86:
332:
862:
797:
1566:
823:
1557:
chemical communication between individuals; 4) pick up trace nutrients in their claws, that are then ingested; and 5) favor a mutualistic relationship with populations of fur moths. More recently, a new hypothesis has emerged, which presents evidence against the previous ones and proposes that all current sloths are descendants from species that defecated on the ground, and there simply has not been enough selective pressure to abandon this behavior, since cases of predation during defecation are actually very rare.
112:
4370:
1385:
887:
64:
4396:
5575:
1690:
6331:
4384:
291:
1758:
1534:
branches, they usually eat, sleep, and even give birth hanging from branches. They sometimes remain hanging from branches even after death. On the ground, the maximum speed of sloths is 3 m (9.8 ft) per minute. Two-toed sloths are generally better able than three-toed sloths to disperse between clumps of trees on the ground.
1521:. The species of sloths recorded to host arthropods include the pale-throated three-toed sloth, the brown-throated three-toed sloth, and Linnaeus's two-toed sloth. Sloths benefit from their relationship with moths because the moths are responsible for fertilizing algae on the sloth, which provides them with nutrients.
1002:. Though data has been collected on over 33 different species of sloths by analyzing bone structures, many of the relationships between clades on a phylogenetic tree were unclear. Much of the morphological evidence collected to support the hypothesis of diphyly has been based on the structure of the inner ear.
1529:
Their limbs are adapted for hanging and grasping, not for supporting their weight. Muscle mass makes up only 25 to 30 percent of their total body weight. Most other mammals have a muscle mass that makes up 40 to 45 percent of their total body weight. Their specialised hands and feet have long, curved
1646:
Three-toed sloths go to the ground to urinate and defecate about once a week, digging a hole and covering it afterwards. They go to the same spot each time and are vulnerable to predation while doing so. Considering the large energy expenditure and dangers involved in the journey to the ground, this
1556:
Sloths descend about once every eight days to defecate on the ground. The reason and mechanism behind this behavior have long been debated among scientists. There are at least five hypotheses: 1) fertilize trees when feces are deposited at the base of the tree; 2) cover feces and avoid predation; 3)
1537:
Sloths are surprisingly strong swimmers and can reach speeds of 13.5 m (44 ft) per minute. They use their long arms to paddle through the water and can cross rivers and swim between islands. Sloths can reduce their already slow metabolism even further and slow their heart rate to less than
2152:
Presslee, S.; Slater, G. J.; Pujos, F.; Forasiepi, A. M.; Fischer, R.; Molloy, K.; Mackie, M.; Olsen, J. V.; Kramarz, A.; Taglioretti, M.; Scaglia, F.; Lezcano, M.; Lanata, J. L.; Southon, J.; Feranec, R.; Bloch, J.; Hajduk, A.; Martin, F. M.; Gismondi, R. S.; Reguero, M.; de Muizon, C.; Greenwood,
1533:
Sloths move only when necessary and even then, very slowly. They usually move at an average speed of 4 m (13 ft) per minute but can move at a marginally higher speed of 4.5 m (15 ft) per minute if they are in immediate danger from a predator. While they sometimes sit on top of
851:
lifestyle, "one of the most striking examples of convergent evolution known among mammals". The ancient
Xenarthra included a significantly greater variety of species, with a wider distribution, than those of today. Ancient sloths were mostly terrestrial, and some reached sizes that rival those of
1412:
Sloths can be 60 to 80 cm (24 to 31 in) long and, depending on the species, weigh from 3.6 to 7.7 kg (7.9 to 17.0 lb). Two-toed sloths are slightly larger than three-toed sloths. Sloths have long limbs and rounded heads with tiny ears. Three-toed sloths also have stubby tails
1665:
for three-toed, and 12 months for two-toed. Newborns stay with their mother for about five months. In some cases, young sloths die from a fall indirectly because the mothers prove unwilling to leave the safety of the trees to retrieve the young. Females normally bear one baby every year, but
408:(two–toed sloths). Despite this traditional naming, all sloths have three toes on each rear limb-- although two-toed sloths have only two digits on each forelimb. The two groups of sloths are from different, distantly related families, and are thought to have evolved their morphology via
620:, which inhabits tropical rainforests in northern South America. It is similar in appearance to, and often confused with, the brown-throated three-toed sloth, which has a much wider distribution. Genetic evidence indicates the two species diverged around six million years ago.
493:) also mean "lazy" or similar. Their slowness permits their low-energy diet of leaves and avoids detection by predatory hawks and cats that hunt by sight. Sloths are almost helpless on the ground but are able to swim. The shaggy coat has grooved hair that is host to symbiotic
1651:
relationship with sloths, as they live in the fur and promote growth of algae, which the sloths eat. Individual sloths tend to spend the bulk of their time feeding on a single "modal" tree; by burying their excreta near the trunk of that tree, they may also help nourish it.
2010:
O'Leary, Maureen A.; Bloch, Jonathan I.; Flynn, John J.; Gaudin, Timothy J.; Giallombardo, Andres; Giannini, Norberto P.; Goldberg, Suzann L.; Kraatz, Brian P.; Luo, Zhe-Xi (8 February 2013). "The
Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals".
1796:
cares for sloths. It has rehabilitated and returned about 130 individuals to the wild. However, a report in May 2016 featured two former veterinarians from the facility who were critical of the sanctuary's efforts, accusing it of mistreating the animals.
1647:
behaviour has been described as a mystery. Recent research shows that moths, which live in the sloth's fur, lay eggs in the sloth's feces. When they hatch, the larvae feed on the feces, and when mature fly up onto the sloth above. These moths may have a
1458:, but because sloths spend so much time with their limbs above their bodies, their hairs grow away from the extremities to provide protection from the elements while they hang upside down. In most conditions, the fur hosts symbiotic algae, which provide
1660:
The pale- and brown-throated three-toed sloths mate seasonally, while the maned three-toed sloth breeds at any time of the year. The reproduction of pygmy three-toed sloths is currently unknown. Litters are of one newborn only, after six months'
1673:
The average lifespan of two-toed sloths in the wild is currently unknown due to a lack of full-lifespan studies in a natural environment. Median life expectancy in human care is about 16 years, with one individual at the
1013:
sequences fall in line with the diphyly (convergent evolution) hypothesis but have overturned some of the other conclusions obtained from morphology. These investigations consistently place two-toed sloths close to
1530:
claws to allow them to hang upside down from branches without effort, and are used to drag themselves along the ground, since they cannot walk. On three-toed sloths, the arms are 50 percent longer than the legs.
1783:. Their claws also provide another, unexpected deterrent to human hunters; when hanging upside-down in a tree, they are held in place by the claws themselves and often do not fall down even if shot from below.
998:) and a single species of the two-toed type will jointly predominate. Based on morphological comparisons, it was thought the two-toed sloths nested phylogenetically within one of the divisions of the extinct
2722:
667:: ground sloths that existed for about 35 million years and went extinct about 11,000 years ago. This group was formerly thought to include both the two-toed sloths and the extinct Greater Antilles sloths.
1356:
Ground sloths disappeared from both North and South
America shortly after the appearance of humans about 11,000 years ago. Evidence suggests human hunting contributed to the extinction of the American
1376:
and some other
Caribbean sloths survived until about 5,000 years ago, long after ground sloths had died out on the mainland, but then went extinct when humans finally colonized the Greater Antilles.
3019:
Moreno, Ricardo S.; Kays, Roland W.; Samudio, Rafael (24 August 2006). "Competitive
Release in Diets of Ocelot (Leopardus pardalis) and Puma (Puma concolor) after Jaguar (Panthera onca) Decline".
1447:, meaning their body temperature may vary according to the environment, normally ranging from 25 to 35 °C (77 to 95 °F), but able to drop to as low as 20 °C (68 °F), inducing
4255:
2681:
Delsuc, F.; Kuch, M.; Gibb, G. C.; Karpinski, E.; Hackenberger, D.; Szpak, P.; Martínez, J. G.; Mead, J. I.; McDonald, H. G.; MacPhee, R.D.E.; Billet, G.; Hautier, L.; Poinar, H. N. (2019).
979:. They presumably waded and paddled in the water for short period, but over a span of 4 million years, they eventually evolved into swimming creatures, becoming specialist bottom feeders of
526:
believed to have evolved in the continent of South
America around 60 million years ago. One study found that xenarthrans broke off from other placental mammals around 100 million years ago.
1443:
rates (less than half of that expected for a mammal of their size), and low body temperatures: 30 to 34 °C (86 to 93 °F) when active, and still lower when resting. Sloths are
907:
Sloths arose in South
America during a long period of isolation and eventually spread to a number of the Caribbean islands as well as North America. It is thought that swimming led to
3234:
3868:
2954:"Molecular evidence for a diverse green algal community growing in the hair of sloths and a specific association with Trichophilus welckeri(Chlorophyta, Ulvophyceae)"
1553:
Sloths are solitary animals that rarely interact with one another except during breeding season, though female sloths do sometimes congregate, more so than do males.
3653:
Pauli, J. N., Mendoza, J. E., Steffan, S. A., Carey, C. C., Weimer, P. J., & Peery, M. Z. (2014). A syndrome of mutualism reinforces the lifestyle of a sloth.
843:
The common ancestor of the two existing sloth genera dates to about 28 million years ago, with similarities between the two- and three- toed sloths an example of
1631:
browsing. Leaves, their main food source, provide very little energy or nutrients, and do not digest easily, so sloths have large, slow-acting, multi-chambered
5935:
1988:
6254:
5921:
5295:
3778:
2859:
1786:
Sloths are victims of animal trafficking where they are sold as pets. However, they generally make very poor pets, as they have such a specialized ecology.
5785:
1861:"The evolution of armadillos, anteaters and sloths depicted by nuclear and mitochondrial phylogenies: implications for the status of the enigmatic fossil
6289:
6261:
4312:
990:
Both types of extant tree sloth tend to occupy the same forests; in most areas, a particular species of the somewhat smaller and generally slower-moving
1749:. Sloths' lower metabolism confines them to the tropics, and they adopt thermoregulation behaviors of cold-blooded animals such as sunning themselves.
5820:
5348:
5185:
2231:
6268:
6191:
6171:
5473:
5267:
5192:
1436:
Sloths have colour vision but have poor visual acuity. They also have poor hearing. Thus, they rely on their sense of smell and touch to find food.
5914:
5700:
5669:
5508:
5206:
4247:
4225:
3294:
Mendel, Frank C. (1 January 1985). "Use of Hands and Feet of Three-Toed Sloths (Bradypus variegatus) during
Climbing and Terrestrial Locomotion".
2205:
6226:
6205:
5972:
5865:
5677:
2796:
222:
4343:
6233:
5799:
5446:
2744:
Amson, E.; Argot, C.; McDonald, H. G.; de Muizon, C. (2015). "Osteology and functional morphology of the axial postcranium of the marine sloth
426:
around 12,000 years ago, along with most large animals across the
Americas. The extinction correlates in time with the arrival of humans, but
6462:
6024:
5996:
5806:
5771:
5715:
5653:
5487:
5398:
3686:
Voirin, B., Kays, R., Wikelski, M., & Lowman, M. (2013). Why Do Sloths Poop on the Ground? In M. Lowman, S. Devy, & T. Ganesh (eds).
2889:
5384:
5355:
6303:
6101:
5827:
5730:
5391:
942:. The latter development, about 3 million years ago, allowed megatheriids and nothrotheriids to also invade North America as part of the
674:: ground sloths that existed for about 23 million years and went extinct about 11,000 years ago; this family included the largest sloths.
4155:
4104:
3516:
1666:
sometimes sloths' low level of movement actually keeps females from finding males for longer than one year. Sloths are not particularly
6523:
5764:
5057:
1639:
break down the tough leaves. As much as two-thirds of a well-fed sloth's body weight consists of the contents of its stomach, and the
6247:
6150:
4074:
3242:
3856:
1927:
1709:
Although habitat is limited to the tropical rainforests of
Central and South America, in that environment sloths are successful. On
4804:
4044:
4014:
3465:
1678:
3800:
Pauli, Jonathan N.; Mendoza, Jorge E.; Steffan, Shawn A.; Carey, Cayelan C.; Weimer, Paul J.; Peery, M. Zachariah (7 March 2014).
561:
The Folivora are divided into at least eight families, only two of which have living species; the remainder are entirely extinct (
6436:
412:
from terrestrial ancestors. Besides the extant species, many species of ground sloths ranging up to the size of elephants (like
681:: ground sloths that lived from approximately 11.6 million to 11,000 years ago. As well as ground sloths, this family included
2790:; MacPhee, R. D. E.; Jull, A. J. T.; McDonald, H. G.; Woods, C. A.; Iturralde-Vinent, M.; Hodgins, G. W. L. (16 August 2005).
