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Ankarana Special Reserve

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278: 217: 66: 349:) foraged together in mixed bands. Within each band different species seemed to specialise in where and how they searched out their insect prey. Some species concentrated on the trunk and branches of trees, some on slender boughs, others searched beneath the leaves. By acting together in this way they probably increased foraging efficiency as each species could catch others' escaped prey. They were also safer from attack by predators, as the group as a whole was more likely to spot approaching danger. 1260: 73: 269:. Around 100 kilometres (62 mi) of cave passages within the massif have been mapped. La Grotte d'Andrafiabe, one of the most accessible caves, comprises at least 8.035 kilometres (4.993 mi) of horizontal passages. Indeed, the Massif contains the longest cave systems in Madagascar, and probably in the whole of Africa. 50: 300:
During the 1986 expedition, Phil Chapman and Jean-Elie Randriamasy collated a bird list for the reserve and recorded 65 species from 32 families representing nearly a third of all bird species that breed in Madagascar. They also noted one interesting aspect of behaviour. They reported that there was
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and in the scientific press. Discoveries included unexpected sub-fossil remains of large extinct lemurs and surviving but previously undescribed species of blind fish, shrimps and other invertebrates. Several expedition members contributed photos to an illustrated introductory guide to Madagascar
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The plateau slopes gently to the east, but on the west it ends abruptly in the "Wall of Ankarana", a sheer cliff that extends 25 kilometres (16 mi) north to south, and rises up to 280 metres (920 ft). To the south, the limestone mass breaks up into separate spires known as
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Beginning in the 1960s, expatriate Frenchman Jean Duflos (who after marriage changed his name to Jean Radofilao) undertook a huge amount of exploration of the cave systems and subterranean rivers of the Massif, much of it on his own or with visiting
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Wilson, J.M.; et al. (1989). "Ecology and Conservation of the Crowned Lemur at Ankarana, N. Madagascar with notes on Sanford's Lemur, Other Sympatrics and Subfossil Lemurs".
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Peyre, J-C.; Arthaud, G.; Radofilao, J.; et al. (1982). "ExpĂ©dition SpĂ©lĂ©ologique, Madagascar 1982". Club Alpin Francais / Federation Francaise de SpĂ©lĂ©ologie: 55pp.
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Jane M. Wilson (1982). "A review of world Troglopedetini (Insecta, Collembola, Paronellidae), including an identification table and descriptions of new species".
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Wilson, Jane M.; Godfrey, L.R.; Simons, E.L.; Stewart, Paul D.; Vuillaume-Randriamanantena, M. (1995). "Past and Present Lemur Fauna at Ankarana, N. Madagascar".
190:. With an average annual rainfall of about 2,000 millimetres (79 in), the underlying rocks have been eroded to produce caves and feed subterranean rivers—a 1212: 288:
Expeditions that first began cataloguing the animals and plants of the Special Reserve created around the Ankarana Massif in the 1980s are described in Dr
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Fowler, S.V.; et al. (1989). "A survey and management proposals for a tropical deciduous forest reserve at Ankarana in northern Madagascar".
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Godfrey, L.R.; Wilson, Jane M.; Simons, E.L.; Stewart, Paul D.; Vuillaume-Randriamanantena, M. (1996). "Ankarana: window to Madagascar's past".
244:. In places where the calcific upper layers have been completely eroded, the harder base rock has been etched into channels and ridges known in 1104: 116: 1649: 1669: 1205: 65: 1664: 240:. In the center of the plateau, seismic activity and eons of rainfall have eroded the limestone, forming deep gorges and ribbons of 1644: 1239: 1624: 881:
Simons, E.L.; et al. (1990). "Discovery of new giant subfossil lemurs in the Ankarana Mountains of Northern Madagascar".
