256:, the definition of species is a little unusual. It is simply an abstract taxonomic concept that clusters a selected range of genetic variants, helping to distinguish branches in a phylogenetic lineage, but it is not a physical entity like a virus that can infect an animal or be isolated. If the diversity level used to define a species is set very low, many will effectively contain a single virus, and the virus and species may even be given the same name, resulting in confusion between the two concepts in the literature, and marginalizing the phylogenetic role of the species taxon. To counter this problem, the diversity level now recognized for species in the
228:(PPV), which is a major cause of fetal death and infertility in pigs. The genus divides phylogenetically into two branches, one that contains many founder members of the family, such as MVM, CPV and PPV, which have been studied in considerable detail, and a second branch occupied exclusively by predicted viruses whose coding sequences were identified recently in the wild using virus discovery approaches, but whose biology remains minimally explored. This second branch currently contains two species whose members infect humans, called
57:
33:
620:
In following years a series of physically similar viruses, including H1, LuIII, minute virus of mice and tumor virus X, were extracted from cells or tissues in routine use in research laboratories and porcine parvovirus, PPV, one of the major causes of reproductive failure in swine, was isolated from
895:
Van
Regenmortel MHV (2000) Introduction to the species concept in virus taxonomy. In: Van Regenmortel MHV, Fauquet CM, Bishop DHL, Carstens EB, Estes MK, Lemon SM, McGeogh DJ, Maniloff J, Mayo MA, Pringle CR, Wickner RB. (eds) Virus Taxonomy-Seventh Report of the International Committee on Taxonomy
550:
Viral replication is nuclear. Entry into the host cell is achieved by attachment to host receptors, which mediates clathrin-mediated endocytosis. Replication follows the rolling-hairpin model. In some virus/host cell combinations, progeny virions can be trafficked through the cytoplasm in vesicles
424:
have non-enveloped protein capsids around 18–26 nm in diameter, which show T=1 icosahedral symmetry. Genomes are single-stranded linear DNA between 4–6kb in length, with small (100–500b) imperfect palindromic sequences at each terminus that fold to form distinctive duplex hairpin telomeres.
943:
Phan TG, Dreno B, da Costa AC, Li L, Orlandi P, Deng X, Kapusinszky B, Siqueira J, Knol AC, Halary F, Dantal J, Alexander KA, Pesavento PA, Delwart E A new protoparvovirus in human fecal samples and cutaneous T cell lymphomas (mycosis fungoides). Virology 496:299–305. doi:
551:
and released from the parental host cell prior to cell death, while the remaining virions are released following cell lysis. Vertebrates from 6 orders are currently known to serve as natural hosts. Transmission routes are typically fecal-oral and/or respiratory.
405:
Protoparvoviruses that infect humans were first discovered in 2012 in the feces of children from
Burkina Faso, and named using the siglum bufavirus. Three genotypes of bufaviruses have so far been detected, circulating in Tunisia, Finland and Bhutan
703:
Cotmore, SF; Agbandje-McKenna, M; Canuti, M; Chiorini, JA; Eis-Hubinger, A; Hughes, J; Mietzsch, M; Modha, S; Ogliastro, M; Pénzes, JJ; Pintel, DJ; Qiu, J; Soderlund-Venermo, M; Tattersall, P; Tijssen, P; and the ICTV Report
Consortium (2019).
933:
Yahiro T, Wangchuk S, Tshering K, Bandhari P, Zangmo S, Dorji T, Tshering K, Matsumoto T, Nishizono A, Söderlund-Venermo M, Ahmed K. 2014. Novel human bufavirus genotype 3 in children with severe diarrhea, Bhutan. Emerg Infect Dis.
640:, abbreviated to FPV), which had been shown to cause epidemics of enteritis, panleukopenia and congenital cerebellar ataxia in domestic cats. In 1978 a virus from the same species as FPV emerged that was able to infect dogs (called
260:
is relatively broad: species are defined as a cluster of similar viruses that encode a particular replication protein, typically called NS1, that is at least 85% identical to the protein encoded by other members of the species.
500:. This not only allows for VP to be expressed, but also produces multiple alternatively spliced versions of each ORF, typically called NS1, NS2(P,Y,L), VP1, and VP2. Some examples of annotated proteins from UniProt are:
953:
Phan TG, Sdiri-Loulizi K, Aouni M, Ambert-Balay K, Pothier P, Deng X, Delwart E (2014) New parvovirus in child with unexplained diarrhea, Tunisia. Emerg Infect Dis 20(11):1911–1913. doi: 10.3201/eid2011.140428
914:
Phan TG, Vo NP, Bonkoungou IJ, Kapoor A, Barro N, O’Ryan M, Kapusinszky B, Wang C, Delwart E. 2012. Acute diarrhea in West
African children: diverse enteric viruses and a novel parvovirus genus. J Virol.
974:
Tattersall, P. 2006. The evolution of parvovirus taxonomy. In: ‘‘The
Parvoviruses’’. J. Kerr, S. F. Cotmore, M. E. Bloom, R. M. Linden, and C. R. Parrish (eds), Chap. 1, pp. 5–4. Hodder Arnold, London.
924:
Väisänen E, Kuisma I, Phan TG, Delwart E, Lappalainen M, Tarkka E, Hedman K, Söderlund-Venermo M. 2014. Bufavirus in feces of patients with gastroenteritis, Finland. Emerg Infect Dis. 20(6):1078–80.
886:
Cotmore SF, Agbandje-McKenna M, Chiorini JA, Mukha DV, Pintel DJ, Qiu J, Soderlund-Venermo M, Tattersall P, Tijssen P, Gatherer D, Davison AJ. 2014. The family
Parvoviridae. Arch. Virol. 159: 1239–47.
412:
A third potential human protoparvovirus —tusavirus 1— has been reported in the feces of a single human, but whether or not it is able to infect humans or was simply ingested remains to be clarified.
852:
318:
310:
303:
296:
289:
397:
350:
342:
334:
326:
1055:
377:
369:
496:
genome has two ORFs. The non-structural ORF (NS) is the "first" on the 5' side, with the structural (VP) on the 3' end. The genome is expressed by extensive
3352:
3826:
860:
671:
4403:
3925:
3120:
1399:
4222:
3890:
3833:
3805:
3777:
3447:
3131:
3062:
2322:
2315:
1406:
1364:
1357:
4937:
4417:
4271:
3798:
3791:
2611:
1427:
1413:
1371:
4509:
4460:
4410:
4389:
3840:
1350:
1048:
4950:
4229:
3302:
3295:
617:
Kilham rat virus, isolated in 1959, was the first member of this family of small, linear, single-stranded DNA viruses to be identified.
1041:
4924:
252:
15 species are currently recognized, many containing several named viruses, virus strains, genotypes or serotypes. When applied to
983:
Ren X, Tao Y, Cui J, Suo S, Cong Y, Tijssen P. 2013. Phylogeny and evolution of porcine parvovirus. Virus
Research. 178:392–397.
829:
409:
A second virus in this genus that infects humans —cutavirus— was initially isolated from the feces of children with diarrhea.
905:
Van
Regenmortel MHV (2003) Viruses are real, virus species are man-made taxonomic constructions. Arch. Virol. 148: 2481–2488.
4955:
56:
240:, but it was renamed to eliminate confusion between members of this genus and members of the entire family
2841:
273:
4585:
4090:
3164:
385:
4902:
644:
or CPV), which rapidly spread globally, causing pandemics of severe intestinal and coronary disease.
1064:
357:
212:
625:
in the First Report of the newly created
International Committee on Taxonomy of Viruses (ICTV)
4942:
4864:
965:
and
Olivier, L. J. 1959. A latent virus of rats isolated in tissue culture. Virology 7: 428–37
757:
4988:
4983:
4978:
621:
infected pigs. In 1971 these viruses were all recognized as part of a taxonomic genus called
636:
and contained all of the aforementioned viruses plus feline panleukopenia virus (now called
1068:
505:
497:
362:
217:
51:
1006:
Hoelzer K, Parrish CR. 2010. The emergence of parvoviruses of carnivores. Vet. Res. 41:39.