2427:
Muizon, C. de; McDonald, H. G.; Salas, R.; Urbina, M. (June 2004). "The evolution of feeding adaptations of the aquatic sloth
6454:
4141:
3502:
2558:
2249:
1832:
1454:
The outer hairs of sloth fur grow in a direction opposite from that of other mammals. In most mammals, hairs grow toward the
1424:. Two-toed sloths have five to seven, while three-toed sloths have eight or nine. The other mammals not having seven are the
6441:
3205:
2564:
554:. These names are from the Latin 'leaf eater' and 'hairy', respectively. Pilosa is one of the smallest of the orders of the
3552:
1368:
may have also played a role, although previous similar glacial retreats were not associated with similar extinction rates.
3603:
Montgomery, G. G., & Sunquist, M. E. (1975). Impact of Sloths on Neotropical Forest Energy Flow and Nutrient Cycling.
1360:. Ground sloth remains found in both North and South America indicate that they were killed, cooked, and eaten by humans.
1345:
The marine sloths of South America's Pacific coast became extinct at the end of the Pliocene following the closing of the
3272:
4281:
3726:
1545:. Three-toed sloths are mostly nocturnal but can be active in the day. They spend 90 per cent of their time motionless.
3752:
2279:"Morphology, molecular phylogeny, and taxonomic inconsistencies in the study of Bradypus sloths (Pilosa: Bradypodidae)"
879:
365:. Noted for their slowness of movement, tree sloths spend most of their lives hanging upside down in the trees of the
3582:
3165:"Sloth biology: an update on their physiological ecology, behavior and role as vectors of arthropods and arboviruses"
3113:"Sloth biology: an update on their physiological ecology, behavior and role as vectors of arthropods and arboviruses"
2926:
1624:(plant eaters), with a limited diet of leaves from only a few trees, and no other mammal digests its food as slowly.
1486:
3331:"Natal dispersal of tree sloths in a human-dominated landscape: Implications for tropical biodiversity conservation"
1980:
1822:
6467:
4428:
900:
3774:
1789:
1361:
810:
427:
4304:
3892:"Social behavior between mothers × young of sloths Bradypus variegatus SCHINZ, 1825 (Xenarthra: Bradypodidae)"
1793:
4797:
4611:
4602:
1577:
743:
705:
4374:
2235:
758:
in the north to western Ecuador in the south, and the other in eastern Peru, western Brazil, and northern
6449:
4217:
4133:
3494:
2241:
1859:
Delsuc, Frédéric; Catzeflis, François M.; Stanhope, Michael J.; Douzery, Emmanuel J. P. (7 August 2001).
1776:
1620:, ranging over up to 140 hectares (350 acres). Three-toed sloths, on the other hand, are almost entirely
1569:
423:
1353:(and which would have also made thermoregulation difficult for the sloths, with their slow metabolism).
6508:
2158:
1055:
The following sloth family phylogenetic tree is based on collagen and mitochondrial DNA sequence data.
943:
17:
3864:
2882:"Sticking their necks out for evolution: Why sloths and manatees have unusually long (or short) necks"
2588:"Phylogenetic Relationships among Sloths (Mammalia, Xenarthra, Tardigrada): The Craniodental Evidence"
2388:"Phylogenetic relationships among sloths (Mammalia, Xenarthra, Tardigrada): the craniodental evidence"
4388:
4335:
1792:
is known for caring, rehabilitating and releasing sloths back into the wild. Also in Costa Rica, the
912:
574:
442:
but became extinct after humans settled the archipelago in the mid-Holocene, around 6,000 years ago.
111:
4201:
2851:
2334:
4790:
3390:
2881:
1675:
1346:
836:
651:
3699:
Monge Nájera, J. (2021). Why sloths defecate on the ground: rejection of the mutualistic model.
1587:
Baby sloths learn what to eat by licking the lips of their mother. All sloths eat the leaves of
938:
were able to colonise North America about 9 million years ago, well before the formation of the
771:: ground sloths that existed for about 23 million years and went extinct about 11,000 years ago.
6350:
4530:
1738:
1694:
1349:; the closing caused a cooling trend in the coastal waters which killed off much of the area's
861:
639:
431:
331:
35:
2532:
85:
4127:
4096:
3488:
1765:
1746:
1710:
1455:
1397:
1044:
578:
6480:
1541:
Wild brown-throated three-toed sloths sleep on average 9.6 hours a day. Two-toed sloths are
298:
Red: two-toed sloth,Blue: three-toed sloth,Purple: both two-toed sloth and three-toed sloth
6423:
6397:
4557:
4421:
4129:
Mammals of the Neotropics, Volume 3: The Central Neotropics: Ecuador, Peru, Bolivia, Brazil
3952:
3490:
Mammals of the Neotropics, Volume 3: The Central Neotropics: Ecuador, Peru, Bolivia, Brazil
3342:
2965:
2805:
2632:
2440:
2173:
2020:
1962:
1474:. Because of the algae, sloth fur is a small ecosystem of its own, hosting many species of
1389:
844:
598:
318:
4066:
497:
which camouflage the animal in the trees and provide it nutrients. The algae also nourish
8:
4548:
3633:
Liberg, O. (1980). Spacing patterns in a population of rural free roaming domestic cats.
2787:
1923:
613:
366:
358:
94:
72:
4036:
4006:
3956:
3461:
3346:
2969:
2809:
2636:
2444:
2177:
2024:
6513:
6388:
5574:
5302:
4400:
3983:
3940:
3832:
3801:
3442:
3434:
3311:
3093:
3044:
2996:
2953:
2828:
2791:
2765:
2655:
2620:
2508:
2484:"Gradual adaptation of bone structure to aquatic lifestyle in extinct sloths from Peru"
2483:
2464:
2456:
2255:
2197:
2116:
2081:
2062:
1897:
1860:
1636:
1538:
a third of normal, allowing them to hold their breath underwater for up to 40 minutes.
1421:
409:
303:
106:
31:
30:
This article is about the Central and South American mammal. For the cardinal sin, see
1565:
796:
453:, literally means "laziness", and their common names in several other languages (e.g.
6518:
6475:
6410:
5813:
4772:
4748:
4739:
4147:
4137:
3988:
3970:
3921:
3913:
3837:
3508:
3498:
3197:
3189:
3142:
3134:
3085:
3064:"Food Habits of the Harpy Eagle, a Top Predator from the Amazonian Rainforest Canopy"
3036:
3001:
2983:
2833:
2714:
2682:
2660:
2604:
2587:
2554:
2513:
2409:
2404:
2387:
2245:
2201:
2189:
2121:
2103:
2066:
2054:
2046:
1902:
1884:
1838:
1828:
1667:
1010:
939:
908:
606:
581:
group that arose about 32 million years ago and became extinct about 5,000 years ago.
478:
343:
6330:
4406:
4178:
3908:
3891:
3446:
3184:
3129:
3112:
3097:
3048:
2782:
2769:
2468:
2311:
822:
6094:
5623:
5274:
5126:
4513:
4503:
4196:
3978:
3960:
3903:
3827:
3817:
3426:
3350:
3303:
3179:
3124:
3075:
3028:
2991:
2973:
2823:
2813:
2757:
2704:
2694:
2650:
2640:
2599:
2546:
2503:
2495:
2448:
2399:
2366:
2329:
2290:
2181:
2111:
2093:
2036:
2028:
1955:
1892:
1876:
1761:
1114:
991:
916:
775:
591:
539:
510:
486:
470:
445:
Sloths are so named because of their very low metabolism and deliberate movements.
439:
278:
228:
77:
6503:
6415:
6402:
6275:
6015:
5983:
5737:
5439:
5009:
4942:
4688:
4414:
3965:
3164:
2536:
2098:
1734:
1702:
1256:
947:
832:
678:
632:
462:
454:
374:
252:
50:
3544:
2683:"Ancient Mitogenomes Reveal the Evolutionary History and Biogeography of Sloths"
6373:
6282:
6219:
6198:
6080:
5907:
5900:
5838:
5723:
5691:
5605:
5592:
5542:
5418:
5333:
5309:
5253:
5246:
5199:
5164:
5112:
4951:
4931:
4921:
4707:
4698:
4670:
4661:
4585:
4575:
3062:
Aguiar-Silva, F. Helena; Sanaiotti, Tânia M.; Luz, Benjamim B. (1 March 2014).
2949:
2783:
2154:
1209:
1033:
925:
871:
827:
699:
664:
234:
99:
3396:
3264:
3032:
2761:
2699:
2645:
2185:
1039:, with both two-toed sloths and Greater Antilles sloths being moved away from
6497:
6182:
6116:
6087:
6059:
6038:
6031:
5755:
5480:
5453:
5171:
5135:
5094:
4187:
4151:
3974:
3917:
3813:
3718:
3512:
3193:
3138:
3089:
3040:
2987:
2952:; Friedl, Thomas; Chiarello, Adriano G.; Blomster, Jaanika (1 January 2010).
2792:"Asynchronous extinction of late Quaternary sloths on continents and islands"
2550:
2413:
2320:
2237:
Mammals of South America, Volume 1: Marsupials, Xenarthrans, Shrews, and Bats
2107:
2050:
1888:
1818:
1444:
1271:
1151:
1029:
960:
934:
806:
671:
370:
266:
246:
4277:
3748:
3355:
3330:
2978:
2818:
2295:
2278:
2032:
1842:
750:) which inhabits tropical forests. It has two separate ranges, split by the
6212:
6164:
6157:
6129:
6066:
6003:
5643:
5409:
5362:
5340:
5320:
5232:
5149:
5027:
4912:
4901:
4651:
3992:
3925:
3841:
3822:
3201:
3146:
3080:
3063:
3005:
2837:
2718:
2664:
2517:
2499:
2193:
2125:
2058:
1906:
1880:
1475:
1224:
1136:
1077:
1036:
999:
951:
930:
896:
768:
737:
683:
570:
435:
362:
272:
214:
202:
3574:
2918:
2482:
Amson, E.; Muizon, C. de; Laurin, M.; Argot, C.; Buffrénil, V. de (2014).
1384:
534:
are also included among Xenarthra. The earliest xenarthrans were arboreal
6382:
6296:
6240:
6141:
6045:
5928:
5893:
5858:
5792:
5744:
5515:
5501:
5464:
5425:
5376:
5369:
5281:
5260:
5156:
4971:
4961:
4891:
4539:
2621:"Complex body size trends in the evolution of sloths (Xenarthra: Pilosa)"
1726:
1621:
1498:
1471:
1025:
956:
891:
854:
801:
717:
624:
520:
494:
419:
414:
4369:
2709:
2426:
886:
63:
6052:
5962:
5879:
5851:
5778:
5661:
5178:
5050:
5036:
3438:
3315:
2460:
1772:
1597:
1581:
1479:
1459:
1440:
1401:
1372:
1015:
562:
546:, stubby teeth, and small brains. Sloths are in the taxonomic suborder
498:
3670:
Chiarello, A. G. (2008). Sloth ecology: an overview of field studies.
2041:
605:
is the most common of the extant species of sloth, which inhabits the
6073:
5872:
5288:
5043:
4844:
4475:
2542:
2533:"The sloths of the West Indies: a systematic and phylogenetic review"
1662:
1648:
1640:
1542:
1482:
1357:
1020:
920:
866:
786:
713:
535:
531:
516:
404:
347:
163:
123:
6344:
3307:
3111:
Gilmore, D. P.; Da Costa, C. P.; Duarte, D. P. F. (1 January 2001).
1689:
6367:
5886:
5707:
5633:
5432:
5213:
5078:
4978:
4875:
4832:
4722:
4639:
4469:
4463:
4457:
4451:
4395:
4336:"Famous Sloth Sanctuary Is A Nightmare For Animals, Ex-Workers Say"
3941:"Unexpected Strong Polygyny in the Brown-Throated Three-Toed Sloth"
3430:
3417:
Britton, S. W. (1 January 1941). "Form and Function in the Sloth".
2948:
Suutari, Milla; Majaneva, Markus; Fewer, David P.; Voirin, Bryson;
1780:
1632:
1628:
1589:
1490:
1350:
1006:
980:
848:
778:: collagen sequence data indicates this group is more distant from
755:
721:
547:
527:
430:
has also been suggested to have contributed. Members of an endemic
398:
378:
143:
54:
3616:
Bailey, T. N. (1974). Social organization in a bobcat population.
2452:
2371:
2350:
1827:(3rd ed.). Johns Hopkins University Press. pp. 100–101.
