49:
33:
69:
910:
668:
613:
780:
1239:
Antal, L., László, B., Kotlík, P., Mozsár, A., Czeglédi, I., Oldal, M., Kemenesi, G., Jakab, F. & Nagy, S.A. (2015): Phylogenetic evidence for a new species of
672:
1093:
1006:
813:
654:
887:
621:
609:
789:
1104:
1076:
1472:
997:
902:
617:
1143:
Smith, 1841 and the generic status of southern
African tetraploid cyprinids (Teleostei, Cyprinidae). European Journal of Taxonomy, (410).
1420:
1446:
1451:
1184:
1399:
1524:
1477:
1412:
640:
181:
324:" were usually placed here by default. More recently, many "barbs" have been reclassified into genera such as
1459:
540:
The small barbs from Africa, by contrast, are quite distinct. They might even warrant establishment of a new
296:
fish. Some are quite significant, but as a whole, the genus is not yet as well represented in aquaria as the
1159:(Actinopterygii, Cyprinidae, Smiliogastrinae) from the Awash River, Ethiopia, and the re-establishment of
1342:
479:, and many African species (particularly the small ones) do not seem to belong here, either. Eventually,
1347:
1557:
1516:
1216:
1529:
699:
507:
again. They all seem to be close relatives – perhaps the closest living relatives – of
68:
739:
1503:
1282:
1263:
854:
1498:
266:
1490:
548:
are not included in the
Cyprininae –, as they seem to be as distinct from barbels and
1368:
1329:
946:
938:
933:
734:
681:
574:
and might even be included therein. In particular, the group called "redfins" may well be
8:
1199:(Pisces, Cyprinidae) of East and Central Africa. Studies on African Cyprinidae. Part II.
1016:
897:
873:
753:
714:
863:
766:
723:
560:. Within the small African barbs, several lineages can be recognized. These are mostly
529:
63:
1485:
1355:
1257:
1180:
979:
919:
748:
686:
604:
317:
1320:
1060:
1049:
1027:
924:
906:
824:
663:
463:
is for the time being restricted to typical barbels, and only contains fishes from
422:
274:
on their mouths, which they can use to search for food at the bottom of the water.
271:
204:
1360:
1552:
1071:
1038:
968:
775:
708:
200:
171:
1464:
1407:
1394:
1334:
984:
957:
849:
838:
804:
694:
297:
110:
48:
32:
1011:
Lorenzoni, Carosi, Quadroni, De Santis, Vanetti, Delmastro & Zaccara, 2021
1546:
1305:
1212:
992:
878:
636:
630:
499:
488:
392:
219:
177:
120:
1164:
1139:
Skelton, P. H., Swartz, E. R., & Vreven, E. J. (2018). The identity of
799:
649:
575:
570:
557:
513:
476:
446:
440:
380:
374:
211:
163:
556:(which are part of the disputed Labeoninae), rendering the old "Barbinae"
1433:
1314:
761:
519:
493:
428:
416:
410:
404:
398:
362:
332:
326:
321:
270:– refer to the fact that most members of the genera have a pair of
260:
1425:
1144:
565:
545:
386:
350:
243:
207:
130:
1259:
Catalogue of the Fossil Fishes in the
British Museum (Natural History)
1438:
1381:
1108:
1101:
541:
524:
434:
356:
344:
281:
for food; in some locations they are of commercial significance. The
235:
80:
1276:
1386:
1299:
1225:
1120:
829:
549:
338:
293:
140:
100:
1373:
561:
452:
368:
301:
278:
523:
are probably closely related to this group, too, and some large
1155:
Englmaier GK, Tesfaye G, Bogutskaya NG (2020) A new species of
468:
464:
90:
1511:
568:
group largely restricted to southern Africa is very close to
553:
475:. However, the genus even in the reduced version is probably
246:
at least for the largest part (including the type species of
239:
196:
472:
598:
There are currently 36 recognized species of this genus:
282:
1111:., but it probably should be placed in another genus.
1245:
503:are highly similar and might better be included in
289:barbs are also often eaten in their native range.
1544:
1249:
1195:Banister, K.E. (1973): A revision of the large
483:is likely to be restricted to the group around
1255:
1201:Bulletin of the British Museum, 26 (1): 3-148.
