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Labrador Retriever

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437: 1184:"Production of yellow versus black pigment in dogs is controlled by three genes: MC1R, Agouti, and CBD103. Dogs carrying wild-type alleles for all three genes have a yellow coat resulting from Agouti antagonism of MC1R signaling in melanocytes (yellow Great Dane, top). Dogs carrying a loss-of-function mutation at MC1R have a yellow coat, regardless of their genotype at Agouti or CBD103 (yellow Labrador retriever, middle). Dogs carrying wild-type alleles for MC1R and Agouti, together with the dominant black allele of CBD103 (KB) have a black coat resulting from the interaction between a β-defensin and MC1R (black Curly Coated Retriever, 52: 583: 778: 604: 429: 338: 504: 570: 869: 460:
24.5 in) for dogs and 55 to 60 centimetres (21.5 to 23.5 in) for bitches, The Kennel Club which advises that dogs should be 56 to 57 centimetres (22 to 22.5 in) with bitches between 55 and 56 centimetres (21.5 and 22 in), and the FCI which quotes a range of 56 to 57 centimetres (22 to 22.5 in) for dogs with bitches ideal at 54 to 56 centimetres (21.5 to 22 in).
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German Shepherds had the highest chance of not completing it. Labrador Retrievers and the Labrador Retriever/Golden Retriever Crossbreeds had the highest success rate. However, German Shepherds and Golden Retrievers had a higher success rate after going through longer training than the training required for Labrador Retrievers.
820:. In 2006 both the United Kingdom and the United States, there are well over twice as many registered Labradors as the next most popular breed. If the comparison is limited to dog breeds of a similar size, then there are around 3–5 times as many Labradors registered in both countries as the next most popular breeds, the 396:, more often black than other colours, long in its head and nose with a deep chest, fine legs, and short and smooth coat, and did not carry its tail as highly as the Newfoundland. Hawker distinguishes the Newfoundland from both the "proper Labrador" and St. John's breed of these dogs in the fifth edition of his book 707:
A UK study found the breed's average life expectancy to be 13.1 years. A joint study by the Royal Veterinary College study and The University of Sydney, concluded that chocolate-coloured Labradors have a shorter average life expectancy than other colours of Labrador (by about 10%) and are more likely
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By 1870 the name Labrador Retriever had become common in England. The liver (now usually called chocolate) Labrador emerged in the late 1800s, with liver-coloured pups documented at the Buccleuch kennels in 1892; the first yellow Labrador on record was born in 1899 (Ben of Hyde, kennels of Major C.J.
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to suffer some health problems. A 2024 Italian study found a life expectancy of 11 years for the breed compared to 10 years overall. A 2005 Swedish study of insurance data found that 25% of Labrador Retrievers died by the age of 10, less than the overall rate of 35% of dogs dying by the age of 10.
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As a result of specialised breeding, there are significant differences between field and trial-bred and show-bred lines of Labradors. In the United States, the former are sometimes mistakenly referred to as "American" and the latter as "English", in fact, both field and show types are bred in both
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at all three loci, the dog will have a yellow coat. If a dog has a loss-of-function mutation at MC1R, it will also have a yellow coat, regardless of the genotypes at the other two loci. Dogs carrying wild-type alleles for MC1R and Agouti, together with the black allele of CBD103, will have a black
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collaborated to develop and establish the Labrador Retriever breed. The dogs Buccleuch Avon and Buccleuch Ned, given by Malmesbury to Buccleuch, were mated with bitches carrying blood from those originally imported by the 5th Duke and the 10th Earl of Home. The offspring are the ancestors of all
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Roccaro, Mariana; Salini, Romolo; Pietra, Marco; Sgorbini, Micaela; Gori, Eleonora; Dondi, Maurizio; Crisi, Paolo E.; Conte, Annamaria; Dalla Villa, Paolo; Podaliri, Michele; Ciaramella, Paolo; Di Palma, Cristina; Passantino, Annamaria; Porciello, Francesco; Gianella, Paola; Guglielmini, Carlo;
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Labrador Retrievers have proven to have a high success rate at becoming guide dogs. A study published in 2006, tested the suitability of four different breeds (Labrador Retriever, Golden Retriever, Labrador Retriever/Golden Retriever Mix, and German Shepherds) as guide dogs. In this experiment,
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to the base of the tail as they are from the floor to the withers. The AKC standard includes an ideal weight for dogs of 25–36 kg (55–80 lb) and for bitches as 25–32 kg (55–70 lb). The guidelines for height vary between the AKC, which gives 55 to 62 centimetres (21.5 to
466:: The Labrador Retriever's coat should be short and dense, but not wiry. The coat is water-resistant, so the dog does not get cold when taking to the water in the winter. That means that the dog naturally has a slightly dry, oily coat. Acceptable colours are black, yellow, and chocolate. 689:. One UK study found a 4.14 odds ratio. Some studies report a predilection for Labrador Retriever bitches. The reason for the predilection is unknown. Ideas proposed include it being hereditary or environmental. The Labrador Retriever also shows a predisposition to other 472:: The head should be broad with slightly pronounced eyebrows. The eyes should be kind and expressive. Appropriate eye colours are brown and hazel. The lining around the eyes should be black. The ears should hang close to the head and be set slightly above the eyes. 634:, who was trained to, if need be, put his human who uses a wheelchair in the recovery position, cover him with a blanket, and activate an emergency phone. A number of Labradors have also been taught to assist their owner in removing money and credit cards from 835:
in the United States, Australia, and many other countries, as well as being widely used by police and other official bodies for their detection and working abilities. Approximately 60–70% of all guide dogs in the United States are Labradors.
