538:
4273:
517:
skull and partial postcranium), and SC 57256 (isolated rib), all held at the MCSNT. Each specimen was deposited in a different level of the geologic lens, so they each lived at different times; the lens is thought to have been deposited over less than 10,000 years, meaning the individuals lived a maximum of a few thousand years apart from each other. Della
Vacchia published an additional description of the species, focusing on its tail anatomy specifically, in 2020, which was mostly confuted in following studies. Bruno, along with five other specimens, was formally described in 2021 by Alfio Alessandro Chiarenza and colleagues.
4268:
3358:
639:
3372:
4255:
3365:
142:
4279:
472:
120:
526:
454:
specimen was publicly presented. It was at this point that the specimen gained the nickname "Antonio", given to it in press reports. In 2004, a book on dinosaurs in Italy by
Cristiano Dal Sasso was published which dedicated a chapter to the subject of the hadrosaurs from Trieste. The Villagio del Pescatore site is one of very traces of dinosaurs ever found in the country. The only others are the single exceptional specimen of the tiny theropod
4262:
587:
2312:
2300:
368:
569:, or anatomical features entirely unique to this species. Specifically, it was originally described with the skull possessed seven such traits, the vertebral column seven, and the limbs six. Much of this peculiar anatomy was thought to have evolved as adaptations for the unique type of environment it lived in. Among other European relatives, Romanian specimens potentially belonging to
393:
Many other fossils, including various other hadrosaur specimens, were uncovered in the process. The main skeleton was not extracted until 1999. The significance of the find was immediately recognized, being the oldest hadrosaur known from the Europe and the most complete large dinosaur skeleton found on the continent since the 19th century discovery of
392:
skeleton was discovered in 1994. Lying on a vertical rockface, the specimen required a difficult excavation process, involving the removal of over 300 cubic metres of mineral and use of large equipment. Palaeontologist Fabio Della
Vacchia among others served as scientific director for the excavation.
553:
of its bones. Bruno, which is 15–20% larger than
Antonio, was found to be an adult, and it shows features related to maturity such as a more robust skull and stouter proportions. Antonio has been estimated at 338 kilograms (745 lb) in weight, while Bruno has been estimated at 514–584 kilograms
516:
through the winter of that year and 2020. Numerous other specimens from the site were also referred to the species; SC 57022 (partial forelimbs; potentially from a more extensive unrecovered skeleton), SC 57023 (isolated left pubis), SC 57025 (isolated cervical vertebra and rib), SC 57026 (complete
548:
was originally thought to be a dwarf hadrosauroid. The preserved portion of the holotype measures 3.6 metres (12 ft), with only the end of the tail being missing. The length of the tail including the unpreserved portion was later estimated to be 2.5 metres (8.2 ft), resulting in a total
453:
capable of dissolving the surrounding rock but not the fossil, which had a different chemical composition. Such spraying treatments were done in hour long sessions, followed by rinsing and preservation of removed bones with a special resin. This process was finished in
December 2000, when the
444:
Upon the completion of excavation, the five tonne block containing the specimen proved too large for transportation. As such, it was separated into six separate blocks; these blocks were then further reduced in size by cutting off sections of rock containing none of the fossil. Once finally
603:
anatomy is its intermediary nature – it possesses a mixture of primitive features and derived features. This is thought to be because these features were acquired gradually in the hadrosaur lineage, rather than all appearing around the same time. As such,
449:, though it was catalogued as property of the Italian State (SI) rather than under the museum. There, the specimen underwent 2800 hours of preparation (the process of removing the fossil from the rock matrix). This done was done through spraying diluted
549:
estimated body length of 4.5 metres (15 ft) for the animal, though it's been argued this is an overestimate. Although the holotype
Antonio was initially thought to represent an adult, it was later assessed to be a subadult individual based on the
511:
was found at this site, SC 57247, nicknamed "Bruno". It consists of a complete skull and extensive postcranial remains. "Bruno" was not prepared until 2019, at which point images were posted online and the specimen was publicly displayed at the
383:
Sometime in the 1980s, Alceo Tarlao and
Giorgio Rimoli reported finding fragments of dinosaur bone while prospecting for rare bones. The abandoned quarry these were found in was only 100m inland, at Villagio del Pescatore,
2253:"The late-surviving early diverging Ibero-Armorican 'duck-billed' dinosaur Fylax and the role of the Late Cretaceous European Archipelago in hadrosauroid biogeography"
541:"Antonio" is the most complete Tethyshadros insularis specimen known, while "Bruno" is the largest (as of May 26th 2024, these facts may change in the future).
1969:
Chiarenza, Alfio
Alessandro; Fabbri, Matteo; Consorti, Lorenzo; Muscioni, Marco; Evans, David C.; Cantalapiedra, Juan L.; Fanti, Federico (2 December 2021).
270:
408:
342:
608:
split form the main lineage at a point where only some of the advanced traits had evolved. In some ways the species more resembles earlier relatives like
1169:
as its closest relative. Other close relatives were generally Asian or North
American species. Modelling of body size evolution suggested that
2365:
4769:
437:
boundary, around 70 million years ago., an age recently re-evaluated by more recent studies to 81.5-80.5 million years ago and to the
416:
537:
4774:
4272:
3357:
2344:
4267:
3371:
616:
503:, a reference to the fact that species would have lived on one of the larger islands of the European Archipelago. The
2044:
4254:
4633:
507:
of the genus is SC 57021 (Antonio), a mostly complete but crushed skeleton. An additional extensive skeleton of
4656:
4643:
1931:
4784:
4648:
668:, making it one of the latest surviving non-hadrosaurid hadrosauromorphs. Their cladogram is shown below:
4278:
2035:
Dal Sasso, Cristiano; Brillante, Giuseppe (2004). "'Antonio' and the
Hadrosaurs of the Triestine Karst".
141:
2290:
4779:
2337:
407:, measuring ten metres thick and seventy metres in diameter. This lens were tougth to pertain to the
3364:
2444:
622:
bone, and comparatively short neck. In others, it is more comparable to the condition seen in true
445:
recovered, it was brought to the Museo Civico di Storia Naturale di Trieste (MCSNT) in the city of
1971:"An Italian dinosaur Lagerstätte reveals the tempo and mode of hadrosauriform body size evolution"
646:
Before the species was described and named, the specimens were assumed to belong to a species of
496:
4741:
4691:
2803:
2655:
4736:
4613:
3169:
2471:
1158:
294:
575:
show the most similar anatomy. Despite this, many differences between the two genera exist;
4723:
4033:
2330:
2223:
2142:
2096:
1982:
1936:
379:
was noted as the most complete large dinosaur skeleton discovered in Europe since this find
4261:
8:
4764:
638:
627:
438:
338:
2227:
2146:
2100:
1986:
4385:
3996:
2581:
2274:
2112:
2003:
1970:
136:
4431:
2154:
4392:
4131:
4052:
4012:
2896:
2782:
2728:
2699:
2278:
2116:
2040:
2008:
565:
has overall unique anatomy compared to its relatives. It possesses a large number of
2183:(Dinosauria, Hadrosauroidea) from the Adriatic island of the European archipelago".
4533:
4526:
4336:
4329:
3858:
3769:
3607:
3508:
3494:
3208:
3176:
2811:
2720:
2634:
2569:
2304:
2264:
2231:
2192:
2150:
2104:
1998:
1990:
1336:
675:
665:
590:
385:
250:
2087:, a new hadrosauroid dinosaur (Ornithischia) from the Upper Cretaceous of Italy".