728:Kotlík, Tsigenopoulos, Ráb & Berrebi, 2002
320:". Historically, most fish commonly known as "
292:The smaller barbs are in some cases traded as
238:Barbinae, but given their relationships, that
1179:. Chartwell Books, Edison, New Jersey, USA.
818:Khaefi, Esmaeili, Geiger & Eagderi, 2017
285:of barbels is poisonous, however. The large
1189:
308:
1169:
47:
31:
1165:https://doi.org/10.3897/zookeys.902.39606
582:entirely, instead of being split between
1233:
1211:
253:
1545:
1281:
1280:
491:species commonly known as "barbels".
1517:A247C93C-FFC2-A309-C1A4-FB62FA06F8D1
1413:7cce67a4-233c-4595-b98a-7c06b29894a6
1145:https://doi.org/10.5852/ejt.2018.410
13:
14:
1569:
1262:. order of the Trustees. p.
1086:
67:
1205:
1149:
1133:
487:– the large European to
1:
1126:
544:– in particular if the
234:is the namesake genus of the
7:
1243:in the Danube River basin.
1114:
258:Their common names –
10:
1574:
593:
316:has a long history as a "
242:is better included in the
1289:
169:
162:
64:Scientific classification
62:
55:
46:
39:
30:
23:
1215:; Pauly, Daniel (eds.).
1177:Freshwater Aquarium Fish
1163:. ZooKeys 902: 107-150.
552:, as these are from the
309:Systematics and taxonomy
1256:Woodward, A.S. (1901).
1002:(Western Balkan barbel)
1175:Lambert, D.J. (1997):
883:(Mediterranean barbel)
471:, as well as adjacent
1499:Paleobiology Database
533:) may well belong in
222:, first described as
1408:Fauna Europaea (new)
1268:barbus megacephalus.
1229:. July 2024 version.
1100:) is known from the
947:Barbus pergamonensis
934:Barbus peloponnesius
735:Barbus ciscaucasicus
682:Barbus borysthenicus
254:Description and uses
1094:Barbus megacephalus
1056:(Thessalian barbel)
1032:S. L. Karaman, 1955
1017:Barbus sperchiensis
951:M. S. Karaman, 1971
898:Barbus niluferensis
874:Barbus meridionalis
868:S. L. Karaman, 1928
809:(Catalonian barbel)
730:(Carpathian barbel)
1217:"Species in genus
1091:A fossil species (
1023:(Sperchios barbel)
953:(Anatolian barbel)
864:Barbus macedonicus
724:Barbus carpathicus
677:(Biharian barbel)
675:& Kotlík, 2016
659:(Bulgarian barbel)
277:Barbels are often
1558:Cyprinidae genera
1540:
1539:
1486:Open Tree of Life
1283:Taxon identifiers
1099:
1080:
1066:
1055:
1054:Stephanidis, 1971
1044:
1034:(Strumica barbel)
1033:
1022:
1021:Stephanidis, 1950
1012:
1007:Barbus samniticus
1001:
988:
980:Barbus prespensis
974:
964:(Romanian barbel)
963:
952:
942:
928:
920:Barbus oligolepis
914:
893:
882:
869:
859:(Lortet's barbel)
858:
844:
833:
819:
814:Barbus karunensis
808:
795:
784:
770:
757:
749:Barbus cyclolepis
743:
729:
718:
703:
690:
676:
658:
644:
625:
605:Barbus balcanicus
509:Aulopyge huegelii
497:and particularly
318:wastebasket taxon
188:
187:
158:
1565:
1533:
1532:
1520:
1519:
1507:
1506:
1494:
1493:
1481:
1480:
1468:
1467:
1465:NHMSYS0000544620
1455:
1454:
1442:
1441:
1429:
1428:
1416:
1415:
1403:
1402:
1390:
1389:
1377:
1376:
1364:
1363:
1351:
1350:
1338:
1337:
1325:
1324:
1323:
1310:
1309:
1308:
1278:
1277:
1271:
1270:
1253:
1247:
1237:
1231:
1230:
1209:
1203:
1193:
1187:
1173:
1167:
1153:
1147:
1137:
1097:
1081:(Vistula barbel)
1075:
1064:
1061:Barbus tyberinus
1053:
1050:Barbus thessalus
1045:(Crimean barbel)
1042:
1031:
1028:Barbus strumicae
1020:
1010:
996:
983:
975:(Italian barbel)
972:
961:
950:
937:
929:(Marmara barbel)
923:
901:
892:De Filippi, 1863
891:
877:
867:
853:
842:
828:
825:Barbus kubanicus
817:
803:
793:
779:
765:
752:
738:
727:
712:
698:
685:
667:
664:Barbus biharicus
653:
634:
608:
423:Mesopotamichthys
156:
72:
71:
51:
35:
21:
20:
1573:
1572:
1568:
1567:
1566:
1564:
1563:
1562:
1543:
1542:
1541:
1536:
1528:
1523:
1515:
1510:
1502:
1497:
1489:
1484:
1476:
1471:
1463:
1458:
1450:
1445:
1437:
1432:
1424:
1419:
1411:
1406:
1398:
1393:
1385:
1380:
1372:
1367:
1359:
1354:
1346:
1341:
1333:
1328:
1319:
1318:
1313:
1304:
1303:
1298:
1285:
1275:
1274:
1254:
1250:
1238:
1234:
1210:
1206:
1194:
1190:
1174:
1170:
1154:
1150:
1141:Barbus capensis
1138:
1134:
1129:
1117:
1089:
1084:
1072:Barbus waleckii
1065:Bonaparte, 1839
1039:Barbus tauricus
973:Bonaparte, 1839
969:Barbus plebejus
888:Barbus miliaris
845:(lizard barbel)
776:Barbus euboicus
709:Barbus carottae
645:(Common barbel)
626:(Danube barbel)
596:
311:
298:Southeast Asian
256:
224:Cyprinus barbus
201:ray-finned fish
184:
175:
172:Cyprinus barbus
155:
66:
57:Barbus plebejus
17:
16:Genus of fishes
12:
11:
5:
1571:
1561:
1560:
1555:
1538:
1537:
1535:
1534:
1521:
1508:
1495:
1482:
1469:
1456:
1443:
1430:
1417:
1404:
1395:Fauna Europaea
1391:
1378:
1365:
1352:
1339:
1326:
1311:
1295:
1293:
1287:
1286:
1273:
1272:
1248:
1232:
1213:Froese, Rainer
1204:
1188:
1168:
1148:
1131:
1130:
1128:
1125:
1124:
1123:
1116:
1113:
1088:
1087:Fossil species
1085:
1083:
1082:
1068:
1067:(Horse barbel)
1057:
1046:
1035:
1024:
1013:
1003:
989:
976:
965:
958:Barbus petenyi
954:
943:
930:
916:
915:(Simav barbel)
894:
884:
870:
860:
850:Barbus lorteti
846:
839:Barbus lacerta
835:
834:(Kuban barbel)
821:
820:(Karun barbel)
810:
796:
786:
772:
758:
745:
744:(Terek barbel)
731:
720:
719:(Yliki barbel)
705:
704:(brook barbel)
695:Barbus caninus
691:
678:
660:
646:
627:
600:
595:
592:
578:and belong in
310:
307:
255:
252:
226:and now named
186:
185:
176:
167:
166:
160:
159:
148:
144:
143:
138:
134:
133:
128:
124:
123:
118:
114:
113:
111:Actinopterygii
108:
104:
103:
98:
94:
93:
88:
84:
83:
78:
74:
73:
60:
59:
53:
52:
44:
43:
37:
36:
28:
27:
15:
9:
6:
4:
3:
2:
1570:
1559:
1556:
1554:
1551:
1550:
1548:
1531:
1526:
1522:
1518:
1513:
1509:
1505:
1500:
1496:
1492:
1487:
1483:
1479:
1474:
1470:
1466:
1461:
1457:
1453:
1448:
1444:
1440:
1435:
1431:
1427:
1422:
1418:
1414:
1409:
1405:
1401:
1396:
1392:
1388:
1383:
