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Benthos

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In the long-term or at steady-state, i.e., the biomass of benthic organisms does not change, the benthic community can be considered a black box diverting organic matter into either metabolites or the geosphere (burial). The macrobenthos also indirectly impacts carbon cycling on the seafloor through
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which killed the dinosaurs 66 million years ago also rendered extinct three-quarters of all other animal and plant species. However, deep-sea benthic forams flourished in the aftermath. In 2020 it was reported that researchers have examined the chemical composition of thousands of samples of these
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Westerhold, Thomas; Marwan, Norbert; Drury, Anna Joy; Liebrand, Diederik; Agnini, Claudia; Anagnostou, Eleni; Barnet, James S. K.; Bohaty, Steven M.; De Vleeschouwer, David; Florindo, Fabio; Frederichs, Thomas; Hodell, David A.; Holbourn, Ann E.; Kroon, Dick; Lauretano, Vittoria; Littler, Kate;
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can attach themselves securely to hard, rocky substrates. As adults they can remain at the same site, shaping depressions and crevices where mobile animals find refuge. This greater diversity in benthic habitats has resulted in a higher diversity of benthic species. The number of benthic animal
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Willerslev, Eske; Froese, Duane; Gilichinsky, David; Rønn, Regin; Bunce, Michael; Zuber, Maria T.; Gilbert, M. Thomas P.; Brand, Tina; Munch, Kasper; Nielsen, Rasmus; Mastepanov, Mikhail; Christensen, Torben R.; Hebsgaard, Martin B.; Johnson, Sarah Stewart (4 September 2007).
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in the shells, since lighter isotopes evaporate more readily in warmer water leaving the heavier isotopes in the shells. Information about past climates can be inferred further from the abundance of forams and diatoms, since they tend to be more abundant in warm
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have extremely long lives. In 2013 researchers reported evidence of endoliths in the ocean floor, perhaps millions of years old, with a generation time of 10,000 years. These are slowly metabolizing and not in a dormant state. Some
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and organic detrital matter. In coastal locations, organic run off from land provides an additional food source. Meiofauna and bacteria consume and recycle organic matter in the sediments, playing an important role in returning
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comes from Greek, meaning "depth of the sea". Microbenthos are found everywhere on or about the seafloor of continental shelves, as well as in deeper waters, with greater diversity in or on seafloor sediments. In
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of river beds, where many benthos reside. Benthos are highly sensitive to contamination, so their close proximity to high pollutant concentrations make these organisms ideal for studying water contamination.
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Organic matter produced in the sunlit layer of the ocean and delivered to the sediments is either consumed by organisms or buried. The organic matter consumed by organisms is used to synthesize
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response of macro-invertebrates' internal tissues can be studied extensively in the laboratory. The concentration of a chemical can cause many changes, including changing feeding behaviors,
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The depth of water, temperature and salinity, and type of local substrate all affect what benthos is present. In coastal waters and other places where light reaches the bottom, benthic
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species exceeds one million. This far exceeds the number of pelagic animal species (about 5000 larger zooplankton species, 22,000 pelagic fish species and 110 marine mammal species).
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before it can reach deep ocean water, the energy source for deep benthic ecosystems is often organic matter from higher up in the water column that drifts down to the depths. This
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Damásio, Joana; Fernández-Sanjuan, Maria; Sánchez-Avila, Juan; Lacorte, Silvia; Prat, Narcís; Rieradevall, Maria; Soares, Amadeu M.V.M.; Barata, Carlos (June 2011).
865: 3181: 577:, the benthic zone offers physically diverse habitats. There is a huge range in how much light and warmth is available, and in the depth of water or extent of 2430:"Multi-biochemical responses of benthic macroinvertebrate species as a complementary tool to diagnose the cause of community impairment in polluted rivers" 1087:
or live on sediment at the bottom of the ocean. Regardless of form, their shells sink to the seafloor after they die. These shells are widely used as
2565:"Relative influences of bioturbation and physical mixing on degradation of bloom-derived particulate organic matter: Clue from microcosm experiments" 3211: 789: 'less', comprises tiny benthic organisms that are less than about 1 mm but greater than about 0.1 mm in size. Some examples are 2288:"Combining biomarker and community approaches using benthic macroinvertebrates can improve the assessment of the ecological status of rivers" 2703: 3785: 3051: 641: 'long', comprises the larger, visible to the naked eye, benthic organisms greater than about 1 mm in size. In shallow waters, 1841: 1000: 2796: 1942:
Lourens, Lucas J.; Lyle, Mitchell; Pälike, Heiko; Röhl, Ursula; Tian, Jun; Wilkens, Roy H.; Wilson, Paul A.; Zachos, James C. (2020).
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of the ocean – that live near or on the seafloor, or within or on surface seafloor sediments. The word
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data can be collected on the amount of and direct effect of specific pollutants in a waterbody. The
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Earth barreling toward 'Hothouse' state not seen in 50 million years, epic new climate record shows
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Haeckel, E. 1891. Plankton-Studien. Jenaische Zeitschrift fĂĽr Naturwissenschaft 25 / (Neue Folge)
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benthic forams and used their findings to build the most detailed climate record of Earth ever.
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Many organisms adapted to deep-water pressure cannot survive in the upper parts of the
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Benthos und das redundant Benton: Neologismen in der deutschsprachigen Limnologie
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US Department of Commerce, National Oceanic and Atmospheric Administration.
3814: 3727: 3506: 3486: 3390: 3128: 3011: 2931: 2891: 2881: 2851: 2530: 2506:"Reviews and syntheses: to the bottom of carbon processing at the seafloor" 2505: 2453: 2106: 2022: 1985: 1668: 1653: 1598: 1593: 1499: 1457: 1453: 1437: 1384: 1290: 1259: 1186: 1084: 974: 893: 756: 674: 628: 574: 525: 498: 491: 471: 348: 278: 216: 183: 149: 120: 110: 78: 3819: 3640: 3615: 3610: 3580: 3363: 3339: 3304: 3113: 3016: 2936: 2896: 2749: 1976: 1673: 1388: 1239: 1049: 1024: 983: 885: 843: 802: 778: 773: 658: 561:, such as lakes, rivers, and streams. There is also a redundant synonym, 173: 115: 2676: 2286:
Rodrigues, Carolina; GuimarĂŁes, Laura; Vieira, Natividade (2019-08-01).