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Rossi, G. (1974). "Morphologie et Evolution d'un karst en milieu tropical. L'Ankarana (Extreme Nord de Madagascar)".
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an unusual strategy used by many of the small insect-eating songbirds. Species such as the paradise flycatcher (
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created in 1956. It is a small, partially vegetated plateau composed of 150-million-year-old middle Jurassic
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Wilson, Jane, ed. (1987). "The Crocodile Caves of Ankarana : Expedition to Northern Madagascar, 1986".
1563: 1522: 1344: 1294: 1639: 1517: 1502: 1457: 1452: 1416: 1324: 1314: 1274: 425: 327: 1437: 1299: 1619: 1578: 1507: 1289: 167: 642: 1527: 1472: 1380: 1375: 1284: 361: 315: 1593: 1185: 1133: 194:. The rugged relief and the dense vegetation have helped protect the region from human intrusion. 1512: 1411: 1334: 369: 1497: 1390: 1360: 1329: 385: 1339: 629: 345: 198: 1190: 1553: 1482: 1319: 440: 401: 333: 277: 8: 1583: 1349: 1309: 1249: 461: 409: 289: 1109:, a new genus and species of cave collembolan from Madagascar with notes on its ecology" 216: 558: 303: 1548: 1487: 1244: 1162: 894: 834: 800: 727: 675: 566: 393: 377: 245: 1573: 1125: 1066: 1029: 925: 890: 861: 830: 792: 760: 702: 309: 256:
boulders and basalt has also flowed deep into the canyons that dissect the Massif.
191: 368:) and other mammal species. The following lemurs are also recorded from the area: 1598: 1588: 724:
Lemurs of the Lost World: exploring the forests and Crocodile Caves of Madagascar
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Duflos, J. (1966). "Bilan des explorations biospeleologique pour l'annee 1965".
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The Ankarana Reserve is an important refuge for significant populations of the
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Duflos, J. (1968). "Bilan des explorations speleologique pour l'annee 1966".
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MĂ©moires et Documents Centre National de la Recherche Scientifique
749:"Ankarana - a rediscovered nature reserve in northern Madagascar" 417: 253: 197:
The southern entrance of the park is in Mahamasina (commune of
1220: 448: 236: 969: 947: 993:, a new species of blind cave goby from Madagascar". 623: 439:
of the following lemurs have been found at Ankarana:
816: 814: 213:. There are also some hotels found at the entrance. 778: 776: 557: 1161:. Aston Publications, Bourne End, UK. p. 96. 1103:JosĂ© G. Palacios-Vargas & Jane Wilson (1990). 811: 1611: 773: 297:which features the Crocodile Caves of Ankarana. 72: 1049:Gurney, A.R. (1984). "Freshwater shrimp genera 1014:"Conservation and ecology of a new blind fish, 1083: 1206: 1156: 982: 691:"The Crocodile Caves of Ankarana, Madagascar" 995:Journal of Ichthyology & Aquatic Biology 531: 529: 284:photographed at the Ankarana Special Reserve 1042: 1213: 1199: 209:and 29 km (18 mi) north-east of 1033: 929: 865: 764: 706: 526: 988: 276: 215: 847: 14: 1612: 1048: 1011: 907: 880: 820: 782: 746: 721: 688: 655: 604: 585: 1194: 1150: 535: 27:Protected area in northern Madagascar 1018:from the Ankarana Caves, Madagascar" 914:, and the evolution of sloth lemurs" 747:Wilson, Jane M; et al. (1988). 613:. UniversitĂ© de Madagascar: 121–129. 594:. UniversitĂ© de Madagascar: 225–252. 553: 551: 42:IV (habitat/species management area) 1650:Protected areas established in 1956 1117:International Journal of Speleology 24: 1670:Important Bird Areas of Madagascar 908:Simons, E.L.; et al. (1992). 25: 1681: 1179: 548: 1665:Madagascar dry deciduous forests 1258: 509:Madagascar dry deciduous forests 151:182 km (70 sq mi) 71: 64: 48: 1645:Protected areas in Diana Region 1096: 1077: 1005: 901: 874: 841: 740: 726:. Impact, London. p. 216. 467:Mesopropithicus dolichobrachion 322:Phyllastrephus madagascariensis 205:some 108 km south-west of 1625:Special reserves of Madagascar 910:"A new giant subfossil lemur, 715: 682: 668: 649: 617: 598: 579: 430:Hapalemur griseus occidentalis 259: 13: 1: 519: 1564:Ankeniheny-Zahamena Corridor 895:10.1016/0047-2484(90)90072-J 835:10.1016/0006-3207(89)90072-4 563:Natural Wonders of the World 491:madagascariensis/grandidieri 422:Daubentonia madagascariensis 414:Propithicus diadema perrieri 252:. The area is littered with 80:Location of Ankarana Reserve 7: 1660:Madagascar subhumid forests 1222:National Parks and Reserves 1157:Bradt, Hilary, ed. (1988). 