327:
162:
35:
314:
702:
598:
1280:
771:
719:
1297:
1260:
1243:
1226:
1722:
1563:
1492:
1475:
1139:
864:
1735:
1441:
950:
916:
293:
279:
150:
136:
1776:
1709:
197:(VLPs). VLPs are not infectious like normal virions, but they nevertheless make up an essential part of the pseudoviral lifecycle.
326:
161:
626:
1195:
1614:
1093:
838:
890:
568:
1748:
1740:
760:
745:
1535:
1619:
1410:
1189:
1171:
1165:
1118:
1375:
1061:
1056:
1002:
350:
212:
34:
1400:
1067:
1032:
604:
1052:
619:
341:
of viruses from this family is unsegmented, -RT, positive-sense, single-stranded RNA and is 4200–9700
1753:
1430:
1370:
1027:
755:
1405:
1183:
1781:
1674:
691:
1636:
1380:
1365:
686:
381:
is unenveloped and looks roughly spherical. The capsid is round with icosahedral symmetry with
1786:
1727:
1591:
1128:
1113:
1073:
639:
612:
362:
358:
1696:
519:
Krupovic M, Blomberg J, Coffin JM, Dasgupta I, Fan H, Geering AD, et al. (June 2018).
382:
190:
29:
8:
933:
286:
194:
143:
593:
1306:
1269:
1007:
545:
520:
477:
446:
234:
521:"Ortervirales: New Virus Order Unifying Five Families of Reverse-Transcribing Viruses"
1683:
1311:
1215:
1200:
1084:
982:
750:
736:
550:
482:
418:
1688:
1133:
540:
532:
472:
462:
394:
1177:
1104:
1043:
993:
677:
644:
366:
95:
83:
71:
1659:
1345:
1334:
1018:
807:
666:
583:
445:
Llorens, C; Soriano, B; Krupovic, M; ICTV Report
Consortium (2 February 2021).
267:
505:
1770:
818:
655:
409:
The genome integrates into the host genome and gets transcribed by host cell
249:
220:
444:
1509:
1391:
1123:
796:
554:
486:
426:
261:
107:
1668:
1546:
1421:
853:
829:
634:
536:
467:
342:
255:
1604:
1526:
1356:
1150:
971:
414:
1458:
881:
422:
354:
59:
1714:
1630:
1653:
390:
386:
346:
410:
345:
long. The genome encodes structural proteins and non-structural
1599:
398:
378:
338:
518:
193:
of the Ty1-copia family. They replicate via structures called
1701:
635:
216:
183:
46:
429:
and assembly can occur in the cytoplasm, or in the nucleus.
1609:
1159:
209:
1768:
1140:Xenotropic murine leukemia virus-related virus
1343:
664:
620:
421:. Genome replication takes place in the host
500:
498:
496:
229:comprises highly divergent members and most
827:
627:
613:
274:The family includes the following genera:
208:is unofficially classified under group VI
544:
493:
476:
466:
304:species not classified into a genus are:
325:
160:
1769:
1635:
1634:
608:
1196:Caprine arthritis encephalitis virus
1615:Human teratocarcinoma-derived virus
1094:Walleye epidermal hyperplasia virus
189:Viruses of the family are actually
13:
839:Caenorhabditis elegans Cer13 virus
569:Penicillium camemberti virus - GP1
332:Saccharomyces cerevisiae Ty1 virus
309:Penicillium camemberti virus - GP1
173:) virion with its building blocks.
14:
1798:
891:Caenorhabditis elegans Cer1 virus
577:
1777:RNA reverse-transcribing viruses
1162:(human immunodeficiency viruses)
33:
1166:Simian immunodeficiency viruses
761:Woolly monkey hepatitis B virus
746:Ground squirrel hepatitis virus
455:The Journal of General Virology
315:Phaseolus vulgaris Tpv2-6 virus
233:encode Gag and Pol on a single
186:, which includes three genera.