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Benthos (organisms that live at the ocean floor) can be contrasted with
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Harris, Peter T. (2020-01-01), Harris, Peter T.; Baker, Elaine (eds.),
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Flows of Energy and Materials in Marine Ecosystems: Theory and Practice
1916:, Carleton College. Modified 23 July 2020. Retrieved 10 September 2020. 1689: 1361: 1357: 1341: 1202: 1175: 1076: 923: 871: 806: 798: 748: 733: 719: 678: 654: 646: 578: 521: 517: 479: 228: 178: 144: 3736: 3645: 3550: 3334: 3098: 3091: 3066: 2976: 2681: 2393:"Online Submission and Review for Science of the Total Environment". 2036:"It's Okay to be Smart • the oldest living thing in the world: These" 1559: 1515: 1445: 1345: 1315: 1007: 814: 752: 501:. The pressure difference can be very significant (approximately one 435: 251: 208: 42: 2649:
Te Ara - the Encyclopædia of New Zealand, updated 21 September 2007.
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Material was copied from this source, which is available under a
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through ecological population assessments or through analyzing
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can find protection and food in soft, loose sediments such as
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Seafloor Geomorphology as Benthic Habitat (Second Edition)
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Feeding and trophic structure of the deep-sea macrobenthos
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benthic diatoms dominate as photosynthetic organisms. In
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Lalli, Carol M.; Parsons, Timothy R. (1997). "Benthos".
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Marine microbenthos are microorganisms that live in the
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Creative Commons Attribution 4.0 International License
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immersion. The seafloor varies widely in the types of
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Benthic macro-invertebrates play a critical role in
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Community of organisms that live in the benthic zone
1909:"Paleoclimatology: How Can We Infer Past Climates?" 1486:(i.e. growth) converted to carbon dioxide through 557:to refer to organisms at the bottom of freshwater 2499: 2497: 2120:Smetacek, Victor (1984), Fasham, M. J. R. (ed.), 994:strongly control opportunities for microbenthos. 3873: 1714:from the Census of Antarctic Marine Life website 1587:(organisms that can swim against water currents) 1270:Hyperbenthos (or hyperbenthic), prefix from 2067:Proceedings of the National Academy of Sciences 1583:(organisms that drift with water currents) and 2494: 2063:"Ancient bacteria show evidence of DNA repair" 1934: 1796:"North American Benthological Society website" 1391:, play an important role as a food source for 1222:Endobenthos (or endobenthic), prefix from 1130:are estimated to be half a million years old. 3779: 2987:Stable isotope analysis in aquatic ecosystems 2697: 2128:, Boston, MA: Springer US, pp. 517–547, 1011:a widespread abundant genus of benthic forams 399: 1839: 1742: 1726:"How does pressure change with ocean depth?" 1250:Epibenthos (or epibenthic), prefix from 3052:Freshwater environmental quality parameters 1579:(organisms that live at the ocean surface) 1411: 3786: 3772: 2704: 2690: 2503: 2466: 1305:The main food sources for the benthos are 454:that live on, in, or near the bottom of a 406: 392: 244:        2529: 2161: 2096: 2086: 1975: 1344:, populate softer bottoms. Fish, such as 520:; most organisms in the benthic zone are 2563:Sun, Ming-Yi; Dai, Jihong (2005-09-01). 2335:"Indicators: Benthic Macroinvertebrates" 2219: 2119: 1874: 1843:Biological Oceanography: An Introduction 1833: 1569: 1364:are important predators and scavengers. 1289: 1137: 1098: 29: 2504:Middelburg, Jack J. (19 January 2018). 1819:Nehring, S. & Albrecht, U. (1997). 573:Compared to the relatively featureless 314:Hydrothermal vent microbial communities 14: 3874: 2711: 2652:Yip, Maricela and Madl, Pierre (1999) 2601: 2562: 2332: 1707: 1705: 1083:forms, that is, they can drift in the 3767: 3240: 3239: 2723: 2685: 1469:Carbon processing in marine sediments 505:for every 10 metres of water depth). 1463: 1028:, an extinct genus of benthic forams 902:Microbenthos, prefix from the Greek 442: 'the depths '), also known as 91:       3315:Oceanic physical-biological process 3177:List of freshwater ecoregions (WWF) 2122:"The Supply of Food to the Benthos" 1879:. Enfield, NH: Science Publishers. 