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Johnston, Andrea N.; Center, Sharon A.; McDonough, Sean P.; Wakshlag, Joseph J.; Warner, Karen L. (1 February 2013). "Hepatic copper concentrations in Labrador Retrievers with and without chronic hepatitis: 72 cases (1980–2010)".
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or motorised scooter in the final years of his life. Among the services that Sully was able to perform for Bush were retrieving dropped items, opening and closing doors, pushing an emergency button and supporting him when
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The tail and coat are designated "distinctive features" of the Labrador by both the Kennel Club and AKC. The AKC adds that the "true Labrador Retriever temperament is as much a hallmark of the breed as the 'otter' tail."
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The Labrador Retriever is a gun dog bred to retrieve on land and water. As a dog specially bred for water retrieving, the Labrador has developed various traits for this job. For retrieving the Labrador Retriever has a
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in 2007. Following the multimillion-dollar, 6 arrest Malaysian detection, they became the first dogs to be awarded Malaysia's "outstanding service award" and software pirates were stated to have put a £30,000
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Candille, Sophie I.; Kaelin, Christopher B.; Cattanach, Bruce M.; Yu, Bin; Thompson, Darren A.; Nix, Matthew A.; Kerns, Julie A.; Schmutz, Sheila M.; Millhauser, Glenn L.; Barsh, Gregory S. (2007).
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Taylor, SM; Shmon, CL; Shelton, GD; Patterson, EE; Minor, K; Mickelson, JR (2008). "Exercise Induced Collapse of Labrador Retrievers: Survey results and preliminary investigation of heritability".
924:. By Endal's death in March 2009, he and his owner/handler Allen Parton had been filmed almost 350 times by crews from several countries, and a film of a year in Endal's life was in production. 747: 627:, a trait that allows it to carry game and waterfowl without damaging it. And for swimming, the Labrador is aided by fully webbed paws, an otter-like tail, and a waterproof coat. 1056:
for conspicuous gallantry or devotion to duty while serving in military conflict. They led their blind owners down more than 70 flights of stairs to escape from the damaged
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bred (bench-bred) lines of this breed in the United States and are based on the American Kennel Club standard. Significant differences between UK and US standards are noted.
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The Labrador is an exceptionally popular dog. As of 2006, it was widely considered the most popular breed in the world, and it is the most popular dog by ownership in
478:: The jaws should be strong and powerful. The muzzle should be of medium length and should not be too tapered. The jaws should hang slightly and curve gracefully back. 2086:
Bandara, Y.; Bayton, W. A.; Williams, T. L.; Scase, T.; Bexfield, N. H. (2021). "Histopathological frequency of canine hepatobiliary disease in the United Kingdom".
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for conspicuous gallantry or devotion to duty while serving in military conflict. She detected explosive devices, which were subsequently disarmed, while serving in
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In 2022 Labrador Retrievers were the second most popular breed in the United States. In 2020 the Labrador was the most popular registered dog breed in New Zealand.
4568: 2722: 1780: 1078: 3063: 735:. They can suffer from exercise induced collapse, which causes hyperthermia, weakness, collapse, and disorientation after short bouts of exercise, or from 321: 298: 1640: 1167:'s labradors Charlie and Bear. They were represented in numerous magazines with the singer and her family, becoming pets celebrities in their own names. 305:, imported progenitors of the breed from Newfoundland to Europe for use as gun dogs. Another early advocate of these Newfoundland fishing dogs was the 590:
appearance: field (left), and show (right), showing the shorter muzzle length, more solid appearance of the head, and "pronounced" stop of the latter
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and 600 detonators. His biggest contribution to the police force and the city was the detection of 3,329 kg of RDX. He also helped detect
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Ennik, Irma; Liinamo, Anna-Elisa; Leighton, Eldin; Van Arendonk, Johan (2006). "Suitability for field service in 4 breeds of guide dogs".
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https://www.seattletimes.com/nation-world/mission-complete-sully-bushs-service-dog-stays-at-former-presidents-side-for-one-last-journey/
4583: 3008: 360:. These were then bred with British hunting dogs to create what became known as the Labrador Retriever. Its early patrons included the 201: 2121:
Hoffmann, G.; van den Ingh, T.S.G.A.M.; Bode, P.; Rothuizen, J. (2006). "Copper‐Associated Chronic Hepatitis in Labrador Retrievers".
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Oliver, James A.C.; Mellersh, Cathryn S. (2020). "Genetics". In Cooper, Barbara; Mullineaux, Elizabeth; Turner, Lynn (eds.).
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region of that colony. It is among the most commonly kept dogs in several countries, particularly in the European world.
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Conant, E.K.; Juras, R.; Cothran, E.G. (2011). "Incidence of the mask phenotype M264V mutation in Labrador retrievers".
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The Labrador is friendly, energetic, and playful. It was bred as a sporting and hunting dog but is widely kept as a
3145: 3113: 2968: 995:, twin Black Labrador counterfeit-detection dogs who became famous in 2007 for "sniffing out nearly 2 million 630:
The high intelligence, initiative and self-direction of Labradors in working roles is exemplified by dogs such as
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The Labrador breed dates back to at least the 1830s, when St. John's water dogs bred by European settlers in
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The most common places where pigmentation is visible are the nose, lips, gums, and the rims of the eyes.
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during the last six months of his life; noted for his role during the president's funeral. A form of
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There is a great deal of variety among Labradors. The following characteristics are typical of the
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Labrador Retrievers: Everything about purchase, care, nutrition, diseases, breeding, and behavior
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The Labrador Retriever Club, Inc. (1995). "The Labrador Dog". In Ziessow, Dr. Bernard W. (ed.).
