108:
517:. The overall shape of the nutlet is circular to elliptic with the average length of the samara up to 4.0 centimetres (1.6 in) and a wing width of 1.2 centimetres (0.47 in). The paired samaras of the species have a 25° to 30° attachment angle and the distal region of the nutlet and wing forming distinct u-shaped shallow
489:
and are generally oblate to widely elliptical in shape. The leaves are five-lobed with the basal two lobes small while the upper lateral lobes are about two-thirds as long as the median lobe and all lobes being triangular in outline. The leaves have five primary veins and range between 3.7 to 7.2
363:
produced one paratype leaf of middle
Miocene age. The last paratype leaf is from the species' northernmost location, having been recovered from an outcrop of the late early Miocene Skonun Formation on the northern side of Graham Island in the
494:
has small teeth while the lobes have a distinct and complex bracing of veins formed by the joining of two external secondary veins. The combination of morphological features is not found in any modern species besides
595:
DeVore, M. L.; Pigg, K. B. (2007). "A brief review of the fossil history of the family
Rosaceae with a focus on the Eocene Okanogan Highlands of eastern Washington State, USA, and British Columbia, Canada".
657:
Champigny, N.; Henderson, C. M.; Rouse, G. E. (1981). "New evidence for the age of the Skonun
Formation, Queen Charlotte Islands, British Columbia".
372:. In addition to the leaf specimens a series of paratype specimens were described for the associated samaras recovered at the Collawash site and
404:
569:
717:
707:
727:
635:
412:
702:
377:
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centimetres (1.5 to 2.8 in) long by 2.9 to 6.0 centimetres (1.1 to 2.4 in) wide in overall dimensions.
107:
439:
373:
339:
were recovered from five different sites. The largest group of specimens was recovered from the early
Miocene,
420:
213:
722:
712:
381:
360:
241:
666:
408:
356:
323:
8:
443:
670:
578:
Journal of the
Faculty of Science, Hokkaido University. Series 4, Geology and Mineralogy
613:
102:
469:
is in recognition of paleobotanist Roland Wilbur Brown for his many contributions to
617:
674:
605:
518:
428:
369:
348:
310:
497:
352:
344:
145:
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525:, the two related species can be separated by the sulcus which is developed in
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340:
302:
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132:
609:
691:
435:
432:
314:
298:
260:
61:
321:, Canada. It is one of several extinct species placed in the living section
636:"Geology and coal resources of the Centralia-Chehalis district, Washington"
351:
Oregon. Two additional paratype leaves were recovered from outcrops of the
416:
411:. Three other paratype leaves and one paratype samara are housed in the
400:
365:
318:
283:
191:
36:
513:
have a notably inflated nutlet and acutely diverging veins which rarely
514:
181:
81:
46:
459:
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396:
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158:
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56:
51:
41:
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279:
91:
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171:
276:
201:
119:
486:
634:
Snavely, P. D.; Brown, R. D. Jr; Roberts, A. E. (1958).
656:
456:
Journal of the
Faculty of Science, Hokkaido University
359:. Another Washington site, on Bunker Creek west of
633:
391:are placed into three different repositories. The
574:(maples) in the Cenozoic of Western North America"
689:
405:University of California Museum of Paleontology
440:United States Geological Survey, Denver office
629:
627:
570:"Systematics, Phylogeny, and Distribution of
330:
650:
419:while the last paratype leaf is part of the
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551:
549:
547:
545:
335:The eleven leaf specimens used to describe
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567:
588:
640:United States Geological Survey Bulletin
542:
485:are simple in structure, with perfectly
446:. Wolfe and Tanai published their 1987
690:
584:(1): 23, 74, 75, 240, & plate 4.
286:described from a series of isolated
13:
659:Canadian Journal of Earth Sciences
521:. While similar in morphology to
413:National Museum of Natural History
343:, Collawash site exposed near the
14:
739:
431:. The specimens were studied by
297:. The species is known from the
106:
34:
598:Plant Systematics and Evolution
568:Wolfe, J.A.; Tanai, T. (1987).