1717: 1702: 1510:Benthos are negatively impacted by 1075:Both foraminifera and diatoms have 890:Marine sediment § Microbenthos 24: 2234:10.1016/B978-0-12-409548-9.11087-5 1406: 25: 3903: 2670: 2469:"The Biological Pump in the Past" 2467:Sigman, D.M.; Haug, G.H. (2003), 3748: 3747: 3735: 3721: 3353: 3025: 2842:Colored dissolved organic matter 2544: 2395:Science of the Total Environment 1852:10.1016/b978-075063384-0/50063-3 1473: 1057: 1038: 1016: 999: 959: 945: 931: 864: 850: 836: 822: 740: 726: 712: 698: 373: 63: 3187:Latin America and the Caribbean 2595: 2556: 2516:(2). Copernicus GmbH: 413–427. 2460: 2421: 2415:10.1016/j.scitotenv.2004.06.001 2386: 2351: 2326: 2279: 2255: 2213: 2189: 2155: 2113: 2053: 2028: 2012: 2000: 1919: 1285: 1265: 1181: 879: 622: 3669:Ecological values of mangroves 3212:North Pacific Subtropical Gyre 2475:, Elsevier, pp. 491–528, 2228:, Elsevier, pp. 660–662, 2226:Encyclopedia of Ocean Sciences 2170:, Elsevier, pp. 105–115, 1901: 1868: 1846:. Elsevier. pp. 177–195. 1813: 1788: 1754: 1736: 1217: 1212: 1156:by M. J. Schleiden (1804–1881) 767: 633:Macrobenthos, prefix from 13: 1: 2633: 2581:10.1016/j.marchem.2004.11.003 2372:10.1016/S0043-1354(05)00684-6 2220:Nunnally, Clifton C. (2019), 2162:Snelgrove, Paul V.R. (2013), 1245: 1161: 3689:Marine conservation activism 3674:Fisheries and climate change 2610:, Elsevier, pp. 35–61, 2446:10.1016/j.watres.2011.04.006 2168:Encyclopedia of Biodiversity 2134:10.1007/978-1-4757-0387-0_20 1166:Zoobenthos, prefix from 7: 3892:Oceanographical terminology 3684:Human impact on marine life 3561:Davidson Seamount § Ecology 2797:Aquatic population dynamics 1565: 1550:) and construction such as 1428:, and metals—settle in the 1367:Benthic organisms, such as 568: 51:Part of a series related to 10: 3908: 2304:10.1007/s10750-019-03991-7 1762: 1522:, oil and gas activities, 1505: 1133: 883: 771: 626: 617: 545: 428: 3839:Pechton / Pecton / Pekton 3805: 3715: 3654: 3526: 3462: 3424: 3371: 3362: 3351: 3300:Marine primary production 3252: 3248: 3235: 3194:List of marine ecoregions 3169: 3034: 3023: 2737: 2733: 2719: 2333:US EPA, OW (2013-11-21). 1538:(and its impacts such as 1178:, starfish and anemones. 1148:A variety of marine worms 3546:Coastal biogeomorphology 3541:Marine coastal ecosystem 2473:Treatise on Geochemistry 1907:Bruckner, Monica (2020) 1875:Sokolova, M. N. (2000). 1695: 1684:Sediment Profile Imagery 1412:Benthos as bioindicators 1067:Pliciloricus enigmaticus 539:in 1891, comes from the 514:dead and decaying matter 476:sedimentary environments 3454:Paradox of the plankton 3265:Diel vertical migration 3159:Freshwater swamp forest 2877:GIS and aquatic science 2725:General components and 2401:(1–3): 1. August 2004. 2201:Encyclopedia Britannica 2088:10.1073/pnas.0706787104 1968:10.1126/science.aba6853 1778:A Greek–English Lexicon 1436:Benthos can be used as 886:Seabed § Sediments 490:, and then down to the 3280:Large marine ecosystem 2972:Shoaling and schooling 2664:University of Salzburg 2531:10.5194/bg-15-413-2018 2366:(20): II. 2005-12-01. 2019:Sussman: Oldest Plants 1743: 1589: 1302: 1300:on marine benthic life 1158: 1106: 380:Marine life portal 45: 3699:Marine protected area 3626:Salt pannes and pools 3401:Marine larval ecology 3376:Census of Marine Life 3260:Deep scattering layer 3217:San Francisco Estuary 3182:Africa and Madagascar 3007:Underwater camouflage 2787:Aquatic biomonitoring 2727:freshwater ecosystems 2647:"Benthic communities" 1769:Liddell, Henry George 1730:oceanservice.NOAA.gov 1573: 1293: 1141: 1102: 516:sustains the benthic 339:Van Veen grab sampler 33: 3434:Marine bacteriophage 3396:Marine invertebrates 1931:, 10 September 2020. 1624:Deep sea communities 1397:California sheephead 549:'depth of the sea'. 470:, also known as the 299:Deep sea communities 3887:Ecology terminology 3795:Groups of organisms 3310:Ocean fertilization 3119:Trophic state index 3077:Lake stratification 2807:Aquatic respiration 2645:Ryan, Paddy (2007) 2522:2018BGeo...15..413M 2407:2004ScTEn.329....1. 2079:2007PNAS..10414401J 2073:(36): 14401–14405. 1960:2020Sci...369.1383W 1954:(6509): 1383–1387. 1552:coastal development 1540:ocean acidification 1234:top layer, e.g., a 1034:Marine microanimals 3799:aquatic ecosystems 3576:Intertidal wetland 3571:Intertidal ecology 3439:Marine prokaryotes 3381:Deep-sea community 3275:Iron fertilization 3198:Specific examples 3124:Upland and lowland 3042:Freshwater biology 2907:Microbial food web 2817:Aquatic toxicology 2760:Aquatic adaptation 2713:Aquatic ecosystems 2659:2019-07-20 at the 2358:"Water Research". 