340:
2621:
3315:
2584:
119:
312:
2547:
2423:
2823:
2658:
2460:
3497:
1475:
2218:
347:
319:
2276:
268:
1699:
532:. The presence of mostly articulated skeletons, exceptional preservation of tissue and lack of any other signs of transportation may indicate that these are demic and autochthonous entities (meaning that they lived and died in the same place where they fossilized).
4277:
447:' crest. The iron carbonate depositions, a result from bacterial metabolism which covered the dinosaur's crest enhanced the preservation of those soft tissues. Evidence of these mats comes from the studies on microfacies and the fossils themselves.
4253:"The stratigraphic record of the Late Jurassic-Early Cretaceous rifting in the Alto Tajo-Serranía de Cuenca region (Iberian Ranges, Spain): genetic and structural evidences for a revision and a new lithostratigraphic proposal"
450:
Obruption is notable in the formation, due to the presence of highly articulated specimens. From actuotaphonomy studies on several different organisms it can be estimated that the burial of most entities was quick.
4257:
4439:
Pérez Moreno, B. P., Sanz, J. L., Buscalioni, A. D., Moratalla, J. J., Ortega, F., and Raskin-Gutman, D. (1994). "A unique multitoothed ornithomimosaur from the Lower
Cretaceous of Spain." Nature, 30: 363–367.
3020:
585:
shows both derived ("avian") and primitive ("dinosaurian") characters. Within the derived characters we can underline the presence of a pygostile, although it is still very large compared to that of modern
4355:
Thomas Martin, Jesús Marugán-Lobón, Romain Vullo, Hugo Martín-Abad, Zhe-Xi Luo & Angela D. Buscalioni (2015). A Cretaceous eutriconodont and integument evolution in early mammals. Nature 526, 380–384.
621:
3129:
3063:
4028:"The ecology of the Lower Cretaceous coelacanths from Las Hoyas Konservat- Lagerstätte (Cuenca, Spain): A new insight after the integration of palaeontological and sedimentological data"
3206:
2921:
3528:
3388:
3352:
3325:
3368:
3145:
2870:
3029:
3583:
3444:
3431:
564:
presents two unique features: very tall neural spines on the vertebrae near the hip, which look like a hump, and a structure on its forearm, which if homologous to
3359:
3579:
3238:
3159:
3077:
3466:
3539:
3514:
3095:
2988:
2914:
2848:
3482:
3086:
2984:
2907:
2830:
2789:
283:
160:
142:
3661:
3630:
3610:
3503:
3106:
4415:
Sanz, José L.; Chiappe, Luis M.; Pérez Moreno, Bernardino P.; Buscalioni, Ángela D.; Moratalla, José J.; Ortega, Francisco & Poyato Ariza, Francisco J.
3709:
3296:
3013:
3006:
2876:
2842:
558:, shows some characters previously unknown in these dinosaurs which enhanced the knowledge on the evolution of the group, such as a high number of teeth.
4538:
3420:
3409:
3332:
3263:
875:
810:
3166:
3507:
3173:
3117:
3099:
3057:
3052:
2964:
2955:
2903:
2884:
2782:
2775:
2768:
2761:
2740:
1501:
3566:
3474:
3395:
3215:
2968:
864:
339:
2994:
4469:
3040:
1558:
150:
3451:
3440:
3427:
2754:
854:
830:
216:
2836:
1820:
1336:
885:
1049:
840:
4548:
1969:
1174:
1077:
479:
1359:
1308:
1139:
4513:
3987:"A new genus and species for the amiiform fishes previously assigned to Amiopsis from the Early Cretaceous of Las Hoyas, Cuenca, Spain"
3838:
Gallego, Oscar F.; Monferran, Mateo D.; Zacarías, Iracema A.; Jiménez, Victoria C.; Buscalioni, Angela D.; Liao, Huanyu (June 2020).
617:
3726:
2796:
4278:"An articulated hindlimb of a basal iguanodont (Dinosauria, Ornithopoda) from the Early Cretaceous Las Hoyas Lagerstätte (Spain)"
4508:
311:
3840:"Clam shrimp fauna (Diplostraca-Spinicaudata and Estheriellina) from the Lower Cretaceous of Las Hoyas, Cuenca (Spain)"
4503:
3887:"A presumed spelaeogriphacean crustacean from an upper Barremian wetland (Las Hoyas; Lower Cretaceous; Central Spain)"
4400:
3457:
613:
3252:
609:
440:, the preservation is exceptional. This may be a result of three factors: Microbial mats, Obruption and Stagnation.