5494:
5142:
5071:
5064:
4782:
3230:
1869:
Proceedings of the Royal Society of London B: Biological Sciences
1824:
Mammal Species of the World: A Taxonomic and Geographic Reference
1722:
1718:
1617:
1605:
1601:
1494:
1425:
1365:
984:
976:
780:
759:
725:
647:
389:
6428:
4248:"Sloths: Hottest-Selling Animal in Colombia's Illegal Pet Trade"
4859:
4838:
4826:
4813:
4438:
4383:
1714:
1613:
1609:
1514:
1467:
1463:
1448:
1417:
875:
733:
655:
555:
551:
543:
523:
393:
382:
350:
173:
153:
133:
4176:
3372:
International Series of Monographs in Pure and Applied Biology
2080:
Svartman, Marta; Stone, Gary; Stanyon, Roscoe (21 July 2006).
1858:
290:
3802:"A syndrome of mutualism reinforces the lifestyle of a sloth"
1502:
968:
751:
3939:
Pauli, Jonathan N.; Peery, M. Zachariah (19 December 2012).
2151:
3392:
Swimming sloth - Planet Earth II: Islands Preview - BBC One
2748:(Mammalia, Tardigrada) with paleobiological implications".
2743:
2619:
Raj Pant, Sara; Goswami, Anjali; Finarelli, John A (2014).
2082:"The Ancestral Eutherian Karyotype Is Present in Xenarthra"
2009:
1757:
1737:, is classified as "vulnerable", while the island-dwelling
1518:
1510:
1506:
964:
729:
4511:
3163:
Gilmore, D. P.; Da Costa, C. P.; Duarte, D. P. F. (2001).
3158:
3156:
2947:
2538:
Biogeography of the West Indies: Patterns and Perspectives
4583:
4125:
3486:
3328:
3799:
3329:
Garcés-Restrepo, M.F.; Pauli, J.N.; Peery, M.Z. (2018).
3061:
2618:
1670:
and several zoos have received sloths of the wrong sex.
1043:. Greater Antilles sloths are now placed in a separate,
687:, a genus of either semiaquatic or fully aquatic sloths.
3775:"The greatest mystery of sloth pooping has been solved"
3655:
Proceedings of the Royal Society B: Biological Sciences
3153:
2680:
2488:
Proceedings of the Royal Society B: Biological Sciences
1513:. Sloths have a highly specific community of commensal
1489:. These include biting and blood-sucking flies such as
955:
of the west coast of South America became adapted to a
790:
is, so it has been elevated back to full family status.
558:
class; its only other suborder contains the anteaters.
3370:
Goffart, M. (1971). "Function and Form in the sloth".
2541:. Boca Raton, London, New York, and Washington, D.C.:
2481:
2309:
1717:, sloths have been estimated to constitute 70% of the
1018:
and three-toed sloths within Megatherioidea, close to
377:. Sloths are considered to be most closely related to
4436:
3162:
3110:
5959:
5688:
5649:
5639:
5629:
5229:
5123:
5024:
4967:
4957:
4927:
4897:
4887:
4126:
Eisenberg, John F.; Redford, Kent H. (15 May 2000).
3487:
Eisenberg, John F.; Redford, Kent H. (15 May 2000).
3172:
Brazilian Journal of Medical and Biological Research
3117:
Brazilian Journal of Medical and Biological Research
2079:
418:) inhabited both North and South America during the
2276:
1954:
975:entered the coastal habitat beginning in the late
4333:
4305:"Sloth sanctuary nurtures animals back to health"
4302:
3235:"Can Moths Explain Why Sloths Poo on the Ground?"
3018:
2270:
1978:
27:Group of tree dwelling mammals noted for slowness
6495:
4170:
4007:"Manly secret of non-mating sloth at London Zoo"
1981:"The Strange Symbiosis Between Sloths and Moths"
1413:about 5 to 6 cm (2.0 to 2.4 in) long.
501:, some species of which exist solely on sloths.
3749:"The 'Busy' Life of the Sloth | BBC Earth"
2159:"Palaeoproteomics resolves sloth relationships"
422:Epoch. However, they became extinct during the
4177:Chiarello, A. & Moraes-Barros, N. (2014).
4119:
4037:"Same-sex sloths dash Drusillas breeding plan"
3889:
3542:
2147:
2145:
2143:
2141:
2139:
2137:
2135:
1681:reaching an age of 49 years before her death.
188:Delsuc, Catzeflis, Stanhope, and Douzery, 2001
5603:
5541:
4798:
4422:
4202:10.2305/IUCN.UK.2014-1.RLTS.T3036A47436575.en
3890:Soares, C. A.; Carneiro, R. S. (1 May 2002).
3661:(1778), 20133006. DOI: 10.1098/rspb.2013.3006
3480:
2676:
2674:
2475:
2335:10.2305/IUCN.UK.2014-1.RLTS.T4778A47439751.en
1979:Bennington-Castro, Joseph (23 January 2014).
1032:. They make the previously recognized family
5023:
2530:
1854:
1852:
1721:of arboreal mammals. Four of the six living
609:in the forests of South and Central America.
4637:
2348:
2342:
2132:
1949:
1947:
1945:
4874:
4805:
4791:
4429:
4415:
3938:
3538:
3536:
3534:
2671:
2420:
2379:
381:, together making up the xenarthran order
289:
84:
62:
4200:
3982:
3964:
3907:
3848:
3831:
3821:
3354:
3322:
3183:
3128:
3079:
2995:
2977:
2827:
2817:
2737:
2708:
2698:
2654:
2644:
2603:
2592:Zoological Journal of the Linnean Society
2507:
2403:
2392:Zoological Journal of the Linnean Society
2370:
2333:
2294:
2277:Moraes-Barros, M.C.; et al. (2011).
2115:
2097:
2040:
1974:
1972:
1896:
1849:
1771:The majority of recorded sloth deaths in
1725:are currently rated "least concern"; the
1407:
504:
3607:, 69–98. DOI:10.1007/978-3-642-88533-4_7
2776:
2612:
2535:. In Woods, C.A.; Sergile, F.E. (eds.).
1942:
1756:
1688:
1564:
1383:
885:
860:
821:
795:
330:
4659:
4215:
3531:
3462:"Article "Sloth's Lazy Image 'A Myth'""
3416:
3369:
3229:
2229:
2157:; Collins, M.; MacPhee, R.D.E. (2019).
1816:
1047:branch of the sloth evolutionary tree.
959:and, eventually, perhaps fully aquatic
754:. One population is found from eastern
14:
6496:
4228:from the original on 12 September 2019
4158:from the original on 19 September 2020
4017:from the original on 19 September 2020
3854:
3716:
3690:(pp. 195-199). Springer, New York, NY.
3519:from the original on 19 September 2020
3459:
3293:
2913:
2911:
2909:
2907:
2585:
2579:
2385:
2223:
2211:from the original on 12 September 2020
1969:
1957:The Land and Wildlife of South America
1812:
1810:
1643:can take a month or more to complete.
1005:Recently obtained molecular data from
6349:
6348:
5590:
4857:
4786:
4489:
4410:
3755:from the original on 16 February 2021
3682:
3680:
3649:
3647:
3545:"Sloths: The World's Slowest Mammals"
2531:White, J.L.; MacPhee, R.D.E. (2001).
2524:
1926:. The Sloth Conservation Foundation.
1918:
1916:
1733:), which inhabits Brazil's dwindling
4346:from the original on 18 January 2021
4077:from the original on 16 January 2021
4047:from the original on 5 December 2020
3781:from the original on 27 January 2021
3585:from the original on 8 November 2020
3555:from the original on 4 December 2020
3497:. pp. 624 (see pp. 94–95, 97).
3211:from the original on 6 November 2020
2303:
1991:from the original on 1 December 2017
1930:from the original on 1 December 2017
361:tree sloths and extinct terrestrial
4303:Sevcenko, Melanie (17 April 2013).
4216:Dowling, Stephen (29 August 2019).
4188:IUCN Red List of Threatened Species
3543:Alina Bradford (26 November 2018).
3468:from the original on 1 January 2021
3388:
2904:
2321:IUCN Red List of Threatened Species
1807:
1364:that came with the end of the last
24:
5591:
4812:
4284:from the original on 31 March 2021
3861:Cecropia: Supplemental Information
3677:
3644:
3618:The Journal of Wildlife Management
3265:"What Does It Mean to Be a Sloth?"
2892:from the original on 26 April 2019
2862:from the original on 11 April 2019
2725:from the original on 23 March 2022
2433:Journal of Vertebrate Paleontology
1913:
1752:
1227:(three-toed sloths)
25:
6535:
6524:Extant Rupelian first appearances
4362:
4334:Schelling, Ameena (19 May 2016).
4107:from the original on 17 July 2019
3729:from the original on 6 March 2021
3275:from the original on 15 June 2019
2310:Plese, T.; Chiarello, A. (2014).
1487:arthropods associated with sloths
599:brown-throated three-toed sloth (
6329:
5573:
4394:
4382:
4368:
4315:from the original on 13 May 2015
4258:from the original on 6 July 2020
3871:from the original on 24 May 2009
3857:"Community Ecology of the Sloth"
3806:Proceedings of the Royal Society
3719:"The woman who got 'slothified'"
2929:from the original on 19 May 2017
2605:10.1111/j.1096-3642.2003.00100.x
2567:from the original on 24 May 2021
2405:10.1111/j.1096-3642.2003.00100.x
2386:Gaudin, T.J. (1 February 2004).
614:pale-throated three-toed sloth (
594:, contains four extant species:
515:Sloths belong to the superorder
110:
4327:
4296:
4270:
4240:
4218:"Why do sloths move so slowly?"
4209:
4089:
4059:
4029:
3999:
3932:
3909:10.1590/S1519-69842002000200008
3883:
3793:
3767:
3741:
3717:Venema, Vibeke (4 April 2014).
3710:
3693:
3664:
3627:
3610:
3597:
3567:
3453:
3419:The Quarterly Review of Biology
3410:
3382:
3363:
3287:
3257:
3223:
3185:10.1590/S0100-879X2001000100002
3130:10.1590/S0100-879X2001000100002
3104:
3055:
3012:
2941:
2874:
2844:
2234:. In Gardner, Alfred L. (ed.).
1684:
1655:
1390:brown-throated three-toed sloth
852:elephants, as was the case for
702:, contains two extant species:
2750:Journal of Mammalian Evolution
2166:Nature Ecology & Evolution
2073:
2003:
1790:The Sloth Institute Costa Rica
1627:They have made adaptations to
1485:. There are a large number of
1340:
13:
1:
4373:The dictionary definition of
4278:"The Sloth Institute website"
4071:Sloth Conservation Foundation
3460:Briggs, Helen (13 May 2008).
2244:. pp. 157–168 (p. 161).
1800:
1431:
4136:. pp. 624 (see p. 96).
3966:10.1371/journal.pone.0051389
3896:Brazilian Journal of Biology
3814:The Royal Society Publishing
3672:The biology of the Xenarthra
2099:10.1371/journal.pgen.0020109
1578:Manuel Antonio National Park
1050:
923:, and that the megalonychid
817:
7:
4612:Hoffmann's two-toed sloth (
4603:Linnaeus's two-toed sloth (
4134:University of Chicago Press
3495:University of Chicago Press
2242:University of Chicago Press
2230:Gardner, Alfred L. (2007).
1961:. Time Inc. 1964. pp.
1548:
1524:
438:also formerly lived in the
424:Quaternary extinction event
388:There are six extant sloth
10:
6540:
4858:
4101:Smithsonian's National Zoo
3335:Journal of Applied Ecology
3068:Journal of Raptor Research
2797:Proc. Natl. Acad. Sci. USA
1379:
1080:(Greater Antilles sloths)
944:Great American Interchange
638:The critically endangered
508:
353:constituting the suborder
29:
6357:
6327:
6181:
6140:
6124:
6115:
6014:
5991:
5982:
5957:
5846:
5837:
5754:
5687:
5618:
5614:
5599:
5586:
5571:
5537:
5463:
5408:
5328:
5319:
5241:
5227:
5122:
5107:
5103:
5092:
5019:
4996:
4941:
4911:
4883:
4870:
4866:
4853:
4821:
4769:
4720:
4696:
4687:
4650:
4636:
4632:
4574:
4502:
4488:
4484:
4446:
4097:"Southern two-toed sloth"
3033:10.1644/05-MAMM-A-360R2.1
2762:10.1007/s10914-014-9280-7
2700:10.1016/j.cub.2019.05.043
2646:10.1186/s12862-014-0184-1
2355:(Pilosa: Megalonychidae)"
2186:10.1038/s41559-019-0909-z
1676:Smithsonian Institution's
1600:, with a diverse diet of
1416:Sloths are unusual among
1294:
1269:
1254:
1247:
1222:
1207:
1200:
1193:
1175:
1149:
1134:
1127:
1112:
1105:
1098:
1091:
1075:
1068:
828:Nothrotheriops shastensis
744:Hoffmann's two-toed sloth
706:Linnaeus's two-toed sloth
314:Latham & Davies, 1795
309:
302:
297:
288:
198:
193:
107:Scientific classification
105:
92:
83:
70:
61:
45:
4531:Pygmy three-toed sloth (
4043:. BBC. 5 December 2013.