224:(CPV), which causes gastrointestinal tract damage in puppies that is about 80% fatal, and
8:
4050:
1809:
1284:
40:
2368:
2003:
1989:
1581:
734:
705:
390:
225:
1018:
807:
4911:
4525:
3484:
3359:
3338:
3231:
2887:
2545:
2187:
1750:
1343:
1270:
739:
641:
637:
525:
476:
282:
278:
221:
44:
4315:
4243:
4136:
3711:
3653:
3616:
3345:
3076:
3034:
2982:
2945:
2915:
2876:
2730:
2656:
2513:
1835:
1798:
1772:
1730:
1531:
1521:
1336:
1188:
1159:
1033:
729:
721:
4916:
210:. Vertebrates serve as natural hosts. There are 15 species in the genus including
4887:
4775:
4646:
4481:
4429:
4342:
4298:
4287:
4125:
3977:
3918:
3742:
3463:
3393:
3281:
3264:
3199:
2796:
2751:
2472:
1979:
1761:
1672:
1592:
1488:
962:
4818:
4796:
4764:
4729:
4635:
4600:
4536:
4379:
4368:
4331:
4173:
4057:
3947:
3876:
3866:
3627:
3419:
3382:
3288:
3083:
2667:
2640:
2287:
2276:
2180:
2145:
2138:
2124:
2117:
2066:
2038:
1968:
1957:
1922:
1706:
1509:
1378:
1315:
1249:
1227:
1149:
4972:
4786:
4681:
4667:
4495:
4488:
4467:
4308:
4257:
4250:
4114:
4010:
3988:
3961:
3937:
3897:
3854:
3819:
3812:
3784:
3770:
3763:
3664:
3600:
3571:
3550:
3529:
3518:
3498:
3430:
3252:
3221:
3210:
3188:
3055:
2922:
2908:
2898:
2865:
2789:
2741:
2684:
2677:
2524:
2424:
2298:
2212:
2194:
2173:
2159:
2059:
1996:
1946:
1868:
1846:
1819:
1782:
1684:
1651:
1499:
1477:
1277:
1260:
1238:
1216:
1138:
1127:
993:
105:
4808:
4753:
4688:
4657:
4624:
4474:
4446:
4264:
4236:
4215:
4201:
4194:
4184:
4020:
3954:
3883:
3731:
3701:
3690:
3637:
3593:
3583:
3564:
3540:
3491:
3403:
3316:
3274:
3113:
3045:
3023:
3003:
2993:
2803:
2719:
2630:
2601:
2589:
2579:
2567:
2557:
2535:
2493:
2483:
2456:
2435:
2413:
2389:
2379:
2357:
2254:
2205:
2166:
2152:
2131:
1900:
1878:
1856:
1723:
1625:
1613:
1570:
1559:
1466:
1455:
1420:
1326:
1178:
743:
632:, which at that time included three genera, one of which retained the name
206:
200:
153:
141:
129:
117:
93:
81:
1026:
4896:
4674:
4453:
4439:
4352:
4208:
3998:
3911:
3904:
3847:
3753:
3674:
3557:
3474:
3440:
3366:
3328:
3309:
3245:
3238:
3069:
2929:
2618:
2446:
2346:
2241:
2091:
2073:
2049:
1912:
1890:
1716:
1694:
1662:
1603:
1392:
1304:
1166:
1116:
1092:
725:
651:
continued to accrue new viruses until 2014, when its name was changed to
538:
530:
518:
510:
4929:
4546:
4502:
4396:
4157:
2966:
2956:
2708:
2308:
1635:
1385:
480:
4742:
4613:
4147:
4103:
3177:
2854:
2826:
2265:
1105:
1077:
702:
32:
4858:
4881:
781:. International Committee on Taxonomy of Viruses (ICTV). March 2021
628:
The Second ICTV Report, published in 1976, established the family
195:
778:
361:(which includes H-1 parvovirus, Kilham rat virus, LuIII virus,
253:
68:
365:, mouse parvovirus, tumor virus X, and rat minute virus);
994:"International Committee on Taxonomy of Viruses (ICTV)"
474:
The capsid is thought to be made up of 60 VP proteins (
853:"ICTV Official Taxonomy: Updates since the 8th Report"
683:
International Committee on Taxonomy of Viruses (ICTV)
1063:
2366:
4970:
3032:
2728:
1770:
4773:
4644:
4340:
4296:
4123:
3391:
1977:
1486:
664:
4762:
4633:
4534:
3625:
2665:
2285:
2274:
1966:
1955:
1313:
1247:
1225:
1049:
4112:
3662:
3428:
3186:
2896:
2863:
2739:
2296:
1475:
1258:
1136:
1125:
896:of Viruses. Elsevier Academic Press, London.
4751:
4655:
4622:
4182:
3699:
3043:
2991:
2717:
2481:
2377:
2355:
1464:
1324:
802:
800:
798:
796:
3472:
2444:
1114:
1056:
1042:
31:
4727:
2954:
882:
880:
878:
733:
4740:
4611:
4598:
4145:
4101:
3175:
3162:
2852:
947:
937:
793:
4088:
847:
845:
843:
696:
487:
338:(the monkey virus Wuharv parvovirus 1);
4971:
2839:
875:
773:
771:
4863:
4862:
4726:
4597:
4087:
3161:
2838:
2239:
2238:
1089:
1037:
1090:
840:
236:. Until 2014, the genus was called
768:
13:
14:
5000:
1012:
597:early vesicular export/cell lysis
401:(protoparvovirus Zsana/2013/HUN);
220:(MVM). This genus also includes
216:for which the exemplar virus is
55:
1000:
986:
977:
968:
956:
927:
918:
908:
899:
889:
822:
779:"Virus Taxonomy: 2020 Release"
750:
706:"ICTV Virus Taxonomy Profile:
319:Eulipotyphla protoparvovirus 1
204:subfamily of the virus family
1:
658:
545:
311:Chiropteran protoparvovirus 1
180:
484:), both VP1 and VP2.