4442:
Sanz, J. L. & Bonaparte, José F. (1992): A New Order of Birds (Class Aves) from the Lower
Cretaceous of Spain.
4276:
Serrano, Mercedes
Llandres; Vullo, Romain; Marugán-Lobón, Jesús; Ortega, Francisco; Buscalioni, Ángela D. (2013),
4237:
sp." in "15.9 Provincia de Quenca, Spain; 4. Calizas de La
Huergina Formation," in Weishampel et al., 2004, p.561
4224:
sp." in "15.9 Provincia de Quenca, Spain; 4. Calizas de La
Huergina Formation," in Weishampel et al., 2004, p.561
4184:
sp." in "15.9 Provincia de Quenca, Spain; 4. Calizas de La
Huergina Formation," in Weishampel et al., 2004, p.561
2999:
4543:
4484:
3651:
3552:
437:
4159:"15.9 Provincia de Quenca, Spain; 4. Calizas de La Huergina Formation," in Weishampel et al., 2004, pp.561-562
4523:
4320:
4251:
Martínez, Marian
Fregenal; Meléndez, Nieves; Muñoz García, M. Belén; Elez, Javier; de la Horra, Raúl (2017),
3070:
3047:
4533:
443:
Microbial mats may be responsible for the preservation of soft tissue in many fossils from Las Hoyas, like
209:
3778:"The late Barremian ecosystem of Las Hoyas sustained by fishes and shrimps as inferred from coprofabrics"
4372:
2862:
4068:
4518:
3136:
4459:(1992): A new bird from the Early Cretaceous of Las Hoyas, Spain, and the early radiation of birds.
4417:(1996): An Early Cretaceous bird from Spain and its implications for the evolution of avian flight.
4528:
2143:
Exceptionally complete specimen, preserving soft tissues such as pelage, ears and internal organs.
463:
Las Hoyas was an inland lacustrine environment which presents an important aquatic and terrestrial
600:
or "bastard wing", which means it had a flight manoeuvrability analogous to that of modern birds.
4461:
4069:"Quasimodichthys gen. nov. (Neopterygii: Semionotiformes): A morphological and ontogenetic study"
4027:
3839:
3777:
560:
3340:
2930:
581:
554:
197:
4362:(2010): A bizarre, humped Carcharodontosauria (Theropoda) from the Lower Cretaceous of Spain
3617:
2415:
3639:
4080:
3898:
3851:
3789:
3289:
3231:
132:
8:
4378:
3244:
2620:
579:, the most diverse Cretaceous bird clade, which became extinct at the end of the period.
417:
385:
4386:
4084:
3902:
3855:
3793:
3270:
3185:
4104:
3867:
3815:
251:
4315:
608:
The Las Hoyas site has been studied for more than two decades by researchers from the
4396:
4339:
4297:
4252:
4108:
4096:
4049:
4008:
3967:
3916:
3911:
3886:
3871:
3819:
3597:
3223:
3123:
2947:
2938:
2650:
1445:
592:
255:
4446:: Jonathan J. Becker (ed.): Papers in Avian Paleontology Honoring Pierce Brodkorb.
4428:
4419:
4364:
4329:
4289:
4088:
4039:
3998:
3957:
3947:
3936:"The Las Hoyas Lagerstätte: a palaeontological view of an Early Cretaceous wetland"
3906:
3859:
3805:
3797:
3737:
3545:
2803:
2583:
2576:
2489:
2452:
2114:
2084:
1809:
1610:
905:
755:
455:
may have been buried in less than 15 days, after a period of sub-areal exposition.
393:
3863:
3801:
118:
4488:
4390:
4092:
3281:
3274:
2747:
2365:
2335:
1708:
1532:
549:
535:
4316:"A New Freshwater Basal Eucryptodiran Turtle from the Early Cretaceous of Spain"
4003:
3986:
3415:
3320:
3731:
3191:
3179:
2691:
2594:
2256:
2149:
1943:
1067:
923:
576:
413:
4293:
3676:
3314:
2822:
2546:
2422:
401:
4497:
4343:
4301:
4100:
4053:
4012:
3971:
3934:
Marugán-Lobón, Jesús; Martín-Abad, Hugo; Buscalioni, Ángela D. (2023-04-03).