3751:. YouTube. 18 May 2009.
2958:BMC Evolutionary Biology
2625:BMC Evolutionary Biology
2586:Gaudin, Timothy (2004).
2551:10.1201/9781420039481-14
1794:Aviarios Sloth Sanctuary
1775:are due to contact with
1570:Hoffman's two-toed sloth
983:, similar to the extant
631:, now found only in the
625:maned three-toed sloth (
402:(three–toed sloths) and
4013:. BBC. 19 August 2010.
3865:Encyclopædia Britannica
3389:BBC (4 November 2016),
3356:10.1111/1365-2664.13138
2979:10.1186/1471-2148-10-86
2923:Encyclopedia Britannica
2819:10.1073/pnas.0502777102
2296:10.1644/10-MAMM-A-086.1
2033:10.1126/science.1229237
1821:; Reeder, D.M. (eds.).
1560:
1347:Central American Seaway
1000:Greater Antilles sloths
652:Isla Escudo de Veraguas
650:to the small island of
635:of southeastern Brazil.
575:Greater Antilles sloths
357:, including the extant
4558:Brown-throated sloth (
3823:10.1098/rspb.2013.3006
3081:10.3356/JRR-13-00017.1
2500:10.1098/rspb.2014.0192
1881:10.1098/rspb.2001.1702
1768:
1739:pygmy three-toed sloth
1727:maned three-toed sloth
1706:
1695:pygmy three-toed sloth
1584:
1408:Morphology and anatomy
1404:
904:
883:
840:
814:
802:Megatherium americanum
640:pygmy three-toed sloth
505:Taxonomy and evolution
474:
466:
458:
336:
36:Sloth (disambiguation)
34:. For other uses, see
4549:Pale-throated sloth (
3701:UNED Research Journal
1766:Dallas World Aquarium
1760:
1747:critically endangered
1711:Barro Colorado Island
1692:
1568:
1439:Sloths have very low
1398:Cahuita National Park
1387:
889:
864:
825:
799:
334:
4391:at Wikimedia Commons
4195:: e.T3036A47436575.
3296:Journal of Mammalogy
3021:Journal of Mammalogy
2693:(12): 2031–2042.e6.
2545:. pp. 201–235.
2349:Hayssen, V. (2011).
2328:: e.T4778A47439751.
2283:Journal of Mammalogy
1817:Gardner, A. (2005).
1596:Two-toed sloths are
1420:in not having seven
946:. Additionally, the
845:convergent evolution
710:Choloepus didactylus
616:Bradypus tridactylus
367:tropical rainforests
6255:Perezfontanatherium
5296:Paroctodontotherium
4749:Southern tamandua (
4740:Northern tamandua (
4521:(Three-toed sloths)
4441:species by suborder
3957:2012PLoSO...751389P
3777:. 23 January 2014.
3347:2018JApEc..55.2253G
3269:natureinstitute.org
3233:(21 January 2014).
2970:2010BMCEE..10...86S
2856:National Geographic
2810:2005PNAS..10211763S
2804:(33): 11763–11768.
2637:2014BMCEE..14..184R
2445:2004JVPal..24..398D
2353:Choloepus hoffmanni
2314:Choloepus hoffmanni
2258:on 27 November 2018
2232:"Suborder Folivora"
2178:2019NatEE...3.1121P
2025:2013Sci...339..662O
1875:(1476): 1605–1615.
1574:Choloepus hoffmanni
1394:Bradypus variegatus
892:Paramylodon harlani
867:Megalonyx wheatleyi
748:Choloepus hoffmanni
698:Choloepodidae, the
601:Bradypus variegatus
95:Choloepus hoffmanni
73:Bradypus variegatus
5786:Megalonychotherium
5303:Pseudoprepotherium
4181:Bradypus torquatus
3605:Ecological Studies
3399:on 30 October 2021
2494:(1782): 20140192.
2153:A.; Chait, B. T.;
1769:
1731:Bradypus torquatus
1707:
1668:sexually dimorphic
1637:symbiotic bacteria
1585:
1422:cervical vertebrae
1405:
1154:(two-toed sloths)
905:
884:
841:
815:
627:Bradypus torquatus
590:Bradypodidae, the
410:parallel evolution
337:
32:Sloth (deadly sin)
6509:Clawed herbivores
6491:
6490:
6476:Open Tree of Life
6351:Taxon identifiers
6342:
6341:
6338:
6337:
6325:
6324:
6321:
6320:
6317:
6316:
6313:
6312:
6290:Pyramiodontherium
6262:Plesiomegatherium
6111:
6110:
5953:
5952:
5949:
5948:
5945:
5944:
5814:Proschismotherium
5582:
5581:
5569:
5568:
5565:
5564:
5561:
5560:
5533:
5532:
5529:
5528:
5525:
5524:
5223:
5222:
5088:
5087:
4992:
4991:
4988:
4987:
4780:
4779:
4765:
4764:
4761:
4760:
4733:
4683:
4682:
4643:
4628:
4627:
4624:
4623:
4596:
4593:(Two-toed sloths)
4570:
4569:
4524:
4495:
4387:Media related to
4143:978-0-226-19542-1
4103:. 25 April 2016.
4067:"About the Sloth"
3504:978-0-226-19542-1
2560:978-0-8493-2001-9
2359:Mammalian Species
2251:978-0-226-28240-4
2019:(6120): 662–667.
1834:978-0-8018-8221-0
1699:Bradypus pygmaeus
1641:digestive process
1337:
1336:
1328:
1327:
1319:
1318:
1310:
1309:
1291:
1290:
1282:
1281:
1236:
1235:
1172:
1171:
1163:
1162:
1011:mitochondrial DNA
940:Isthmus of Panama
909:oceanic dispersal
882:) and restoration
654:off the coast of
644:Bradypus pygmaeus
607:Neotropical realm
592:three-toed sloths
540:vertebral columns
329:
328:
323:
315:
189:
16:(Redirected from
6531:
6484:
6483:
6471:
6470:
6458:
6457:
6445:
6444:
6432:
6431:
6419:
6418:
6406:
6405:
6393:
6392:
6391:
6378:
6377:
6376:
6346:
6345:
6333:
6122:
6121:
6095:Pronothrotherium
5989:
5988:
5980:
5979:
5961:
5844:
5843:
5821:Pseudortotherium
5752:
5751:
5690:
5651:
5641:
5631:
5616:
5615:
5612:
5611:
5601:
5600:
5588:
5587:
5577:
5539:
5538:
5349:Glossotheriopsis
5326:
5325:
5275:Octodontotherium
5239:
5238:
5231:
5186:Scelidotheridium
5127:Scelidotheriidae
5125:
5105:
5104:
5101:
5100:
5026:
5021:
5020:
5017:
5016:
4969:
4959:
4929:
4899:
4889:
4881:
4880:
4872:
4871:
4868:
4867:
4855:
4854:
4807:
4800:
4793:
4784:
4783:
4732:
4731:
4727:
4708:Giant anteater (
4694:
4693:
4671:Silky anteater (
4657:
4656:
4641:
4634:
4633:
4595:
4594:
4590:
4581:
4580:
4523:
4522:
4518:
4509:
4508:
4493:
4486:
4485:
4431:
4424:
4417:
4408:
4407:
4399:Data related to
4398:
4386:
4372:
4356:
4355:
4353:
4351:
4331:
4325:
4324:
4322:
4320:
4300:
4294:
4293:
4291:
4289:
4274:
4268:
4267:
4265:
4263:
4244:
4238:
4237:
4235:
4233:
4213:
4207:
4206:
4204:
4174:
4168:
4167:
4165:
4163:
4123:
4117:
4116:
4114:
4112:
4093:
4087:
4086:
4084:
4082:
4063:
4057:
4056:
4054:
4052:
4033:
4027:
4026:
4024:
4022:
4003:
3997:
3996:
3986:
3968:
3936:
3930:
3929:
3911:
3887:
3881:
3880:
3878:
3876:
3855:Montgomery, Sy.
3852:
3846:
3845:
3835:
3825:
3797:
3791:
3790:
3788:
3786:
3771:
3765:
3764:
3762:
3760:
3745:
3739:
3738:
3736:
3734:
3714:
3708:
3697:
3691:
3688:Treetops at Risk
3684:
3675:
3668:
3662:
3651:
3642:
3631:
3625:
3614:
3608:
3601:
3595:
3594:
3592:
3590:
3571:
3565:
3564:
3562:
3560:
3540:
3529:
3528:
3526:
3524:
3484:
3478:
3477:
3475:
3473:
3457:
3451:
3450:
3414:
3408:
3407:
3406:
3404:
3395:, archived from
3386:
3380:
3379:
3367:
3361:
3360:
3358:
3341:(5): 2253–2262.
3326:
3320:
3319:
3291:
3285:
3284:
3282:
3280:
3261:
3255:
3254:
3252:
3250:
3241:. Archived from
3227:
3221:
3220:
3218:
3216:
3210:
3187:
3169:
3160:
3151:
3150:
3132:
3108:
3102:
3101:
3083:
3059:
3053:
3052:
3016:
3010:
3009:
2999:
2981:
2945:
2939:
2938:
2936:
2934:
2915:
2902:
2901:
2899:
2897:
2878:
2872:
2871:
2869:
2867:
2848:
2842:
2841:
2831:
2821:
2780:
2774:
2773:
2741:
2735:
2734:
2732:
2730:
2712:
2702:
2678:
2669:
2668:
2658:
2648:
2616:
2610:
2609:
2607:
2583:
2577:
2576:
2574:
2572:
2528:
2522:
2521:
2511:
2479:
2473:
2472:
2424:
2418:
2417:
2407:
2383:
2377:
2376:
2374:
2346:
2340:
2339:
2337:
2307:
2301:
2300:
2298:
2274:
2268:
2267:
2265:
2263:
2254:. Archived from
2227:
2221:
2220:
2218:
2216:
2210:
2172:(7): 1121–1130.
2163:
2149:
2130:
2129:
2119:
2101:
2077:
2071:
2070:
2044:
2007:
2001:
2000:
1998:
1996:
1976:
1967:
1966:
1960:
1951:
1940:
1939:
1937:
1935:
1920:
1911:
1910:
1900:
1856:
1847:
1846:
1814:
1777:electrical lines
1762:Three-toed sloth
1250:
1249:
1203:
1202:
1196:
1195:
1130:
1129:
1115:Scelidotheriidae
1108:
1107:
1101:
1100:
1094:
1093:
1071:
1070:
1059:
1058:
992:three-toed sloth
917:Greater Antilles
776:Scelidotheriidae
511:List of pilosans
440:Greater Antilles
436:Caribbean sloths
321:
313:
293:
279:Scelidotheriidae
187:
115:
114:
88:
78:three-toed sloth
66:
49:Temporal range:
43:
42:
21:
6539:
6538:
6534:
6533:
6532:
6530:
6529:
6528:
6494:
6493:
6492:
6487:
6479:
6474:
6466:
6461:
6453:
6448:
6440:
6435:
6427:
6422:
6414:
6409:
6401:
6396:
6387:
6386:
6381:
6372:
6371:
6366:
6353:
6343:
6334:
6309:
6276:Proeremotherium
6269:Pliomegatherium
6192:Anisodontherium
6177:
6172:Proprepotherium
6136:
6107:
6016:Nothrotheriinae
6010:
5984:Nothrotheriidae
5978:
5965:
5941:
5833:
5750:
5738:Proplatyarthrus
5683:
5608:
5595:
5578:
5557:
5521:
5474:Glossotheridium
5459:
5440:Magdalenabradys
5404:
5315:
5268:Octodontobradys
5235:
5219:
5193:Scelidotheriops
5118:
5097:
5084:
5015:
5010:Pseudoglyptodon
5001:
4984:
4943:Myrmecophagidae
4937:
4907:
4862:
4849:
4817:
4811:
4781:
4776:
4757:
4751:T. tetradactyla
4729:
4728:
4726:
4716:
4689:Myrmecophagidae
4679:
4646:
4620:
4592:
4591:
4589:
4566:
4520:
4519:
4517:
4498:
4480:
4442:
4435:
4365:
4360:
4359:
4349:
4347:
4332:
4328:
4318:
4316:
4301:
4297:
4287:
4285:
4276:
4275:
4271:
4261:
4259:
4254:. 29 May 2013.