529:but not seen in the samaras of
399:are currently preserved in the
374:United States Geological Survey
718:Extinct flora of North America
476:
421:University of British Columbia
1:
708:Fossil taxa described in 1987
536:
305:sediments exposed in Western
728:Fossils of the United States
487:actinodromous vein structure
32:Early Miocene–Middle Miocene
7:
505:is placed into the section
10:
744:
403:collections housed at the
331:History and classification
610:10.1007/s00606-007-0540-3
395:leaf and thirteen of the
247:
240:
103:Scientific classification
101:
23:
703:Plants described in 1987
376:site USGS 8904 near the
16:Extinct species of maple
442:and Toshimasa Tanai of
387:The type specimens for
382:Jackson County, Oregon
355:at the Vera site near
361:Centralia, Washington
409:Berkeley, California
357:Veradale, Washington
671:1981CaJES..18.1900C
444:Hokkaido University
313:, USA and Northern
509:. The samaras of
378:Little Butte Creek
263:& Tanai, 1987
723:Fossils of Canada
665:(12): 1900–1903.
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631:
622:
621:
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448:type description
429:British Columbia
370:British Columbia
349:Clackamas County
311:Washington state
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33:
29:Temporal range:
21:
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679:10.1139/e81-176
655:
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632:
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479:
423:collections in
353:Latah Formation
345:Collawash River
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264:
258:
250:
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713:Miocene plants
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623:
604:(1–2): 45–57.
587:
540:
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478:
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462:of the chosen
433:paleobotanists
415:, part of the
401:paleobotanical
332:
329:
303:middle Miocene
282:in the family
275:is an extinct
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233:A. browni
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541:
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531:A. nipponicum
528:
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523:A. nipponicum
520:
516:
512:
508:
504:
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498:A. nipponicum
493:
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473:paleobotany.
472:
468:
465:
464:specific name
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457:
453:
449:
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441:
437:
436:Jack A. Wolfe
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315:Graham Island
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242:Binomial name
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133:Tracheophytes
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26:
22:
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571:
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501:and as such
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214:
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165:
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139:
126:
24:
18:
483:Acer browni
477:Description
417:Smithsonian
389:Acer browni
366:Haida Gwaii
337:Acer browni
319:Haida Gwaii
284:Sapindaceae
272:Acer browni
254:Acer browni
192:Sapindaceae
146:Angiosperms
25:Acer browni
692:Categories
537:References
515:anastomise
507:Parviflora
481:Leaves of
341:Aquitanian
324:Parviflora
219:Parviflora
182:Sapindales
527:A. browni
511:A. browni
503:A. browni
492:A. browni
460:etymology
452:A. browni
425:Vancouver
397:paratypes
226:Species:
210:Section:
116:Kingdom:
618:10169419
471:Tertiary
393:holotype
188:Family:
159:Eudicots
667:Bibcode
458:. The
454:in the
438:of the
295:samaras
280:species
251:†
230:†
198:Genus:
178:Order:
120:Plantae
616:
519:sulcus
467:browni
307:Oregon
291:leaves
288:fossil
172:Rosids
698:Maple
646:: 65.
614:S2CID
407:, in
299:early
277:maple
261:Wolfe
217:sect.
215:Acer
166:Clade
153:Clade
140:Clade
127:Clade
644:1053
572:Acer
450:for
293:and
203:Acer
37:PreꞒ
675:doi
606:doi
602:266
380:in
368:of
347:in
301:to
694::
673:.
663:18
661:.
642:.
638:.
626:^
612:.
600:.
582:22
580:.
576:.
544:^
533:.
427:,
384:.
327:.
317:,
309:,
168::
155::
142::
129::
87:Pg
681:.
677::
669::
620:.
608::
92:N
82:K
77:J
72:T
67:P
62:C
57:D
52:S
47:O
42:Ꞓ
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