2164:"Marine Sediments" 1644:Intertidal ecology 1604:Benthopelagic fish 1590: 1418:aquatic ecosystems 1303: 1189:, prefix from 1159: 1144:Example zoobenthos 1107: 781:, prefix from 555:freshwater biology 309:Intertidal ecology 284:Benthopelagic fish 205:    46: 3869: 3868: 3761: 3760: 3742:Oceans portal 3711: 3710: 3707: 3706: 3586:Hydrothermal vent 3522: 3521: 3411:Seashore wildlife 3242:Marine ecosystems 3231: 3230: 3227: 3226: 2997:Thermal pollution 2962:Ramsar Convention 2902:Microbial ecology 2862:Fisheries science 2802:Aquatic predation 2617:978-0-12-814960-7 2482:978-0-08-043751-4 2440:(12): 3599–3613. 2243:978-0-12-813082-7 2177:978-0-12-384720-1 2143:978-1-4757-0387-0 1886:978-1-57808-090-8 1548:ocean circulation 1464:Carbon processing 1328:can proliferate. 1323:photosynthesizing 587:Burrowing animals 508:Because light is 488:continental shelf 416: 415: 260:hydrothermal vent 16:(Redirected from 3899: 3882:Marine organisms 3788: 3781: 3774: 3765: 3764: 3751: 3750: 3744: 3740: 3739: 3730: 3728:Lakes portal 3726: 3725: 3724: 3694:Marine pollution 3386:Deep-water coral 3369: 3368: 3357: 3290:Marine chemistry 3250: 3249: 3237: 3236: 3154:Freshwater marsh 3047:Freshwater biome 3029: 2745:Acoustic ecology 2735: 2734: 2721: 2720: 2706: 2699: 2692: 2683: 2682: 2627: 2626: 2625: 2624: 2599: 2593: 2592: 2569:Marine Chemistry 2560: 2554: 2548: 2543: 2533: 2501: 2492: 2491: 2490: 2489: 2464: 2458: 2457: 2425: 2419: 2418: 2390: 2384: 2383: 2355: 2349: 2348: 2346: 2345: 2330: 2324: 2323: 2283: 2277: 2276: 2274: 2273: 2259: 2253: 2252: 2251: 2250: 2217: 2211: 2210: 2208: 2207: 2193: 2187: 2186: 2185: 2184: 2159: 2153: 2152: 2151: 2150: 2117: 2111: 2110: 2100: 2090: 2057: 2051: 2050: 2048: 2047: 2038:. Archived from 2032: 2026: 2016: 2010: 2004: 1998: 1997: 1979: 1938: 1932: 1923: 1917: 1905: 1899: 1898: 1872: 1866: 1865: 1837: 1831: 1817: 1811: 1810: 1808: 1807: 1798:. Archived from 1792: 1786: 1765: 1764: 1758: 1752: 1746: 1740: 1734: 1733: 1721: 1715: 1709: 1532:invasive species 1477: 1318:to the pelagic. 1262:or a sea snail. 1176:polychaete worms 1111:extinction event 1061: 1042: 1020: 1003: 988:intertidal zones 963: 949: 935: 868: 854: 840: 826: 744: 730: 716: 702: 647:polychaete worms 643:seagrass meadows 548: 547: 439: 432: 408: 401: 394: 378: 377: 376: 258: 245: 235: 223: 206: 92: 77: 67: 66: 48: 47: 21: 3907: 3906: 3902: 3901: 3900: 3898: 3897: 3896: 3872: 3871: 3870: 3865: 3801: 3792: 3762: 3757: 3734: 3733: 3722: 3720: 3719: 3703: 3664:Coral bleaching 3650: 3631:Seagrass meadow 3528:Marine habitats 3518: 3492:Coral reef fish 3458: 3444:Marine protists 3420: 3358: 3349: 3320:Ocean turbidity 3295:Marine food web 3244: 3223: 3165: 3104:River ecosystem 3057:Freshwater fish 3030: 3021: 2827:Bioluminescence 2812:Aquatic science 2729: 2715: 2710: 2673: 2661:Wayback Machine 2636: 2631: 2630: 2622: 2620: 2618: 2600: 2596: 2561: 2557: 2502: 2495: 2487: 2485: 2483: 2465: 2461: 2426: 2422: 2392: 2391: 2387: 2357: 2356: 2352: 2343: 2341: 2331: 2327: 2284: 2280: 2271: 2269: 2267:archive.epa.gov 2261: 2260: 2256: 2248: 2246: 2244: 2218: 2214: 2205: 2203: 2195: 2194: 2190: 2182: 2180: 2178: 2160: 2156: 2148: 2146: 2144: 2118: 2114: 2058: 2054: 2045: 2043: 2034: 2033: 2029: 2017: 2013: 2005: 2001: 1939: 1935: 1924: 1920: 1906: 1902: 1887: 1873: 1869: 1862: 1838: 1834: 1818: 1814: 1805: 1803: 1794: 1793: 1789: 1783:Perseus Project 1759: 1755: 1741: 1737: 1722: 1718: 1710: 1703: 1698: 1639:Epibenthic sled 1629:Deep sea mining 1588: 1568: 1556:undersea cables 1546:and changes to 1520:deep-sea mining 1508: 1492:biological pump 1478: 1466: 1442:water pollution 1414: 1409: 1407:Ecological role 1301: 1299: 1288: 1268: 1248: 1220: 1215: 1184: 1164: 1157: 1155: 1150: 1146: 1136: 1105: 1093:oxygen isotopes 1089:climate proxies 1071: 1062: 1053: 1046:Darkfield photo 1043: 1029: 1021: 1012: 1004: 969: 967:Dinoflagellates 964: 955: 950: 941: 936: 900: 882: 875: 869: 860: 855: 846: 841: 832: 827: 776: 770: 763: 745: 736: 731: 722: 717: 708: 703: 631: 625: 620: 603:Sessile species 571: 559:bodies of water 412: 374: 372: 365: 364: 363: 359:Marine sediment 334:Epibenthic sled 328: 320: 319: 318: 273: 265: 264: 263: 262: 256: 254: 243: 240: 239: 237:seagrass meadow 233: 231: 225:mangrove forest 221: 219: 204: 200: 192: 191: 190: 166: 158: 157: 156: 137: 129: 128: 127: 103: 90: 81: 75: 64: 28: 23: 22: 15: 12: 11: 5: 3905: 3895: 3894: 3889: 3884: 3867: 3866: 3864: 3863: 3858: 3853: 3850: 3845: 3840: 3837: 3832: 3827: 