4618:
4413:
4364:
4227:
4158:
4061:
3881:
3600:
3586:
3565:
3183:
3056:
2823:
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1314:
882:
617:
32:
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4623:
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4371:
4220:
4183:
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3480:
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2982:
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2404:
2196:
1994:
1726:
1525:
1494:
1180:
827:
566:
492:
461:
404:
334:
218:
561:
was a beaked animal with a hand formed into a fleshy pad and three-toed feet.
4758:
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3120:
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2795:
2706:
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2555:
2210:
Benson, R. B. J.; Hunt, G.; Carrano, M.T.; Campione, N.; Mannion, P. (2018).
1742:
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899:
651:
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70:
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3228:
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3070:
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2742:
2735:
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2456:
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2316:
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2252:
2012:
1680:
1470:
1022:
858:
803:
513:
488:
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421:
412:
326:
202:
4665:
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4519:
4357:
4199:
4172:
4124:
4117:
3950:
3936:
3908:
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3811:
3790:
3783:
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3710:
3703:
3656:
3621:
3593:
3529:
3522:
3291:
3141:
3127:
3099:
3077:
3019:
2882:
2713:
2677:
2641:
2537:
2524:
2483:
2353:
2108:
1632:
1398:
1374:
1242:
1046:
737:
623:
580:
479:
The genus was named and described by Dalla Vecchia in 2009. Its only and
450:
234:
119:
45:
2212:"Cope's rule and the adaptive landscape of dinosaur body size evolution"
4607:
4582:
4575:
4547:
4540:
4420:
4306:
4005:
3985:
3943:
3818:
3670:
3537:
3487:
3452:
3445:
3416:
3256:
3134:
3106:
2975:
2948:
2923:
2763:
2756:
2749:
2627:
2236:
2211:
1542:
1422:
1297:
941:
400:
90:
55:
4589:
4505:
4484:
4322:
4019:
3957:
3466:
3438:
3298:
3162:
3092:
3063:
2889:
2875:
2618:
1218:
610:
550:
525:
456:
430:
395:
389:
153:
95:
39:
4685:
1173:
was not significantly different in size from its closest relatives.
4708:
4469:
4138:
4026:
3826:
3797:
3724:
3649:
3614:
3515:
3501:
3459:
3283:
3084:
3049:
2955:
2941:
2903:
2413:
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1350:
1266:
720:
696:
504:
425:
186:
173:
85:
80:
65:
60:
50:
2039:. Bloomington, Indiana: Indina University Press. pp. 99–113.
586:
429:, in addition to geologic data, the hadrosaur fossil dated to the
4728:
3762:
3197:
465:
446:
403:, Belgium. The find was embedded in a fossil-bearing chalkstone
346:
128:
100:
75:
3628:
3221:
2371:
1968:
1446:
779:
163:
2322:
3473:
3003:
972:
660:
650:, due to its lack of crest and assumed nature as a member of
500:
367:
350:
330:
2250:
2841:
2133:
Dal Sasso, Cristiano (January 2003). "Dinosaurs of Italy".
2209:
388:, Italy. It was from this quarry that a nearly complete
2251:
Prieto-Marquez, Albert; Carrera Farias, Miguel (2021).
654:. In 2021, Prieto-Márquez and Carrera Farias recovered
626:, such as is seen with its hand morphology, lack of an
614:, such as in its general skull shape, possession of a
475:
Duino Castle, where the "Bruno" specimen was displayed
2288:
2178:
127:
Cast of the holotype skeleton at Museo Capellini of
468:, and some footprints belonging to various groups.
2179:Della Vacchia, F. M. (2020). "The unusual tail of
2034:
4756:
2185:Rivista Italiana di Paleontologia e Stratigrafia
18:Hadrosauroid dinosaur from Late Cretaceous Italy
1964:
1962:
1960:
1958:
1956:
1954:
1952:
375:skeletons discovered in 1882; the discovery of
2428:
2078:
2076:
3400:
2338:
2082:
2074:
2072:
2070:
2068:
2066:
2064:
2062:
2060:
2058:
2056:
1949:
460:(discovered one year earlier than Antonio),
341:(Previously tought to come from the younger
2128:
2126:
2535:
2345:
2331:
2053:
2030:
2028:
2026:
2024:
2022:
118:
2268:
2235:
2203:
2132:
2002:
2123:
637:
585:
536:
524:
470:
366:
2019:
362:
4757:
2174:
2172:
2170:
2168:
2166:
2164:
499:means "insular" or "of the island" in
4690:
4689:
4294:
3387:
2522:
2364:
2326:
630:, and teeth arranged into batteries.
4295:
4770:Late Cretaceous dinosaurs of Europe
2161:
1165:in a similar position, except with
13:
3388:
2089:Journal of Vertebrate Paleontology
14:
4796:
1157:Chiarenza et al 2021 performed a
633:
4277:
4271:
4266:
4260:
4253:
3370:
3363:
3356:
2310:
2298:
140:
43:
4634:List of informal dinosaur names
2523:
2352:
664:, in a derived position in the
487:. The genus name refers to the
353:. The type and only species is
4644:Timeline of hadrosaur research
2244:
2083:Dalla Vecchia, F. M. (2009). "
1932:Timeline of hadrosaur research
1161:of Bruno. This analysis found
520:
1:
4775:Fossil taxa described in 2009
2257:Acta Palaeontologica Polonica
2155:10.1016/S1631-0683(03)00007-1
1942:
596:Another prominent aspect of
7:
1925:
411:. Based on the presence of
10:
4801:
1995:10.1038/s41598-021-02490-x
4698:
4599:
4450:
4430:
4305:
4301:
4290:
4251:
4182:
4148:
4092:
4051:
3995:
3980:
3971:
3898:
3857:
3750:
3741:
3691:
3680:
3411:
3396:
3383:
3354:
3316:
3273:
3216:
3207:
3044:
3033:
2965:
2936:
2932:
2921:
2858:
2822:
2790:
2781:
2617:
2580:
2550:
2546:
2531:
2518:
2482:
2464:
2455:
2437:
2424:
2360:
1740:
1723:
1717:
1700:
1694:
1677:
1670:
1653:
1646:
1629:
1622:
1596:
1579:
1572:
1565:
1539:
1522:
1515:
1508:
1491:
1484:
1467:
1460:
1443:
1436:
1419:
1412:
1395:
1388:
1371:
1364:
1347:
1340:
1311:
1294:
1287:
1280:
1263:
1256:
1239:
1232:
1215:
1208:
1191:
1184:
1060:
1043:
1036:
1019:
1012:
986:
969:
962:
955:
938:
931:
896:
879:
872:
855:
848:
841:
824:
817:
800:
793:
776:
769:
734:
717:
710:
693:
686:
679:
373:Iguanodon bernissartensis
300:
293:
137:Scientific classification
135:
126:
117:
25:
2197:10.13130/2039-4942/14075
658:as the sister taxon to
554:(1,133–1,287 lb).