415:
277:(which includes the viruses
264:Recognized species in genus
7:
944:10.1016/j.virol.2016.06.013
714:Journal of General Virology
304:Carnivore protoparvovirus 4
297:Carnivore protoparvovirus 3
290:Carnivore protoparvovirus 2
274:Carnivore protoparvovirus 1
247:
10:
5005:
612:
606:fecal-oral or respiratory
398:Ungulate protoparvovirus 2
386:Ungulate protoparvovirus 1
4871:
4806:
4784:
4736:
4722:
4607:
4593:
4584:
4523:
4427:
4377:
4366:
4329:
4285:
4171:
4134:
4097:
4083:
4038:
4008:
3986:
3975:
3935:
3864:
3751:
3740:
3729:
3688:
3651:
3614:
3581:
3538:
3527:
3516:
3461:
3417:
3380:
3326:
3262:
3219:
3208:
3197:
3171:
3157:
3101:
3021:
2980:
2943:
2885:
2874:
2848:
2834:
2825:
2777:
2706:
2654:
2628:
2599:
2577:
2555:
2533:
2522:
2511:
2470:
2433:
2422:
2411:
2344:
2263:
2252:
2248:
2234:
2105:
2083:
2047:
2036:
2029:
1944:
1910:
1888:
1866:
1844:
1833:
1807:
1796:
1759:
1748:
1704:
1682:
1660:
1649:
1623:
1601:
1590:
1579:
1568:
1557:
1519:
1497:
1453:
1302:
1236:
1214:
1176:
1147:
1103:
1099:
1085:
1076:
830:"ICTV 10th Report (2018)
758:"ICTV 10th Report (2018)
351:Primate protoparvovirus 4
343:Primate protoparvovirus 3
335:Primate protoparvovirus 2
327:Primate protoparvovirus 1
234:Primate protoparvovirus 3
230:Primate protoparvovirus 1
178:
173:
50:
39:
30:
23:
381:(rat bufavirus SY-2015);
378:Rodent protoparvovirus 3
370:Rodent protoparvovirus 2
358:Rodent protoparvovirus 1
330:(the human bufaviruses);
213:Rodent protoparvovirus 1
857:ICTV Official Taxonomy
314:(megabat bufavirus 1);
446:Genomic segmentation
322:(Mpulungu bufavirus);
1069:virus classification
726:10.1099/jgv.0.001212
506:Minute virus of mice
498:alternative splicing
488:Genomic organization
363:minute virus of mice
218:minute virus of mice
52:Virus classification
4051:Permutotetraviridae
1810:Orthopolintovirales
1285:Malacoherpesviridae
443:Genomic arrangement
41:Electron micrograph
2369:Malgrandaviricetes
2004:Sphaerolipoviridae
1990:Matshushitaviridae
1582:Nucleocytoviricota
391:porcine parvovirus
373:(rat parvovirus 1)
226:porcine parvovirus
4966:
4965:
4865:Taxon identifiers
4856:
4855:
4852:
4851:
4848:
4847:
4844:
4843:
4840:
4839:
4836:
4835:
4832:
4831:
4828:
4827:
4718:
4717:
4714:
4713:
4710:
4709:
4706:
4705:
4702:
4701:
4698:
4697:
4580:
4579:
4576:
4575:
4572:
4571:
4568:
4567:
4564:
4563:
4560:
4559:
4556:
4555:
4526:Yunchangviricetes
4519:
4518:
4362:
4361:
4325:
4324:
4281:
4280:
4167:
4166:
4079:
4078:
4075:
4074:
4071:
4070:
4067:
4066:
4034:
4033:
4030:
4029:
3971:
3970:
3725:
3724:
3721:
3720:
3684:
3683:
3647:
3646:
3610:
3609:
3512:
3511:
3508:
3507:
3485:Carmotetraviridae
3457:
3456:
3413:
3412:
3376:
3375:
3360:Gammaflexiviridae
3353:Deltaflexiviridae
3339:Alphaflexiviridae
3232:Alphatetraviridae
3153:
3152:
3149:
3148:
3145:
3144:
3141:
3140:
3097:
3096:
3093:
3092:
3017:
3016:
3013:
3012:
2976:
2975:
2939:
2938:
2888:Chrymotiviricetes
2821:
2820:
2817:
2816:
2813:
2812:
2773:
2772:
2769:
2768:
2765:
2764:
2761:
2760:
2702:
2701:
2698:
2697:
2694:
2693:
2650:
2649:
2546:Bacilladnaviridae
2507:
2506:
2503:
2502:
2466:
2465:
2407:
2406:
2403:
2402:
2399:
2398:
2340:
2339:
2336:
2335:
2332:
2331:
2230:
2229:
2226:
2225:
2222:
2221:
2188:Portogloboviridae
2101:
2100:
2025:
2024:
2021:
2020:
2017:
2016:
2013:
2012:
1940:
1939:
1936:
1935:
1932:
1931:
1829:
1828:
1792:
1791:
1751:Preplasmiviricota
1744:
1743:
1740:
1739:
1645:
1644:
1553:
1552:
1549:
1548:
1545:
1544:
1541:
1540:
1449:
1448:
1445:
1444:
1441:
1440:
1437:
1436:
1344:Autographiviridae
1298:
1297:
1294:
1293:
1271:Alloherpesviridae
1210:
1209:
1206:
1205:
1202:
1201:
1198:
1197:
1022:: Protoparvovirus
642:canine parvovirus
638:feline parvovirus
610:
609:
526:canine parvovirus
472:
471:
420:Viruses in genus
283:feline parvovirus
279:canine parvovirus
222:canine parvovirus
187:
186:
45:canine parvovirus
4996:
4959:
4958:
4946:
4945:
4933:
4932:
4920:
4919:
4907:
4906:
4905:
4892:
4891:
4890:
4860:
4859:
4782:
4781:
4771:
4770:
4760:
4759:
4749:
4748:
4738:
4737:
4730:dsDNA-RT viruses
4724:
4723:
4653:
4652:
4642:
4641:
4631:
4630:
4620:
4619:
4609:
4608:
4601:ssRNA-RT viruses
4595:
4594:
4591:
4590:
4532:
4531:
4375:
4374:
4343:Serpentovirales
4338:
4337:
4316:Orthomyxoviridae
4294:
4293:
4244:Peribunyaviridae
4180:
4179:
4143:
4142:
4137:Chunqiuviricetes
4132:
4131:
4121:
4120:
4110:
4109:
4099:
4098:
4091:(–)ssRNA viruses
4085:
4084:
3984:
3983:
3827:Nanghoshaviridae
3749:
3748:
3738:
3737:
3712:Botourmiaviridae
3697:
3696:
3660:
3659:
3654:Howeltoviricetes
3623:
3622:
3617:Amabiliviricetes
3536:
3535:
3525:
3524:
3470:
3469:
3426:
3425:
3389:
3388:
3346:Betaflexiviridae
3217:
3216:
3206:
3205:
3195:
3194:
3184:
3183:
3173:
3172:
3165:(+)ssRNA viruses
3159:
3158:
3077:Picobirnaviridae
3041:
3040:
3035:Duplopiviricetes
3030:
3029:
2989:
2988:
2983:Vidaverviricetes
2952:
2951:
2946:Resentoviricetes
2916:Megabirnaviridae
2894:
2893:
2883:
2882:
2877:Duplornaviricota
2872:
2871:
2861:
2860:
2850:
2849:
2836:
2835:
2832:
2831:
2737:
2736:
2731:Huolimaviricetes
2726:
2725:
2715:
2714:
2663:
2662:
2657:Repensiviricetes
2531:
2530:
2520:
2519:
2514:Cressdnaviricota
2479:
2478:
2442:
2441:
2431:
2430:
2420:
2419:
2375:
2374:
2364:
2363:
2353:
2352:
2294:
2293:
2283:
2282:
2272:
2271:
2261:
2260:
2250:
2249:
2236:
2235:
2045:
2044:
2034:
2033:
1975:
1974:
1964:
1963:
1953:
1952:
1842:
1841:
1836:Tectiliviricetes
1805:
1804:
1799:Polintoviricetes
1773:Priklausovirales