3824: 3823: 3822: 3817: 3806: 3803: 3802: 3791: 3790: 3783: 3776: 3768: 3759: 3758: 3756: 3755: 3745: 3731: 3716: 3713: 3712: 3709: 3708: 3705: 3704: 3702: 3701: 3696: 3691: 3686: 3681: 3676: 3671: 3666: 3660: 3658: 3652: 3651: 3649: 3648: 3643: 3638: 3633: 3628: 3623: 3618: 3613: 3608: 3603: 3598: 3593: 3588: 3583: 3578: 3573: 3568: 3563: 3558: 3553: 3548: 3543: 3538: 3532: 3530: 3524: 3523: 3520: 3519: 3517: 3516: 3511: 3510: 3509: 3504: 3499: 3494: 3489: 3482:Saltwater fish 3479: 3477:Marine reptile 3474: 3468: 3466: 3460: 3459: 3457: 3456: 3451: 3449:Marine viruses 3446: 3441: 3436: 3430: 3428: 3426:Microorganisms 3422: 3421: 3419: 3418: 3416:Wild fisheries 3413: 3408: 3403: 3398: 3393: 3388: 3383: 3378: 3372: 3366: 3360: 3359: 3352: 3350: 3348: 3347: 3342: 3337: 3332: 3330:Thorson's rule 3327: 3322: 3317: 3312: 3307: 3302: 3297: 3292: 3287: 3285:Marine biology 3282: 3277: 3272: 3267: 3262: 3256: 3254: 3246: 3245: 3233: 3232: 3229: 3228: 3225: 3224: 3222: 3221: 3220: 3219: 3214: 3209: 3204: 3196: 3191: 3190: 3189: 3184: 3173: 3171: 3167: 3166: 3164: 3163: 3162: 3161: 3156: 3151: 3146: 3144:Brackish marsh 3141: 3131: 3126: 3121: 3116: 3111: 3106: 3101: 3096: 3095: 3094: 3084: 3079: 3074: 3072:Lake ecosystem 3069: 3064: 3062:Hyporheic zone 3059: 3054: 3049: 3044: 3038: 3036: 3032: 3031: 3024: 3022: 3020: 3019: 3014: 3009: 3004: 2999: 2994: 2989: 2984: 2979: 2974: 2969: 2964: 2959: 2954: 2949: 2944: 2939: 2934: 2929: 2924: 2919: 2914: 2912:Microbial loop 2909: 2904: 2899: 2894: 2889: 2884: 2879: 2874: 2869: 2864: 2859: 2857:Eutrophication 2854: 2849: 2844: 2839: 2837:Cascade effect 2834: 2829: 2824: 2819: 2814: 2809: 2804: 2799: 2794: 2789: 2784: 2783: 2782: 2777: 2772: 2765:Aquatic animal 2762: 2757: 2752: 2747: 2741: 2739: 2731: 2730: 2717: 2716: 2709: 2708: 2701: 2694: 2686: 2680: 2679: 2672: 2671:External links 2669: 2668: 2667: 2650: 2643: 2635: 2632: 2629: 2628: 2616: 2594: 2575:(3): 201–218. 2555: 2510:Biogeosciences 2493: 2481: 2459: 2434:Water Research 2420: 2385: 2360:Water Research 2350: 2325: 2278: 2254: 2242: 2212: 2188: 2176: 2154: 2142: 2112: 2052: 2027: 2011: 1999: 1933: 1918: 1900: 1885: 1867: 1860: 1832: 1812: 1787: 1753: 1735: 1716: 1700: 1699: 1697: 1694: 1693: 1692: 1687: 1681: 1679:Profundal zone 1676: 1671: 1666: 1661: 1656: 1651: 1646: 1641: 1636: 1631: 1626: 1621: 1616: 1611: 1606: 1601: 1596: 1574: 1567: 1564: 1562:construction. 1536:climate change 1507: 1504: 1480: 1479: 1472: 1470: 1465: 1462: 1426:surface runoff 1413: 1410: 1408: 1405: 1395:, such as the 1330:Filter feeders 1297:eutrophication 1294: 1287: 1284: 1267: 1264: 1247: 1244: 1219: 1216: 1214: 1211: 1183: 1180: 1163: 1160: 1142: 1135: 1132: 1124:Actinomycetota 1104:Benthic diatom 1103: 1073: 1072: 1063: 1056: 1054: 1044: 1037: 1035: 1031: 1030: 1022: 1015: 1013: 1005: 998: 971: 970: 965: 958: 956: 951: 944: 942: 937: 930: 881: 878: 877: 876: 870: 863: 861: 856: 849: 847: 842: 835: 833: 828: 821: 795:foraminiferans 772:Main article: 769: 766: 765: 764: 746: 739: 737: 732: 725: 723: 718: 711: 709: 704: 697: 627:Main article: 624: 621: 619: 616: 570: 567: 492:abyssal depths 414: 413: 411: 410: 403: 396: 388: 385: 384: 383: 382: 367: 366: 362: 361: 356: 351: 346: 341: 336: 330: 329: 326: 325: 322: 321: 317: 316: 311: 306: 301: 296: 291: 286: 281: 275: 274: 271: 270: 267: 266: 255: 242: 241: 232: 220: 203: 202: 201: 198: 197: 194: 193: 189: 188: 187: 186: 181: 176: 168: 167: 164: 163: 160: 159: 155: 154: 153: 152: 147: 139: 138: 135: 134: 131: 130: 126: 125: 124: 123: 118: 113: 105: 104: 101: 100: 97: 96: 69: 68: 60: 59: 53: 52: 26: 9: 6: 4: 3: 2: 3904: 3893: 3890: 3888: 3885: 3883: 3880: 3879: 3877: 3862: 3859: 3857: 3854: 3851: 3849: 3846: 3844: 3841: 3838: 3836: 3833: 3831: 3828: 3825: 3821: 3818: 3816: 3813: 3812: 3811: 3808: 3807: 3804: 3800: 3796: 3789: 3784: 3782: 3777: 3775: 3770: 3769: 3766: 3754: 3746: 3743: 3738: 3732: 3729: 3718: 3717: 3714: 3700: 3697: 3695: 3692: 3690: 3687: 3685: 3682: 3680: 3677: 3675: 3672: 3670: 3667: 3665: 3662: 3661: 3659: 3657: 3653: 3647: 3644: 3642: 3639: 3637: 3636:Sponge ground 3634: 3632: 3629: 3627: 3624: 3622: 3619: 3617: 3614: 3612: 3609: 3607: 3604: 3602: 3601:Marine biomes 3599: 3597: 3594: 3592: 3589: 3587: 3584: 3582: 3579: 3577: 3574: 3572: 3569: 3567: 3564: 3562: 3559: 3557: 3554: 3552: 3549: 3547: 3544: 3542: 3539: 3537: 3534: 3533: 3531: 3529: 3525: 3515: 3512: 3508: 3505: 3503: 3502:Demersal fish 3500: 3498: 3497:Deep-sea fish 3495: 3493: 3490: 3488: 3485: 3484: 3483: 3480: 3478: 3475: 3473: 3472:Marine mammal 3470: 3469: 3467: 3465: 3461: 3455: 3452: 3450: 3447: 3445: 