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This article is about the breed of dog. For the single by Japanese idol girl group AKB48, see
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Please help update this article to reflect recent events or newly available information.
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Alborali, Giovanni L.; Rota Nodari, Sara; Sabatelli, Sonia; Peli, Angelo (2024).
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Article written for The Field, 30 May 1896, 'Labrador Dogs', by John S Kerss
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The three colour varieties, from front to back: Yellow, black, and chocolate
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A Soldier's Best Friend; Scout dogs and their handlers in the Vietnam War
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for Animal Gallantry and Devotion to Duty), the first dog to ride on the
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Instructions to Young Sportsmen in all that relates to Guns and Shooting
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countries and all Labrador Retrievers are descended from British lines.
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The Labrador Retriever is one of the more commonly affected breeds for
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even referring to the Lesser Newfoundland as the Newfoundland. Colonel
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were first introduced to Britain from ships trading between Canada and
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Egenvall, A; Bonnett, Bn; Hedhammar, å; Olson, P (30 September 2005).
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The Domestic Dog: its Evolution, Behavior and Interactions With People
455:: Labradors are a medium-large breed. They should be as long from the 4465: 4400: 3972: 3814: 3698: 3668: 724: 674: 554:
According to a 2011 study, 13 out of 245 Labradors studied were
547: 275: 245: 3436:. North Pomfret, VT: Trafalgar Square Publishing. pp. 299–301. 2400: 2193:"Chocolate Labradors have a shorter lifespan than rest of the breed" 3140: 3108: 2938: 2881: 1216:"Longevity of companion dog breeds: those at risk from early death" 1000: 921: 264: 1851:
Journal of Veterinary Behavior: Clinical Applications and Research
1028:(MIA) in Afghanistan before being recovered safe and well in 2009. 569: 4527: 4523: 4349: 4090: 4012: 4002: 3741: 2120: 1679:"A β-defensin mutation causes black coat colour in domestic dogs" 1103: 979: 736: 666: 456: 248: 3168:"British army dog awarded bravery medal for work in Afghanistan" 2905:"Fresh off Malaysian triumph, DVD-sniffing dogs tackle New York" 1848: 503: 376:. Early writers have confused the Labrador with the much larger 162:
Black, chocolate, or yellow (ranges from pale yellow to fox red)
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dogs; they are used at the Italian School of Canine Lifeguard.
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Bexfield, Nick (2017). "Canine Idiopathic Chronic Hepatitis".
947: 3949: 2253: 2213:"Factors related to longevity and mortality of dogs in Italy" 1074: 1011: 897: 872: 854:'s 2000–2005 appointees to its 'hall of fame' are Labradors. 631: 587: 530:. Coat colour is determined primarily by three genes, called 353: 2961:"Australian dog returns home after a year in the wilderness" 1672: 1670: 2386: 2209: 1618:. Fédération Cynologique Internationale. Accessed May 2020. 1404:
British dogs, their points, selection, and show preparation
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to track down dead or wounded soldiers or enemy positions.
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Gundogs: Their past, their performance and their prospects
2934:"DVD pirates put out hits on Lucky and Flo the crime dogs" 2085: 2058:
Veterinary Clinics of North America: Small Animal Practice
1956:"Assistance dogs are trained as partners for the disabled" 4410: 3036: 2684:
Labrador retriever: Most popular breed of assistance dog.
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describes the first Labrador as being not larger than an
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American Kennel Club, The Labrador Retriever Club, Inc.
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Marley, "The World's Worst Dog", featured in journalist
3136:"Army dog killed in Afghanistan given posthumous medal" 1877:"Why the Labrador Is the World's Most Popular Bird Dog" 1079:
Royal Gloucestershire, Berkshire and Wiltshire Regiment
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Dogs: The ultimate dictionary of over 1,000 dog breeds
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Journal of the American Veterinary Medical Association
1213: 719:, especially in larger dogs. Eye diseases may include 2319:. Canine Inherited Disorders Database. Archived from 1472:. Future Publishing Limited Quay House. 19 April 2022 882:
Notable labradors within various categories include:
484:: The body should have a powerful and muscular build. 3315:. Ramsbury, Marlborough, UK: The Crowood Press Ltd. 2877:"DVD-sniffing dogs awarded medals, returning to NYC" 1953: 1466:"Best labrador colour: does coat make a difference?" 1214:
McMillan, K. M.; Bielby, J.; Williams, C.L. (2024).
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gene is responsible for the condition in the breed.