2270:10.4202/app.00821.2020
2181:Tethyshadros insularis
2135:Comptes Rendus Palevol
2085:Tethyshadros insularis
643:
593:
542:
534:
485:Tethyshadros insularis
476:
380:
307:Tethyshadros insularis
4737:Paleobiology Database
4614:Basturs Poble bonebed
1159:phylogenetic analysis
641:
589:
540:
533:compared with a human
528:
474:
370:
4785:Ornithischian genera
4636:(contains hadrosaur
4034:Probrachylophosaurus
2109:10.1671/039.029.0428
1937:2009 in paleontology
615:
417:Murciella cuvillieri
363:Discovery and naming
329:hadrosauroid") is a
2228:2018Palgy..61...13B
2147:2003CRPal...2...45D
2101:2009JVPal..29.1100D
1987:2021NatSR..1123295C
628:antorbital fenestra
579:is more similar to
439:Calcare di Aurisina
339:Calcare di Aurisina
313:Dalla Vecchia, 2009
4386:Ornithopodichnites
3997:Brachylophosaurini
2582:Hypsilophodontidae
2237:10.1111/pala.12329
2037:Dinosaurs of Italy
1975:Scientific Reports
644:
594:
543:
535:
477:
409:Liburnia Formation
381:
343:Liburnia Formation
337:dinosaur from the
4752:
4751:
4692:Taxon identifiers
4683:
4682:
4679:
4678:
4675:
4674:
4393:Shiraminesauropus
4286:
4285:
4249:
4248:
4245:
4244:
4241:
4240:
4237:
4236:
4132:Naashoibitosaurus
4053:Austrokritosauria
4013:Brachylophosaurus
3967:
3966:
3379:
3378:
3352:
3351:
3348:
3347:
3344:
3343:
3340:
3339:
3336:
3335:
3029:
3028:
2917:
2916:
2913:
2912:
2897:Transylvanosaurus
2783:Rhabdodontomorpha
2729:Notohypsilophodon
2700:Macrogryphosaurus
2514:
2513:
2510:
2509:
2506:
2505:
2502:
2501:
1922:
1921:
1913:
1912:
1904:
1903:
1895:
1894:
1886:
1885:
1877:
1876:
1868:
1867:
1859:
1858:
1850:
1849:
1841:
1840:
1832:
1831:
1823:
1822:
1814:
1813:
1805:
1804:
1796:
1795:
1787:
1786:
1778:
1777:
1769:
1768:
1761:
1760:
1753:
1752:
1611:
1610:
1554:
1553:
1326:
1325:
1154:
1153:
1145:
1144:
1136:
1135:
1127:
1126:
1118:
1117:
1109:
1108:
1100:
1099:
1091:
1090:
1082:
1081:
1073:
1072:
1001:
1000:
920:
919:
911:
910:
758:
757:
749:
748:
642:specimen SC 57247
318:
317:
286:T. insularis
274:
4792:
4780:Fossils of Italy
4745:
4744:
4732:
4731:
4719:
4718:
4717:
4687:
4686:
4660:mummy S.M. R4036
4534:Microhadrosaurus
4527:Mandschurosaurus
4337:Camptosaurichnus
4330:Bonaparteichnium
4303:
4302:
4296:Related articles
4292:
4291:
4281:
4275:
4270:
4265:
4264:
4257:
3978:
3977:
3859:Parasaurolophini
3770:Angulomastacator
3748:
3747:
3689:
3688:
3608:Shuangmiaosaurus
3509:Huehuecanauhtlus
3495:Gongpoquansaurus
3409:
3408:
3398:
3397:
3385:
3384:
3374:
3368:
3367:
3360:
3214:
3213:
3209:Hadrosauriformes
3177:Ratchasimasaurus
3042:
3041:
2934:
2933:
2930:
2929:
2812:Muttaburrasaurus
2788:
2787:
2721:Muttaburrasaurus
2635:Atlascopcosaurus
2570:Thescelosauridae
2548:
2547:
2544:
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2130:
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2120:
2095:(4): 1100–1116.
2080:
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2050:
2032:
2017:
2016:
2006:
1966:
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1342:
1337:Hadrosauromorpha
1290:
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1235:
1234:
1211:
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1038:
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712:
689:
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682:
681:
676:Hadrosauromorpha
672:
671:
666:Hadrosauromorpha
619:
602:
591:Life restoration
557:As a hadrosaur,
386:Trieste Province
309:
305:
269:
262:
251:Hadrosauromorpha
249:
233:
217:
201:
145:
144:
122:
112:
42:
31:Temporal range:
23:
22:
4800:
4799:
4795:
4794:
4793:
4791:
4790:
4789:
4755:
4754:
4753:
4748:
4740:
4735:
4727:
4722:
4713:
4712:
4707:
4694:
4684:
4671:
4652:mummy AMNH 5060
4619:Dakota (fossil)
4595:
4446:
4426:
4414:Tetrapodosaurus
4365:Hadrosaurichnus
4297:
4282:
4259:
4233:
4228:Shantungosaurus
4178:
4159:Augustynolophus
4144:
4088:
4062:Bonapartesaurus
4047:
3991:
3963:
3894:
3882:Parasaurolophus
3853:
3737:
3685:
3676:
3601:Qianjiangsaurus
3587:Probactrosaurus
3566:Penelopognathus
3405:
3392:
3375:
3362:
3332:
3312:
3269:
3203:
3184:Riabininohadros
3057:Bayannurosaurus
3038:
3025:
2961:
2926:
2909:
2854:
2824:Tenontosauridae
2818:
2777:
2613:
2592:Gideonmantellia
2576:
2540:
2527:
2498:
2478:
2451:
2445:Avemetatarsalia
2430:Avemetatarsalia
2420:
2367:
2366:
2356:
2351:
2321:
2311:
2309:
2299:
2297:
2289:
2287:
2286:
2249:
2245:
2208:
2204:
2177:
2162:
2131:
2124:
2081:
2054:
2047:
2033:
2020:
1967:
1950:
1945:
1928:
1923:
1914:
1905:
1896:
1887:
1878:
1869:
1860:
1851:
1842:
1833:
1824:
1815:
1806:
1797:
1788:
1779:
1770:
1612:
1555:
1327:
1315:Probactrosaurus
1155:
1146:
1137:
1128:
1119:
1110:
1101:
1092:
1083:
1074:
1002:
921:
912:
883:Penelopognathus
759:
750:
636:
621:
600:
523:
371:One of several
365:
314:
311:
303:
302:
289:
268:
260:
247:
231:
215:
199:
139:
113:
111:
110:
109:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
38:81.5–80.5
37:
36:
33:Late Cretaceous
29:
19:
12:
11:
5:
4798:
4788:
4787:
4782:
4777:
4772:
4767:
4750:
4749:
4747:
4746:
4733:
4720:
4704:
4702:
4696:
4695:
4681:
4680:
4677:
4676:
4673:
4672:
4670:
4669:
4662:
4654:
4646:
4641:
4631:
4629:Hypsilophodont
4626:
4624:Hadrosaur diet
4621:
4616:
4611:
4603:
4601:
4600:Other articles
4597:
4596:
4594:
4593:
4586:
4579:
4572:
4565:
4562:Stephanosaurus
4558:
4551:
4544:
4537:
4530:
4523:
4516:
4509:
4502:
4499:Fulgurotherium
4495:
4488:
4481:
4473:
4466:
4458:
4456:
4448:
4447:
4445:
4444:
4436:
4434:
4428:
4427:
4425:
4424:
4417:
4410:
4403:
4396:
4389:
4382:
4375:
4372:Iguanodontipus
4368:
4361:
4354:
4347:
4340:
4333:
4326:
4319:
4311:
4309:
4299:
4298:
4288:
4287:
4284:
4283:
4252:
4250:
4247:
4246:
4243:
4242:
4239:
4238:
4235:
4234:
4232:
4231:
4224:
4221:Laiyangosaurus
4217:
4210:
4203:
4196:
4188:
4186:
4184:Edmontosaurini
4180:
4179:
4177:
4176:
4169:
4166:Prosaurolophus
4162:
4154:
4152:
4146:
4145:
4143:
4142:
4135:
4128:
4121:
4114:
4111:Coahuilasaurus
4107:
4098:
4096:
4090:
4089:
4087:
4086:
4079:
4072:
4065:
4057:
4055:
4049:
4048:
4046:
4045:
4037:
4030:
4023:
4016:
4009:
4001:
3999:
3993:
3992:
3990:
3989:
3981:
3975:
3969:
3968:
3965:
3964:
3962:
3961:
3954:
3947:
3940:
3933:
3926:
3919:
3912:
3904:
3902:
3896:
3895:
3893:
3892:
3885:
3878:
3871:
3863:
3861:
3855:
3854:
3852:
3851:
3844:
3837:
3830:
3823:
3815:
3808:
3801:
3794:
3787:
3780:
3773:
3766:
3759:
3751:
3745:
3743:Lambeosaurinae
3739:
3738:
3736:
3735:
3728:
3721:
3714:
3707:
3700:
3692:
3686:
3681:
3678:
3677:
3675:
3674:
3667:
3660:
3653:
3646:
3639:
3632:
3625:
3618:
3611:
3604:
3597:
3590:
3583:
3576:
3569:
3562:
3559:Nanyangosaurus
3555:
3552:Nanningosaurus
3548:
3541:
3534:
3526:
3519:
3512:
3505:
3498:
3491:
3484:
3481:Gilmoreosaurus
3477:
3470:
3463:
3456:
3449:
3442:
3435:
3427:
3420:
3412:
3406:
3403:Hadrosauroidea
3401:
3394:
3393:
3390:Hadrosauroidea
3381:
3380:
3377:
3376:
3355:
3353:
3350:
3349:
3346:
3345:
3342:
3341:
3338:
3337:
3334:
3333:
3331:
3330:
3329:
3328:
3322:
3320:
3318:Hadrosauroidea
3314:
3313:
3311:
3310:
3306:Mantellisaurus
3302:
3295:
3288:
3279:
3277:
3275:Iguanodontidae
3271:
3270:
3268:
3267:
3260:
3253:
3250:Mantellisaurus
3246:
3239:
3232:
3225:
3217:
3211:
3205:
3204:
3202:
3201:
3194:
3187:
3180:
3173:
3166:
3159:
3152:
3145:
3138:
3131:
3124:
3117:
3114:Iguanacolossus
3110:
3103:
3096:
3089:
3081:
3074:
3067:
3060:
3053:
3045:
3039:
3036:Ankylopollexia
3034:
3031:
3030:
3027:
3026:
3024:
3023:
3016:
3008:
3000:
2997:Eousdryosaurus
2993:
2986:
2983:Dysalotosaurus
2979:
2971:
2969:
2963:
2962:
2960:
2959:
2952:
2945:
2937:
2927:
2922:
2919:
2918:
2915:
2914:
2911:
2910:
2908:
2907:
2900:
2893:
2886:
2879:
2872:
2864:
2862:
2860:Rhabdodontidae
2856:
2855:
2853:
2852:
2845:
2838:
2829:
2827:
2820:
2819:
2817:
2816:
2808:
2800:
2791:
2785:
2779:
2778:
2776:
2775:
2771:Weewarrasaurus
2767:
2760:
2753:
2746:
2739:
2732:
2725:
2717:
2710:
2703:
2696:
2693:Leaellynasaura
2689:
2681:
2674:
2671:Gasparinisaura
2667:
2664:Galleonosaurus
2660:
2652:
2645:
2638:
2631:
2623:
2621:
2615:
2614:
2612:
2611:
2603:
2596:
2587:
2585:
2578:
2577:
2575:
2574:
2566:
2559:
2551:
2541:
2536:
2529:
2528:
2516:
2515:
2512:
2511:
2508:
2507:
2504:
2503:
2500:
2499:
2497:
2496:
2495:
2494:
2488:
2486:
2480:
2479:
2477:
2476:
2475:
2474:
2465:
2459:
2453:
2452:
2450:
2449:
2448:
2447:
2438:
2432:
2422:
2421:
2417:
2416:
2407:
2405:Neornithischia
2398:
2389:
2380:
2374:
2358:
2357:
2350:
2349:
2342:
2335:
2327:
2320:
2319:
2307:
2285:
2284:
2243:
2202:
2160:
2122:
2052:
2045:
2018:
1947:
1946:
1944:
1941:
1940:
1939:
1934:
1927:
1924:
1920:
1919:
1916:
1915:
1911:
1910:
1907:
1906:
1902:
1901:
1898:
1897:
1893:
1892:
1889:
1888:
1884:
1883:
1880:
1879:
1875:
1874:
1871:
1870:
1866:
1865:
1862:
1861:
1857:
1856:
1853:
1852:
1848:
1847:
1844:
1843:
1839:
1838:
1835:
1834:
1830:
1829:
1826:
1825:
1821:
1820:
1817:
1816:
1812:
1811:
1808:
1807:
1803:
1802:
1799:
1798:
1794:
1793:
1790:
1789:
1785:
1784:
1781:
1780:
1776:
1775:
1772:
1771:
1767:
1766:
1763:
1762:
1759:
1758:
1755:
1754:
1751:
1750:
1747:
1746:
1739:
1736:
1735:
1732:
1731:
1727:Nanyangosaurus
1722:
1716:
1713:
1712:
1709:
1708:
1699:
1693:
1690:
1689:
1686:
1685:
1676:
1671:
1669:
1666:
1665:
1662:
1661:
1652:
1647:
1645:
1642:
1641:
1638:
1637:
1628:
1623:
1621:
1618:
1617:
1614:
1613:
1609:
1608:
1605:
1604:
1595:
1592:
1591:
1588:
1587:
1578:
1573:
1571:
1566:
1564:
1561:
1560:
1557:
1556:
1552:
1551:
1548:
1547:
1538:
1535:
1534:
1531:
1530:
1526:Nanningosaurus
1521:
1516:
1514:
1509:
1507:
1504:
1503:
1500:
1499:
1495:Gilmoreosaurus
1490:
1485:
1483:
1480:
1479:
1476:
1475:
1466:
1461:
1459:
1456:
1455:
1452:
1451:
1442:
1437:
1435:
1432:
1431:
1428:
1427:
1418:
1413:
1411:
1408:
1407:
1404:
1403:
1394:
1389:
1387:
1384:
1383:
1380:
1379:
1370:
1365:
1363:
1360:
1359:
1356:
1355:
1346:
1341:
1339:
1333:
1332:
1329:
1328:
1324:
1323:
1320:
1319:
1310:
1307:
1306:
1303:
1302:
1293:
1288:
1286:
1281:
1279:
1276:
1275:
1272:
1271:
1262:
1257:
1255:
1252:
1251:
1248:
1247:
1238:
1233:
1231:
1228:
1227:
1224:
1223:
1214:
1209:
1207:
1204:
1203:
1200:
1199:
1190:
1185:
1183:
1181:Hadrosauroidea
1175:
1152:
1151:
1148:
1147:
1143:
1142:
1139:
1138:
1134:
1133:
1130:
1129:
1125:
1124:
1121:
1120:
1116:
1115:
1112:
1111:
1107:
1106:
1103:
1102:
1098:
1097:
1094:
1093:
1089:
1088:
1085:
1084:
1080:
1079:
1076:
1075:
1071:
1070:
1067:
1066:
1059:
1056:
1055:
1052:
1051:
1042:
1037:
1035:
1032:
1031:
1028:
1027:
1018:
1013:
1011:
1008:
1007:
1004:
1003:
999:
998:
995:
994:
985:
982:
981:
978:
977:
968:
963:
961:
956:
954:
951:
950:
947:
946:
937:
932:
930:
927:
926:
923:
922:
918:
917:
914:
913:
909:
908:
905:
904:
895:
892:
891:
888:
887:
878:
873:
871:
868:
867:
864:
863:
854:
849:
847:
842:
840:
837:
836:
833:
832:
828:Gilmoreosaurus
823:
818:
816:
813:
812:
809:
808:
799:
794:
792:
789:
788:
785:
784:
775:
770:
768:
765:
764:
761:
760:
756:
755:
752:
751:
747:
746:
743:
742:
733:
730:
729:
726:
725:
716:
711:
709:
706:
705:
702:
701:
692:
687:
685:
680:
678:
670:
635:
634:Classification
632:
567:autapomorphies
522:
519:
493:Hadrosauroidea
364:
361:
316:
315:
312:
298:
297:
291:
290:
282:
280:
276:
275:
258:
254:
253:
245:
238:
237:
229:
222:
221:
219:Neornithischia
213:
206:
205:
197:
190:
189:
184:
177:
176:
171:
167:
166:
161:
157:
156:
151:
147:
146:
133:
132:
124:
123:
115:
114:
106:
105:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
30:
17:
9:
6:
4:
3:
2:
4797:
4786:
4783:
4781:
4778:
4776:
4773:
4771:
4768:
4766:
4763:
4762:
4760:
4743:
4738:
4734:
4730:
4725:
4721:
4716:
4710:
4706:
4705:
4703:
4701:
4697:
4693:
4688:
4668:
4667:
4663:
4661:
4659:
4658:Edmontosaurus
4655:
4653:
4651:
4650:Edmontosaurus
4647:
4645:
4642:
4639:
4635:
4632:
4630:
4627:
4625:
4622:
4620:
4617:
4615:
4612:
4610:
4609:
4605:
4604:
4602:
4598:
4592:
4591:
4587:
4585:
4584:
4580:
4578:
4577:
4573:
4571:
4570:
4569:Syngonosaurus
4566:
4564:
4563:
4559:
4557:
4556:
4552:
4550:
4549:
4545:
4543:
4542:
4538:
4536:
4535:
4531:
4529:
4528:
4524:
4522:
4521:
4517:
4515:
4514:
4513:Koutalisaurus
4510:
4508:
4507:
4503:
4501:
4500:
4496:
4494:
4493:
4492:Eucercosaurus
4489:
4487:
4486:
4482:
4479:
4478:
4474:
4472:
4471:
4467:
4465:
4464:
4463:Arstanosaurus
4460:
4459:
4457:
4455:
4454:
4449:
4443:
4442:
4438:
4437:
4435:
4433:
4429:
4423:
4422:
4418:
4416:
4415:
4411:
4409:
4408:
4404:
4402:
4401:
4397:
4395:
4394:
4390:
4388:
4387:
4383:
4381:
4380:
4379:Orcauichnites
4376:
4374:
4373:
4369:
4367:
4366:
4362:
4360:
4359:
4355:
4353:
4352:
4348:
4346:
4345:
4341:
4339:
4338:
4334:
4332:
4331:
4327:
4325:
4324:
4320:
4318:
4317:
4316:Amblydactylus
4313:
4312:
4310:
4308:
4304:
4300:
4293:
4289:
4280:
4276:
4274:
4269:
4263:
4256:
4230:
4229:
4225:
4223:
4222:
4218:
4216:
4215:
4214:Kundurosaurus
4211:
4209:
4208:
4207:Kerberosaurus
4204:
4202:
4201:
4197:
4195:
4194:
4193:Edmontosaurus
4190:
4189:
4187:
4185:
4181:
4175:
4174:
4170:
4168:
4167:
4163:
4161:
4160:
4156:
4155:
4153:
4151:
4147:
4141:
4140:
4136:
4134:
4133:
4129:
4127:
4126:
4122:
4120:
4119:
4115:
4113:
4112:
4108:
4105:
4104:
4103:Anasazisaurus
4100:
4099:
4097:
4095:
4091:
4085:
4084:
4083:Secernosaurus
4080:
4078:
4077:
4076:Kelumapusaura
4073:
4071:
4070:
4066:
4064:
4063:
4059:
4058:
4056:
4054:
4050:
4043:
4042:
4038:
4036:
4035:
4031:
4029:
4028:
4024:
4022:
4021:
4017:
4015:
4014:
4010:
4008:
4007:
4003:
4002:
4000:
3998:
3994:
3988:
3987:
3983:
3982:
3979:
3976:
3974:
3973:Saurolophinae
3970:
3960:
3959:
3955:
3953:
3952:
3948:
3946:
3945:
3941:
3939:
3938:
3934:
3932:
3931:
3927:
3925:
3924:
3923:Hypacrosaurus
3920:
3918:
3917:
3916:Corythosaurus
3913:
3911:
3910:
3906:
3905:
3903:
3901:
3900:Lambeosaurini
3897:
3891:
3890:
3886:
3884:
3883:
3879:
3877:
3876:
3875:Charonosaurus
3872:
3870:
3869:
3865:
3864:
3862:
3860:
3856:
3850:
3849:
3848:Tsintaosaurus
3845:
3843:
3842:
3841:Pararhabdodon
3838:
3836:
3835:
3834:Nipponosaurus
3831:
3829:
3828:
3824:
3821:
3820:
3816:
3814:
3813:
3809:
3807:
3806:
3805:Jaxartosaurus
3802:
3800:
3799:
3795:
3793:
3792:
3788:
3786:
3785:
3781:
3779:
3778:
3774:
3772:
3771:
3767:
3765:
3764:
3760:
3758:
3757:
3756:Adynomosaurus
3753:
3752:
3749:
3746:
3744:
3740:
3734:
3733:
3729:
3727:
3726:
3722:
3720:
3719:
3718:Lapampasaurus
3715:
3713:
3712:
3708:
3706:
3705:
3701:
3699:
3698:
3694:
3693:
3690:
3687:
3684:
3683:Hadrosauridae
3679:
3673:
3672:
3668:
3666:
3665:
3664:Zhanghenglong
3661:
3659:
3658:
3654:
3652:
3651:
3647:
3645:
3644:
3640:
3638:
3637:
3636:Telmatosaurus
3633:
3631:
3630:
3626:
3624:
3623:
3619:
3617:
3616:
3612:
3610:
3609:
3605:
3603:
3602:
3598:
3596:
3595:
3591:
3589:
3588:
3584:
3582:
3581:
3580:Portellsaurus
3577:
3575:
3574:
3570:
3568:
3567:
3563:
3561:
3560:
3556:
3554:
3553:
3549:
3547:
3546:
3542:
3540:
3539:
3535:
3532:
3531:
3527:
3525:
3524:
3520:
3518:
3517:
3513:
3511:
3510:
3506:
3504:
3503:
3499:
3497:
3496:
3492:
3490:
3489:
3485:
3483:
3482:
3478:
3476:
3475:
3471:
3469:
3468:
3464:
3462:
3461:
3457:
3455:
3454:
3450:
3448:
3447:
3443:
3441:
3440:
3436:
3433:
3432:
3428:
3426:
3425:
3421:
3419:
3418:
3414:
3413:
3410:
3407:
3404:
3399:
3395:
3391:
3386:
3382:
3373:
3369:
3366:
3359:
3326:
3325:
3324:
3323:
3321:
3319:
3315:
3308:
3307:
3303:
3301:
3300:
3296:
3294:
3293:
3289:
3286:
3285:
3281:
3280:
3278:
3276:
3272:
3266:
3265:
3261:
3259:
3258:
3254:
3252:
3251:
3247:
3245:
3244:
3243:Jinzhousaurus
3240:
3238:
3237:
3236:Hypselospinus
3233:
3231:
3230:
3226:
3224:
3223:
3219:
3218:
3215:
3212:
3210:
3206:
3200:
3199:
3195:
3193:
3192:
3191:Theiophytalia
3188:
3186:
3185:
3181:
3179:
3178:
3174:
3172:
3171:
3167:
3165:
3164:
3160:
3158:
3157:
3153:
3151:
3150:
3146:
3144:
3143:
3139:
3137:
3136:
3132:
3130:
3129:
3125:
3123:
3122:
3121:Lanzhousaurus
3118:
3116:
3115:
3111:
3109:
3108:
3104:
3102:
3101:
3097:
3095:
3094:
3090:
3087:
3086:
3082:
3080:
3079:
3075:
3073:
3072:
3068:
3066:
3065:
3061:
3059:
3058:
3054:
3052:
3051:
3047:
3046:
3043:
3040:
3037:
3032:
3022:
3021:
3017:
3014:
3013:
3009:
3006:
3005:
3001:
2999:
2998:
2994:
2992:
2991:
2990:Elrhazosaurus
2987:
2985:
2984:
2980:
2978:
2977:
2973:
2972:
2970:
2968:
2964:
2958:
2957:
2953:
2951:
2950:
2946:
2944:
2943:
2939:
2938:
2935:
2931:
2928:
2925:
2920:
2906:
2905:
2901:
2899:
2898:
2894:
2892:
2891:
2887:
2885:
2884:
2880:
2878:
2877:
2873:
2871:
2870:
2866:
2865:
2863:
2861:
2857:
2851:
2850:
2849:Tenontosaurus
2846:
2844:
2843:
2839:
2836:
2835:
2834:Convolosaurus
2831:
2830:
2828:
2825:
2821:
2814:
2813:
2809:
2806:
2805:
2801:
2798:
2797:
2796:Ampelognathus
2793:
2792:
2789:
2786:
2784:
2780:
2773:
2772:
2768:
2766:
2765:
2761:
2759:
2758:
2754:
2752:
2751:
2747:
2745:
2744:
2740:
2738:
2737:
2733:
2731:
2730:
2726:
2723:
2722:
2718:
2716:
2715:
2711:
2709:
2708:
2707:Mahuidacursor
2704:
2702:
2701:
2697:
2695:
2694:
2690:
2687:
2686:
2682:
2680:
2679:
2675:
2673:
2672:
2668:
2666:
2665:
2661:
2658:
2657:
2653:
2651:
2650:
2646:
2644:
2643:
2639:
2637:
2636:
2632:
2630:
2629:
2625:
2624:
2622:
2620:
2616:
2609:
2608:
2604:
2602:
2601:
2600:Hypsilophodon
2597:
2594:
2593:
2589:
2588:
2586:
2583:
2579:
2572:
2571:
2567:
2565:
2564:
2563:Callovosaurus
2560:
2558:
2557:
2556:Burianosaurus
2553:
2552:
2549:
2545:
2542:
2539:
2534:
2530:
2526:
2521:
2517:
2492:
2491:
2490:
2489:
2487:
2485:
2481:
2473:
2469:
2468:
2467:
2466:
2463:
2460:
2458:
2454:
2446:
2442:
2441:
2440:
2439:
2436:
2433:
2431:
2427:
2423:
2415:
2411:
2408:
2406:
2402:
2399:
2397:
2393:
2390:
2388:
2384:
2381:
2379:
2375:
2373:
2369:
2368:
2363:
2359:
2355:
2348:
2343:
2341:
2336:
2334:
2329:
2328:
2325:
2318:
2308:
2306:
2296:
2295:
2292:
2280:
2276:
2271:
2266:
2262:
2258:
2254:
2247:
2238:
2233:
2229:
2225:
2221:
2217:
2216:Palaeontology
2213:
2206:
2198:
2194:
2190:
2186:
2182:
2175:
2173:
2171:
2169:
2167:
2165:
2156:
2152:
2148:
2144:
2140:
2136:
2129:
2127:
2118:
2114:
2110:
2106:
2102:
2098:
2094:
2090:
2086:
2079:
2077:
2075:
2073:
2071:
2069:
2067:
2065:
2063:
2061:
2059:
2057:
2048:
2046:0-253-34514-6
2042:
2038:
2031:
2029:
2027:
2025:
2023:
2014:
2010:
2005:
2000:
1996:
1992:
1988:
1984:
1980:
1976:
1972:
1965:
1963:
1961:
1959:
1957:
1955:
1953:
1948:
1938:
1935:
1933:
1930:
1929:
1918:
1917:
1909:
1908:
1900:
1899:
1891:
1890:
1882:
1881:
1873:
1872:
1864:
1863:
1855:
1854:
1846:
1845:
1837:
1836:
1828:
1827:
1819:
1818:
1810:
1809:
1801:
1800:
1792:
1791:
1783:
1782:
1774:
1773:
1765:
1764:
1757:
1756:
1749:
1748:
1745:
1744:
1743:Hadrosauridae
1738:
1737:
1734:
1733:
1730:
1729:
1728:
1721:
1720:
1715:
1714:
1711:
1710:
1707:
1706:
1705:
1704:Adynomosaurus
1698:
1697:
1692:
1691:
1688:
1687:
1684:
1683:
1682:
1675:
1674:
1668:
1667:
1664:
1663:
1660:
1659:
1658:
1657:Zhanghenglong
1651:
1650:
1644:
1643:
1640:
1639:
1636:
1635:
1634:
1627:
1626:
1620:
1619:
1616:
1615:
1607:
1606:
1603:
1602:
1601:
1594:
1593:
1590:
1589:
1586:
1585:
1584:
1583:Telmatosaurus
1577:
1576:
1570:
1569:
1563:
1562:
1559:
1558:
1550:
1549:
1546:
1545:
1544:
1537:
1536:
1533:
1532:
1529:
1528:
1527:
1520:
1519:
1513:
1512:
1506:
1505:
1502:
1501:
1498:
1497:
1496:
1489:
1488:
1482:
1481:
1478:
1477:
1474:
1473:
1472:
1465:
1464:
1458:
1457:
1454:
1453:
1450:
1449:
1448:
1441:
1440:
1434:
1433:
1430:
1429:
1426:
1425:
1424:
1417:
1416:
1410:
1409:
1406:
1405:
1402:
1401:
1400:
1393:
1392:
1386:
1385:
1382:
1381:
1378:
1377:
1376:
1369:
1368:
1362:
1361:
1358:
1357:
1354:
1353:
1352:
1345:
1344:
1338:
1335:
1334:
1331:
1330:
1322:
1321:
1318:
1317:
1316:
1309:
1308:
1305:
1304:
1301:
1300:
1299:
1292:
1291:
1285:
1284:
1278:
1277:
1274:
1273:
1270:
1269:
1268:
1261:
1260:
1254:
1253:
1250:
1249:
1246:
1245:
1244:
1237:
1236:
1230:
1229:
1226:
1225:
1222:
1221:
1220:
1213:
1212:
1206:
1205:
1202:
1201:
1198:
1197:
1196:
1195:Jinzhousaurus
1189:
1188:
1182:
1179:
1178:
1174:
1172:
1168:
1167:Telmatosaurus
1164:
1160:
1150:
1149:
1141:
1140:
1132:
1131:
1123:
1122:
1114:
1113:
1105:
1104:
1096:
1095:
1087:
1086:
1078:
1077:
1069:
1068:
1065:
1064:
1063:Hadrosauridae
1058:
1057:
1054:
1053:
1050:
1049:
1048:
1041:
1040:
1034:
1033:
1030:
1029:
1026:
1025:
1024:
1017:
1016:
1010:
1009:
1006:
1005:
997:
996:
993:
992:
991:
984:
983:
980:
979:
976:
975:
974:
967:
966:
960:
959:
953:
952:
949:
948:
945:
944:
943:
936:
935:
929:
928:
925:
924:
916:
915:
907:
906:
903:
902:
901:
900:Telmatosaurus
894:
893:
890:
889:
886:
885:
884:
877:
876:
870:
869:
866:
865:
862:
861:
860:
853:
852:
846:
845:
839:
838:
835:
834:
831:
830:
829:
822:
821:
815:
814:
811:
810:
807:
806:
805:
798:
797:
791:
790:
787:
786:
783:
782:
781:
774:
773:
767:
766:
763:
762:
754:
753:
745:
744:
741:
740:
739:
732:
731:
728:
727:
724:
723:
722:
715:
714:
708:
707:
704:
703:
700:
699:
698:
691:
690:
684:
683:
677:
674:
673:
669:
667:
663:
662:
657:
653:
652:Hadrosauridae
649:
640:
631:
629:
625:
620:
613:
612:
607:
599:
592:
588:
584:
582:
578:
577:Telmatosaurus
574:
573:
572:Telmatosaurus
568:
564:
560:
555:
552:
547:
539:
532:
527:
518:
515:
510:
506:
502:
498:
497:specific name
494:
490:
486:
482:
473:
469:
467:
463:
459:
458:
452:
448:
442:
440:
436:
435:Maastrichtian
432:
428:
427:
423:
419:
418:
414:
413:foraminiferan
410:
406:
402:
398:
397:
391:
387:
378:
374:
369:
360:
358:
357:
352:
348:
344:
340:
336:
332:
328:
324:
323:
310:
308:
299:
296:
295:Binomial name
292:
288:
287:
281:
278:
277:
272:
271:Dalla Vecchia
267:
266:
259:
256:
255:
252:
246:
243:
240:
239:
236:
230:
227:
224:
223:
220:
214:
211:
208:
207:
204:
198:
195:
192:
191:
188:
185:
182:
179:
178:
175:
172:
169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
143:
138:
134:
130:
125:
121:
116:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
41:
34:
28:
24:
21:
16:
4700:Tethyshadros
4699:
4664:
4657:
4649:
4637:
4606:
4588:
4581:
4574:
4567:
4560:
4553:
4546:
4539:
4532:
4525:
4518:
4511:
4504:
4497:
4490:
4483:
4477:Claorhynchus
4475:
4468:
4461:
4453:Nomina dubia
4451:
4441:Guegoolithus
4439:
4419:
4412:
4407:Staurichnium
4405:
4400:Sousaichnium
4398:
4391:
4384:
4377:
4370:
4363:
4356:
4351:Dakotasaurus
4349:
4344:Caririchnium
4342:
4335:
4328:
4321:
4314:
4258:
4226:
4219:
4212:
4205:
4198:
4191:
4171:
4164:
4157:
4150:Saurolophini
4137:
4130:
4123:
4116:
4109:
4101:
4094:Kritosaurini
4081:
4074:
4069:Huallasaurus
4067:
4060:
4041:Wulagasaurus
4039:
4032:
4025:
4018:
4011:
4004:
3984:
3956:
3949:
3942:
3935:
3930:Lambeosaurus
3928:
3921:
3914:
3907:
3887:
3880:
3873:
3866:
3846:
3839:
3832:
3825:
3817:
3810:
3803:
3796:
3789:
3782:
3775:
3768:
3761:
3754:
3732:Yamatosaurus
3730:
3723:
3716:
3709:
3702:
3697:Aquilarhinus
3695:
3669:
3662:
3655:
3648:
3643:Tethyshadros
3642:
3641:
3634:
3627:
3620:
3613:
3606:
3599:
3592:
3585:
3578:
3573:Plesiohadros
3571:
3564:
3557:
3550:
3545:Lophorhothon
3543:
3536:
3528:
3521:
3514:
3507:
3500:
3493:
3486:
3479:
3472:
3465:
3458:
3451:
3444:
3437:
3431:Batyrosaurus
3429:
3424:Bactrosaurus
3422:
3415:
3361:
3304:
3297:
3290:
3282:
3264:Ouranosaurus
3262:
3255:
3248:
3241:
3234:
3229:Brighstoneus
3227:
3220:
3196:
3189:
3182:
3175:
3168:
3161:
3156:Osmakasaurus
3154:
3149:Oblitosaurus
3147:
3140:
3133:
3126:
3119:
3112:
3105:
3098:
3091:
3083:
3076:
3071:Camptosaurus
3069:
3062:
3055:
3048:
3018:
3012:Kangnasaurus
3010:
3002:
2995:
2988:
2981:
2974:
2967:Dryosauridae
2954:
2947:
2940:
2902:
2895:
2888:
2881:
2874:
2869:Matheronodon
2867:
2847:
2840:
2832:
2810:
2802:
2794:
2769:
2762:
2755:
2748:
2743:Sektensaurus
2741:
2736:Qantassaurus
2734:
2727:
2719:
2712:
2705:
2698:
2691:
2685:Kangnasaurus
2683:
2676:
2669:
2662:
2654:
2649:Diluvicursor
2647:
2640:
2633:
2626:
2607:Vectidromeus
2605:
2598:
2590:
2568:
2561:
2554:
2472:Ornithischia
2457:Ornithischia
2409:
2400:
2396:Ornithischia
2391:
2382:
2260:
2256:
2246:
2222:(1): 13–48.
2219:
2215:
2205:
2188:
2184:
2180:
2141:(1): 45–66.
2138:
2134:
2092:
2088:
2084:
2036:
1981:(1): 23295.
1978:
1974:
1741:
1725:
1724:
1702:
1701:
1681:Plesiohadros
1679:
1678:
1655:
1654:
1631:
1630:
1600:Tethyshadros
1599:
1598:
1597:
1581:
1580:
1541:
1540:
1524:
1523:
1493:
1492:
1471:Bactrosaurus
1469:
1468:
1445:
1444:
1421:
1420:
1397:
1396:
1373:
1372:
1349:
1348:
1313:
1312:
1296:
1295:
1265:
1264:
1241:
1240:
1217:
1216:
1193:
1192:
1171:Tethyshadros
1170:
1166:
1163:Tethyshadros
1162:
1156:
1061:
1045:
1044:
1023:Plesiohadros
1021:
1020:
990:Tethyshadros
989:
988:
987:
971:
970:
940:
939:
898:
897:
881:
880:
859:Lophorhothon
857:
856:
826:
825:
804:Bactrosaurus
802:
801:
778:
777:
736:
735:
719:
718:
695:
694:
659:
656:Tethyshadros
655:
648:hadrosaurine
645:
624:hadrosaurids
618:supraorbital
609:
606:T. insularis
605:
598:Tethyshadros
597:
595:
583:, overall.