3442: 3440: 3437: 3435: 3432: 3431: 3429: 3427: 3423: 3417: 3414: 3412: 3409: 3407: 3404: 3402: 3399: 3397: 3394: 3392: 3389: 3387: 3384: 3382: 3379: 3377: 3374: 3373: 3370: 3367: 3365: 3361: 3356: 3346: 3343: 3341: 3338: 3336: 3333: 3331: 3328: 3326: 3323: 3321: 3318: 3316: 3313: 3311: 3308: 3306: 3303: 3301: 3298: 3296: 3293: 3291: 3288: 3286: 3283: 3281: 3278: 3276: 3273: 3271: 3268: 3266: 3263: 3261: 3258: 3257: 3255: 3251: 3247: 3243: 3238: 3234: 3218: 3215: 3213: 3210: 3208: 3205: 3203: 3200: 3199: 3197: 3195: 3192: 3188: 3185: 3183: 3180: 3179: 3178: 3175: 3174: 3172: 3168: 3160: 3157: 3155: 3152: 3150: 3147: 3145: 3142: 3140: 3137: 3136: 3135: 3132: 3130: 3127: 3125: 3122: 3120: 3117: 3115: 3112: 3110: 3107: 3105: 3102: 3100: 3097: 3093: 3090: 3089: 3088: 3085: 3083: 3080: 3078: 3075: 3073: 3070: 3068: 3065: 3063: 3060: 3058: 3055: 3053: 3050: 3048: 3045: 3043: 3040: 3039: 3037: 3033: 3028: 3018: 3015: 3013: 3010: 3008: 3005: 3003: 3002:Trophic level 3000: 2998: 2995: 2993: 2990: 2988: 2985: 2983: 2980: 2978: 2975: 2973: 2970: 2968: 2967:Sediment trap 2965: 2963: 2960: 2958: 2955: 2953: 2950: 2948: 2945: 2943: 2942:Phytoplankton 2940: 2938: 2935: 2933: 2930: 2928: 2925: 2923: 2920: 2918: 2915: 2913: 2910: 2908: 2905: 2903: 2900: 2898: 2895: 2893: 2890: 2888: 2885: 2883: 2880: 2878: 2875: 2873: 2870: 2868: 2865: 2863: 2860: 2858: 2855: 2853: 2850: 2848: 2845: 2843: 2840: 2838: 2835: 2833: 2830: 2828: 2825: 2823: 2820: 2818: 2815: 2813: 2810: 2808: 2805: 2803: 2800: 2798: 2795: 2793: 2792:Aquatic plant 2790: 2788: 2785: 2781: 2778: 2776: 2773: 2771: 2768: 2767: 2766: 2763: 2761: 2758: 2756: 2755:Anoxic waters 2753: 2751: 2748: 2746: 2743: 2742: 2740: 2736: 2732: 2728: 2722: 2718: 2714: 2707: 2702: 2700: 2695: 2693: 2688: 2687: 2684: 2678: 2675: 2674: 2665: 2662: 2658: 2655: 2651: 2648: 2644: 2641: 2638: 2637: 2619: 2613: 2609: 2605: 2598: 2590: 2586: 2582: 2578: 2574: 2570: 2566: 2559: 2552: 2547: 2541: 2537: 2532: 2527: 2523: 2519: 2515: 2511: 2507: 2500: 2498: 2484: 2478: 2474: 2470: 2463: 2455: 2451: 2447: 2443: 2439: 2435: 2431: 2424: 2416: 2412: 2408: 2404: 2400: 2396: 2389: 2381: 2377: 2373: 2369: 2365: 2361: 2354: 2340: 2336: 2329: 2321: 2317: 2313: 2309: 2305: 2301: 2297: 2293: 2292:Hydrobiologia 2289: 2282: 2268: 2264: 2258: 2245: 2239: 2235: 2231: 2227: 2223: 2216: 2202: 2198: 2192: 2179: 2173: 2169: 2165: 2158: 2145: 2139: 2135: 2131: 2127: 2123: 2116: 2108: 2104: 2099: 2094: 2089: 2084: 2080: 2076: 2072: 2068: 2064: 2056: 2042:on 2018-07-13 2041: 2037: 2031: 2024: 2020: 2015: 2009: 2003: 1995: 1991: 1987: 1983: 1978: 1977:11577/3351324 1973: 1969: 1965: 1961: 1957: 1953: 1949: 1945: 1937: 1930: 1927: 1922: 1915: 1914: 1910: 1904: 1896: 1892: 1888: 1882: 1878: 1871: 1863: 1861:9780750633840 1857: 1853: 1849: 1845: 1844: 1836: 1829: 1826: 1822: 1816: 1802:on 2008-07-05 1801: 1797: 1791: 1784: 1780: 1779: 1774: 1773:Scott, Robert 1770: 1766: 1757: 1750: 1747:18: 232-336. 1745: 1739: 1731: 1727: 1720: 1713: 1708: 1706: 1701: 1691: 1688: 1685: 1682: 1680: 1677: 1675: 1672: 1670: 1667: 1665: 1662: 1660: 1657: 1655: 1652: 1650: 1647: 1645: 1642: 1640: 1637: 1635: 1634:Demersal fish 1632: 1630: 1627: 1625: 1622: 1620: 1617: 1615: 1614:Bottom feeder 1612: 1610: 1609:Bioirrigation 1607: 1605: 1602: 1600: 1597: 1595: 1592: 1591: 1586: 1582: 1578: 1572: 1563: 1561: 1557: 1553: 1549: 1545: 1544:ocean warming 1541: 1537: 1533: 1529: 1525: 1521: 1517: 1513: 1503: 1501: 1495: 1493: 1489: 1485: 1476: 1471: 1468: 1467: 1461: 1459: 1455: 1451: 1447: 1443: 1439: 1438:bioindicators 1434: 1431: 1427: 1423: 1422:concentration 1419: 1404: 1402: 1398: 1394: 1390: 1386: 1385:brittle stars 1382: 1381:sea cucumbers 1378: 1374: 1370: 1365: 1363: 1359: 1355: 1351: 1348:, as well as 1347: 1343: 1339: 1335: 1331: 1327: 1324: 1319: 1317: 1313: 1308: 1307:phytoplankton 1298: 1292: 1283: 1281: 1277: 1273: 1272:Ancient Greek 1263: 1261: 1257: 1253: 1252:Ancient Greek 1243: 1241: 1237: 1233: 1229: 1225: 1224:Ancient Greek 1210: 1208: 1204: 1200: 1196: 1192: 1191:Ancient Greek 1188: 1179: 1177: 1173: 1169: 1168:Ancient Greek 1154: 1149: 1145: 1140: 1131: 1129: 1125: 1120: 1115: 1112: 1101: 1097: 1094: 1090: 1086: 1082: 1078: 1069: 1068: 1060: 1055: 1051: 1047: 1041: 1036: 1033: 1032: 1027: 1026: 1019: 1014: 1010: 1009: 1002: 997: 996: 995: 993: 989: 985: 980: 976: 968: 962: 957: 954: 948: 943: 940: 934: 929: 928: 927: 925: 921: 917: 913: 909: 905: 899: 898:bioirrigation 895: 891: 887: 873: 867: 862: 859: 853: 848: 845: 839: 834: 831: 830:Foraminiferan 825: 820: 819: 818: 816: 812: 808: 804: 803:gastrotriches 800: 796: 792: 788: 