106:
674:
Sendi, H.; Le Tirant, S.; Palková, H.; Chorvát, D.; Šurka, J.; Cumming, R. (2023). "Umenocoleidae (Insecta: Dictyoptera) from
Turonian sediments of Kzyl-Zhar, Kazakhstan and Cenomanian northern Myanmar amber".
543:"Beetle or roach: systematic position of the enigmatic Umenocoleidae based on new material from Zhonggou Formation in Jiuquan, Northwest China, and a morphocladistic analysis"
610:
Luo, Cihang; Beutel, Rolf G.; Engel, Michael S.; Liang, Kun; Li, Liqin; Li, Jiahao; Xu, Chunpeng; Vršanský, Peter; Jarzembowski, Edmund A.; Wang, Bo (February 2022).
300:
403:
243:
514:
500:
492:
435:
421:
339:
332:
760:
465:
412:
391:
379:
483:
367:
293:
278:
443:
430:
347:
308:
236:
474:
456:
263:
505:
747:
814:
541:
Liuo, C.-H.; Beutel, R. G.; Thomson, U. R; Zheng, D.-R.; Li, J.-H.; Zhao, X.-Y.; Zhang, H.-C.; Wang, B. (2021).
649:
434:
Burmese amber, Myanmar, mid
Cretaceous (latest Albian-earliest Cenomanian) (Also placed in Alienopteridae or
105:
222:-like forewings. This was probably an adaptation for living under bark and in other tight spaces.
738:
612:"Life history and evolution of the enigmatic Cretaceous–Eocene Alienopteridae: A critical review"
611:
791:
700:
786:
778:
765:
425:
Sendi et al. 2023 Burmese amber, Myanmar, mid
Cretaceous (latest Albian-earliest Cenomanian)
416:
Sendi et al. 2023 Burmese amber, Myanmar, mid
Cretaceous (latest Albian-earliest Cenomanian)
407:
Sendi et al. 2023 Burmese amber, Myanmar, mid
Cretaceous (latest Albian-earliest Cenomanian)
819:
215:. They were originally considered to be beetles due to their beetle-like morphology, with
8:
631:
564:
324:
320:
251:
100:
773:
635:
568:
627:
680:
623:
554:
285:
723:
559:
542:
270:
684:
583:
316:
212:
160:
808:
355:
178:
59:
398:
Burmese amber, Myanmar, mid
Cretaceous (latest Albian-earliest Cenomanian)
386:
Burmese amber, Myanmar, mid
Cretaceous (latest Albian-earliest Cenomanian)
374:
Burmese amber, Myanmar, mid
Cretaceous (latest Albian-earliest Cenomanian)
732:
204:
34:
752:
359:
208:
79:
44:
216:
137:
117:
84:
694:
717:
460:
Sendi et al. 2023 Kzyl-Zhar, Kazakhstan, Late
Cretaceous (Turonian)
448:
74:
69:
54:
49:
39:
219:
89:
64:
584:"A new family of Coleoptera from the Lower Cretaceous of Kansu"
255:
247:
147:
127:
673:
447:
Sendi et al. 2023 Kzyl-Zhar, Kazakhstan, Late
Cretaceous (
540:
609:
354:Crato Formation, Brazil, Early Cretaceous (Aptian)
211:. They are considered to be closely related to the
358:, Myanmar, mid Cretaceous (latest Albian-earliest
667:
806:
558:
581:
807:
699:
698:
396:Luo, Beutel, Xu et Jarzembowski, 2021
605:
603:
601:
536:
534:
521:have been excluded from the family.
323:, Japan, Early Cretaceous (Aptian)
273:, Brazil, Early Cretaceous (Aptian)
13:
327:, China, Early Cretaceous (Albian)
288:, China, Early Cretaceous (Aptian)
14:
831:
598:
531:
104:
32:
628:10.1016/j.earscirev.2021.103914
519:Nel, Prokop et Kirejtshuk, 2014
372:Mlynský, Vršanský et Wang, 2018
315:Dzun-Bain Formation, Mongolia,
642:
575:
225:
1:
582:Chen, S.; Tan, C. C. (1973).