4246:
4245:
4241:
4231:
4229:
4214:
4210:
4175:
4171:
4161:
4159:
4144:
4124:
4120:
4110:
4108:
4095:
4094:
4090:
4080:
4078:
4065:
4064:
4060:
4050:
4048:
4035:
4034:
4030:
4020:
4018:
4005:
4004:
4000:
3937:
3933:
3888:
3884:
3874:
3872:
3853:
3849:
3798:
3794:
3784:
3782:
3773:
3772:
3768:
3758:
3756:
3747:
3746:
3742:
3732:
3730:
3715:
3711:
3698:
3694:
3685:
3678:
3669:
3665:
3652:
3645:
3632:
3628:
3615:
3611:
3602:
3598:
3588:
3586:
3573:
3572:
3568:
3558:
3556:
3541:
3532:
3522:
3520:
3505:
3485:
3481:
3471:
3469:
3458:
3454:
3415:
3411:
3402:
3400:
3387:
3383:
3368:
3364:
3327:
3323:
3308:10.2307/1381249
3292:
3288:
3278:
3276:
3263:
3262:
3258:
3248:
3246:
3228:
3224:
3214:
3212:
3208:
3167:
3161:
3154:
3109:
3105:
3060:
3056:
3017:
3013:
2950:Aiello, Annette
2946:
2942:
2932:
2930:
2917:
2916:
2905:
2895:
2893:
2880:
2879:
2875:
2865:
2863:
2850:
2849:
2845:
2784:Steadman, D. W.
2781:
2777:
2742:
2738:
2728:
2726:
2687:Current Biology
2679:
2672:
2617:
2613:
2584:
2580:
2570:
2568:
2561:
2529:
2525:
2480:
2476:
2425:
2421:
2384:
2380:
2347:
2343:
2308:
2304:
2275:
2271:
2261:
2259:
2252:
2228:
2224:
2214:
2212:
2208:
2161:
2150:
2133:
2078:
2074:
2008:
2004:
1994:
1992:
1977:
1970:
1953:
1952:
1943:
1933:
1931:
1922:
1921:
1914:
1857:
1850:
1835:
1815:
1808:
1803:
1755:
1753:Human relations
1735:Atlantic Forest
1703:Thomas Landseer
1693:Depiction of a
1687:
1658:
1563:
1551:
1527:
1499:triatomine bugs
1462:from predatory
1434:
1410:
1382:
1343:
1338:
1329:
1320:
1311:
1302:
1301:
1300:
1292:
1283:
1257:Nothrotheriidae
1237:
1183:
1182:
1181:
1173:
1164:
1053:
833:Nothrotheriidae
820:
700:two-toed sloths
679:Nothrotheriidae
633:Atlantic Forest
513:
507:
375:Central America
253:Nothrotheriidae
186:
109:
57:
51:Early Oligocene
47:
39:
28:
23:
22:
15:
12:
11:
5:
6537:
6527:
6526:
6521:
6516:
6511:
6506:
6489:
6488:
6486:
6485:
6472:
6459:
6446:
6433:
6420:
6407:
6394:
6379:
6363:
6361:
6355:
6354:
6340:
6339:
6336:
6335:
6328:
6326:
6323:
6322:
6319:
6318:
6315:
6314:
6311:
6310:
6308:
6307:
6300:
6293:
6286:
6283:Promegatherium
6279:
6272:
6265:
6258:
6251:
6244:
6237:
6230:
6223:
6220:Megathericulus
6216:
6209:
6202:
6199:Diabolotherium
6195:
6187:
6185:
6179:
6178:
6176:
6175:
6168:
6161:
6154:
6146:
6144:
6138:
6137:
6135:
6134:
6125:
6119:
6113:
6112:
6109:
6108:
6106:
6105:
6098:
6091:
6084:
6081:Nothrotheriops
6077:
6070:
6063:
6056:
6049:
6042:
6035:
6028:
6020:
6018:
6012:
6011:
6009:
6008:
6000:
5992:
5986:
5977:
5976:
5968:
5966:
5958:
5955:
5954:
5951:
5950:
5947:
5946:
5943:
5942:
5940:
5939:
5932:
5925:
5918:
5915:Protomegalonyx
5911:
5908:Pliometanastes
5904:
5901:Pattersonocnus
5897:
5890:
5883:
5876:
5869:
5862:
5855:
5847:
5841:
5839:Megalonychinae
5835:
5834:
5832:
5831:
5824:
5817:
5810:
5803:
5796:
5789:
5782:
5775:
5768:
5760:
5758:
5749:
5748:
5741:
5734:
5727:
5724:Mesopotamocnus
5720:
5712:
5704:
5701:Deseadognathus
5696:
5694:
5692:Megalonychidae
5685:
5684:
5682:
5681:
5673:
5670:Schismotherium
5665:
5657:
5647:
5637:
5627:
5619:
5609:
5606:Megatherioidea
5604:
5597:
5596:
5593:Megatherioidea
5584:
5583:
5580:
5579:
5572:
5570:
5567:
5566:
5563:
5562:
5559:
5558:
5556:
5555:
5554:
5553:
5547:
5545:
5543:Megatherioidea
5535:
5534:
5531:
5530:
5527:
5526:
5523:
5522:
5520:
5519:
5512:
5509:Pleurolestodon
5505:
5498:
5491:
5484:
5477:
5469:
5467:
5461:
5460:
5458:
5457:
5450:
5443:
5436:
5429:
5422:
5419:Bolivartherium
5414:
5412:
5406:
5405:
5403:
5402:
5395:
5388:
5381:
5373:
5366:
5359:
5352:
5345:
5337:
5334:Archaeomylodon
5329:
5323:
5317:
5316:
5314:
5313:
5310:Urumacotherium
5306:
5299:
5292:
5285:
5278:
5271:
5264:
5257:
5254:Eionaletherium
5250:
5247:Baraguatherium
5242:
5236:
5228:
5225:
5224:
5221:
5220:
5218:
5217:
5210:
5207:Sibyllotherium
5203:
5200:Scelidotherium
5196:
5189:
5182:
5175:
5168:
5165:Neonematherium
5161:
5153:
5146:
5139:
5131:
5129:
5120:
5119:
5117:
5116:
5108:
5098:
5093:
5090:
5089:
5086:
5085:
5083:
5082:
5075:
5068:
5061:
5054:
5047:
5040:
5032:
5030:
5014:
5013:
5004:
5002:
4997:
4994:
4993:
4990:
4989:
4986:
4985:
4983:
4982:
4975:
4965:
4955:
4947:
4945:
4939:
4938:
4936:
4935:
4932:Palaeomyrmidon
4925:
4917:
4915:
4909:
4908:
4906:
4905:
4895:
4884:
4878:
4864:
4863:
4851:
4850:
4848:
4847:
4841:
4835:
4829:
4822:
4819:
4818:
4810:
4809:
4802:
4795:
4787:
4778:
4777:
4770:
4767:
4766:
4763:
4762:
4759:
4758:
4756:
4755:
4746:
4736:
4734:
4718:
4717:
4715:
4714:
4704:
4702:
4691:
4685:
4684:
4681:
4680:
4678:
4677:
4667:
4665:
4654:
4648:
4647:
4630:
4629:
4626:
4625:
4622:
4621:
4619:
4618:
4609:
4599:
4597:
4578:
4572:
4571:
4568:
4567:
4565:
4564:
4555:
4551:B. tridactylus
4546:
4537:
4527:
4525:
4506:
4500:
4499:
4482:
4481:
4479:
4478:
4472:
4466:
4460:
4454:
4447:
4444:
4443:
4434:
4433:
4426:
4419:
4411:
4405:
4404:
4403:at Wikispecies
4392:
4380:
4364:
4363:External links
4361:
4358:
4357:
4326:
4309:Deutsche Welle
4295:
4269:
4239:
4208:
4169:
4142:
4118:
4088:
4058:
4028:
3998:
3951:(12): e51389.
3931:
3902:(2): 249–252.
3882:
3847:
3792:
3766:
3740:
3709:
3692:
3676:
3663:
3643:
3626:
3609:
3596:
3566:
3530:
3503:
3479:
3452:
3431:10.1086/394620
3409:
3381:
3362:
3321:
3302:(2): 359–366.
3286:
3256:
3245:on 29 May 2018
3222:
3152:
3103:
3054:
3027:(4): 808–816.
3011:
2940:
2903:
2873:
2858:. March 2014.
2843:
2775:
2756:(4): 473–518.
2736:
2670:
2611:
2598:(2): 255–305.
2578:
2559:
2523:
2474:
2439:(2): 398–410.
2419:
2398:(2): 255–305.