784: 783:Ancient Greek 780: 775: 762: 758: 754: 750: 743: 738: 735: 729: 724: 721: 715: 710: 707: 701: 696: 695: 694: 692: 688: 684: 680: 676: 675:turbellarians 672: 668: 664: 660: 656: 652: 648: 644: 640: 636: 635:Ancient Greek 630: 615: 612: 608: 604: 600: 596: 592: 588: 584: 580: 576: 566: 564: 560: 556: 552: 542: 538: 534: 529: 527: 523: 519: 515: 511: 506: 504: 500: 495: 493: 489: 486:, out to the 485: 481: 477: 473: 469: 465: 461: 457: 453: 449: 445: 441: 438: 431: 427: 424: 423:Ancient Greek 420: 409: 404: 402: 397: 395: 390: 389: 387: 386: 381: 371: 370: 369: 368: 360: 357: 355: 354:Demersal zone 352: 350: 347: 345: 344:Bioirrigation 342: 340: 337: 335: 332: 331: 324: 323: 315: 312: 310: 307: 305: 304:Demersal fish 302: 300: 297: 295: 292: 290: 289:Bottom feeder 287: 285: 282: 280: 277: 276: 269: 268: 261: 253: 249: 238: 230: 226: 218: 214: 210: 196: 195: 185: 182: 180: 177: 175: 172: 171: 170: 169: 162: 161: 151: 148: 146: 143: 142: 141: 140: 133: 132: 122: 119: 117: 114: 112: 109: 108: 107: 106: 99: 98: 95: 88: 84: 83:Benthopelagic 80: 74: 71: 70: 62: 61: 58: 55: 54: 50: 49: 44: 40: 36: 32: 19: 3815:Macrobenthos 3809: 3656:Conservation 3507:Pelagic fish 3487:Coastal fish 3391:Marine fungi 3129:Water garden 3012:Water column 2957:Productivity 2932:Pelagic zone 2892:Macrobenthos 2882:Hydrobiology 2852:Ecohydrology 2821: 2621:, retrieved 2607: 2597: 2572: 2568: 2558: 2513: 2509: 2486:, retrieved 2472: 2462: 2437: 2433: 2423: 2398: 2394: 2388: 2363: 2359: 2353: 2342:. Retrieved 2338: 2328: 2295: 2291: 2281: 2270:. Retrieved 2266: 2257: 2247:, retrieved 2225: 2215: 2204:. Retrieved 2200: 2191: 2181:, retrieved 2167: 2157: 2147:, retrieved 2125: 2115: 2070: 2066: 2055: 2044:. Retrieved 2040:the original 2030: 2025:, 2 May 2010 2023:The Guardian 2014: 2002: 1951: 1947: 1936: 1928: 1921: 1911: 1903: 1876: 1870: 1842: 1835: 1825:Lauterbornia 1824: 1820: 1815: 1804:. Retrieved 1800:the original 1790: 1776: 1756: 1738: 1729: 1719: 1669:Pelagic zone 1654:Neritic zone 1599:Benthic fish 1594:Aphotic zone 1518:and litter, 1509: 1500:bioturbation 1496: 1481: 1458:inflammation 1450:quantitative 1435: 1415: 1389:sea anemones 1366: 1320: 1304: 1286:Food sources 1275: 1269: 1266:Hyperbenthos 1260:sea cucumber 1255: 1249: 1227: 1221: 1194: 1187:Phytobenthos 1185: 1182:Phytobenthos 1171: 1165: 1152: 1147: 1143: 1116: 1108: 1085:water column 1074: 1065: 1023: 1006: 984:photic zones 978: 975:benthic zone 972: 903: 901: 894:bioturbation 880:Microbenthos 874:(water bear) 805:and smaller 786: 777: 659:sea anemones 638: 632: 629:Macrobenthos 623:Macrobenthos 575:pelagic zone 572: 562: 550: 535:, coined by 532: 530: 526:detritivores 507: 499:water column 496: 472:benthic zone 443: 436: 433: 426: 418: 417: 349:Bioturbation 279:Benthic fish 217:bivalve reef 184:hyperbenthos 150:phytobenthos 121:microbenthos 111:macrobenthos 79:Benthic zone 72: 57:Benthic life 56: 3820:Meiobenthos 3641:Sponge reef 3616:Rocky shore 3611:Oyster reef 3581:Kelp forest 3464:Vertebrates 3364:Marine life 3340:Viral shunt 3305:Marine snow 3207:Maharashtra 3114:Stream pool 3017:Zooplankton 2937:Photic zone 2897:Meiobenthos 2750:Algal bloom 2339:www.epa.gov 2298:(1): 1–24. 2008:29 Aug 2013 1929:LiveScience 1827:31: 17-30, 1674:Photic zone 1488:respiration 1454:biochemical 1362:crustaceans 1358:cephalopods 1342:polychaetes 1240:sand dollar 1218:Endobenthos 1213:By location 1151:Plate from 1109:The sudden 1050:gastrotrich 1025:Heterohelix 924:flagellates 844:Gastrotrich 807:crustaceans 799:tardigrades 779:Meiobenthos 774:Meiobenthos 768:Meiobenthos 759:larvae and 734:Sea squirts 720:Echinoderms 679:crustaceans 677:and larger 671:sea squirts 655:echinoderms 585:it offers. 553:is used in 480:tidal pools 272:Communities 174:endobenthos 165:By location 116:meiobenthos 3876:Categories 3621:Salt marsh 3556:Coral reef 3345:Whale fall 3325:Photophore 3202:Everglades 3170:Ecoregions 3109:Stream bed 3082:Macrophyte 3035:Freshwater 2867:Food chain 2780:Water bird 2640:"Benthos". 