:Cronquist 536:Peucideae 501:Coniferae 482:Cycadales 458:Coniferae 455:Gnetaceae 443:Gnetaceae 426:into two 374:Pinophyta 322:Araucaria 307:Pinopsida 300:Pinophyta 176:Families 122:Pinopsida 112:Pinophyta 72:Kingdom: 3600:VicFlora 3589:VASCAN: 3572:50324779 3567:Tropicos 3390:BioLib: 3354:Q1000370 3348:Wikidata 3231:see also 3176:Websites 3160:24565948 3117:41809857 3064:52110440 3056:30153503 3025:29925623 2978:25222863 2937:PLoS ONE 2891:31892953 2759:(1966). 2741:43144314 2707:86797396 2632:(2009). 2618:39793752 2519:Articles 2487:6 August 2378:see also 2370:(1904). 2252:(1926). 2120:. Heath. 1503:conifer- 1447:Taxaceae 1416:Pinaceae 1249:Taxaceae 1137:Gnetidae 979:Taxaceae 939:Pinaceae 931:Pinales 880:families 790:taxonomy 773:Zimmerm. 630:Gneticae 558:(1960). 550:Gnetales 505:Gnetales 475:splitter 391:Taxonomy 385:Gnetales 314:conifers 297:division 269:Synonyms 247:Taxaceae 220:Pinaceae 35:Pinales 3654:Pinales 3533:4736806 3510:1446380 3369:Pinales 3339:Pinales 3319:Pinales 3307:Pinales 3264:30 July 3223:28 July 3168:4662571 3125:5403322 3016:6030518 2969:4164646 2946:Bibcode 2882:6935586 2846:4111081 2790:1217531 2580:2419600 2509:27 July 2404:Longman 2399:economy 2215:Botanik 1410:redwood 1406:juniper 1107:Pinidae 948:Gorozh. 934:Gorozh. 882:and 83 876:species 627:Pinicae 591:Pinales 520:Gorozh. 465:In the 420:Lindley 396:History 358:redwood 342:juniper 334:cypress 295:in the 293:Pinales 147:Gorozh. 142:Pinales 138:Order: 132:Pinidae 118:Class: 76:Plantae 3631:886455 3582:471121 3579:uBio: 3517:NZOR: 3484:500028 3383:215529 3166:  3158:  3123:  3115:  3062:  3054:  3023:  3013:  2976:  2966:  2889:  2879:  2844:  2788:  2739:  2705:  2616:  2578:  2439:  2356:  2331:  2306:  2278:  2197:  2135:  2103:  1436:spruce 1428:cedars 1386:Pilger 884:genera 806:extant 771:& 769:Takht. 709:Reveal 707:& 684:Reveal 571:class 428:tribes 405:ovules 370:fossil 364:, and 362:spruce 311:extant 3626:WoRMS 3554:Plazi 3546:82140 3471:12236 3466:IRMNG 3458:47375 3432:1CONO 3406:623FD 3164:S2CID 3121:S2CID 3113:JSTOR 3060:S2CID 2915:(1). 2842:JSTOR 2786:JSTOR 2769:Taxon 2764:(PDF) 2737:S2CID 2703:S2CID 2681:(PDF) 2614:S2CID 2592:(PDF) 2576:JSTOR 2458:, in 2350:Brill 2325:Brill 2258:, in 2083:Books 1489:Notes 1432:larch 1424:pines 818:basal 749:Engl. 705:Chase 701:sensu 680:Chase 678:, by 401:Brown 350:larch 346:kauri 326:cedar 304:class 290:order 163:Pinus 96:Clade 83:Clade 3505:NCBI 3479:ITIS 3440:GBIF 3427:EPPO 3419:4064 3393:2307 3378:APNI 3286:2020 3266:2020 3225:2020 3196:2020 3156:PMID 3052:PMID 3021:PMID 2974:PMID 2887:PMID 2511:2020 2489:2020 2437:ISBN 2354:ISBN 2329:ISBN 2304:ISBN 2276:ISBN 2195:ISBN 2133:ISBN 2101:ISBN 