1379:
1375:
1370:
1366:
1362:
1357:
1353:
1349:
1344:
1340:
1336:
1331:
1327:
1322:
1316:
1312:
1307:
1301:
1297:
1296:
1294:
1292:
1288:
1284:
1279:
1269:
1265:
1261:
1260:
1252:
1246:
1242:
1236:
1228:
1227:
1222:
1220:
1214:
1208:
1202:
1198:
1192:
1186:
1185:0-7858-0867-1
1182:
1178:
1172:
1166:
1162:
1158:
1152:
1146:
1142:
1136:
1132:
1122:
1119:
1118:
1112:
1110:
1106:
1103:
1098:Günther, 1876
1096:
1095:
1078:
1074:
1073:
1069:
1063:
1062:
1058:
1052:
1051:
1047:
1043:Kessler, 1877
1041:
1040:
1036:
1030:
1029:
1025:
1019:
1018:
1014:
1009:
1008:
1004:
999:
995:
994:
993:Barbus rebeli
990:
986:
985:S. L. Karaman
982:
981:
977:
971:
970:
966:
960:
959:
955:
949:
948:
944:
940:
936:
935:
931:
926:
922:
921:
917:
912:
908:
904:
900:
899:
895:
890:
889:
885:
880:
876:
875:
871:
866:
865:
861:
856:
852:
851:
847:
841:
840:
836:
831:
827:
826:
822:
816:
815:
811:
806:
802:
801:
797:
792:
791:
790:Barbus fucini
787:
785:(Evia barbel)
782:
778:
777:
773:
771:(Kura barbel)
768:
764:
763:
759:
755:
751:
750:
746:
741:
737:
736:
732:
726:
725:
721:
716:
711:
710:
706:
701:
697:
696:
692:
688:
684:
683:
679:
674:
670:
666:
665:
661:
656:
652:
651:
647:
642:
638:
633:
632:
631:Barbus barbus
628:
623:
619:
615:
614:Tsigenopoulos
611:
607:
606:
602:
601:
599:
591:
589:
585:
581:
577:
573:
572:
567:
563:
559:
555:
551:
550:typical carps
547:
543:
538:
536:
532:
531:
526:
522:
521:
516:
515:
510:
506:
502:
501:
500:Messinobarbus
496:
495:
490:
489:Ponto-Caspian
486:
482:
478:
474:
470:
466:
462:
457:
455:
454:
449:
448:
443:
442:
437:
436:
431:
430:
425:
424:
419:
418:
413:
412:
407:
406:
401:
400:
395:
394:
393:Hypselobarbus
389:
388:
383:
382:
377:
376:
371:
370:
365:
364:
359:
358:
353:
352:
347:
346:
341:
340:
335:
334:
329:
328:
323:
319:
315:
306:
304:
303:
299:
295:
290:
288:
284:
280:
275:
273:
269:
268:
263:
262:
251:
249:
245:
241:
237:
233:
229:
228:Barbus barbus
225:
221:
220:common barbel
217:
213:
209:
206:
202:
198:
194:
193:
183:
179:
174:
173:
168:
165:
161:
154:
153:
149:
146:
145:
142:
139:
136:
135:
132:
129:
126:
125:
122:
121:Cypriniformes
119:
116:
115:
112:
109:
106:
105:
102:
99:
96:
95:
92:
89:
86:
85:
82:
79:
76:
75:
70:
65:
61:
58:
54:
50:
45:
42:
41:Barbus barbus
38:
34:
29:
26:
22:
19:
1290:
1267:
1258:
1251:
1244:
1240:
1235:
1224:
1218:
1207:
1200:
1196:
1191:
1176:
1171:
1161:E. akakianus
1160:
1156:
1151:
1140:
1135:
1105:Sipang Fauna
1092:
1090:
1070:
1059:
1048:
1037:
1026:
1015:
1005:
991:
978:
967:
962:Heckel, 1852
956:
945:
939:Valenciennes
932:
918:
896:
886:
872:
862:
848:
843:Heckel, 1843
837:
823:
812:
800:Barbus haasi
798:
788:
774:
760:
747:
733:
722:
707:
693:
680:
662:
650:Barbus bergi
648:
629:
603:
597:
587:
584:Pseudobarbus
583:
580:Pseudobarbus
579:
576:monophyletic
571:Pseudobarbus
569:
558:paraphyletic
539:
534:
528:
527:barbs (e.g.