3336:. Shrewsbury, UK: Swan Hill Press. pp. 28–32. 3334:
The Shooting Man's Dog: A complete guide to gundogs
3240:"Céline Dion Shared a Rare Photo of Her Three Sons" 2659: 1630:, 'The American Kennel Club', Retrieved 29 May 2014 523:(medium to dark brown; originally called "liver"). 3401: 2661: 2558: 2530: 1997:"Italian school teaches dogs to become lifeguards" 1954:Blystone, Richard; Mallary Gelb (10 August 2000). 1821: 1743: 645:. It continues in that role today, along with the 3399: 3357:. New York, NY: Galahad Books. pp. 238–243. 2953: 511:Labradors are registered in three colours: Solid 4540: 2660:Devantier, Alecia T.; Turkington, Carol (2007). 2081: 2079: 1778: 4569:Provincial symbols of Newfoundland and Labrador 3400:Kern, Kerry V.; Earle-Bridges, Michele (1995). 2197:VetCompassTM and the University of Sydney Study 3104:"In pictures: Sadie the hero dog gets a medal" 2987: 2821:"Murió Frida, perrita rescatista de la Marina" 2584: 2582: 2504: 2470: 2464: 1397: 1209: 1207: 251:. It was developed in the United Kingdom from 3928: 3467: 3353:Jones, Arthur F.; Hamilton, Ferelith (1971). 3352: 3160: 2363:"Retinal Dysplasia/Oculoskeletal Dysplasia 1" 2076: 2028:. New York, NY: Sterling Publishing. p.  1077:, in November 2005; she served with the 1034:, an American black Labrador who served as a 974:who detected arms and ammunition used in the 685:Labrador Retrievers have a predisposition to 526:Puppies of all colours can occur in the same 519:(anything from creamy white to fox-red), and 4098: 3294:. New York, NY: DK Publishing. p. 195. 2621: 2619: 2199:. Royal Veterinary College. 22 October 2018. 2051: 2049: 1929:"8 Things To Know About Labrador Retrievers" 1805:: CS1 maint: multiple names: authors list ( 1415: 1413: 404:Radclyffe). The breed was recognised by the 3422:– via Internet Archive (archive.org). 2579: 1406:(3 ed.). Upcot Gill. pp. 356–358. 1393: 1391: 1350:The Official Book of the Labrador Retriever 1204: 948:Police, military, rescue and detection dogs 558:for the M264V mutation responsible for the 154:Short, dense, weather-resistant double coat 3935: 3921: 3474: 3460: 3032:"Jake, the rescue dog: An impressive life" 1569:"Labrador Retriever Dog Breed Information" 50: 2616: 2471:Freytas-tamura, Kimiko De (5 July 2016). 2447: 2289: 2271: 2238: 2228: 2046: 1720: 1410: 1247: 693:diseases, including: reactive hepatitis, 424:The head is broad with a pronounced stop. 2995:"Handler never gave up on lost army dog" 2341:Newton, C. D.; Nunamaker, D. M. (Eds.) " 2055: 1869: 1388: 867: 602: 581: 568: 502: 435: 427: 419: 336: 3310: 3265: 3029: 2380: 2311: 2309: 2123:Journal of Veterinary Internal Medicine 1994: 1895: 1819: 1621: 1490: 1333: 1331: 1312:. Akc.org. 4 April 2012. Archived from 577: 499:Labrador Retriever coat colour genetics 309:, who bred them for their expertise in 14: 4541: 3943:Dogs originating in the United Kingdom 3428: 3331: 2827:(in Spanish). Mexico. 15 November 2022 2653: 2421: 2415: 2023: 2017: 1903:"Your guide to the Labrador Retriever" 1419: 1120:Former President of the United States 1094:, mines, and weapons while serving in 976:1993 Mumbai (Bombay) serial explosions 4226:English Toy Terrier (Black & Tan) 3916: 3455: 3375:"Retriever (Labrador) breed standard" 3286: 3011:from the original on 15 November 2009 2965:defence.gov.au, Defence Media Release 2931: 2874: 2343:Textbook of Small Animal Orthopaedics 1995:Manetti, Francesco (23 August 2010). 1921: 1592: 1590: 1278: 1276: 1274: 1272: 1270: 1268: 202:Fédération Cynologique Internationale 4416:Dumfriesshire Black and Tan Foxhound 2902: 2847:"Police dogs sniff for pirated DVDs" 2720: 2507:BSAVA Textbook of Veterinary Nursing 2306: 1628:"Get to Know the Labrador Retriever" 1328: 1302: 771: 3270:. Shrewsbury, UK: Swan Hill Press. 3268:The Complete Gundog Training Manual 3056: 3005:Australian Broadcasting Corporation 1609: 1597:Retriever (Labrador) Breed Standard 1538: 1494:The NEW Complete Labrador Retriever 1284:"Labrador Retriever Breed Standard" 938:confined the former president to a 916:and the first dog known to work a ' 857: 831:They are the most popular breed of 24: 3732:Nova Scotia Duck Tolling Retriever 2971:. 