2634:References 2623:2024-09-24 2488:2024-09-22 2344:2022-10-14 2272:2022-10-12 2249:2024-09-23 2206:2024-09-23 2183:2024-09-23 2149:2024-09-23 2046:2018-07-13 2006:Bob Yirka 1806:2008-08-16 1690:Stream bed 1446:biomarkers 1332:, such as 1295:Effect of 1246:Epibenthos 1232:oxygenated 1203:macroalgae 1162:Zoobenthos 1077:planktonic 884:See also: 872:Tardigrade 815:ostracodes 579:intertidal 522:scavengers 518:food chain 503:atmosphere 482:along the 421:(from 229:coral reef 199:By habitat 179:epibenthos 145:zoobenthos 18:Zoobenthos 3646:Tide pool 3551:Cold seep 3335:Upwelling 3099:Rheotaxis 3092:Fish pond 3067:Limnology 2992:Substrate 2977:Siltation 2847:Dead zone 2677:"Benthos" 2654:"Benthos" 2589:0304-4203 2540:1726-4189 2380:0043-1354 2320:186207664 2312:1573-5117 2197:"Benthos" 1994:221593388 1560:wind farm 1516:pollution 1369:sea stars 1350:sea stars 1346:dragonets 1316:phosphate 1126:found in 1119:endoliths 1064:Armoured 1008:Elphidium 990:changing 791:nematodes 761:amphipods 691:cumaceans 611:barnacles 531:The term 484:foreshore 452:organisms 448:community 446:, is the 252:cold seep 209:tide pool 207:Shallow: 43:tide pool 3848:Pleuston 3843:Plankton 3753:Category 3679:HERMIONE 3596:Mangrove 3406:Seagrass 2952:Pleuston 2947:Plankton 2927:Particle 2872:Food web 2657:Archived 2454:21571352 2107:17728401 1986:32913105 1895:46724477 1664:Plankton 1649:Littoral 1619:Deep sea 1581:plankton 1566:See also 1528:shipping 1430:sediment 1338:bivalves 1280:rock cod 1153:Das Meer 916:ciliates 908:bacteria 811:copepods 809:such as 747:Aquatic 706:Seagrass 687:lobsters 681:such as 651:bivalves 605:such as 583:sediment 569:Overview 510:absorbed 294:Deep sea 248:seamount 87:coupling 37:and two 3861:Tripton 3835:Neuston 3810:Benthos 3606:Mudflat 3566:Estuary 3536:Bay mud 3514:Seabird 3270:f-ratio 3253:General 3134:Wetland 2922:Neuston 2887:Hypoxia 2832:Biomass 2822:Benthos 2738:General 2518:Bibcode 2403:Bibcode 2098:1958816 2075:Bibcode 1956:Bibcode 1948:Science 1781:at the 1712:Benthos 1577:neuston 1524:tourism 1512:fishing 1506:Threats 1484:biomass 1373:oysters 1334:sponges 1326:diatoms 1312:nitrate 1236:sea pen 1207:seaweed 1199:diatoms 1134:By type 1128:Siberia 1081:benthic 979:benthos 953:Ciliate 939:Diatoms 912:diatoms 858:Copepod 667:sponges 618:By size 607:oysters 551:Benthos 537:Haeckel 533:benthos 478:, from 444:benthon 437:bĂ©nthos 419:Benthos 327:Related 213:bay mud 136:By type 102:By size 73:Benthos 39:chitons 35:Seaweed 3856:Seston 3852:Plocon 3830:Nekton 3826:Herpon 3591:Lagoon 2917:Nekton 2775:Mammal 2770:Insect 2614:  2587:  2538:  2479:  2452:  2378:  2318:  2310:  2240:  2174:  2140:  2105:  2095:  1992:  1984:  1893:  1883:  1858:  1763:βένθος 1659:Nekton 1585:nekton 1558:, and 1401:humans 1399:, and 1360:, and 1354:snails 1195:phutĂłn 1096:water. 920:amoeba 904:mikrĂłs 896:, and 663:corals 639:makrĂłs 563:Benton 546:βένθος 468:stream 430:βένθος 246:Deep: 94:Seabed 2982:Spawn 2316:S2CID 1990:S2CID 1696:Notes 1686:(SPI) 1377:clams 1276:hupĂ©r 1274: 1254: 1238:or a 1228:Ă©ndon 1226: 1193: 1172:zĂ´ion 1170: 1117:Some 1048:of a 992:tides 787:meĂ®on 785: 757:midge 753:snail 683:crabs 637: 543:noun 541:Greek 466:, or 460:river 425: 41:in a 3087:Pond 2612:ISBN 2585:ISSN 2536:ISSN 2477:ISBN 2450:PMID 2376:ISSN 2308:ISSN 2238:ISBN 2172:ISBN 2138:ISBN 2103:PMID 1982:PMID 1913:SERC 1891:OCLC 1881:ISBN 1856:ISBN 1393:fish 1387:and 1336:and 1314:and 1201:and 1079:and 813:and 749:worm 689:and 609:and 599:sand 597:and 595:clay 464:lake 3797:in 3149:Fen 3139:Bog 2577:doi 2526:doi 2442:doi 2411:doi 2399:329 2368:doi 2300:doi 2296:839 2230:doi 2130:doi 2093:PMC 2083:doi 2071:104 1972:hdl 1964:doi 1952:369 1848:doi 1749:BHL 1440:of 1256:epĂ­ 1209:). 591:mud 524:or 456:sea 450:of 3878:: 2606:, 2583:. 2573:96 2571:. 2567:. 2534:. 2524:. 2514:15 2512:. 2508:. 2496:^ 2471:, 2448:. 2438:45 2436:. 2432:. 2409:. 2397:. 2374:. 2364:39 2362:. 2337:. 2314:. 2306:. 2294:. 2290:. 2265:. 2236:, 2224:, 2199:. 2166:, 2136:, 2124:, 2101:. 2091:. 2081:. 2069:. 2065:. 2021:, 1988:. 1980:. 1970:. 1962:. 1950:. 1946:. 1889:. 1854:. 1823:. 1775:; 1771:; 1767:. 1728:. 1704:^ 1554:, 1542:, 1534:, 1530:, 1526:, 1514:, 1502:. 1494:. 1403:. 1383:, 1379:, 1375:, 1371:, 1356:, 1352:, 1282:. 1242:. 926:. 922:, 918:, 914:, 910:, 892:, 888:, 817:. 801:, 797:, 793:, 755:, 751:, 693:. 685:, 673:, 669:, 665:, 661:, 657:, 653:, 649:, 601:. 593:, 565:. 528:. 494:. 462:, 458:, 250:• 227:• 215:• 211:• 89:) 3787:e 3780:t 3773:v 2705:e 2698:t 2691:v 2666:. 2591:. 2579:: 2553:. 2542:. 2528:: 2520:: 2456:. 2444:: 2417:. 2413:: 2405:: 2382:. 2370:: 2347:. 2322:. 2302:: 2275:. 2232:: 2209:. 2132:: 2109:. 2085:: 2077:: 2049:. 1996:. 1974:: 1966:: 1958:: 1897:. 1864:. 1850:: 1830:. 1809:. 1785:. 1751:. 1732:. 1205:( 440:) 434:( 407:e 400:t 393:v 85:( 20:)

Index

Zoobenthos

Seaweed
chitons
tide pool
Benthic life
Benthos
Benthic zone
Benthopelagic
coupling
Seabed
macrobenthos
meiobenthos
microbenthos
zoobenthos
phytobenthos
endobenthos
epibenthos
hyperbenthos
tide pool
bay mud
bivalve reef
mangrove forest
coral reef
seagrass meadow
seamount
cold seep
hydrothermal vent
Benthic fish
Benthopelagic fish

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