1768:
1767:
1757:
1756:
1731:Marseilleviridae
1658:
1657:
1599:
1598:
1588:
1587:
1577:
1576:
1566:
1565:
1532:Papillomaviridae
1522:Zurhausenvirales
1495:
1494:
1484:
1483:
1473:
1472:
1462:
1461:
1337:Ackermannviridae
1322:
1321:
1311:
1310:
1256:
1255:
1245:
1244:
1234:
1233:
1223:
1222:
1189:Tristromaviridae
1160:Lipothrixviridae
1145:
1144:
1134:
1133:
1123:
1122:
1112:
1111:
1101:
1100:
1087:
1086:
1083:
1082:
1058:
1051:
1044:
1035:
1034:
1007:
1004:
998:
997:
990:
984:
981:
975:
972:
966:
960:
954:
951:
945:
941:
935:
931:
925:
922:
916:
912:
906:
903:
897:
893:
887:
884:
873:
872:
870:
868:
859:. Archived from
849:
838:
837:
826:
820:
819:
817:
815:
804:
791:
790:
788:
786:
775:
766:
765:
754:
748:
747:
737:
700:
694:
693:
691:
689:
680:
668:
572:Replication site
554:
553:
541:
533:
521:
513:
483:
428:
427:
60:
59:
35:
21:
20:
16:Genus of viruses
5004:
5003:
4999:
4998:
4997:
4995:
4994:
4993:
4969:
4968:
4967:
4962:
4954:
4949:
4941:
4936:
4928:
4923:
4915:
4910:
4903:Protoparvovirus
4901:
4900:
4895:
4886:
4885:
4880:
4873:Protoparvovirus
4867:
4857:
4824:
4802:
4776:Revtraviricetes
4732:
4694:
4647:Revtraviricetes
4603:
4552:
4515:
4482:Paramyxoviridae
4430:Mononegavirales
4423:
4404:Crepuscuviridae
4358:
4321:
4299:Articulavirales
4288:Insthoviricetes
4277:
4163:
4126:Negarnaviricota
4093:
4063:
4026:
4004:
3978:Stelpaviricetes
3967:
3931:
3926:Polycipiviridae
3919:Dicistroviridae
3860:
3743:Pisoniviricetes
3717:
3680:
3643:
3606:
3577:
3504:
3464:Tolucaviricetes
3453:
3409:
3394:Amarillovirales
3372:
3322:
3282:Closteroviridae
3265:Martellivirales
3258:
3200:Kitrinoviricota
3167:
3137:
3121:Polymycoviridae
3089:
3009:
2972:
2935:
2844:
2809:
2797:Finnlakeviridae
2757:
2752:Pleolipoviridae
2690:
2646:
2624:
2595:
2573:
2551:
2499:
2473:Quintoviricetes
2462:
2395:
2328:
2244:
2218:
2097:
2079:
2009:
1980:Halopanivirales
1928:
1906:
1884:
1862:
1825:
1788:
1762:Maveriviricetes
1736:
1700:
1678:
1673:Phycodnaviridae
1641:
1619:
1593:Pokkesviricetes
1537:
1515:
1489:Papovaviricetes
1433:
1400:Rountreeviridae
1290:
1194:
1172:
1095:
1072:
1062:
1015:
1010:
1005:
1001:
992:
991:
987:
982:
978:
973:
969:
961:
957:
952:
948:
942:
938:
932:
928:
923:
919:
913:
909:
904:
900:
894:
890:
885:
876:
866:
864:
851:
850:
841:
832:Protoparvovirus
828:
827:
823:
813:
811:
806:
805:
794:
784:
782:
777:
776:
769:
756:
755:
751:
701:
697:
687:
685:
678:
674:Protoparvovirus
670:
669:
665:
661:
653:Protoparvovirus
615:
584:Protoparvovirus
569:Release details
548:
537:
529:
517:
509:
494:Protoparvovirus
490:
475:
452:Protoparvovirus
422:Protoparvovirus
418:
266:Protoparvovirus
250:
191:Protoparvovirus
167:Protoparvovirus
119:Quintoviricetes
54:
25:Protoparvovirus
17:
12:
11:
5:
5002:
4992:
4991:
4986:
4981:
4964:
4963:
4961:
4960:
4947:
4934:
4921:
4908:
4893:
4877:
4875:
4869:
4868:
4854:
4853:
4850:
4849:
4846:
4845:
4842:
4841:
4838:
4837:
4834:
4833:
4830:
4829:
4826:
4825:
4823:
4822:
4819:Caulimoviridae
4814:
4812:
4804:
4803:
4801:
4800:
4797:Hepadnaviridae
4792:
4790:
4779:
4768:
4765:Artverviricota
4757:
4746:
4734:
4733:
4720:
4719:
4716:
4715:
4712:
4711:
4708:
4707:
4704:
4703:
4700:
4699:
4696:
4695:
4693:
4692:
4685:
4678:
4671:
4663:
4661:
4650:
4639:
4636:Artverviricota
4628:
4617:
4605:
4604:
4588:
4582:
4581:
4578:
4577:
4574:
4573:
4570:
4569:
4566:
4565:
4562:
4561:
4558:
4557:
4554:
4553:
4551:
4550:
4542:
4540:
4537:Goujianvirales
4529:
4521:
4520:
4517:
4516:
4514:
4513:
4506:
4499:
4492:
4485:
4478:
4471:
4464:
4457:
4450:
4443:
4435:
4433:
4425:
4424:
4422:
4421:
4414:
4407:
4400:
4393:
4385:
4383:
4380:Jingchuvirales
4372:
4369:Monjiviricetes
4364:
4363:
4360:
4359:
4357:
4356:
4348:
4346:
4335:
4332:Milneviricetes
4327:
4326:
4323:
4322:
4320:
4319:
4312:
4304:
4302:
4291:
4283:
4282:
4279:
4278:
4276:
4275:
4268:
4261:
4254:
4247:
4240:
4233:
4226:
4223:Leishbuviridae
4219:
4212:
4205:
4198:
4190:
4188:
4177:
4174:Ellioviricetes
4169:
4168:
4165:
4164:
4162:
4161:
4153:
4151:
4140:
4129:
4118:
4107:
4095:
4094:
4081:
4080:
4077:
4076:
4073:
4072:
4069:
4068:
4065:
4064:
4062:
4061:
4058:Sarthroviridae
4054:
4042:
4040:
4036:
4035:
4032:
4031:
4028:
4027:
4025:
4024:
4016:
4014:
4006:
4005:
4003:
4002:
3994:
3992:
3981:
3973:
3972:
3969:
3968:
3966:
3965:
3958:
3951:
3948:Alvernaviridae
3943:
3941:
3933:
3932:
3930:
3929:
3922:
3915:
3908:
3901:
3894:
3891:Solinviviridae
3887:
3880:
3877:Picornaviridae
3872:
3870:
3867:Picornavirales
3862:
3861:
3859:
3858:
3851:
3844:
3837:
3834:Nanhypoviridae
3830:
3823:
3816:
3809:
3806:Medioniviridae
3802:
3795:
3788:
3781:
3778:Cremegaviridae
3774:
3767:
3759:
3757:
3746:
3735:
3727:
3726:
3723:
3722:
3719:
3718:
3716:
3715:
3707:
3705:
3694:
3686:
3685:
3682:
3681:
3679:
3678:
3670:
3668:
3657:
3649:
3648:
3645:
3644:
3642:
3641:
3633:
3631:
3628:Wolframvirales
3620:
3612:
3611:
3608:
3607:
3605:
3604:
3597:
3589:
3587:
3579:
3578:
3576:
3575:
3568:
3561:
3554:
3546:
3544:
3533:
3522:
3514:
3513:
3510:
3509:
3506:
3505:
3503:
3502:
3495:
3488:
3480:
3478:
3467:
3459:
3458:
3455:
3454:
3452:
3451:
3448:Sinhaliviridae
3444:
3436:
3434:
3423:
3420:Magsaviricetes
3415:
3414:
3411:
3410:
3408:
3407:
3399:
3397:
3386:
3383:Flasuviricetes
3378:
3377:
3374:
3373:
3371:
3370:
3363:
3356:
3349:
3342:
3334:
3332:
3324:
3323:
3321:
3320:
3313:
3306:
3299:
3292:
3289:Endornaviridae
3285:
3278:
3270:
3268:
3260:
3259:
3257:
3256:
3249:
3242:
3235:
3227:
3225:
3214:
3203:
3192:
3181:
3169:
3168:
3155:
3154:
3151:
3150:
3147:
3146:
3143:
3142:
3139:
3138:
3136:
3135:
3132:Botybirnavirus
3124:
3117:
3105:
3103:
3099:
3098:
3095:
3094:
3091:
3090:
3088:
3087:
3084:Partitiviridae
3080:
3073:
3066:
3063:Curvulaviridae
3059:
3051:
3049:
3038:
3027:
3019:
3018:
3015:
3014:
3011:
3010:
3008:
3007:
2999:
2997:
2986:
2978:
2977:
2974:
2973:
2971:
2970:
2962:
2960:
2949:
2941:
2940:
2937:
2936:
2934:
2933:
2926:
2919:
2912:
2904:
2902:
2891:
2880:
2869:
2858:
2846:
2845:
2829:
2823:
2822:
2819:
2818:
2815:
2814:
2811:
2810:
2808:
2807:
2800:
2793:
2781:
2779:
2775:
2774:
2771:
2770:
2767:
2766:
2763:
2762:
2759:
2758:
2756:
2755:
2747:
2745:
2734:
2723:
2712:
2704:
2703:
2700:
2699:
2696:
2695:
2692:
2691:
2689:
2688:
2681:
2673:
2671:
2668:Geplafuvirales
2660:
2652:
2651:
2648:
2647:
2645:
2644:
2641:Redondoviridae
2636:
2634:
2626:
2625:
2623:
2622:
2615:
2607:
2605:
2597:
2596:
2594:
2593:
2585:
2583:
2575:
2574:
2572:
2571:
2563:
2561:
2553:
2552:
2550:
2549:
2541:
2539:
2528:
2517:
2509:
2508:
2505:
2504:
2501:
2500:
2498:
2497:
2489:
2487:
2476:
2468:
2467:
2464:
2463:
2461:
2460:
2452:
2450:
2439:
2428:
2417:
2409:
2408:
2405:
2404:
2401:
2400:
2397:
2396:
2394:
2393:
2385:
2383:
2372:
2361:
2350:
2342:
2341:
2338:
2337:
2334:
2333:
2330:
2329:
2327:
2326:
2323:Plectroviridae
2319:
2316:Paulinoviridae
2312:
2304:
2302:
2291:
2288:Faserviricetes
2280:
2277:Hofneiviricota
2269:
2258:
2246:
2245:
2232:
2231:
2228:
2227:
2224:
2223:
2220:
2219:
2217:
2216:
2209:
2198:
2191:
2184:
2181:Polydnaviridae
2177:
2170:
2163:
2156:
2149:
2146:Globuloviridae
2142:
2139:Fuselloviridae
2135:
2128:
2125:Bicaudaviridae
2121:
2118:Ampullaviridae
2109:
2107:
2103:
2102:
2099:
2098:
2096:
2095:
2087:
2085:
2081:
2080:
2078:
2077:
2070:
2067:Hytrosaviridae
2063:
2055:
2053:
2042:
2039:Naldaviricetes
2031:
2027:
2026:
2023:
2022:
2019:
2018:
2015:
2014:
2011:
2010:
2008:
2007:
2000:
1993:
1985:
1983:
1972:
1969:Laserviricetes
1961:
1958:Dividoviricota
1950:
1942:
1941:
1938:
1937:
1934:
1933:
1930:
1929:
1927:
1926:
1923:Corticoviridae
1918:
1916:
1908:
1907:
1905:
1904:
1896:
1894:
1886:
1885:
1883:
1882:
1874:
1872:
1864:
1863:
1861:
1860:
1852:
1850:
1839:
1831:
1830:
1827:
1826:
1824:
1823:
1815:
1813:
1802:
1794:
1793:
1790:
1789:
1787:
1786:
1778:
1776:
1765:
1754:
1746:
1745:
1742:
1741:
1738:
1737:
1735:
1734:
1727:
1720:
1712:
1710:
1707:Pimascovirales
1702:
1701:
1699:
1698:
1690:
1688:
1680:
1679:
1677:
1676:
1668:
1666:
1655:
1647:
1646:
1643:
1642:
1640:
1639:
1631:
1629:
1621:
1620:
1618:
1617:
1609:
1607:
1596:
1585:
1574:
1563:
1555:
1554:
1551:
1550:
1547:
1546:
1543:
1542:
1539:
1538:
1536:
1535:
1527:
1525:
1517:
1516:
1514:
1513:
1510:Polyomaviridae
1505:
1503:
1492:
1481:
1470:
1459:
1451:
1450:
1447:
1446:
1443:
1442:
1439:
1438:
1435:
1434:
1432:
1431:
1424:
1417:
1410:
1407:Salasmaviridae
1403:
1396:
1389:
1382:
1379:Herelleviridae
1375:
1368:
1365:Drexlerviridae
1361:
1358:Demerecviridae
1354:
1347:
1340:
1332:
1330:
1319:
1316:Caudoviricetes
1308:
1300:
1299:
1296:
1295:
1292:
1291:
1289:
1288:
1281:
1274:
1266:
1264:
1253:
1250:Herviviricetes
1242:
1231:
1228:Heunggongvirae
1220:
1212:
1211:
1208:
1207:
1204:
1203:
1200:
1199:
1196:
1195:
1193:
1192:
1184:
1182:
1174:
1173:
1171:
1170:
1163:
1155:
1153:
1150:Ligamenvirales
1142:
1131:
1120:
1109:
1097:
1096:
1080:
1074:
1073:
1061:
1060:
1053:
1046:
1038:
1032:
1031:
1024:
1014:
1013:External links
1011:
1009:
1008:
999:
985:
976:
967:
955:
946:
936:
926:
917:
907:
898:
888:
874:
863:on 19 May 2014
839:
821:
792:
767:
749:
720:(3): 367–368.
695:
662:
660:
657:
614:
611:
608:
607:
604:
601:
598:
595:
592:
589:
586:
580:
579:
576:
573:
570:
567:
564:
563:Tissue tropism
561:
558:
547:
544:
543:
542:
534:
522:
514:
489:
486:
470:
469:
466:
463:
460:
457:
454:
448:
447:
444:
441:
438:
435:
432:
417:
414:
403:
402:
394:
382:
374:
366:
354:
347:
339:
331:
323:
315:
307:
300:
293:
286:
249:
246:
194:is a genus of
185:
184:
176:
175:
171:
170:
163:
159:
158:
151:
147:
146:
139:
135:
134:
127:
123:
122:
115:
111:
110:
103:
99:
98:
91:
87:
86:
79:
72:
71:
66:
62:
61:
48:
47:
37:
36:
28:
27:
15:
9:
6:
4:
3:
2:
5001:
4990:
4987:
4985:
4982:
4980:
4977:
4976:
4974:
4957:
4952:
4948:
4944:
4939:
4935:
4931:
4926:
4922:
4918:
4913:
4909:
4904:
4898:
4894:
4889:
4883:
4879:
4878:
4876:
4874:
4870:
4866:
4861:
4821:
4820:
4816:
4815:
4813:
4811:
4810:
4805:
4799:
4798:
4794:
4793:
4791:
4789:
4788:
4787:Blubervirales
4783:
4780:
4778:
4777:
4772:
4769:
4767:
4766:
4761:
4758:
4756:
4755:
4750:
4747:
4745:
4744:
4739:
4735:
4731:
4725:
4721:
4691:
4690:
4686:
4684:
4683:
4682:Pseudoviridae
4679:
4677:
4676:
4672:
4670:
4669:
4668:Belpaoviridae
4665:
4664:
4662:
4660:
4659:
4654:
4651:
4649:
4648:
4643:
4640:
4638:
4637:
4632:
4629:
4627:
4626:
4621:
4618:
4616:
4615:
4610:
4606:
4602:
4596:
4592:
4589:
4587:
4583:
4549:
4548:
4544:
4543:
4541:
4539:
4538:
4533:
4530:
4528:
4527:
4522:
4512:
4511:
4507:
4505:
4504:
4500:
4498:
4497:
4496:Rhabdoviridae
4493:
4491:
4490:
4489:Pneumoviridae
4486:
4484:
4483:
4479:
4477:
4476:
4472:
4470:
4469:
4468:Mymonaviridae
4465:
4463:
4462:
4458:
4456:
4455:
4451:
4449:
4448:
4444:
4442:
4441:
4437:
4436:
4434:
4432:
4431:
4426:
4420:
4419:
4418:Natareviridae
4415:
4413:
4412:
4408:
4406:
4405:
4401:
4399:
4398:
4394:
4392:
4391:
4387:
4386:
4384:
4382:
4381:
4376:
4373:
4371:
4370:
4365:
4355:
4354:
4350:
4349:
4347:
4345:
4344:
4339:
4336:
4334:
4333:
4328:
4318:
4317:
4313:
4311:
4310:
4309:Amnoonviridae
4306:
4305:
4303:
4301:
4300:
4295:
4292:
4290:
4289:
4284:
4274:
4273:
4272:Wupedeviridae
4269:
4267:
4266:
4262:
4260:
4259:
4258:Phenuiviridae
4255:
4253:
4252:
4251:Phasmaviridae
4248:
4246:
4245:
4241:
4239:
4238:
4234:
4232:
4231:
4227:
4225:
4224:
4220:
4218:
4217:
4213:
4211:
4210:
4206:
4204:
4203:
4199:
4197:
4196:
4192:
4191:
4189:
4187:
4186:
4181:
4178:
4176:
4175:
4170:
4160:
4159:
4155:
4154:
4152:
4150:
4149:
4144:
4141:
4139:
4138:
4133:
4130:
4128:
4127:
4122:
4119:
4117:
4116:
4115:Orthornavirae
4111:
4108:
4106:
4105:
4100:
4096:
4092:
4086:
4082:
4060:
4059:
4055:
4053:
4052:
4047:
4044:
4043:
4041:
4037:
4023:
4022:
4018:
4017:
4015:
4013:
4012:
4011:Stellavirales
4007:
4001:
4000:
3996:
3995:
3993:
3991:
3990:
3989:Patatavirales
3985:
3982:
3980:
3979:
3974:
3964:
3963:
3962:Solemoviridae
3959:
3957:
3956:
3952:
3950:
3949:
3945:
3944:
3942:
3940:
3939:
3938:Sobelivirales
3934:
3928:
3927:
3923:
3921:
3920:
3916:
3914:
3913:
3909:
3907:
3906:
3902:
3900:
3899:
3898:Caliciviridae
3895:
3893:
3892:
3888:
3886:
3885:
3881:
3879:
3878:
3874:
3873:
3871:
3869:
3868:
3863:
3857:
3856:
3855:Tobaniviridae
3852:
3850:
3849:
3845:
3843:
3842:
3838:
3836:
3835:
3831:
3829:
3828:
3824:
3822:
3821:
3820:Mononiviridae
3817:
3815:
3814:
3813:Mesoniviridae
3810:
3808:
3807:
3803:
3801:
3800:
3799:Gresnaviridae
3796:
3794:
3793:
3792:Euroniviridae
3789:
3787:
3786:
3785:Coronaviridae
3782:
3780:
3779:
3775:
3773:
3772:
3771:Arteriviridae
3768:
3766:
3765:
3764:Abyssoviridae
3761:
3760:
3758:
3756:
3755:
3750:
3747:
3745:
3744:
3739:
3736:
3734:
3733:
3728:
3714:
3713:
3709:
3708:
3706:
3704:
3703:
3698:
3695:
3693:
3692:
3687:
3677:
3676:
3672:
3671:
3669:
3667:
3666:
3665:Cryppavirales
3661:
3658:
3656:
3655:
3650:
3640:
3639:
3635:
3634:
3632:
3630:
3629:
3624:
3621:
3619:
3618:
3613:
3603:
3602:
3601:Steitzviridae
3598:
3596:
3595:
3591:
3590:
3588:
3586:
3585:
3580:
3574:
3573:
3572:Solspiviridae
3569:
3567:
3566:
3562:
3560:
3559:
3555:
3553:
3552:
3551:Atkinsviridae
3548:
3547:
3545:
3543:
3542:
3537:
3534:
3532:
3531:
3530:Leviviricetes
3526:
3523:
3521:
3520:
3519:Lenarviricota
3515:
3501:
3500:
3499:Tombusviridae
3496:
3494:
3493:
3489:
3487:
3486:
3482:
3481:
3479:
3477:
3476:
3471:
3468:
3466:
3465:
3460:
3450:
3449:
3445:
3443:
3442:
3438:
3437:
3435:
3433:
3432:
3431:Nodamuvirales
3427:
3424:
3422:
3421:
3416:
3406:
3405:
3401:
3400:
3398:
3396:
3395:
3390:
3387:
3385:
3384:
3379:
3369:
3368:
3364:
3362:
3361:
3357:
3355:
3354:
3350:
3348:
3347:
3343:
3341:
3340:
3336:
3335:
3333:
3331:
3330:
3325:
3319:
3318:
3314:
3312:
3311:
3307:
3305:
3304:
3300:
3298:
3297:
3293:
3291:
3290:
3286:
3284:
3283:
3279:
3277:
3276:
3272:
3271:
3269:
3267:
3266:
3261:
3255:
3254:
3253:Matonaviridae
3250:
3248:
3247:
3243:
3241:
3240:
3236:
3234:
3233:
3229:
3228:
3226:
3224:
3223:
3222:Hepelivirales
3218:
3215:
3213:
3212:
3211:Alsuviricetes
3207:
3204:
3202:
3201:
3196:
3193:
3191:
3190:
3189:Orthornavirae
3185:
3182:
3180:
3179:
3174:
3170:
3166:
3160:
3156:
3134:
3133:
3128:
3125:
3123:
3122:
3118:
3116:
3115:
3110:
3107:
3106:
3104:
3100:
3086:
3085:
3081:
3079:
3078:
3074:
3072:
3071:
3067:
3065:
3064:
3060:
3058:
3057:
3056:Amalgaviridae
3053:
3052:
3050:
3048:
3047:
3042:
3039:
3037:
3036:
3031:
3028:
3026:
3025:
3020:
3006:
3005:
3001:
3000:
2998:
2996:
2995:
2990:
2987:
2985:
2984:
2979:
2969:
2968:
2964:
2963:
2961:
2959:
2958:
2953:
2950:
2948:
2947:
2942:
2932:
2931:
2927:
2925:
2924:
2923:Quadriviridae
2920:
2918:
2917:
2913:
2911:
2910:
2909:Chrysoviridae
2906:
2905:
2903:
2901:
2900:
2899:Ghabrivirales
2895:
2892:
2890:
2889:
2884:
2881:
2879:
2878:
2873:
2870:
2868:
2867:
2866:Orthornavirae
2862:
2859:
2857:
2856:
2851:
2847:
2843:
2842:dsRNA viruses
2837:
2833:
2830:
2828:
2824:
2806:
2805:
2801:
2799:
2798:
2794:
2792:
2791:
2790:Anelloviridae
2786:
2783:
2782:
2780:
2776:
2754:
2753:
2749:
2748:
2746:
2744:
2743:
2742:Haloruvirales
2738:
2735:
2733:
2732:
2727:
2724:
2722:
2721:
2716:
2713:
2711:
2710:
2705:
2687:
2686:
2685:Genomoviridae
2682:
2680:
2679:
2678:Geminiviridae
2675:
2674:
2672:
2670:
2669:
2664:
2661:
2659:
2658:
2653:
2643:
2642:
2638:
2637:
2635:
2633:
2632:
2627:
2621:
2620:
2616:
2614:
2613:
2612:Metaxyviridae
2609:
2608:
2606:
2604:
2603:
2598:
2592:
2591:
2587:
2586:
2584:
2582:
2581:
2576:
2570:
2569:
2565:
2564:
2562:
2560:
2559:
2554:
2548:
2547:
2543:
2542:
2540:
2538:
2537:
2532:
2529:
2527:
2526:
2525:Arfiviricetes
2521:
2518:
2516:
2515:
2510:
2496:
2495:
2491:
2490:
2488:
2486:
2485:
2480:
2477:
2475:
2474:
2469:
2459:
2458:
2454:
2453:
2451:
2449:
2448:
2443:
2440:
2438:
2437:
2432:
2429:
2427:
2426:
2425:Cossaviricota
2421:
2418:
2416:
2415:
2410:
2392:
2391:
2387:
2386:
2384:
2382:
2381:
2376:
2373:
2371:
2370:
2365:
2362:
2360:
2359:
2354:
2351:
2349:
2348:
2343:
2325:
2324:
2320:
2318:
2317:
2313:
2311:
2310:
2306:
2305:
2303:
2301:
2300:
2299:Tubulavirales
2295:
2292:
2290:
2289:
2284:
2281:
2279:
2278:
2273:
2270:
2268:
2267:
2262:
2259:
2257:
2256:
2251:
2247:
2243:
2242:ssDNA viruses
2237:
2233:
2215:
2214:
2213:Rhizidiovirus
2210:
2208:
2207:
2202:
2199:
2197:
2196:
2195:Thaspiviridae
2192:
2190:
2189:
2185:
2183:
2182:
2178:
2176:
2175:
2174:Plasmaviridae
2171:
2169:
2168:
2164:
2162:
2161:
2160:Halspiviridae
2157:
2155:
2154:
2150:
2148:
2147:
2143:
2141:
2140:
2136:
2134:
2133:
2129:
2127:
2126:
2122:
2120:
2119:
2114:
2111:
2110:
2108:
2104:
2094:
2093:
2089:
2088:
2086:
2082:
2076:
2075:
2071:
2069:
2068:
2064:
2062:
2061:
2060:Baculoviridae
2057:
2056:
2054:
2052:
2051:
2046:
2043:
2041:
2040:
2035:
2032:
2028:
2006:
2005:
2001:
1999:
1998:
1997:Simuloviridae
1994:
1992:
1991:
1987:
1986:
1984:
1982:
1981:
1976:
1973:
1971:
1970:
1965:
1962:
1960:
1959:
1954:
1951:
1949:
1948:
1947:Helvetiavirae
1943:
1925:
1924:
1920:
1919:
1917:
1915:
1914:
1909:
1903:
1902:
1898:
1897:
1895:
1893:
1892:
1887:
1881:
1880:
1876:
1875:
1873:
1871:
1870:
1869:Kalamavirales
1865:
1859:
1858:
1854:
1853:
1851:
1849:
1848:
1847:Belfryvirales
1843:
1840:
1838:
1837:
1832:
1822:
1821:
1820:Adintoviridae
1817:
1816:
1814:
1812:
1811:
1806:
1803:
1801:
1800:
1795:
1785:
1784:
1783:Lavidaviridae
1780:
1779:
1777:
1775:
1774:
1769:
1766:
1764:
1763:
1758:
1755:
1753:
1752:
1747:
1733:
1732:
1728:
1726:
1725:
1721:
1719:
1718:
1714:
1713:
1711:
1709:
1708:
1703:
1697:
1696:
1692:
1691:
1689:
1687:
1686:
1685:Imitervirales
1681:
1675:
1674:
1670:
1669:
1667:
1665:
1664:
1659:
1656:
1654:
1653:
1652:Megaviricetes
1648:
1638:
1637:
1633:
1632:
1630:
1628:
1627:
1622:
1616:
1615:
1611:
1610:
1608:
1606:
1605:
1600:
1597:
1595:
1594:
1589:
1586:
1584:
1583:
1578:
1575:
1573:
1572:
1567:
1564:
1562:
1561:
1556:
1534:
1533:
1529:
1528:
1526:
1524:
1523:
1518:
1512:
1511:
1507:
1506:
1504:
1502:
1501:
1500:Sepolyvirales
1496:
1493:
1491:
1490:
1485:
1482:
1480:
1479:
1478:Cossaviricota
1474:
1471:
1469:
1468:
1463:
1460:
1458:
1457:
1452:
1430:
1429:
1428:Zobellviridae
1425:
1423:
1422:
1418:
1416:
1415:
1414:Schitoviridae
1411:
1409:
1408:
1404:
1402:
1401:
1397:
1395:
1394:
1390:
1388:
1387:
1383:
1381:
1380:
1376:
1374:
1373:
1372:Guelinviridae
1369:
1367:
1366:
1362:
1360:
1359:
1355:
1353:
1352:
1348:
1346:
1345:
1341:
1339:
1338:
1334:
1333:
1331:
1329:
1328:
1323:
1320:
1318:
1317:
1312:
1309:
1307:
1306:
1301:
1287:
1286:
1282:
1280:
1279:
1278:Herpesviridae
1275:
1273:
1272:
1268:
1267:
1265:
1263:
1262:
1261:Herpesvirales
1257:
1254:
1252:
1251:
1246:
1243:
1241:
1240:
1239:Peploviricota
1235:
1232:
1230:
1229:
1224:
1221:
1219:
1218:
1217:Duplodnaviria
1213:
1191:
1190:
1186:
1185:
1183:
1181:
1180:
1175:
1169:
1168:
1164:
1162:
1161:
1157:
1156:
1154:
1152:
1151:
1146:
1143:
1141:
1140:
1139:Tokiviricetes
1135:
1132:
1130:
1129:
1128:Taleaviricota
1124:
1121:
1119:
1118:
1113:
1110:
1108:
1107:
1102:
1098:
1094:
1093:dsDNA viruses
1088:
1084:
1081:
1079:
1075:
1070:
1066:
1059:
1054:
1052:
1047:
1045:
1040:
1039:
1036:
1030:
1029:
1025:
1023:
1021:
1017:
1016:
1003:
995:
989:
980:
971:
964:
959:
950:
940:
934:20(6):1037–9.
930:
921:
911:
902:
892:
883:
881:
879:
862:
858:
854:
848:
846:
844:
835:
833:
825:
809:
803:
801:
799:
797:
780:
774:
772:
763:
761:
753:
745:
741:
736:
731:
727:
723:
719:
715:
711:
709:
699:
684:
677:
675:
667:
663:
656:
654:
650:
645:
643:
639:
635:
631:
626:
624:
618:
605:
602:
599:
596:
593:
590:
587:
585:
582:
581:
578:Transmission
577:
575:Assembly site
574:
571:
568:
566:Entry details
565:
562:
559:
556:
555:
552:
540:
535:
532:
527:
523:
520:
515:
512:
507:
503:
502:
501:
499:
495:
485:
482:
478:
467:
464:
462:Non-enveloped
461:
458:
455:
453:
450:
449:
445:
442:
439:
436:
433:
430:
429:
426:
423:
413:
410:
407:
400:
399:
395:
392:
388:
387:
383:
380:
379:
375:
372:
371:
367:
364:
360:
359:
355:
353:
352:
348:
345:
344:
340:
337:
336:
332:
329:
328:
324:
321:
320:
316:
313:
312:
308:
306:
305:
301:
299:
298:
294:
292:
291:
287:
284:
280:
276:
275:
271:
270:
269:
267:
262:
259:
255:
245:
243:
239:
235:
231:
227:
223:
219:
215:
214:
209:
208:
203:
202:
197:
193:
192:
183:
182:
177:
172:
169:
168:
164:
161:
160:
157:
156:
152:
149:
148:
145:
144:
140:
137:
136:
133:
132:
128:
125:
124:
121:
120:
116:
113:
112:
109:
108:
107:Cossaviricota
104:
101:
100:
97:
96:
92:
89:
88:
85:
84:
80:
77:
74:
73:
70:
67:
64:
63:
58:
53:
49:
46:
42:
38:
34:
29:
26:
22:
19:
4989:Virus genera
4984:Parvoviruses
4979:Parvovirinae
4872:
4817:
4809:Ortervirales
4807:
4795:
4785:
4774:
4763:
4754:Pararnavirae
4752:
4741:
4689:Retroviridae
4687:
4680:
4673:
4666:
4658:Ortervirales
4656:
4645:
4634:
4625:Pararnavirae
4623:
4612:
4545:
4535:
4524:
4510:Xinmoviridae
4508:
4501:
4494:
4487:
4480:
4475:Nyamiviridae
4473:
4466:
4461:Lispiviridae
4459:
4452:
4447:Bornaviridae
4445:
4438:
4428:
4416:
4411:Myriaviridae
4409:
4402:
4395:
4390:Aliusviridae
4388:
4378:
4367:
4351:
4341:
4330:
4314:
4307:
4297:
4286:
4270:
4265:Tospoviridae
4263:
4256:
4249:
4242:
4237:Nairoviridae
4235:
4228:
4221:
4216:Hantaviridae
4214:
4207:
4202:Arenaviridae
4200:
4195:Cruliviridae
4193:
4185:Bunyavirales
4183:
4172:
4156:
4146:
4135:
4124:
4113:
4102:
4056:
4049:
4045:
4021:Astroviridae
4019:
4009:
3997:
3987:
3976:
3960:
3955:Barnaviridae
3953:
3946:
3936:
3924:
3917:
3910:
3903:
3896:
3889:
3884:Marnaviridae
3882:
3875:
3865:
3853:
3846:
3841:Olifoviridae
3839:
3832:
3825:
3818:
3811:
3804:
3797:
3790:
3783:
3776:
3769:
3762:
3752:
3741:
3732:Pisuviricota
3730:
3710:
3702:Ourlivirales
3700:
3691:Miaviricetes
3689:
3673:
3663:
3652:
3638:Narnaviridae
3636:
3626:
3615:
3599:
3594:Blumeviridae
3592:
3584:Timlovirales
3582:
3570:
3565:Fiersviridae
3563:
3556:
3549:
3541:Norzivirales
3539:
3528:
3517:
3497:
3492:Luteoviridae
3490:
3483:
3473:
3462:
3446:
3439:
3429:
3418:
3404:Flaviviridae
3402:
3392:
3381:
3365:
3358:
3351:
3344:
3337:
3327:
3317:Virgaviridae
3315:
3308:
3301:
3294:
3287:
3280:
3275:Bromoviridae
3273:
3263:
3251:
3244:
3237:
3230:
3220:
3209:
3198:
3187:
3176:
3130:
3126:
3119:
3114:Birnaviridae
3112:
3108:
3082:
3075:
3068:
3061:
3054:
3046:Durnavirales
3044:
3033:
3024:Pisuviricota
3022:
3004:Cystoviridae
3002:
2994:Mindivirales
2992:
2981:
2965:
2955:
2944:
2928:
2921:
2914:
2907:
2897:
2886:
2875:
2864:
2853:
2804:Spiraviridae
2802:
2795:
2788:
2784:
2750:
2740:
2729:
2720:Saleviricota
2718:
2707:
2683:
2676:
2666:
2655:
2639:
2631:Recrevirales
2629:
2617:
2610:
2602:Mulpavirales
2600:
2590:Smacoviridae
2588:
2580:Cremevirales
2578:
2568:Circoviridae
2566:
2558:Cirlivirales
2556:
2544:
2536:Baphyvirales
2534:
2523:
2512:
2494:Parvoviridae
2492:
2484:Piccovirales
2482:
2471:
2457:Bidnaviridae
2455:
2445:
2436:Mouviricetes
2434:
2423:
2414:Shotokuvirae
2412:
2390:Microviridae
2388:
2380:Petitvirales
2378:
2367:
2358:Phixviricota
2356:
2345:
2321:
2314:
2307:
2297:
2286:
2275:
2264:
2255:Monodnaviria
2253:
2211:
2206:Dinodnavirus
2204:
2200:
2193:
2186:
2179:
2172:
2167:Ovaliviridae
2165:
2158:
2153:Guttaviridae
2151:
2144:
2137:
2132:Clavaviridae
2130:
2123:
2116:
2112:
2090:
2072:
2065:
2058:
2048:
2037:
2002:
1995:
1988:
1978:
1967:
1956:
1945:
1921:
1911:
1901:Adenoviridae
1899:
1889:
1879:Tectiviridae
1877:
1867:
1857:Turriviridae
1855:
1845:
1834:
1818:
1808:
1797:
1781:
1771:
1760:
1749:
1729:
1724:Iridoviridae
1722:
1715:
1705:
1693:
1683:
1671:
1661:
1650:
1634:
1626:Chitovirales
1624:
1614:Asfarviridae
1612:
1602:
1591:
1580:
1571:Bamfordvirae
1569:
1560:Varidnaviria
1558:
1530:
1520:
1508:
1498:
1487:
1476:
1467:Shotokuvirae
1465:
1456:Monodnaviria
1454:
1426:
1421:Siphoviridae
1419:
1412:
1405:
1398:
1391:
1384:
1377:
1370:
1363:
1356:
1351:Chaseviridae
1349:
1342:
1335:
1327:Caudovirales
1325:
1314:
1303:
1283:
1276:
1269:
1259:
1248:
1237:
1226:
1215:
1187:
1179:Primavirales
1177:
1165:
1158:
1148:
1137:
1126:
1115:
1104:
1027:
1019:
1002:
988:
979:
970:
958:
949:
939:
929:
920:
910:
901:
891:
865:. Retrieved
861:the original
856:
831:
824:
812:. Retrieved
808:"Viral Zone"
783:. Retrieved
760:Parvoviridae
759:
752:
717:
713:
708:Parvoviridae
707:
698:
686:. Retrieved
682:
673:
666:
652:
648:
646:
633:
630:Parvoviridae
629:
627:
622:
619:
616:
583:
560:Host details
549:
536:VP2 of CPV,
516:NS2 of MVM,
493:
491:
473:
465:Linear ssDNA
451:
421:
419:
411:
408:
404:
396:
384:
376:
368:
356:
349:
346:(cutavirus);
341:
333:
325:
317:
309:
302:
295:
288:
272:
265:
263:
258:Parvoviridae
257:
251:
242:Parvoviridae
241:
237:
233:
229:
211:
207:Parvoviridae
205:
201:Parvovirinae
199:
190:
189:
188:
179:
166:
165:
155:Parvovirinae
154:
143:Parvoviridae
142:
131:Piccovirales
130:
118:
106:
95:Shotokuvirae
94:
83:Monodnaviria
82:
75:
65:(unranked):
24:
18:
4897:Wikispecies
4675:Metaviridae
4454:Filoviridae
4440:Artoviridae
4353:Aspiviridae
4230:Mypoviridae
4209:Fimoviridae
3999:Potyviridae
3912:Secoviridae
3905:Iflaviridae
3848:Roniviridae
3754:Nidovirales
3675:Mitoviridae
3558:Duinviridae
3475:Tolivirales
3441:Nodaviridae
3367:Tymoviridae
3329:Tymovirales
3310:Togaviridae
3303:Mayoviridae
3296:Kitaviridae
3246:Hepeviridae
3239:Benyviridae
3070:Hypoviridae
2930:Totiviridae
2619:Nanoviridae
2447:Polivirales
2347:Sangervirae
2092:Nimaviridae
2074:Nudiviridae
2050:Lefavirales
1913:Vinavirales
1891:Rowavirales
1717:Ascoviridae
1695:Mimiviridae
1663:Algavirales
1604:Asfuvirales
1393:Podoviridae
1305:Uroviricota
1167:Rudiviridae
1117:Zilligvirae
915:86:11024–30
594:endocytosis
588:Vertebrates
504:NS1 of the
456:Icosahedral
150:Subfamily:
4973:Categories
4547:Yueviridae
4503:Sunviridae
4397:Chuviridae
4158:Qinviridae
4039:Unassigned
3102:Unassigned
2967:Reoviridae
2957:Reovirales
2778:Unassigned
2709:Trapavirae
2309:Inoviridae
2106:Unassigned
2084:Unassigned
2030:Unassigned
1636:Poxviridae
1386:Myoviridae
963:Kilham, L.
659:References
649:Parvovirus
634:Parvovirus
623:Parvovirus
546:Life cycle
238:Parvovirus
4888:Q18345118
4743:Riboviria
4614:Riboviria
4148:Muvirales
4104:Riboviria
3178:Riboviria
2855:Riboviria
2266:Loebvirae
1106:Adnaviria
1065:Baltimore
1020:Viralzone
688:8 January
434:Structure
416:Structure
268:include:
90:Kingdom:
4943:11907835
4930:10047239
4882:Wikidata
4046:Families
3109:Families
2785:Families
2113:Families
810:. ExPASy
744:30672729
672:"Genus:
591:Variable
437:Symmetry
248:Taxonomy
181:see text
174:Species
138:Family:
102:Phylum:
4956:1506574
867:11 June
814:15 June
735:6537627
613:History
603:Nucleus
600:Nucleus
528:(CPV),
524:VP1 of
508:(MVM),
254:viruses
198:in the
196:viruses
162:Genus:
126:Order:
114:Class:
3127:Genera
2201:Genera
785:10 May
742:
732:
679:(html)
647:Genus
539:P61826
531:P17455
519:P0DJZ2
511:P03134
440:Capsid
4938:IRMNG
4728:VII:
2840:III:
557:Genus
431:Genus
76:Realm
69:Virus
4951:NCBI
4925:GBIF
4917:6XVL
4599:VI:
3163:IV:
2240:II:
1028:ICTV
869:2014
816:2015
787:2021
740:PMID
690:2019
492:The
481:2CAS
281:and
232:and
4912:CoL
4089:V:
2827:RNA
1091:I:
1078:DNA
730:PMC
722:doi
718:100
477:PDB
468:No
459:T=1
43:of
4975::
4953::
4940::
4927::
4914::
4899::
4884::
4586:RT
4048::
3129::
3111::
2787::
2203::
2115::
877:^
855:.
842:^
795:^
770:^
738:.
728:.
716:.
712:.
681:.
655:.
479::
285:);
244:.
78::
1071:)
1067:(
1057:e
1050:t
1043:v
996:.
871:.
836:.
834:"
818:.
789:.
764:.
762:"
746:.
724::
710:"
692:.
676:"
393:)
389:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