2378:
2341:
2302:
2269:
2250:
2222:
2131:
2072:
2002:
1968:
1941:
1912:
1848:
1833:
1805:
1804:
1802:
1799:
1754:
1751:
1686:
1683:
1657:
1654:
1562:
1559:
1550:
1547:
1526:
1523:
1433:
1430:
1409:
1406:
1381:
1378:
1362:Climate change
1342:
1339:
1335:
1334:
1331:
1330:
1326:
1325:
1322:
1321:
1317:
1316:
1313:
1312:
1308:
1307:
1304:
1303:
1298:Megatherioidea
1296:
1295:
1293:
1289:
1288:
1285:
1284:
1280:
1279:
1276:
1275:
1268:
1265:
1264:
1261:
1260:
1253:
1248:
1246:
1243:
1242:
1239:
1238:
1234:
1233:
1230:
1229:
1221:
1218:
1217:
1214:
1213:
1210:Megalonychidae
1206:
1201:
1199:
1194:
1192:
1189:
1188:
1185:
1184:
1177:
1176:
1174:
1170:
1169:
1166:
1165:
1161:
1160:
1157:
1156:
1148:
1145:
1144:
1141:
1140:
1133:
1128:
1126:
1123:
1122:
1119:
1118:
1111:
1106:
1104:
1099:
1097:
1092:
1090:
1087:
1086:
1083:
1082:
1074:
1069:
1067:
1057:
1052:
1049:
1034:Megalonychidae
1030:nothrotheriids
963:lifestyle. In
926:Pliometanastes
872:Megalonychidae
819:
816:
794:
793:
792:
791:
772:
765:
764:
763:
741:
690:
689:
688:
675:
668:
665:Megalonychidae
661:
660:
659:
636:
621:
610:
586:Megatherioidea
582:
506:
503:
428:climate change
327:
326:
325:
324:
316:
307:
306:
300:
299:
295:
294:
286:
285:
284:
283:
282:
281:
275:
269:
259:
258:
257:
256:
255:
249:
237:
235:Megalonychidae
231:
226:
218:
208:Megatherioidea
205:
196:
195:
191:
190:
181:
177:
176:
171:
167:
166:
161:
157:
156:
151:
147:
146:
141:
137:
136:
131:
127:
126:
121:
117:
116:
103:
102:
100:two-toed sloth
90:
89:
81:
80:
68:
67:
59:
58:
48:
26:
9:
6:
4:
3:
2:
6536:
6525:
6522:
6520:
6517:
6515:
6512:
6510:
6507:
6505:
6502:
6501:
6499:
6482:
6477:
6473:
6469:
6464:
6460:
6456:
6451:
6447:
6443:
6438:
6434:
6430:
6425:
6421:
6417:
6412:
6408:
6404:
6399:
6395:
6390:
6384:
6380:
6375:
6369:
6365:
6364:
6362:
6360:
6356:
6352:
6347:
6332:
6306:
6305:
6301:
6299:
6298:
6294:
6292:
6291:
6287:
6285:
6284:
6280:
6278:
6277:
6273:
6271:
6270:
6266:
6264:
6263:
6259:
6257:
6256:
6252:
6250:
6249:
6245:
6243:
6242:
6238:
6236:
6235:
6231:
6229:
6228:
6227:Megatheridium
6224:
6222:
6221:
6217:
6215:
6214:
6210:
6208:
6207:
6206:Eomegatherium
6203:
6201:
6200:
6196:
6194:
6193:
6189:
6188:
6186:
6184:
6183:Megatheriinae
6180:
6174:
6173:
6169:
6167:
6166:
6162:
6160:
6159:
6155:
6153:
6152:
6148:
6147:
6145:
6143:
6139:
6132:
6131:
6127:
6126:
6123:
6120:
6118:
6117:Megatheriidae
6114:
6104:
6103:
6099:
6097:
6096:
6092:
6090:
6089:
6088:Nothrotherium
6085:
6083:
6082:
6078:
6076:
6075:
6071:
6069:
6068:
6064:
6062:
6061:
6060:Mcdonaldocnus
6057:
6055:
6054:
6050:
6048:
6047:
6043:
6041:
6040:
6039:Chasicobradys
6036:
6034:
6033:
6032:Aymaratherium
6029:
6027:
6026:
6022:
6021:
6019:
6017:
6013:
6006:
6005:
6001:
5999:
5998:
5994:
5993:
5990:
5987:
5985:
5981:
5975:
5974:
5973:Analcimorphus
5970:
5969:
5967:
5964:
5956:
5938:
5937:
5933:
5931:
5930:
5926:
5924:
5923:
5919:
5917:
5916:
5912:
5910:
5909:
5905:
5903:
5902:
5898:
5896:
5895:
5891:
5889:
5888:
5884:
5882:
5881:
5877:
5875:
5874:
5870:
5868:
5867:
5866:Megalonychops
5863:
5861:
5860:
5856:
5854:
5853:
5849:
5848:
5845:
5842:
5840:
5836:
5830:
5829:
5825:
5823:
5822:
5818:
5816:
5815:
5811:
5809:
5808:
5804:
5802:
5801:
5797:
5795:
5794:
5790:
5788:
5787:
5783:
5781:
5780:
5776:
5774:
5773:
5769:
5767:
5766:
5762:
5761:
5759:
5757:
5756:Ortotheriinae
5753:
5747:
5746:
5742:
5740:
5739:
5735:
5733:
5732:
5728:
5726:
5725:
5721:
5718:
5717:
5713:
5710:
5709:
5705:
5703:
5702:
5698:
5697:
5695:
5693:
5686:
5680:
5679:
5678:Similhapalops
5674:
5672:
5671:
5666:
5664:
5663:
5658:
5656:
5655:
5648:
5646:
5645:
5638:
5636:
5635:
5628:
5626:
5625:
5621:
5620:
5617:
5613:
5610:
5607:
5602:
5598:
5594:
5589:
5585:
5576:
5551:
5550:
5549:
5548:
5546:
5544:
5540:
5536:
5518:
5517:
5513:
5511:
5510:
5506:
5504:
5503:
5499:
5497:
5496:
5492:
5490:
5489:
5485:
5483:
5482:
5481:Glossotherium
5478:
5476:
5475:
5471:
5470:
5468:
5466:
5462:
5456:
5455:
5454:Thinobadistes
5451:
5449:
5448:
5444:
5442:
5441:
5437:
5435:
5434:
5430:
5428:
5427:
5423:
5421:
5420:
5416:
5415:
5413:
5411:
5407:
5401:
5400:
5396:
5394:
5393:
5389:
5387:
5386:
5382:
5379:
5378:
5374:
5372:
5371:
5367:
5365:
5364:
5360:
5358:
5357:
5353:
5351:
5350:
5346:
5343:
5342:
5338:
5336:
5335:
5331:
5330:
5327:
5324:
5322:
5318:
5312:
5311:
5307:
5305:
5304:
5300:
5298:
5297:
5293:
5291:
5290:
5286:
5284:
5283:
5279:
5277:
5276:
5272:
5270:
5269:
5265:
5263:
5262:
5258:
5256:
5255:
5251:
5249:
5248:
5244:
5243:
5240:
5237:
5234:
5226:
5216:
5215:
5211:
5209:
5208:
5204:
5202:
5201:
5197:
5195:
5194:
5190:
5188:
5187:
5183:
5181:
5180:
5176:
5174:
5173:
5172:Proscelidodon
5169:
5167:
5166:
5162:
5159:
5158:
5154:
5152:
5151:
5147:
5145:
5144:
5140:
5138:
5137:
5136:Analcitherium
5133:
5132:
5130:
5128:
5121:
5115:
5114:
5110:
5109:
5106:
5102:
5099:
5096:
5095:Mylodontoidea
5091:
5081:
5080:
5076:
5074:
5073:
5069:
5067:
5066:
5062:
5060:
5059:
5055:
5053:
5052:
5048:
5046:
5045:
5041:
5039:
5038:
5034:
5033:
5031:
5029:
5022:
5018:
5012:
5011:
5006:
5005:
5003:
5000:
4995:
4981:
4980:
4976:
4974:
4973:
4966:
4964:
4963:
4956:
4954:
4953:
4949:
4948:
4946:
4944:
4940:
4934:
4933:
4926:
4924:
4923:
4919:
4918:
4916:
4914:
4910:
4904:
4903:
4896:
4894:
4893:
4886:
4885:
4882:
4879:
4877:
4873:
4869:
4865:
4861:
4856:
4852:
4846:
4842:
4840:
4836:
4834:
4830:
4828:
4824:
4823:
4820:
4815:
4808:
4803:
4801:
4796:
4794:
4789:
4788:
4785:
4775:
4774:
4768:
4754:
4752:
4747:
4745:
4743:
4738:
4737:
4735:
4725:
4724:
4719:
4713:
4711:
4710:M. tridactyla
4706:
4705:
4703:
4701:
4700:
4695:
4692:
4690:
4686:
4676:
4674:
4673:C. didactylus
4669:
4668:
4666:
4664:
4663:
4658:
4655:
4653:
4649:
4645:
4644:
4635:
4631:
4617:
4615:
4610:
4608:
4606:
4605:C. didactylus
4601:
4600:
4598:
4588:
4587:
4582:
4579:
4577:
4576:Choloepodidae
4573:
4563:
4561:
4560:B. variegatus
4556:
4554:
4552:
4547:
4545:
4543:
4540:Maned sloth (
4538:
4536:
4534:
4529:
4528:
4526:
4516:
4515:
4510:
4507:
4505:
4501:
4497:
4496:
4487:
4483:
4477:
4473:
4471:
4467:
4465:
4461:
4459:
4455:
4453:
4449:
4448:
4445:
4440:
4432:
4427:
4425:
4420:
4418:
4413:
4412:
4409:
4402:
4397:
4393:
4390:
4385:
4381:
4379:at Wiktionary
4378:
4377:
4371:
4367:
4366:
4345:
4341:
4337:
4330:
4314:
4310:
4306:
4299:
4283:
4279:
4273:
4257:
4253:
4249:
4243:
4227:
4223:
4219:
4212:
4203:
4198:
4194:
4190:
4189:
4184:
4182:
4173:
4157:
4153:
4149:
4145:
4139:
4135:
4131:
4130:
4122:
4106:
4102:
4098:
4092:
4076:
4072:
4068:
4062:
4046:
4042:
4038:
4032:
4016:
4012:
4008:
4002:
3994:
3990:
3985:
3980:
3976:
3972:
3967:
3962:
3958:
3954:
3950:
3946:
3942:
3935:
3927:
3923:
3919:
3915:
3910:
3905:
3901:
3897:
3893:
3886:
3870:
3866:
3862:
3858:
3851:
3843:
3839:
3834:
3829:
3824:
3819:
3815:
3811:
3807:
3803:
3796:
3780:
3776:
3770:
3754:
3750:
3744:
3728:
3724:
3720:
3713:
3706:
3702:
3696:
3689:
3683:
3681:
3673:
3667:
3660:
3656:
3650:
3648:
3640:
3636:
3630:
3623:
3619:
3613:
3606:
3600:
3584:
3580:
3579:Animal Corner
3576:
3570:
3554:
3550:
3546:
3539:
3537:
3535:
3518:
3514:
3510:
3506:
3500:
3496:
3492:
3491:
3483:
3467:
3463:
3456:
3448:
3444:
3440:
3436:
3432:
3428:
3424:
3420:
3413:
3398:
3394:
3393:
3385:
3377:
3373:
3366:
3357:
3352:
3348:
3344:
3340:
3336:
3332:
3325:
3317:
3313:
3309:
3305:
3301:
3297:
3290:
3274:
3270:
3266:
3260:
3244:
3240:
3236:
3232:
3226:
3207:
3203:
3199:
3195:
3191:
3186:
3181:
3177:
3173:
3166:
3159:
3157:
3148:
3144:
3140:
3136:
3131:
3126:
3122:
3118:
3114:
3107:
3099:
3095:
3091:
3087:
3082:
3077:
3073:
3069:
3065:
3058:
3050:
3046:
3042:
3038:
3034:
3030:
3026:
3022:
3015:
3007:
3003:
2998:
2993:
2989:
2985:
2980:
2975:
2971:
2967:
2963:
2959:
2955:
2951:
2944:
2928:
2924:
2920:
2914:
2912:
2910:
2908:
2891:
2887:
2883:
2877:
2861:
2857:
2853:
2847:
2839:
2835:
2830:
2825:
2820:
2815:
2811:
2807:
2803:
2799:
2798:
2793:
2789:
2788:Martin, P. S.
2785:
2779:
2771:
2767:
2763:
2759:
2755:
2751:
2747:
2740:
2724:
2720:
2716:
2711:
2706:
2701:
2696:
2692:
2688:
2684:
2677:
2675:
2666:
2662:
2657:
2652:
2647:
2642:
2638:
2634:
2630:
2626:
2622:
2615:
2606:
2601:
2597:
2593:
2589:
2582:
2566:
2562:
2556:
2552:
2548:
2544:
2540:
2539:
2534:
2527:
2519:
2515:
2510:
2505:
2501:
2497:
2493:
2489:
2485:
2478:
2470:
2466:
2462:
2458:
2454:
2453:10.1671/2429b
2450:
2446:
2442:
2438:
2434:
2430:
2423:
2415:
2411:
2406:
2401:
2397:
2393:
2389:
2382:
2373:
2372:10.1644/873.1
2368:
2364:
2360:
2356:
2354:
2345:
2336:
2331:
2327:
2323:
2322:
2317:
2315:
2306:
2297:
2292:
2289:(1): 86–100.