12 November 2009. Archived from 2757:Australian National Kennel Council 2135:10.1111/j.1939-1676.2006.tb01798.x 1587: 1265: 989:rifles and five 9 mm pistols. 885: 852:Australian National Kennel Council 641:The breed is used in water rescue/ 415: 27:British breed of retriever gun dog 25: 4595: 4584:Dog breeds originating in England 2785:. Edition.cnn.com. 10 August 2000 2627:"AKC Dog Registration Statistics" 2532:"New Zealand: Leading dog breeds" 1974:"thought this was Bark-lays bank" 1310:"AKC Dog Registration Statistics" 999:DVDs" on a 6-month secondment to 748:progressive rod-cone degeneration 3146:British Broadcasting Corporation 3114:British Broadcasting Corporation 3064:Life saving: Animals' war photos 2969:Australian Department of Defence 2088:Journal of Small Animal Practice 776: 711:Labradors are somewhat prone to 412:(AKC) registration was in 1917. 3232: 3207: 3128: 3096: 3023: 2925: 2903:Chan, Sewell (28 August 2007). 2896: 2868: 2839: 2813: 2797: 2775: 2741: 2714: 2689: 2664:Extraordinary Jobs with Animals 2551: 2523: 2498: 2422:Raffan, Eleanor (10 May 2016). 2355: 2335: 2247: 2230:10.1016/j.prevetmed.2024.106155 2203: 2185: 2149: 2114: 1988: 1966: 1947: 1842: 1813: 1772: 1737: 1633: 1575:. The American Kennel Club, Inc 1561: 1532: 1484: 1178: 768:List of most popular dog breeds 398:Instructions to Young Sportsmen 125:54–56 cm (21.5–22 in) 115:56–57 cm (22–22.5 in) 3355:The World Encyclopedia of Dogs 3030:Glagola, Nick (27 July 2007). 2875:Blass, Evan (20 August 2007). 2697:"Most Popular Dog Breeds 2022" 2217:Preventive Veterinary Medicine 1828:. Cambridge University Press. 1746:Research in Veterinary Science 1458: 1432: 1402:. In Drury, William D. (ed.). 1340: 1186:bottom)." — Candille, Kaelin, 932:US President George H. W. Bush 274:. It may also be trained as a 13: 1: 4360:Cavalier King Charles Spaniel 3654:Slovakian Roughhaired Pointer 2932:Blass, Evan (22 March 2007). 2260:Acta Veterinaria Scandinavica 1198: 761: 146:25–32 kg (55–70 lb) 136:29–36 kg (65–80 lb) 2825:Excélsior (excelsior.com.mx) 1491:Warwick, Helen (1986). "5". 1171: 1088:improvised explosive devices 1024:that spent almost 14 months 970:("Chain", or "Shackles"), a 665:Labradors have been used as 7: 4205:West Highland White Terrier 3684:Wirehaired Pointing Griffon 3481: 3292:The Encyclopedia of the Dog 3176:. London, UK. 29 April 2014 2759:. 2000–2005. Archived from 2567:. Statistica Research. 2022 2539:. Statistica Research. 2019 1398:Holland-Hibert, A. (1903). 1346: 1100:Royal Army Veterinary Corps 721:progressive retinal atrophy 263:), and was named after the 10: 4600: 4262:Staffordshire Bull Terrier 3599:Irish Red and White Setter 3584:German Shorthaired Pointer 3579:German Roughhaired Pointer 3258: 2440:10.1016/j.cmet.2016.04.012 2070:10.1016/j.cvsm.2016.11.012 1863:10.1016/j.jveb.2006.06.004 1758:10.1016/j.rvsc.2011.02.002 1240:10.1038/s41598-023-50458-w 889: 861: 842:Seven out of the thirteen 765: 496: 332: 29: 4504: 4383: 4342: 4274: 4239: 4218: 4089: 4011: 3948: 3871: 3813: 3740: 3697: 3589:German Wirehaired Pointer 3574:German Longhaired Pointer 3489: 1976:. Metro. 28 February 2007 1958:. CNN.com. Archived from 1146:'s autobiographical book 1016:Australian special forces 785:This section needs to be 680: 660: 492: 216: 205: 200: 193: 187: 182: 178: 169: 166: 161: 158: 153: 150: 140: 129: 119: 108: 103: 99: 87: 79: 71: 61: 49: 44: 32:Labrador Retriever (song) 4041:English Springer Spaniel 3770:English Springer Spaniel 3707:Chesapeake Bay Retriever 2273:10.1186/1751-0147-46-121 2024:Burnam, John C. (2008). 1524:: CS1 maint: location ( 1380:: CS1 maint: location ( 1108:rocket-propelled grenade 4446:Old Welsh Grey Sheepdog 3750:American Cocker Spaniel 3311:Hancock, David (2013). 3266:Douglas, James (2003). 2171:10.2460/javma.242.3.372 1820:Serpell, James (1995). 1705:10.1126/science.1147880 1602:31 October 2011 at the 1400:"Ch.XXX - The Labrador" 1114: 4257:Miniature Bull Terrier 4195:Sporting Lucas Terrier 4160:Parson Russell Terrier 4130:Dandie Dinmont Terrier 4081:Welsh Springer Spaniel 4031:English Cocker Spaniel 4026:Curly-Coated Retriever 3823:American Water Spaniel 3805:Welsh Springer Spaniel 3765:English Cocker Spaniel 3712:Curly-coated Retriever 3332:Hudson, David (1995). 3073:(Report). London, UK: 2637:. 2006. Archived from 2600:. 