1536:Soybean chlorotic mottle virus
1184:Equine infectious anemia virus
561:
512:
447:"ICTV Virus Taxonomy Profile:
438:
404:
1:
1620:Human Endogenous Retrovirus-W
1411:Strawberry vein banding virus
1190:Bovine immunodeficiency virus
1172:Feline immunodeficiency virus
1119:Abelson murine leukemia virus
432:
397:are poorly characterized and
389:to quasi-isometric and has a
1376:Kalanchoe top-spotting virus
1003:Avian sarcoma leukosis virus
372:
351:RNA-dependent DNA polymerase
213:Reverse Transcribing Viruses
167:Saccharomyces cerevisiae Ty1
7:
1401:Carnation etched ring virus
1068:Simian-T-lymphotropic virus
1033:Jaagsiekte sheep retrovirus
506:"ICTV Report Pseudoviridae"
200:
10:
1803:
1053:Human T-lymphotropic virus
385:(T) = 3 and 4. It is also
1643:
1590:
1561:
1544:
1524:
1507:
1490:
1473:
1456:
1439:
1431:Cassava vein mosaic virus
1419:
1389:
1371:Cacao swollen shoot virus
1354:
1332:
1295:
1278:
1258:
1241:
1224:
1213:
1148:
1102:
1082:
1041:
1028:Mouse mammary tumor virus
1016:
991:
980:
969:
948:
931:
914:
903:
879:
862:
851:
816:
805:
794:
769:
756:Woodchuck hepatitis virus
734:
717:
700:
675:
653:
321:
243:is included in the order
132:
127:
28:
21:
1406:Cauliflower mosaic virus
692:Heron hepatitis B virus
1381:Rubus yellow net virus
1366:Banana streak MY virus
687:Duck hepatitis B virus
393:of 30-50 nm. LTR-
334:
174:
1129:Feline leukemia virus
1114:Murine leukemia virus
1074:Bovine leukemia virus
363:reverse transcription
359:reverse transcriptase
329:
164:
640:Reverse transcribing
537:10.1128/JVI.00515-18
468:10.1099/jgv.0.001563
383:triangulation number
247:along with families
195:virus-like particles
191:LTR retrotransposons
30:Virus classification
703:Herpetohepadnavirus
525:Journal of Virology
401:have not reported.
349:which codes for an
1307:Simian foamy virus
1281:Prosimiispumavirus
1270:Feline foamy virus
1008:Rous sarcoma virus
335:
235:open reading frame
175:
165:Reconstruction of
1764:
1763:
1637:Taxon identifiers
1628:
1627:
1586:
1585:
1582:
1581:
1578:
1577:
1328:
1327:
1324:
1323:
1320:
1319:
1312:Human foamy virus
1216:Spumaretrovirinae
1209:
1208:
1201:Visna-maedi virus
1085:Epsilonretrovirus
983:Orthoretrovirinae
965:
964:
899:
898:
847:
846:
790:
789:
786:
785:
751:Hepatitis B virus
737:Orthohepadnavirus
419:RNA polymerase II
159:
158:
16:Family of viruses
1794:
1757:
1756:
1744:
1743:
1731:
1730:
1718:
1717:
1705:
1704:
1692:
1691:
1679:
1678:
1677:
1664:
1663:
1662:
1632:
1631:
1352:
1351:
1341:
1340:
1335:dsDNA-RT viruses
1222:
1221:
1134:Koala retrovirus
989:
988:
978:
977:
912:
911:
860:
859:
825:
824:
814:
813:
808:ssRNA-RT viruses
803:
802:
772:Parahepadnavirus
720:Metahepadnavirus
673:
672:
662:
661:
629:
622:
615:
606:
605:
601:(Darwin Zoology)
572:
565:
559:
558:
548:
516:
510:
509:
502:
491:
490:
480:
470:
442:
395:retrotransposons
38:
37:
19:
18:
1802:
1801:
1797:
1796:
1795:
1793:
1792:
1791:
1767:
1766:
1765:
1760:
1752:
1747:
1739:
1734:
1726:
1721:
1713:
1708:
1700:
1695:
1687:
1682:
1673:
1672:
1667:
1658:
1657:
1652:
1639:
1629:
1624:
1574:
1557:
1540:
1520:
1503:
1486:
1469:
1452:
1435:
1415:
1385:
1316:
1298:Simiispumavirus
1291:
1274:
1254:
1237:
1205:
1178:Puma lentivirus
1144:
1105:Gammaretrovirus
1098:
1078:
1044:Deltaretrovirus
1037:
1012:
994:Alpharetrovirus
961:
944:
927:
895:
875:
843:
782:
765:
730:
713:
696:
678:Avihepadnavirus
649:
645:Revtraviricetes
633:
580:
575:
566:
562:
517:
513:
504:
503:
494:
443:
439:
435:
407:
375:
324:
203:
182:is a family of
97:Revtraviricetes
32:
17:
12:
11:
5:
1800:
1790:
1789:
1784:
1782:Virus families
1779:
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1522:
1521:
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1518:
1515:
1513:
1505:
1504:
1502:
1501:
1498:
1496:
1488:
1487:
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1479:
1471:
1470:
1468:
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1453:
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1450:
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1417:
1416:
1414:
1413:
1408:
1403:
1397:
1395:
1387:
1386:
1384:
1383:
1378:
1373:
1368:
1362:
1360:
1349:
1346:Caulimoviridae
1338:
1330:
1329:
1326:
1325:
1322:
1321:
1318:
1317:
1315:
1314:
1309:
1303:
1301:
1293:
1292:
1290:
1289:
1286:
1284:
1276:
1275:
1273:
1272:
1266:
1264:
1261:Felispumavirus
1256:
1255:
1253:
1252:
1249:
1247:
1244:Equispumavirus
1239:
1238:
1236:
1235:
1232:
1230:
1227:Bovispumavirus
1219:
1211:
1210:
1207:
1206:
1204:
1203:
1198:
1193:
1187:
1181:
1175:
1169:
1163:
1156:
1154:
1146:
1145:
1143:
1142:
1137:
1131:
1126:
1121:
1116:
1110:
1108:
1100:
1099:
1097:
1096:
1090:
1088:
1080:
1079:
1077:
1076:
1071:
1065:
1059:
1049:
1047:
1039:
1038:
1036:
1035:
1030:
1024:
1022:
1019:Betaretrovirus
1014:
1013:
1011:
1010:
1005:
999:
997:
986:
975:
967:
966:
963:
962:
960:
959:
956:
954:
946:
945:
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942:
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929:
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857:
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844:
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833:
822:
811:
800:
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791:
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723:
715:
714:
712:
711:
708:
706:
698:
697:
695:
694:
689:
683:
681:
670:
667:Hepadnaviridae
659:
651:
650:
632:
631:
624:
617:
609:
603:
602:
596:
591:
579:
578:External links
576:
574:
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560:
511:
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436:
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431:
406:
403:
374:
371:
330:Genome map of
323:
320:
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311:
298:
297:
290:
283:
268:Caulimoviridae
202:
199:
157:
156:
155:
154:
147:
140:
130:
129:
125:
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117:
113:
112:
105:
101:
100:
93:
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85:Artverviricota
81:
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57:
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25:
15:
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1720:
1716:
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1707:
1703:
1698:
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1681:
1676:
1675:Pseudoviridae
1670:
1666:
1661:
1655:
1651:
1650:
1648:
1646:
1645:Pseudoviridae
1642:
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1233:
1231:
1229:
1228:
1223:
1220:
1218:
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1212:
1202:
1199:
1197:
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1188:
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1179:
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1087:
1086:
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1054:
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1020:
1015:
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985:
984:
979:
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968:
958:
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953:
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947:
941:
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938:
936:
935:
930:
924:
923:
921:
919:
918:
913:
910:
908:
907:
906:Pseudoviridae
902:
892:
889:
888:
886:
884:
883:
878:
872:
871:
869:
867:
866:
861:
858:
856:
855:
850:
840:
837:
836:
834:
832:
831:
826:
823:
821:
820:
819:Belpaoviridae
815:
812:
809:
804:
801:
799:
798:
793:
779:
778:
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774:
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768:
762:
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688:
685:
684:
682:
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671:
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668:
663:
660:
658:
657:
656:Blubervirales
652:
647:
646:
641:
637:
630:
625:
623:
618:
616:
611:
610:
607:
600:
599:Pseudoviridae
597:
595:
592:
590:
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588:Pseudoviridae
586:
582:
581:
570:
564:
556:
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542:
538:
534:
530:
526:
522:
515:
507:
501:
499:
497:
488:
484:
479:
474:
469:
464:
460:
456:
452:
450:
449:Pseudoviridae
441:
437:
430:
428:
424:
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400:
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380:
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340:
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328:
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302:Pseudoviridae
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250:Belpaoviridae
246:
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241:Pseudoviridae
238:
236:
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231:Pseudoviridae
228:
227:Pseudoviridae
224:
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221:invertebrates
218:
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206:Pseudoviridae
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179:Pseudoviridae
172:
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126:
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121:Pseudoviridae
118:
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94:
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62:
58:
55:
52:
51:
48:
45:
42:
41:
36:
31:
27:
24:
23:Pseudoviridae
20:
1787:Ortervirales
1644:
1564:Vaccinivirus
1562:
1545:
1525:
1510:Solendovirus
1508:
1493:Ruflodivirus
1491:
1476:Rosadnavirus
1474:
1457:
1440:
1420:
1392:Caulimovirus
1390:
1355:
1344:
1296:
1279:
1259:
1242:
1225:
1214:
1149:
1124:Friend virus
1103:
1083:
1042:
1017:
992:
981:
972:Retroviridae
970:
949:
932:
915:
905:
904:
880:
865:Errantivirus
863:
852:
828:
817:
797:Ortervirales
795:
770:
735:
718:
701:
676:
665:
654:
643:
587:
585:ICTV Report:
584:
563:
528:
524:
514:
458:
454:
448:
440:
408:
376:
365:step during
336:
331:
313:
308:
301:
299:
292:
285:
278:
273:
266:
262:Retroviridae
260:
254:
248:
245:Ortervirales
244:
240:
239:
230:
226:
225:
205:
204:
188:
178:
177:
176:
170:
166:
149:
142:
135:
120:
119:
109:Ortervirales
108:
96:
84:
73:Pararnavirae
72:
60:
53:
43:(unranked):
22:
1669:Wikispecies
1547:Tungrovirus
1442:Dioscovirus
1422:Cavemovirus
1070:(types 1-4)
934:Pseudovirus
854:Metaviridae
830:Semotivirus
594:NCBI ICTVdb
405:Replication
367:replication
343:nucleotides
287:Pseudovirus
256:Metaviridae
215:and infect
171:Pseudovirus
144:Pseudovirus
1771:Categories
1592:Endogenous
1527:Soymovirus
1357:Badnavirus
1151:Lentivirus
433:References
415:eukaryotic
377:The viral
1459:Petuvirus
951:Sirevirus
917:Hemivirus
882:Metavirus
642:viruses (
571:(species)
425:, or the
423:cytoplasm
387:isometric
373:Structure
355:replicase
294:Sirevirus
280:Hemivirus
151:Sirevirus
137:Hemivirus
68:Kingdom:
61:Riboviria
1660:Q3772842
1654:Wikidata
555:29618642
487:33528349
417:nuclear
413:such as
391:diameter
361:for the
347:proteins
300:Further
201:Taxonomy
116:Family:
80:Phylum:
1064:, 3, 4)
546:5974489
478:8515867
427:nucleus
411:enzymes
184:viruses
169:(genus
128:Genera
104:Order:
92:Class:
1754:600049
1741:186670
1728:106553
1605:ERVWE1
1600:ERV-Fc
1186:(EIAV)
1136:(KoRV)
1062:HTLV-2
1057:HTLV-1
567:NCBI:
553:
543:
531:(12).
485:
475:
399:lipids
379:capsid
357:, and
339:genome
322:Genome
265:, and
1749:WoRMS
1723:IRMNG
1689:626V3
1192:(BIV)
1180:(PLV)
1174:(FIV)
1168:(SIV)
636:Virus
461:(3).
217:fungi
54:Realm
47:Virus
1736:NCBI
1715:1938
1710:GBIF
1702:8051
1610:HCP5
551:PMID
483:PMID
337:The
219:and
1697:EoL
1684:CoL
1160:HIV
541:PMC
533:doi
473:PMC
463:doi
459:102
210:RNA
1773::
1751::
1738::
1725::
1712::
1699::
1686::
1671::
1656::
638::
549:.
539:.
529:92
527:.
523:.
495:^
481:.
471:.
457:.
453:.
369:.
353:,
271:.
259:,
253:,
237:.
223:.
56::
1337:)
1333:(
1055:(
810:)
806:(
648:)
628:e
621:t
614:v
557:.
535::
508:.
489:.
465::
451:"
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