2288:
2284:
2280:
2273:
2257:
2253:
2247:
2243:
2239:
2238:
2233:
2226:
2207:
2203:
2199:
2195:
2191:
2187:
2183:
2179:
2175:
2171:
2167:
2160:
2156:
2148:
2146:
2144:
2142:
2140:
2138:
2136:
2127:
2123:
2118:
2113:
2109:
2105:
2100:
2095:
2091:
2087:
2086:PLOS Genetics
2083:
2076:
2068:
2064:
2060:
2056:
2052:
2048:
2043:
2038:
2034:
2030:
2026:
2022:
2018:
2014:
2006:
1990:
1986:
1982:
1975:
1973:
1964:
1959:
1958:
1950:
1948:
1946:
1929:
1925:
1919:
1917:
1908:
1904:
1899:
1894:
1890:
1886:
1882:
1878:
1874:
1870:
1866:
1864:
1855:
1853:
1844:
1840:
1836:
1830:
1826:
1825:
1820:
1813:
1811:
1806:
1798:
1795:
1791:
1787:
1784:
1782:
1778:
1774:
1767:
1763:
1759:
1750:
1748:
1744:
1740:
1736:
1732:
1728:
1724:
1720:
1716:
1712:
1704:
1700:
1696:
1691:
1682:
1680:
1677:
1671:
1669:
1664:
1653:
1650:
1644:
1642:
1638:
1634:
1630:
1625:
1623:
1619:
1615:
1611:
1607:
1603:
1599:
1594:
1592:
1591:
1583:
1579:
1576:) feeding in
1575:
1571:
1567:
1558:
1554:
1546:
1544:
1539:
1535:
1531:
1522:
1520:
1516:
1512:
1508:
1504:
1500:
1496:
1492:
1488:
1484:
1481:
1477:
1473:
1469:
1465:
1461:
1457:
1452:
1450:
1446:
1445:heterothermic
1442:
1437:
1429:
1427:
1423:
1419:
1414:
1403:
1399:
1395:
1391:
1386:
1377:
1375:
1374:
1369:
1367:
1363:
1359:
1354:
1352:
1348:
1333:
1332:
1324:
1323:
1315:
1314:
1306:
1305:
1299:
1287:
1286:
1278:
1277:
1274:
1273:
1272:Megatheriidae
1267:
1266:
1263:
1262:
1259:
1258:
1252:
1251:
1245:
1244:
1241:
1240:
1232:
1231:
1228:
1226:
1220:
1219:
1216:
1215:
1212:
1211:
1205:
1204:
1198:
1197:
1191:
1190:
1187:
1186:
1180:
1179:Mylodontoidea
1168:
1167:
1159:
1158:
1155:
1153:
1152:Choloepodidae
1147:
1146:
1143:
1142:
1139:
1138:
1132:
1131:
1125:
1124:
1121:
1120:
1117:
1116:
1110:
1109:
1103:
1102:
1096:
1095:
1089:
1088:
1085:
1084:
1081:
1079:
1073:
1072:
1066:
1065:
1061:
1060:
1056:
1048:
1046:
1042:
1038:
1035:
1031:
1027:
1023:
1022:
1017:
1012:
1008:
1003:
1001:
997:
993:
988:
986:
982:
978:
974:
970:
966:
962:
958:
954:
953:
949:
948:nothrotheriid
945:
941:
937:
936:
935:Thinobadistes
932:
928:
927:
922:
918:
914:
910:
902:
898:
894:
893:
888:
881:
877:
873:
869:
868:
863:
859:
857:
856:
850:
846:
838:
834:
830:
829:
824:
812:
808:
807:Megatheriidae
804:
803:
798:
789:
788:
783:
782:
777:
773:
770:
766:
761:
757:
753:
749:
745:
742:
739:
736:north of the
735:
731:
727:
723:
719:
715:
711:
707:
704:
703:
701:
697:
696:
695:
694:Mylodontoidea
691:
686:
685:
680:
676:
673:
672:Megatheriidae
669:
666:
662:
657:
653:
649:
645:
641:
637:
634:
630:
628:
622:
619:
617:
611:
608:
604:
602:
596:
595:
593:
589:
588:
587:
583:
580:
576:
572:
568:
567:
566:
564:
559:
557:
553:
550:of the order
549:
545:
541:
537:
533:
529:
525:
522:
519:, a group of
518:
512:
502:
500:
496:
492:
488:
484:
480:
476:
472:
468:
464:
460:
456:
452:
449:, related to
448:
443:
441:
437:
433:
429:
425:
421:
417:
416:
411:
407:
406:
401:
400:
395:
391:
386:
384:
380:
376:
372:
371:South America
368:
364:
363:ground sloths
360:
356:
352:
349:
345:
341:
333:
320:
317:
311:
310:
308:
305:
301:
296:
292:
287:
280:
276:
274:
270:
268:
267:Choloepodidae
265:
264:
263:
262:Mylodontoidea
260:
254:
250:
248:
247:Megatheriidae
244:
243:
242:
238:
236:
232:
230:
227:
225:
224:
223:Similhapalops
219:
217:
216:
211:
210:
209:
206:
204:
200:
199:
197:
192:
185:
182:
179:
178:
175:
172:
169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
145:
142:
139:
138:
135:
132:
129:
128:
125:
122:
119:
118:
113:
108:
104:
101:
97:
96:
91:
87:
82:
79:
75:
74:
69:
65:
60:
56:
52:
44:
41:
37:
33:
19:
6358:
6302:
6295:
6288:
6281:
6274:
6267:
6260:
6253:
6246:
6239:
6234:Megatheriops
6232:
6225:
6218:
6213:Eremotherium
6211:
6204:
6197:
6190:
6170:
6165:Prepoplanops
6163:
6158:Prepotherium
6156:
6149:
6130:Thalassocnus
6128:
6100:
6093:
6086:
6079:
6072:
6067:Mionothropus
6065:
6058:
6051:
6044:
6037:
6030:
6023:
6004:Thalassocnus
6002:
5995:
5971:
5934:
5927:
5920:
5913:
5906:
5899:
5892:
5885:
5878:
5871:
5864:
5857:
5850:
5826:
5819:
5812:
5805:
5800:Paranabradys
5798:
5791:
5784:
5777:
5770:
5763:
5743:
5736:
5729:
5722:
5714:
5706:
5699:
5676:
5668:
5660:
5652:
5644:Hiskatherium
5642:
5632:
5622:
5514:
5507:
5500:
5493:
5486:
5479:
5472:
5452:
5447:Sphenotherus
5445:
5438:
5431:
5424:
5417:
5410:Lestodontini
5397:
5390:
5383:
5375:
5368:
5363:Mylodonopsis
5361:
5354:
5347:
5341:Brievabradys
5339:
5332:
5321:Mylodontinae
5308:
5301:
5294:
5287:
5280:
5273:
5266:
5259:
5252:
5245:
5233:Mylodontidae
5212:
5205:
5198:
5191:
5184:
5177:
5170:
5163:
5155:
5150:Chubutherium
5148:
5141:
5134:
5111:
5077:
5070:
5063:
5056:
5049:
5042:
5035:
5028:Megalocnidae
5008:
4998:
4977:
4970:
4960:
4952:Myrmecophaga
4950:
4930:
4920:
4913:Cyclopedidae
4902:Orthoarthrus
4900:
4890:
4843:Superorder:
4771:
4750:
4741:
4721:
4709:
4699:Myrmecophaga
4697:
4672:
4660:
4652:Cyclopedidae
4640:Vermilingua
4638:
4614:C. hoffmanni
4613:
4604:
4584:
4559:
4550:
4542:B. torquatus
4541:
4532:
4512:
4504:Bradypodidae
4491:
4490:
4375:
4348:. Retrieved
4339:
4329:
4317:. Retrieved
4308:
4298:
4286:. Retrieved
4272:
4260:. Retrieved
4251:
4242:
4230:. Retrieved
4224:. BBC News.
4221:
4211:
4192:
4186:
4180:
4172:
4162:25 September
4160:. Retrieved
4128:
4121:
4109:. Retrieved
4100:
4091:
4079:. Retrieved
4070:
4061:
4049:. Retrieved
4040:
4031:
4019:. Retrieved
4010:
4001:
3948:
3944:
3934:
3899:
3895:
3885:
3873:. Retrieved
3860:
3850:
3809:
3805:
3795:
3783:. Retrieved
3769:
3757:. Retrieved
3743:
3731:. Retrieved
3722:
3712:
3704:
3700:
3695:
3687:
3671:
3666:
3658:
3654:
3641:(3),336-349.
3638:
3634:
3629:
3624:(3),435-446.
3621:
3617:
3612:
3604:
3599:
3587:. Retrieved
3578:
3569:
3557:. Retrieved
3549:Live Science
3548:
3523:25 September
3521:. Retrieved
3489:
3482:
3470:. Retrieved
3464:. BBC News.
3455:
3425:(1): 13–34.
3422:
3418:
3412:
3401:, retrieved
3397:the original
3391:
3384:
3375:
3371:
3365:
3338:
3334:
3324:
3299:
3295:
3289:
3277:. Retrieved
3268:
3259:
3247:. Retrieved
3243:the original
3238:
3225:
3213:. Retrieved
3175:
3171:
3120:
3116:
3106:
3074:(1): 24–35.
3071:
3067:
3057:
3024:
3020:
3014:
2961:
2957:
2943:
2931:. Retrieved
2922:
2894:. Retrieved
2886:ScienceDaily
2885:
2876:
2864:. Retrieved
2855:
2846:
2801:
2795:
2778:
2753:
2749:
2746:Thalassocnus
2745:
2739:
2727:. Retrieved
2710:11336/136908
2690:
2686:
2628:
2624:
2614:
2595:
2591:
2581:
2569:. Retrieved
2537:
2526:
2491:
2487:
2477:
2436:
2432:
2429:Thalassocnus
2428:
2422:
2395:
2391:
2381:
2365:(1): 37–55.
2362:
2358:
2352:
2344:
2325:
2319:
2313:
2305:
2286:
2282:
2272:
2260:. Retrieved
2256:the original
2236:
2225:
2215:18 September
2213:. Retrieved
2169:
2165:
2089:
2085:
2075:
2016:
2012:
2005:
1993:. Retrieved
1984:
1956:
1932:. Retrieved
1872:
1868:
1863:Eurotamandua
1862:
1823:
1819:Wilson, D.E.
1788:
1785:
1770:
1742:
1730:
1708:
1698:
1685:Distribution
1679:National Zoo
1672:
1659:
1656:Reproduction
1645:
1626:
1595:
1588:
1586:
1573:
1555:
1552:
1540:
1536:
1532:
1528:
1517:, mites and
1472:harpy eagles
1453:
1438:
1435:
1428:, with six.
1415:
1411:
1393:
1371:
1370:
1355:
1344:
1297:
1270:
1255:
1225:Bradypodidae
1223:
1208:
1178:
1150:
1137:Mylodontidae
1135:
1113:
1078:Megalocnidae
1076:
1063:
1062:
1054:
1040:
1037:polyphyletic
1026:megatheriids
1019:
1004:
995:
989:
973:Thalassocnus
972:
952:Thalassocnus
950:
933:
924:
906:
897:Mylodontidae
890:
865:
853:
842:
826:
800:
785:
779:
769:Mylodontidae
747:
738:Amazon River
709:
693:
692:Superfamily
684:Thalassocnus
682:
643:
626:
615:
600:
585:
584:Superfamily
571:Megalocnidae
560:
538:with sturdy
514:
490:
482:
450:
446:
444:
413:
403:
397:
387:
354:
339:
338:
335:Green sloth
273:Mylodontidae
261:
240:
229:Bradypodidae
221:
215:Hiskatherium
213:
207:
203:Megalocnidae
183:
160:Superorder:
93:
71:
40:
6383:Wikispecies
6297:Sibotherium
6241:Megatherium
6142:Planopsinae
6046:Huilabradys
6025:Amphibradys
5997:Neohapalops
5936:Zacatzontli
5929:Xibalbaonyx
5894:Nohochichak
5859:Australonyx
5807:Pliomorphus
5793:Ortotherium
5772:Eucholoeops
5745:Urumacocnus
5716:Hyperleptus
5654:Hyperleptus
5516:Simomylodon
5502:Paramylodon
5488:Kiyumylodon
5465:Mylodontini
5426:Lestobradys
5399:Strabosodon
5377:Oreomylodon
5370:Ocnotherium
5282:Octomylodon
5261:Nematherium
5157:Nematherium
4972:Protamandua
4962:Neotamandua
4892:Argyromanis
4876:Vermilingua
4742:T. mexicana
4730:(Tamanduas)
4642:(Anteaters)
4533:B. pygmaeus
4474:Superorder
4468:Infraclass
4232:2 September
3875:6 September
3759:11 February
3589:22 November
3559:22 November
3178:(1): 9–25.
3123:(1): 9–25.
2262:29 November
2240:. Chicago:
2155:Penkman, K.
2092:(7): e109.
1934:29 November
1743:B. pygmaeus
1622:herbivorous
1456:extremities
1341:Extinctions
1016:mylodontids
957:semiaquatic
855:Megatherium
718:the Guianas
712:) found in
646:) which is
499:sloth moths
495:green algae
491:laiskiainen
420:Pleistocene
415:Megatherium
344:Neotropical
319:Phyllophaga
312:Tardigrada
6498:Categories
6053:Lakukullus
5963:Megatheria
5922:Sinclairia
5880:Megistonyx
5852:Ahytherium
5779:Hapaloides
5662:Pelecyodon
5552:see below↓
5385:Promylodon
5356:Megabradys
5179:Scelidodon
5051:Megalocnus
5037:Acratocnus
4262:2 December
4222:BBC Future
4111:30 October
4081:31 October
3785:9 February
3733:1 December
3674:, 269-280.
3249:23 January
3215:1 December
2933:1 December
2866:1 December
2631:(1): 184.
2042:11336/7302
1995:1 December
1924:"Overview"
1801:References
1773:Costa Rica
1616:and small
1598:omnivorous
1582:Costa Rica
1491:mosquitoes
1483:arthropods
1460:camouflage
1432:Physiology
1402:Costa Rica
1373:Megalocnus
981:seagrasses
931:mylodontid
536:herbivores
532:armadillos
509:See also:
348:xenarthran
322:Owen, 1842
241:Megatheria
180:Suborder:
18:Megatheria
6514:Folivores
6304:Urumaquia
6102:Xyophorus
6074:Nothropus
5873:Megalonyx
5828:Torcellia
5731:Paulocnus
5392:Ranculcus
5289:Orophodon
5113:Choloepus
5044:Imagocnus
4845:Xenarthra
4825:Kingdom:
4586:Choloepus
4492:Folivora
4476:Xenarthra
4152:493329394
3975:1932-6203
3918:1519-6984
3707:(1), 4-4.
3513:493329394
3239:Phenomena
3194:1678-4510
3139:0100-879X
3090:0892-1016
3041:0022-2372
2988:1471-2148
2964:(1): 86.