502: 426:western lesser bamboo lemur 328:Hypsipetes madagascariensis 10: 1686: 1059:Journal of Natural History 883:Journal of Human Evolution 227: 220:Ankarana Plateau, showing 1579:Mandena Conservation Zone 1541: 1430: 1404: 1267: 1256: 1232: 1186:Madagascar National Parks 1071:10.1080/00222938400770481 1035:10.1017/S0030605300021669 1016:Glossogobius ankaranensis 991:Glossogobius ankaranensis 867:10.1017/S0030605300022390 848:Stewart, Paul D. (1988). 766:10.1017/S0030605300027794 708:10.1017/S0030605300020470 374:Lepilemur septentrionalis 340:Lepopterus madagascarinus 168:Madagascar National Parks 163: 155: 147: 110: 99: 88: 59: 47: 37: 32: 1130:10.5038/1827-806X.19.1.6 1012:Wilson, Jane M. (1996). 989:Banister, K.E. (1994). " 689:Wilson, Jane M. (1987). 316:Phyllastrephus zosterops 294:Lemurs of the Lost World 272: 176:Ankarana Special Reserve 33:Ankarana Special Reserve 1635:Geography of Madagascar 1630:Mountains of Madagascar 823:Biological Conservation 493:, and the baboon lemur 370:northern sportive lemur 1533:Tampoketsa Analamaitso 1233:Strict Nature Reserves 1107:Troglobius coprophagus 637:Cite journal requires 386:fat-tailed dwarf lemur 285: 224: 1655:Geology of Madagascar 1594:Montagne des Français 1542:Other protected areas 1405:Marine National Parks 722:Wilson, Jane (2014). 362:Sanford's brown lemur 346:Xenopirostris polleni 280: 219: 199:Tanambao Marivorahona 132:13.07278°S 48.91472°E 972:Primate Conservation 441:greater bamboo lemur 402:eastern woolly lemur 334:Nectarinia souimanga 307:), the common jery ( 292:'s travel narrative 1584:Makira Natural Park 918:Folia Primatologica 785:Folia Primatologica 607:Revue de GĂ©ographie 588:Revue de GĂ©ographie 462:Babakotia radofilai 290:Jane Wilson-Howarth 164:Governing body 137:-13.07278; 48.91472 128: /  1640:Plateaus of Africa 1396:Zombitse-Vohibasia 850:"Ankarana damaged" 390:Cheirogalus medius 313:), the greenbuls ( 304:Terpsiphone mutata 286: 225: 1607: 1606: 1549:Ambato Atsinanana 1280:Andasibe-Mantadia 931:10.1159/000156629 797:10.1159/000156379 676:"World Cave List" 471:Palaeopropithicus 394:fork-marked lemur 378:brown mouse lemur 358:Eulemur coronatus 337:) and the vagas ( 203:Route nationale 6 173: 172: 100:Nearest city 16:(Redirected from 1677: 1620:IUCN Category IV 1574:Mahavavy-Kinkony 1488:Cap Sainte-Marie 1463:Anjanaharibe-Sud 1431:Special Reserves 1386:Tsimanampetsotsa 1356:Montagne d'Ambre 1262: 1215: 1208: 1201: 1192: 1191: 1173: 1172: 1154: 1148: 1147: 1145: 1144: 1138: 1132:. Archived from 1113: 1100: 1094: 1093: 1081: 1075: 1074: 1046: 1040: 1039: 1037: 1009: 1003: 1002: 986: 980: 979: 967: 958: 957: 945: 936: 935: 933: 905: 899: 898: 878: 872: 871: 869: 845: 839: 838: 818: 809: 808: 780: 771: 770: 768: 744: 738: 737: 719: 713: 712: 710: 686: 680: 679: 672: 666: 665: 653: 647: 646: 640: 635: 633: 625: 621: 615: 614: 602: 596: 595: 583: 577: 576: 555: 546: 545: 533: 410:Perrier's sifaka 382:Microcebus rufus 366:Eulemur sanfordi 331:), the sunbird ( 310:Neomixis tenella 192:karst topography 159:20 February 1956 143: 142: 140: 139: 138: 133: 129: 126: 125: 124: 121: 75: 74: 68: 54:Ankarana reserve 52: 30: 29: 21: 18:Ankarana Reserve 1685: 1684: 1680: 1679: 1678: 1676: 1675: 1674: 1610: 1609: 1608: 1603: 1599:Tsitongambarika 1589:Menabe Antimena 1537: 1426: 1422:Nosy Ve-Androka 1400: 1371:Nosy Ve-Androka 1263: 1254: 1228: 1219: 1182: 1177: 1176: 1169: 1155: 1151: 1142: 1140: 1136: 1111: 1101: 1097: 1082: 1078: 1047: 1043: 1010: 1006: 987: 983: 968: 961: 946: 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Index

Ankarana Reserve
IUCN category

Map showing the location of Ankarana Special Reserve
Diana Region
Antsiranana
13°4′22″S 48°54′53″E / 13.07278°S 48.91472°E / -13.07278; 48.91472
Madagascar National Parks
protected area
Madagascar
limestone
karst topography
Tanambao Marivorahona
Route nationale 6
Antsiranana
Ambilobe

karst
flowstone
malagasy
basalt
speleologists

Crowned lemur
Jane Wilson-Howarth
Terpsiphone mutata
Neomixis tenella
Phyllastrephus zosterops
Phyllastrephus madagascariensis
Hypsipetes madagascariensis

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