518:
514:Carasobarbus
512:
508:
504:
498:
492:
484:
480:
477:paraphyletic
460:
458:
451:
447:Puntioplites
445:
441:Pseudobarbus
439:
433:
427:
421:
415:
409:
403:
397:
391:
385:
381:Clypeobarbus
379:
375:Carasobarbus
373:
367:
361:
355:
349:
343:
337:
331:
325:
313:
312:
300:
291:
286:
276:
265:
259:
257:
247:
231:
227:
223:
215:
212:type species
191:
190:
189:
170:
164:Type species
157:Daudin, 1805
151:
150:
56:
40:
24:
18:
1434:iNaturalist
1315:Wikispecies
794:Costa, 1853
781:Stephanidis
762:Barbus cyri
535:Labeobarbus
520:Labeobarbus
494:Luciobarbus
429:Poropuntius
417:Luciobarbus
411:Leptobarbus
405:Labeobarbus
399:Hypsibarbus
363:Caecobarbus
333:Barbichthys
327:Arabibarbus
137:Subfamily:
1547:Categories
1157:Enteromius
1127:References
830:L. S. Berg
767:De Filippi
566:tetraploid
546:Labeoninae
387:Enteromius
351:Barbonymus
244:Cyprininae
208:Cyprinidae
131:Cyprinidae
1121:Danionins
1109:Indonesia
1102:Paleogene
925:Battalgil
700:Bonaparte
655:Chichkoff
542:subfamily
530:L. reinii
525:hexaploid
485:B. barbus
435:Probarbus
357:Barbopsis
345:Barboides
236:subfamily
87:Kingdom:
81:Eukaryota
1374:46581712
1300:Wikidata
1226:FishBase
1115:See also
907:Kottelat
879:A. Risso
687:Dybowski
637:Linnaeus
339:Barbodes
294:aquarium
178:Linnaeus
141:Barbinae
127:Family:
101:Chordata
97:Phylum:
91:Animalia
77:Domain:
1426:5201936
1306:Q133326
911:Ekmekçi
855:Sauvage
805:Mertens
740:Kessler
717:, 1998)
622:Berrebi
594:Species
562:diploid
453:Puntius
369:Capoeta
302:Puntius
272:barbels
267:barbels
218:is the
203:in the
147:Genus:
117:Order:
107:Class:
1553:Barbus
1530:154291
1491:257936
1452:163618
1400:304389
1387:1BARUG
1335:Barbus
1321:Barbus
1291:Barbus
1241:Barbus
1219:Barbus
1197:Barbus
1183:
1079:, 1970
1000:, 1926
998:Koller
987:, 1924
941:, 1842
927:, 1941
913:, 2009
909:&
881:, 1827
857:, 1882
832:, 1913
807:, 1925
783:, 1950
769:, 1865
756:, 1837
754:Heckel
742:, 1877
715:Bianco
702:, 1839
689:, 1862
673:László
657:, 1935
624:, 2002
620:&
610:Kotlík
588:Barbus
554:garras
505:Barbus
481:Barbus
469:Europe
465:Africa
461:Barbus
459:Thus,
314:Barbus
287:Barbus
279:fished
248:Barbus
232:Barbus
216:Barbus
210:. The
205:family
192:Barbus
152:Barbus
25:Barbus
1525:WoRMS
1512:Plazi
1504:64194
1478:40829
1439:49620
1348:71191
1077:Rolik
903:Turan
669:Antal
322:barbs
261:barbs
240:taxon
197:genus
195:is a
1473:NCBI
1447:ITIS
1421:GBIF
1382:EPPO
1361:382L
1343:BOLD
1181:ISBN
641:1758
586:and
564:; a
517:and
473:Asia
467:and
450:and
264:and
182:1758
1460:NBN
1369:EoL
1356:CoL
1330:ADW
1264:302
1107:of
618:Ráb
283:roe
250:).
214:of
199:of
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444:,
438:,
432:,
426:,
420:,
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390:,
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378:,
372:,
366:,
360:,
354:,
348:,
342:,
336:,
330:,
305:.
230:.
180:,
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713:(
643:)
635:(
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