581:hadrosaurids
576:
570:
563:Tethyshadros
562:
559:Tethyshadros
558:
556:
546:Tethyshadros
545:
544:
531:Tethyshadros
530:
514:Duino Castle
509:Tethyshadros
508:
489:Tethys Ocean
484:
481:type species
478:
455:
443:
424:
422:alligatoroid
415:
394:
382:
377:Tethyshadros
376:
372:
356:T. insularis
355:
354:
335:hadrosauroid
322:Tethyshadros
321:
320:
319:
306:
301:
285:
284:
265:Tethyshadros
264:
263:
241:
225:
209:
203:Ornithischia
193:
180:
27:Tethyshadros
26:
20:
15:
4666:Willinakaqe
4638:nomina nuda
4555:Siluosaurus
4520:Loncosaurus
4358:Dinehichnus
4200:Kamuysaurus
4173:Saurolophus
4125:Kritosaurus
4118:Gryposaurus
3951:Sahaliyania
3937:Magnapaulia
3909:Amurosaurus
3889:Tlatolophus
3868:Adelolophus
3812:Kazaklambia
3791:Blasisaurus
3784:Arenysaurus
3777:Aralosaurus
3711:Hadrosaurus
3704:Eotrachodon
3657:Yunganglong
3622:Sirindhorna
3594:Protohadros
3530:Koshisaurus
3523:Jintasaurus
3292:Comptonatus
3142:Napaisaurus
3128:Lurdusaurus
3100:Fukuisaurus
3078:Cedrorestes
3020:Valdosaurus
2883:Pareisactus
2714:Morrosaurus
2678:Isasicursor
2642:Chakisaurus
2538:Ornithopoda
2525:Ornithopoda
2484:Ornithopoda
2354:Ornithopoda
1633:Eotrachodon
1399:Jintasaurus
1375:Yunganglong
1243:Sirindhorna
1047:Eotrachodon
738:Protohadros
521:Description
451:formic acid
441:formation.
399:remains at
235:Ornithopoda
4765:Hadrosaurs
4759:Categories
4608:Arkharavia
4583:Tichosteus
4576:Thespesius
4548:Pteropelyx
4541:Orthomerus
4421:Wintonopus
4006:Acristavus
3986:Barsboldia
3944:Olorotitan
3819:Latirhinus
3671:Zuoyunlong
3538:Levnesovia
3488:Gobihadros
3453:Datonglong
3446:Claosaurus
3417:Altirhinus
3327:see below↓
3257:Morelladon
3135:Magnamanus
3107:Hippodraco
2976:Dryosaurus
2949:Hesperonyx
2924:Dryomorpha
2764:Trinisaura
2757:Tietasaura
2750:Talenkauen
2628:Anabisetia
2493:see below↓
2387:Dinosauria
1943:References
1543:Claosaurus
1423:Levnesovia
1298:Zuoyunlong
942:Claosaurus
462:a theropod
401:Bernissart
187:Dinosauria
4590:Trachodon
4506:Glishades
4485:Diclonius
4323:Anomoepus
4307:Ichnotaxa
4020:Maiasaura
3958:Velafrons
3467:Equijubus
3439:Choyrodon
3299:Iguanodon
3163:Planicoxa
3093:Dakotadon
3064:Calvarius
2890:Rhabdodon
2876:Mochlodon
2619:Elasmaria
2370:Kingdom:
2305:Dinosaurs
2279:236696588
2117:198128196
1219:Equijubus
611:Iguanodon
551:histology
457:Scipionyx
431:Campanian
396:Iguanodon
390:hadrosaur
279:Species:
160:Kingdom:
154:Eukaryota
4729:47460141
4709:Wikidata
4470:Cionodon
4139:Rhinorex
4027:Ornatops
3827:Minqaria
3798:Canardia
3725:Malefica
3650:Xuwulong
3615:Siamodon
3516:Jeyawati
3502:Gonkoken
3460:Eolambia
3284:Barilium
3085:Cumnoria
3050:Barilium
2956:Owenodon
2942:Draconyx
2904:Zalmoxes
2804:Fostoria
2656:Fostoria
2414:Cerapoda
2378:Chordata
2376:Phylum:
2372:Animalia
2013:34857789
1926:See also
1351:Eolambia
1267:Xuwulong
721:Eolambia
697:Jeyawati
529:Size of
505:holotype
491:and the
426:Acynodon
174:Chordata
170:Phylum:
164:Animalia
150:Domain:
131:, Italy
4715:Q133402
3763:Ajnabia
3198:Uteodon
2291:Portals
2224:Bibcode
2143:Bibcode
2097:Bibcode
2004:8640049
1983:Bibcode
466:Saltrio
447:Trieste
347:Trieste
327:Tethyan
304:†
283:†
257:Genus:
129:Bologna
107:↓
4742:161197
4432:Ootaxa
3629:Tanius
3222:Bolong
2277:
2115:
2043:
2011:
2001:
1447:Tanius
780:Tanius
495:. The
273:, 2009
3474:Fylax
3004:Iyuku
2410:Clade
2401:Clade
2392:Clade
2383:Clade
2317:Italy
2275:S2CID
2191:(3).
2113:S2CID
973:Fylax
661:Fylax
601:'
501:Latin
464:from
351:Italy
345:) of
331:genus
242:Clade
226:Clade
210:Clade
194:Clade
181:Clade
3170:Proa
2842:Iani
2470:see
2443:see
2041:ISBN
2009:PMID
420:and
405:lens
46:PreꞒ
4724:EoL
2265:doi
2232:doi
2193:doi
2189:126
2151:doi
2105:doi
1999:PMC
1991:doi
483:is
333:of
4761::
4739::
4726::
4711::
2412::
2403::
2394::
2385::
2273:.
2263:.
2261:66
2259:.
2255:.
2230:.
2220:61
2218:.
2214:.
2187:.
2163:^
2149:.
2137:.
2125:^
2111:.
2103:.
2093:29
2091:.
2055:^
2021:^
2007:.
1997:.
1989:.
1979:11
1977:.
1973:.
1951:^
359:.
349:,
325:("
244::
228::
212::
196::
183::
96:Pg
40:Ma
35:,
4640:)
4480:?
4106:?
4044:?
3822:?
3533:?
3434:?
3309:?
3287:?
3088:?
3015:?
3007:?
2837:?
2826:?
2815:?
2807:?
2799:?
2774:?
2724:?
2688:?
2659:?
2610:?
2595:?
2584:?
2573:?
2346:e
2339:t
2332:v
2293::
2281:.
2267::
2240:.
2234::
2226::
2199:.
2195::
2157:.
2153::
2145::
2139:2
2119:.
2107::
2099::
2049:.
2015:.
1993::
1985::
433:-
261:†
248:†
232:†
216:†
200:†
101:N
91:K
86:J
81:T
76:P
71:C
66:D
61:S
56:O
51:Ꞓ
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