2006. Archived from 1781:"American vs. English" 1420:Hawker, Peter (1846). 1152:, adapted into a 2008 1062:11 September 2001 879: 615: 598: 591: 574: 508: 441: 433: 425: 345: 318:3rd Earl of Malmesbury 316:During the 1880s, the 307:2nd Earl of Malmesbury 257:colony of Newfoundland 4426:English White Terrier 4421:English Water Spaniel 4391:Black and Tan Terrier 4051:Flat-Coated Retriever 3887:English Water Spaniel 3717:Flat-coated Retriever 3524:Braque du Bourbonnais 3043:National Public Radio 2783:"Endal the super dog" 1262:Supplementary Table 3 1036:search and rescue dog 958:search and rescue dog 930:, served with former 871: 638:with prior training. 606: 585: 572: 506: 439: 431: 423: 400:, published in 1846. 340: 322:6th Duke of Buccleuch 299:5th Duke of Buccleuch 297:and his nephews, the 253:St. John's water dogs 183:Kennel club standards 4436:North Country Beagle 4370:King Charles Spaniel 4309:Pembroke Welsh Corgi 4304:Old English Sheepdog 4294:Cardigan Welsh Corgi 4135:Jack Russell Terrier 3998:West Country Harrier 3853:Portuguese Water Dog 3833:Cantabrian Water Dog 3790:Pont-Audemer Spaniel 3534:Braque Saint-Germain 3509:Blue Picardy Spaniel 3408:. Barron's. p.  3379:thekennelclub.org.uk 3007:. 12 November 2009. 2763:on 19 September 2007 2701:American Kennel Club 2668:. Ferguson. p.  2635:American Kennel Club 2594:thekennelclub.org.uk 2389:J Am Anim Hosp Assoc 2367:pawprintgenetics.com 2323:on 27 September 2007 2317:"Labrador Retriever" 1645:thekennelclub.org.uk 1440:"Labrador Retriever" 1288:American Kennel Club 1040:September 11 attacks 1008:out for their lives. 702:gall bladder disease 578:Show and field lines 410:American Kennel Club 94:St. John's water dog 4554:1830s introductions 4491:Tweed Water Spaniel 4481:Toy Trawler Spaniel 4441:Old English Bulldog 4406:Cumberland Sheepdog 3902:Tweed Water Spaniel 3897:Old Spanish Pointer 3838:Irish Water Spaniel 3659:Small Münsterländer 3649:Saint-Usuge Spaniel 3609:Large Münsterländer 3559:Drentse Patrijshond 3075:Imperial War Museum 2975:on 27 November 2012 2749:"ANKC hall of fame" 2641:on 9 September 2007 2604:on 4 September 2009 1935:. 20 September 2019 1697:2007Sci...318.1418C 1691:(5855): 1418–1423. 1232:2024NatSR..14..531M 936:Parkinson's disease 822:German shepherd dog 752:autosomal recessive 741:proopiomelanocortin 546:. If a dog carries 408:in 1903. The first 382:Lesser Newfoundland 4549:Labrador Retriever 4375:Northern Inuit Dog 4190:Smooth Fox Terrier 4165:Patterdale Terrier 4145:Manchester Terrier 4115:Bedlington Terrier 4066:Labrador Retriever 3988:Scottish Deerhound 3727:Labrador Retriever 3639:Portuguese Pointer 3629:Perdigueiro Galego 3614:Old Danish Pointer 3195:"Friend of Bill". 2910:The New York Times 2727:New Zealand Herald 2477:The New York Times 2348:2007-09-27 at the 2100:10.1111/jsap.13354 2005:. Associated Press 1909:. 26 February 2019 1655:on 10 January 2015 1616:Labrador Retriever 1606:. The Kennel Club. 1539:Ziessow, Bernard. 1220:Scientific Reports 1131:Russian President 1075:Kabul, Afghanistan 1058:World Trade Center 880: 673:they were used as 616: 613:common wood pigeon 592: 575: 509: 442: 434: 426: 362:Earl of Malmesbury 346: 329:modern Labradors. 293:In the 1830s, the 261:province of Canada 255:imported from the 234:Labrador Retriever 45:Labrador Retriever 4536: 4535: 4396:Blue Paul Terrier 4319:Shetland Sheepdog 4299:Lancashire Heeler 4270: 4269: 4231:Yorkshire Terrier 3910: 3909: 3858:Spanish Water Dog 3843:Lagotto Romagnolo 3689:Wirehaired Vizsla 3519:Braque d'Auvergne 3419:978-0-8120-9018-5 3385:Ltd. January 2018 3322:978-1-84797-492-1 3301:978-0-7566-6004-8 3277:978-1-904057-05-5 3215:"Putin's Pooches" 3116:. 6 February 2007 3084:on 8 January 2009 3071:london.iwm.org.uk 2967:(Press release). 2805:The Seattle Times 2679:978-1-4381-1170-4 2516:978-1-910-44339-2 2002:Los Angeles Times 1883:. 