524:
384:Luo, Xu et Jarzembowski, 2021
16:Extinct family of cockroaches
650:"Fossilworks: Umenocoleidae"
560:10.1016/j.palwor.2021.01.003
7:
815:Prehistoric insect families
342:, Early Cretaceous (Albian)
254:, China, Early Cretaceous (
246:, China, Early Cretaceous (
10:
836:
685:10.1007/s11756-023-01356-z
470:Vršanský and Grimaldi 2003
268:Vršanský and Grimaldi 1999
707:
499:have been transferred to
191:
186:
101:Scientific classification
99:
23:
588:Acta Entomologica Sinica
203:is an extinct family of
207:insects known from the
787:Paleobiology Database
616:Earth-Science Reviews
301:Dzun-Bain Formation
404:Archaeospinapteryx
325:Zhonggou Formation
321:Kitadani Formation
303:, Mongolia, Aptian
252:Zhonggou Formation
30:Barremian–Turonian
802:
801:
774:Open Tree of Life
701:Taxon identifiers
520:
511:
498:
489:
480:
471:
397:
385:
373:
353:
338:
314:
299:
284:
269:
242:
241:Chen and Tan 1973
198:
197:
182:
827:
795:
794:
782:
781:
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768:
756:
755:
743:
742:
741:
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538:
518:
509:
496:
487:
478:
469:
395:
383:
371:
351:
336:
312:
297:
286:Xiagou Formation
282:
267:
244:Dalazi Formation
240:
177:
172:
159:
109:
108:
94:
31:
27:Temporal range:
21:
20:
835:
834:
830:
829:
828:
826:
825:
824:
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804:
803:
798:
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772:
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751:
746:
737:
736:
731:
722:
721:
716:
703:
693:
692:
672:
668:
658:
656:
654:fossilworks.org
648:
647:
643:
608:
599:
580:
576:
539:
532:
527:
515:Permoponopterix
501:Cratovitismidae
493:Antophiloblatta
436:Cratovitismidae
422:Poikiloprosopon
340:Jordanian amber
333:Compunctiotypus
271:Crato Formation
228:
176:
170:
157:
103:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
833:
823:
822:
817:
800:
799:
797:
796:
783:
770:
757:
744:
729:
713:
711:
705:
704:
691:
690:
666:
641:
597:
574:
529:
528:
526:
523:
462:
461:
452:
439:
426:
417:
408:
399:
387:
375:
363:
343:
328:
317:Zaza Formation
304:
289:
274:
259:
227:
224:
213:Alienopteridae
196:
195:
189:
188:
184:
183:
168:
164:
163:
161:Umenocoleoidea
155:
151:
150:
145:
141:
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135:
131:
130:
125:
121:
120:
115:
111:
110:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
26:
15:
9:
6:
4:
3:
2:
832:
821:
818:
816:
813:
812:
810:
793:
788:
784:
780:
775:
771:
767:
762:
758:
754:
749:
745:
740:
739:Umenocoleidae
734:
730:
725:
719:
715:
714:
712:
710:
709:Umenocoleidae
706:
702:
697:
686:
682:
678:
670:
655:
651:
645:
637:
633:
629:
625:
621:
617:
613:
606:
604:
602:
594:(2): 169–179.
593:
589:
585:
578:
570:
566:
561:
556:
552:
548:
544:
537:
535:
530:
522:
517:
516:
510:Vršanský 1999
508:
507:
502:
495:
494:
486:
485:
479:Koubová, 2020
477:
476:
468:
467:
466:Jantaropterix
459:
458:
453:
450:
446:
445:
440:
437:
433:
432:
427:
424:
423:
418:
415:
414:
413:Trapezionotum
409:
406:
405:
400:
394:
393:
392:Laticephalana
388:
382:
381:
380:Enervipraeala
376:
370:
369:
364:
361:
357:
356:Burmese amber
350:
349:
344:
341:
335:
334:
329:
326:
322:
318:
313:Vršanský 2003
311:
310:
305:
302:
298:Vršanský 2003
296:
295:
290:
287:
283:Vršanský 2003
281:
280:
275:
272:
266:
265:
260:
257:
253:
249:
245:
239:
238:
233:
232:
231:
223:
221:
218:
214:
210:
206:
202:
201:Umenocoleidae
194:
190:
185:
180:
175:
174:Umenocoleidae
169:
166:
165:
162:
156:
154:Superfamily:
153:
152:
149:
146:
143:
142:
139:
136:
133:
132:
129:
126:
123:
122:
119:
116:
113:
112:
107:
102:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
24:Umenocoleidae
22:
19:
708:
676:
669:
657:. Retrieved
653:
644:
619:
615:
591:
587:
577:
550:
546:
513:
504:
491:
484:Lepidopterix
482:
473:
464:
463:
455:
442:
429:
420:
411:
402:
390:
378:
368:Alienopterix
366:
346:
337:Kaddumi 2005
331:
307:
294:Elytropterix
292:
279:Blattapterix
277:
262:
235:
229:
205:dictyopteran
200:
199:
192:
181:and Tan 1973
173:
18:
820:Dictyoptera
733:Wikispecies
659:17 December
553:: 121–130.
547:Palaeoworld
497:Sendi, 2020
488:Sendi, 2020
444:Nigropterix
431:Vzrkadlenie
348:Umenopterix
309:Petropterix
237:Umenocoleus
226:Systematics
217:sclerotised
809:Categories
622:: 103914.
525:References
475:Perspicuus
457:Umenotypus
360:Cenomanian
319:, Russia,
264:Ponopterix
209:Cretaceous
138:Arthropoda
724:Q21366583
636:245601183
569:234208413
193:See text
124:Kingdom:
118:Eukaryota
718:Wikidata
677:Biologia
503:, while
449:Turonian
352:Lee 2016
167:Family:
134:Phylum:
128:Animalia
114:Domain:
779:4664010
753:4801412
506:Vitisma
187:Genera
148:Insecta
144:Class:
792:227708
766:111841
634:
567:
256:Albian
248:Aptian
230:After
220:elytra
761:IRMNG
632:S2CID
565:S2CID
748:GBIF
661:2021
512:and
490:and
179:Chen
35:Preęž’
681:doi
624:doi
620:225
555:doi
811::
789::
776::
763::
750::
735::
720::
679:.
652:.
630:.
618:.
614:.
600:^
592:16
590:.
586:.
563:.
551:31
549:.
545:.
533:^
481:,
472:,
428:†?
250:)
85:Pg
687:.
683::
663:.
638:.
626::
571:.
557::
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410:â€
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306:â€
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234:â€
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158:â€
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80:K
75:J
70:T
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60:C
55:D
50:S
45:O
40:ęž’
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