1451:yews 1420:firs 1268:s.l. 988:s.l. 966:Link 853:fern 832:and 810:taxa 719:s.l. 690:s.l. 682:and 354:pine 318:cone 288:The 3613:WFO 3492:NBN 3445:640 3414:EoL 3401:CoL 3148:doi 3105:doi 3044:doi 3040:129 3011:PMC 3001:doi 2997:285 2964:PMC 2954:doi 2917:doi 2877:PMC 2867:doi 2834:doi 2807:doi 2778:doi 2729:doi 2695:doi 2648:doi 2644:161 2606:doi 2568:doi 2544:doi 800:to 759:(3) 757:Pax 752:(1) 743:(1) 740:Pax 698:or 366:yew 338:fir 3650:: 3628:: 3615:: 3602:: 3592:52 3569:: 3556:: 3543:: 3530:: 3507:: 3494:: 3481:: 3468:: 3455:: 3442:: 3429:: 3416:: 3403:: 3380:: 3365:: 3350:: 3277:. 3253:. 3216:. 3210:. 3186:. 3162:. 3154:. 3144:75 3142:. 3136:. 3119:. 3111:. 3101:78 3099:. 3086:79 3084:. 3078:. 3058:. 3050:. 3038:. 3019:. 3009:. 2995:. 2989:. 2972:. 2962:. 2952:. 2940:. 2934:. 2913:73 2911:. 2905:. 2885:. 2875:. 2863:13 2861:. 2857:. 2840:. 2830:59 2828:. 2822:. 2803:11 2801:. 2784:. 2774:15 2772:. 2766:. 2755:; 2751:; 2735:. 2725:26 2723:. 2701:. 2691:19 2689:. 2683:. 2668:; 2664:; 2642:. 2636:. 2628:; 2612:. 2600:. 2594:. 2574:. 2564:19 2562:. 2540:67 2538:. 2532:. 2500:. 2478:. 2435:. 2431:. 2376:, 2352:. 2348:. 2327:. 2323:. 2302:. 2230:; 2193:. 2189:. 2151:; 2099:. 2095:. 2060:^ 2033:^ 2004:^ 1989:^ 1946:^ 1871:^ 1842:^ 1827:^ 1800:^ 1773:^ 1758:^ 1703:^ 1640:^ 1623:^ 1596:^ 1581:^ 1434:, 1430:, 1426:, 1422:, 1408:, 1404:, 977:, 973:, 954:, 767:, 376:. 360:, 356:, 352:, 348:, 344:, 340:, 336:, 332:, 328:, 324:, 302:, 249:, 240:, 231:, 222:, 213:, 186:, 169:L. 98:: 85:: 3288:. 3268:. 3244:. 3236:) 3229:( 3227:. 3198:. 3170:. 3150:: 3127:. 3107:: 3066:. 3046:: 3027:. 3003:: 2980:. 2956:: 2948:: 2942:9 2925:. 2919:: 2893:. 2869:: 2848:. 2836:: 2813:. 2809:: 2792:. 2780:: 2743:. 2731:: 2709:. 2697:: 2656:. 2650:: 2620:. 2608:: 2602:1 2582:. 2570:: 2550:. 2546:: 2513:. 2491:. 2445:. 2362:. 2337:. 2312:. 2284:. 2203:. 2141:. 2109:. 2070:. 2055:. 2043:. 2028:. 2016:. 1999:. 1984:. 1972:. 1960:. 1941:. 1929:. 1917:. 1905:. 1893:. 1881:. 1866:. 1854:. 1837:. 1822:. 1810:. 1795:. 1783:. 1768:. 1753:. 1741:. 1729:. 1717:. 1698:. 1686:. 1674:. 1662:. 1650:. 1635:. 1618:. 1591:. 1453:) 1449:( 1438:) 1418:( 1412:) 1400:( 981:) 969:( 958:) 950:( 941:) 937:( 696:) 692:( 261:) 257:( 201:† 192:† 20:)

Index

Coniferales
Pinus sylvestris, scots pine tree
Pinus sylvestris
Scientific classification
Edit this classification
Plantae
Tracheophytes
Gymnospermae
Pinophyta
Pinopsida
Pinidae
Pinales
Gorozh.
Type genus
Pinus
L.
Araucariaceae
Arctopityaceae
Cheirolepidiaceae
Cupressaceae
Pinaceae
Podocarpaceae
Sciadopityaceae
Taxaceae
Synonyms
order
division
Pinophyta
class
Pinopsida

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