2 November 2019 1050:Salty and Roselle 1044:Hurricane Katrina 1026:missing in action 910:PDSA's Gold Medal 864:List of Labradors 806: 805: 733:retinal dysplasia 729:corneal dystrophy 687:chronic hepatitis 548:wild type alleles 446:conformation show 366:Duke of Buccleuch 344:, whelped in 1885 326:12th Earl of Home 295:10th Earl of Home 230: 229: 212: 211: 174: 173: 16:(Redirected from 4591: 4247:Bull and terrier 4210:Wire Fox Terrier 4180:Sealyham Terrier 4175:Scottish Terrier 4140:Lakeland Terrier 4110:Airedale Terrier 4096: 4095: 4056:Golden Retriever 3968:English Foxhound 3937: 3930: 3923: 3914: 3913: 3722:Golden Retriever 3664:Spinone Italiano 3476: 3469: 3462: 3453: 3452: 3447: 3423: 3407: 3394: 3392: 3390: 3368: 3347: 3326: 3305: 3281: 3253: 3251: 3249: 3247: 3236: 3230: 3229: 3227: 3225: 3211: 3205: 3204: 3192: 3186: 3185: 3183: 3181: 3164: 3158: 3157: 3155: 3153: 3132: 3126: 3125: 3123: 3121: 3100: 3094: 3093: 3091: 3089: 3083: 3077:. Archived from 3068: 3060: 3054: 3053: 3051: 3049: 3027: 3021: 3020: 3018: 3016: 2991: 2985: 2984: 2982: 2980: 2957: 2951: 2950: 2948: 2946: 2929: 2923: 2922: 2920: 2918: 2900: 2894: 2893: 2891: 2889: 2872: 2866: 2865: 2863: 2861: 2854:(abcnews.go.com) 2843: 2837: 2836: 2834: 2832: 2817: 2811: 2801: 2795: 2794: 2792: 2790: 2779: 2773: 2772: 2770: 2768: 2745: 2739: 2738: 2736: 2734: 2721:Kirkness, Luke. 2718: 2712: 2711: 2709: 2707: 2693: 2687: 2686: 2667: 2657: 2651: 2650: 2648: 2646: 2623: 2614: 2613: 2611: 2609: 2586: 2577: 2576: 2574: 2572: 2562: 2555: 2549: 2548: 2546: 2544: 2534: 2527: 2521: 2520: 2502: 2496: 2495: 2493: 2491: 2468: 2462: 2461: 2451: 2419: 2413: 2412: 2384: 2378: 2377: 2375: 2373: 2359: 2353: 2339: 2333: 2332: 2330: 2328: 2313: 2304: 2303: 2293: 2275: 2251: 2245: 2244: 2242: 2232: 2207: 2201: 2200: 2189: 2183: 2182: 2153: 2147: 2146: 2118: 2112: 2111: 2083: 2074: 2073: 2053: 2044: 2043: 2021: 2015: 2014: 2012: 2010: 1992: 1986: 1985: 1983: 1981: 1970: 1964: 1963: 1951: 1945: 1944: 1942: 1940: 1925: 1919: 1918: 1916: 1914: 1899: 1893: 1892: 1890: 1888: 1873: 1867: 1866: 1846: 1840: 1839: 1827: 1817: 1811: 1810: 1804: 1796: 1794: 1792: 1783:. Archived from 1776: 1770: 1769: 1741: 1735: 1734: 1724: 1674: 1665: 1664: 1662: 1660: 1651:. Archived from 1637: 1631: 1625: 1619: 1613: 1607: 1594: 1585: 1584: 1582: 1580: 1565: 1559: 1558: 1556: 1554: 1545: 1536: 1530: 1529: 1523: 1515: 1513: 1511: 1488: 1482: 1481: 1479: 1477: 1462: 1456: 1455: 1453: 1451: 1444:vcahospitals.com 1436: 1430: 1429: 1417: 1408: 1407: 1395: 1386: 1385: 1379: 1371: 1369: 1367: 1344: 1338: 1335: 1326: 1325: 1323: 1321: 1306: 1300: 1299: 1297: 1295: 1280: 1263: 1261: 1251: 1211: 1192: 1191: 1182: 1163:Canadian singer 1158:of the same name 988: 858:Famous Labradors 826:Golden retriever 801: 798: 792: 780: 779: 772: 754:mutation in the 655:Golden Retriever 607:Retrieving shot 386:Charles St. John 225: 219: 180: 179: 101: 100: 89:Foundation stock 72:Common nicknames 64: 54: 42: 41: 21: 4599: 4598: 4594: 4593: 4592: 4590: 4589: 4588: 4539: 4538: 4537: 4532: 4500: 4451:Paisley Terrier 4431:Norfolk Spaniel 4379: 4365:English Mastiff 4338: 4266: 4235: 4214: 4170:Plummer Terrier 4155:Norwich Terrier 4150:Norfolk Terrier 4102: 4085: 4021:Clumber Spaniel 4007: 3944: 3941: 3911: 3906: 3892:Norfolk Spaniel 3867: 3809: 3795:Russian Spaniel 3760:Clumber Spaniel 3736: 3693: 3624:Picardy Spaniel 3549:Ca Mè Mallorquí 3529:Braque Français 3514:Bracco Italiano 3493: 3485: 3480: 3450: 3444: 3430:Morris, Desmond 3420: 3388: 3386: 3383:The Kennel Club 3373: 3365: 3344: 3323: 3302: 3278: 3261: 3256: 3245: 3243: 3238: 3237: 3233: 3223: 3221: 3219:russianlife.org 3213: 3212: 3208: 3194: 3193: 3189: 3179: 3177: 3166: 3165: 3161: 3151: 3149: 3148:. 29 April 2014 3134: 3133: 3129: 3119: 3117: 3102: 3101: 3097: 3087: 3085: 3081: 3066: 3062: 3061: 3057: 3047: 3045: 3028: 3024: 3014: 3012: 2993: 2992: 2988: 2978: 2976: 2959: 2958: 2954: 2944: 2942: 2930: 2926: 2916: 2914: 2901: 2897: 2887: 2885: 2873: 2869: 2859: 2857: 2845: 2844: 2840: 2830: 2828: 2819: 2818: 2814: 2802: 2798: 2788: 2786: 2781: 2780: 2776: 2766: 2764: 2747: 2746: 2742: 2732: 2730: 2719: 2715: 2705: 2703: 2695: 2694: 2690: 2680: 2658: 2654: 2644: 2642: 2625: 2624: 2617: 2607: 2605: 2588: 2587: 2580: 2570: 2568: 2557: 2556: 2552: 2542: 2540: 2529: 2528: 2524: 2517: 2503: 2499: 2489: 2487: 2469: 2465: 2428:Cell Metabolism 2420: 2416: 2401:10.5326/0440295 2385: 2381: 2371: 2369: 2361: 2360: 2356: 2350:Wayback Machine 2340: 2336: 2326: 2324: 2315: 2314: 2307: 2252: 2248: 2208: 2204: 2191: 2190: 2186: 2154: 2150: 2119: 2115: 2084: 2077: 2054: 2047: 2040: 2022: 2018: 2008: 2006: 1993: 1989: 1979: 1977: 1972: 1971: 1967: 1962:on 17 May 2013. 