2543:CRC Press
2414:0024-4082
2202:174813630
2108:1553-7404
2067:206544776
2051:0036-8075
1889:0962-8452
1663:gestation
1649:symbiotic
1635:in which
1543:nocturnal
1495:sandflies
1480:parasitic
1476:commensal
1441:metabolic
1358:megafauna
1051:Phylogeny
1041:Megalonyx
1021:Megalonyx
985:sirenians
921:Oligocene
901:San Diego
818:Evolution
787:Choloepus
714:Venezuela
528:Anteaters
521:placental
517:Xenarthra
467:paresseux
432:radiation
405:Choloepus
379:anteaters
346:group of
194:Families
164:Xenarthra
130:Kingdom:
124:Eukaryota
6519:Pilosans
6455:11800002
6403:Folivora
6389:Folivora
6374:Q2274076
6368:Wikidata
6359:Folivora
5887:Meizonyx
5765:Diodomus
5708:Hapalops
5634:Hapalops
5624:Bradypus
5433:Lestodon
5214:Valgipes
5079:Parocnus
5058:Mesocnus
4999:Folivora
4979:Tamandua
4922:Cyclopes
4839:Mammalia
4833:Chordata
4831:Phylum:
4827:Animalia
4773:Category
4723:Tamandua
4662:Cyclopes
4514:Bradypus
4494:(Sloths)
4470:Eutheria
4464:Mammalia
4458:Chordata
4452:Animalia
4450:Kingdom
4401:Folivora
4389:Folivora
4344:Archived
4340:The Dodo
4319:18 April
4313:Archived
4288:23 March
4282:Archived
4256:Archived
4252:ABC News
4226:Archived
4156:Archived
4105:Archived
4075:Archived
4051:30 April
4045:Archived
4041:BBC News
4021:30 April
4015:Archived
4011:BBC News
3993:23284687
3945:PLOS ONE
3926:12489397
3869:Archived
3842:24452028
3812:(1778).
3779:Archived
3753:Archived
3727:Archived
3723:BBC News
3583:Archived
3553:Archived
3517:Archived
3466:Archived
3447:85162387
3403:17 April
3378:: 94–95.
3273:Archived
3206:Archived
3202:11151024
3147:11151024
3098:86270583
3049:37859321
3006:20353556
2927:Archived
2896:26 April
2890:Archived
2860:Archived
2838:16085711
2770:16700349
2723:Archived
2719:31178321
2665:25319928
2565:Archived
2518:24621950
2469:83859607
2206:Archived
2194:31171860
2126:16848642
2059:23393258
1989:Archived
1928:Archived
1907:11487408
1843:62265494
1781:poachers
1633:stomachs
1629:arboreal
1590:Cecropia
1549:Behavior
1525:Activity
1426:manatees
1388:Feeding
1351:seagrass
1064:Folivora
1007:collagen
996:Bradypus
929:and the
913:pilosans
849:arboreal
756:Honduras
722:Colombia
548:Folivora
544:pelvises
542:, fused
479:Romanian
475:perezoso
459:Faultier
399:Bradypus
359:arboreal
355:Folivora
304:Synonyms
184:Folivora
154:Mammalia
144:Chordata
140:Phylum:
134:Animalia
120:Domain:
55:Holocene
6429:2852810
6248:Ocnopus
6151:Planops
5495:Mylodon
5143:Catonyx
5072:Neocnus
5065:Miocnus
4837:Class:
4814:Pilosan
4456:Phylum
4437:Extant
3984:3526605
3953:Bibcode
3833:3906947
3575:"Sloth"
3439:2808832
3343:Bibcode
3316:1381249
3279:29 June
3231:Ed Yong
2997:2858742
2966:Bibcode
2919:"Sloth"
2852:"Sloth"
2829:1187974
2806:Bibcode
2729:11 June
2656:4243956
2633:Bibcode
2509:3973278
2461:4524727
2441:Bibcode
2174:Bibcode
2117:1513266
2021:Bibcode
2013:Science
1985:Gizmodo
1898:1088784
1764:in the
1723:species
1719:biomass
1705:, 1825)
1618:lizards
1606:carrion
1602:insects
1515:beetles
1468:ocelots
1464:jaguars
1418:mammals
1380:Biology
1366:ice age
977:Miocene
919:by the
915:to the
837:La Brea
781:Mylodon
760:Bolivia
726:Ecuador
648:endemic
524:mammals
487:Finnish
471:Spanish
392:in two
390:species
351:mammals
170:Order:
150:Class:
6504:Sloths
6481:495015
6468:948953
6442:727503
4860:Pilosa
4816:genera
4462:Class
4439:Pilosa
4350:20 May
4150:
4140:
3991:
3981:
3973:
3924:
3916:
3840:
3830:
3511:
3501:
3472:21 May
3445:
3437:
3314:
3200:
3192:
3145:
3137:
3096:
3088:
3047:
3039:
3004:
2994:
2986:
2836:
2826:
2768:
2717:
2663:
2653:
2571:9 June
2557:
2516:
2506:
2467:
2459:
2412:
2248:
2200:
2192:
2124:
2114:
2106:
2065:
2057:
2049:
1905:
1895:
1887:
1841:
1831:
1715:Panama
1614:leaves
1610:fruits
1470:, and
1449:torpor
961:marine
876:fossil
847:to an
811:London
734:Brazil
732:, and
656:Panama
573:: the
556:mammal
552:Pilosa
463:French
455:German
394:genera
383:Pilosa
342:are a
340:Sloths
174:Pilosa
46:Sloths
4376:sloth
3635:Oikos
3443:S2CID
3435:JSTOR
3312:JSTOR
3209:(PDF)
3168:(PDF)
3094:S2CID
3045:S2CID
2766:S2CID
2465:S2CID
2457:JSTOR
2209:(PDF)
2198:S2CID
2162:(PDF)
2063:S2CID
1965:, 54.
1745:) is
1519:moths
1511:mites
1507:ticks
1045:basal
969:Chile
784:than
752:Andes
579:basal
483:leneș
447:Sloth
6463:NCBI
6437:ITIS
4352:2016
4321:2013
4290:2022
4264:2017
4234:2019
4193:2014
4164:2016
4148:OCLC
4138:ISBN
4113:2019
4083:2019
4053:2015
4023:2015
3989:PMID
3971:ISSN
3922:PMID
3914:ISSN
3877:2009
3838:PMID
3787:2021
3761:2022
3735:2017
3591:2020
3561:2020
3525:2016
3509:OCLC
3499:ISBN
3474:2010
3405:2017
3281:2017
3251:2014
3217:2017
3198:PMID
3190:ISSN
3143:PMID
3135:ISSN
3086:ISSN
3037:ISSN
3002:PMID
2984:ISSN
2935:2017
2898:2019
2868:2017
2834:PMID
2731:2020
2715:PMID
2661:PMID
2573:2020
2555:ISBN
2514:PMID
2410:ISSN
2326:2014
2264:2017
2246:ISBN
2217:2020
2190:PMID
2122:PMID
2104:ISSN
2055:PMID
2047:ISSN
1997:2017
1936:2017
1903:PMID
1885:ISSN
1839:OCLC
1829:ISBN
1779:and
1561:Diet
1509:and
1503:lice
1493:and
1478:and
1028:and
1009:and
967:and
965:Peru
880:AMNH
730:Peru
623:The
612:The
597:The
577:, a
530:and
451:slow
373:and
98:, a
76:, a
6450:MSW
6424:EoL
6416:4DM
6411:CoL
6398:ADW
4197:doi
3979:PMC
3961:doi
3904:doi
3828:PMC
3818:doi
3810:281
3659:281
3427:doi
3351:doi
3304:doi
3180:doi
3125:doi
3076:doi
3029:doi
2992:PMC
2974:doi
2824:PMC
2814:doi
2802:102
2758:doi
2705:hdl
2695:doi
2651:PMC
2641:doi
2600:doi
2596:140
2547:doi
2504:PMC
2496:doi
2492:281
2449:doi
2431:".
2400:doi
2396:140
2367:doi
2330:doi
2291:doi
2182:doi
2112:PMC
2094:doi
2037:hdl
2029:doi
2017:339
1893:PMC
1877:doi
1873:268
1713:in
1701:) (
1580:in
1396:),
911:of
565:):
434:of
369:of
53:to
6500::
6478::
6465::
6452::
6439::
6426::
6413::
6400::
6385::
6370::
4342:.
4338:.
4311:.
4307:.
4280:.
4250:.
4220:.
4191:.
4185:.
4154:.
4146:.
4132:.
4099:.
4073:.
4069:.
4039:.
4009:.
3987:.
3977:.
3969:.
3959:.
3947:.
3943:.
3920:.
3912:.
3900:62
3898:.
3894:.
3867:.
3863:.
3859:.
3836:.
3826:.
3816:.
3808:.
3804:.
3725:.
3721:.
3705:13
3703:,
3679:^
3657:,
3646:^
3639:32
3637:,
3622:38
3620:,
3581:.
3577:.
3551:.
3547:.
3533:^
3515:.
3507:.
3493:.
3441:.
3433:.
3423:16
3421:.
3376:34
3374:.
3349:.
3339:55
3337:.
3333:.
3310:.
3300:66
3298:.
3271:.
3267:.
3237:.
3204:.
3196:.
3188:.
3176:34
3174:.
3170:.
3155:^
3141:.
3133:.
3121:34
3119:.
3115:.
3092:.
3084:.
3072:48
3070:.
3066:.
3043:.
3035:.
3025:87
3023:.
3000:.
2990:.
2982:.
2972:.
2962:10
2960:.
2956:.
2925:.
2921:.
2906:^
2888:.
2884:.
2854:.
2832:.
2822:.
2812:.
2800:.
2794:.
2786:;
2764:.
2754:22
2752:.
2721:.
2713:.
2703:.
2691:29
2689:.
2685:.
2673:^
2659:.
2649:.
2639:.
2629:14
2627:.
2623:.
2594:.
2590:.
2563:.
2553:.
2512:.
2502:.
2490:.
2486:.
2463:.
2455:.
2447:.
2437:24
2435:.
2408:.
2394:.
2390:.
2363:43
2361:.
2357:.
2324:.
2318:.
2287:92
2285:.
2281:.
2204:.
2196:.
2188:.
2180:.
2168:.
2164:.
2134:^
2120:.
2110:.
2102:.
2088:.
2084:.
2061:.
2053:.
2045:.
2035:.
2027:.
2015:.
1987:.
1983:.
1971:^
1963:15
1944:^
1915:^
1901:.
1891:.
1883:.
1871:.
1867:.
1851:^
1837:.
1809:^
1612:,
1608:,
1604:,
1593:.
1505:,
1501:,
1497:,
1466:,
1451:.
1400:,
1024:,
987:.
971:,
899:,
874:)
858:.
835:,
809:,
728:,
724:,
720:,
716:,
489::
485:,
481::
477:,
473::
469:,
465::
461:,
457::
396:–
385:.
6133:?
6007:?
5960:†
5719:?
5711:?
5689:†
5675:†
5667:†
5659:†
5650:†
5640:†
5630:†
5380:?
5344:?
5230:†
5160:?
5124:†
5025:†
5007:†
4968:†
4958:†
4928:†
4898:†
4888:†
4806:e
4799:t
4792:v
4753:)
4744:)
4712:)
4675:)
4616:)
4607:)
4562:)
4553:)
4544:)
4535:)
4430:e
4423:t
4416:v
4354:.
4323:.
4292:.
4266:.
4236:.
4205:.
4199::
4183:"
4179:"
4166:.
4115:.
4085:.
4055:.
4025:.
3995:.
3963::
3955::
3949:7
3928:.
3906::
3879:.
3844:.
3820::
3789:.
3763:.
3737:.
3593:.
3563:.
3527:.
3476:.
3449:.
3429::
3359:.
3353::
3345::
3318:.
3306::
3283:.
3253:.
3219:.
3182::
3149:.
3127::
3100:.
3078::
3051:.
3031::
3008:.
2976::
2968::
2937:.
2900:.
2870:.
2840:.
2816::
2808::
2772:.
2760::
2733:.
2707::
2697::
2667:.
2643::
2635::
2608:.
2602::
2575:.
2549::
2520:.
2498::
2471:.
2451::
2443::
2416:.
2402::
2375:.
2369::
2351:"
2338:.
2332::
2316:"
2312:"
2299:.
2293::
2266:.
2219:.
2184::
2176::
2170:3
2128:.
2096::
2090:2
2069:.
2039::
2031::
2023::
1999:.
1938:.
1909:.
1879::
1865:"
1845:.
1741:(
1729:(
1697:(
1572:(
1392:(
994:(
903:)
895:(
878:(
870:(
839:)
831:(
813:)
805:(
774:†
767:†
762:.
746:(
740:.
708:(
677:†
670:†
663:†
658:.
642:(
629:)
618:)
603:)
569:†
563:†
277:†
271:†
251:†
245:†
239:†
233:†
220:†
212:†
201:†
38:.
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.