1952: 1948: 1938: 1936: 1927: 1926: 1922: 1912: 1910: 1901: 1900: 1896: 1886: 1884: 1875: 1874: 1870: 1847: 1843: 1836: 1818: 1814: 1798: 1797: 1790: 1788: 1777: 1773: 1742: 1738: 1675: 1668: 1658: 1656: 1649:The Kennel Club 1641:"Breed colours" 1639: 1638: 1634: 1626: 1622: 1614: 1610: 1604:Wayback Machine 1595: 1588: 1578: 1576: 1567: 1566: 1562: 1552: 1550: 1543: 1537: 1533: 1517: 1516: 1509: 1507: 1505: 1489: 1485: 1475: 1473: 1464: 1463: 1459: 1449: 1447: 1438: 1437: 1433: 1418: 1411: 1396: 1389: 1373: 1372: 1365: 1363: 1361: 1345: 1341: 1336: 1329: 1319: 1317: 1308: 1307: 1303: 1293: 1291: 1290:. 31 March 1994 1282: 1281: 1266: 1212: 1205: 1201: 1196: 1195: 1185: 1183: 1179: 1174: 1149:Marley & Me 1117: 983: 950: 894: 888: 886:Assistance dogs 877:PDSA Gold Medal 866: 860: 802: 796: 793: 790: 781: 777: 770: 764: 717:elbow dysplasia 683: 663: 601: 580: 560:melanistic mask 501: 495: 418: 416:Characteristics 394:English Pointer 335: 303:Lord John Scott 221: 217: 189:The Kennel Club 62: 57: 40: 35: 28: 23: 22: 15: 12: 11: 5: 4597: 4587: 4586: 4581: 4576: 4571: 4566: 4561: 4556: 4551: 4534: 4533: 4531: 4530: 4505: 4502: 4501: 4499: 4498: 4493: 4488: 4483: 4478: 4473: 4468: 4463: 4458: 4456:Southern Hound 4453: 4448: 4443: 4438: 4433: 4428: 4423: 4418: 4413: 4408: 4403: 4398: 4393: 4387: 4385: 4381: 4380: 4378: 4377: 4372: 4367: 4362: 4357: 4352: 4346: 4344: 4340: 4339: 4337: 4336: 4334:Welsh Sheepdog 4331: 4326: 4321: 4316: 4311: 4306: 4301: 4296: 4291: 4286: 4284:Bearded Collie 4280: 4278: 4272: 4271: 4268: 4267: 4265: 4264: 4259: 4254: 4249: 4243: 4241: 4237: 4236: 4234: 4233: 4228: 4222: 4220: 4216: 4215: 4213: 4212: 4207: 4202: 4197: 4192: 4187: 4182: 4177: 4172: 4167: 4162: 4157: 4152: 4147: 4142: 4137: 4132: 4127: 4122: 4120:Border Terrier 4117: 4112: 4106: 4104: 4093: 4087: 4086: 4084: 4083: 4078: 4076:Sussex Spaniel 4073: 4068: 4063: 4058: 4053: 4048: 4043: 4038: 4036:English Setter 4033: 4028: 4023: 4017: 4015: 4009: 4008: 4006: 4005: 4000: 3995: 3990: 3985: 3980: 3975: 3970: 3965: 3960: 3954: 3952: 3946: 3945: 3940: 3939: 3932: 3925: 3917: 3908: 3907: 3905: 3904: 3899: 3894: 3889: 3884: 3878: 3876: 3869: 3868: 3866: 3865: 3860: 3855: 3850: 3845: 3840: 3835: 3830: 3825: 3819: 3817: 3811: 3810: 3808: 3807: 3802: 3800:Sussex Spaniel 3797: 3792: 3787: 3782: 3780:German Spaniel 3777: 3772: 3767: 3762: 3757: 3755:Boykin Spaniel 3752: 3746: 3744: 3738: 3737: 3735: 3734: 3729: 3724: 3719: 3714: 3709: 3703: 3701: 3695: 3694: 3692: 3691: 3686: 3681: 3676: 3671: 3666: 3661: 3656: 3651: 3646: 3641: 3636: 3631: 3626: 3621: 3619:Pachón Navarro 3616: 3611: 3606: 3601: 3596: 3591: 3586: 3581: 3576: 3571: 3569:French Spaniel 3566: 3564:English Setter 3561: 3556: 3551: 3546: 3544:Burgos Pointer 3541: 3536: 3531: 3526: 3521: 3516: 3511: 3506: 3504:Ariège Pointer 3500: 3498: 3487: 3486: 3479: 3478: 3471: 3464: 3456: 3449: 3448: 3442: 3425: 3424: 3418: 3396: 3395: 3370: 3369: 3363: 3349: 3348: 3342: 3328: 3327: 3321: 3307: 3306: 3300: 3283: 3282: 3276: 3262: 3260: 3257: 3255: 3254: 3231: 3206: 3187: 3159: 3127: 3095: 3055: 3022: 2986: 2952: 2924: 2895: 2867: 2838: 2812: 2796: 2774: 2740: 2713: 2688: 2678: 2652: 2615: 2598:UK Kennel Club 2578: 2550: 2522: 2515: 2497: 2463: 2434:(5): 893–900. 2414: 2395:(6): 295–301. 2379: 2354: 2334: 2305: 2246: 2202: 2184: 2165:(3): 372–380. 2148: 2129:(4): 856–861. 2113: 2094:(9): 730–736. 2075: 2064:(3): 645–663. 2045: 2038: 2016: 1987: 1965: 1946: 1920: 1907:gundog journal 1894: 1868: 1841: 1834: 1812: 1787:on 5 July 2017 1771: 1752:(3): e98–e99. 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Index

Black Labrador
Labrador Retriever (song)
side view of a yellow dog in a grassy field, facing left
Foundation stock
St. John's water dog
The Kennel Club
standard
Fédération Cynologique Internationale
standard
domestic dog
breed
retriever
gun dog
St. John's water dogs
colony of Newfoundland
province of Canada
Labrador
companion dog
guide
assistance dog
rescue
therapy
10th Earl of Home
5th Duke of Buccleuch
Lord John Scott
2nd Earl of Malmesbury
waterfowling
3rd Earl of Malmesbury
6th Duke of Buccleuch
12th Earl of Home

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