3920:
2721:
2657:
2631:
2557:
2246:
2146:
1729:
1282:
1219:
405:
231:
218:
70:
27:
4382:
4044:
3575:
3113:
3035:
2668:
2613:
2498:
2396:
2228:
2198:
2081:
2051:
1888:
1692:
1428:
1229:
989:
146:
4334:
3776:
Barrios-de Pedro, Sandra; Chin, Karen; Buscalioni, Ángela D. (June 2020).
298:
3691:
3496:
3403:
3308:
2539:
2459:
2286:
2127:
2033:
1845:
1584:
1158:
826:
688:
525:
485:
468:
45:
4314:
Pérez García, Adán; Fuente, Marcelo S. de la; Ortega, Francisco (2012),
3962:
3952:
3935:
3810:
548:
from Las Hoyas (avian and non avian) are unique in many ways. The first
3624:
3604:
3490:
3347:
3258:
2816:
2521:
2470:
1898:
1870:
1790:
1548:
1519:
1129:
1112:
1039:
954:
901:
674:
587:
565:
514:
498:
473:
90:
55:
4448:
Natural
History Museum of Los Angeles County Contributions in Science
568:
would push back the origin of feathers earlier in theropod evolution.
4432:
3697:
3591:
3153:
2728:
2433:
2311:
2096:
1916:
1673:
1474:
1455:
1397:
1326:
1255:
1197:
1163:
785:
545:
510:
420:
389:
183:
95:
39:
31:
3885:
Jaume, Damià; Pinardo-Moya, Eva; Boxshall, Geoff A. (January 2013).
4481:
2063:
1759:
1100:
971:
775:
539:
518:
506:
502:
409:
85:
80:
65:
60:
50:
3933:
2217:
3684:
2275:
1020:
681:
100:
75:
4250:
4275:
3837:
2978:
2897:
2002:
494:
3985:
MartÍn-Abad, Hugo; Poyato-Ariza, Francisco JosÉ (2017-06-05).
3775:
1095:
799:
652:
597:
572:
490:
464:
273:
4395:, Berkeley: University of California Press, pp. 1–880,
4067:
Paiva, Hanna
Carolina Lins de; Gallo, Valéria (2018-12-01).
4026:
Martín-Abad, Hugo; Martínez, Marian
Fregenal- (2021-12-02).
1698:
624:, which is responsible for the fossil record from the area.
1522:
529:
187:
3884:
408:. The site is mostly known for its exquisitely preserved
4377:
4313:
3984:
3760:
416:. The lithology of the formation mostly consists of
4025:
2610:"incomplete but fully-articulated right hindlimb"
123:Lithographic limestones of the Las Hoyas locality
4167:
4165:
4495:
4133:
4193:"Table 11.1," in Weishampel et al., 2004, p.213
4171:"Table 11.1," in Weishampel et al. ,2004, p.212
423:deposited in a freshwater wetland environment.
404:within the formation, located near the city of
4211:"Table 6.1," in Weishampel et al., 2004, p.138
4187:
4162:
346:
318:
4205:
2656:
2619:
2458:
2421:
2412:"Nearly complete and articulated skeleton."
1697:
1473:
647:
622:Museo de las Ciencias de Castilla-La Mancha
354:La Huérguina Formation (Castilla-La Mancha)
4539:Cretaceous paleontological sites of Europe
4258:Revista de la Sociedad Geológica de España
4155:
4153:
4151:
4149:
4147:
4145:
4066:
117:
4333:
4043:
4002:
3991:Zoological Journal of the Linnean Society
3961:
3951:
3910:
3809:
3734:, contemporaneous Lagerstätte of Colombia
618:National University of Distance Education
493:, with specimens of at least five or six
4360:Ortega, F., Escaso, F. & Sanz, J. L.
4073:Journal of South American Earth Sciences
3727:List of dinosaur-bearing rock formations
2364:
2168:
2001:
1758:
1643:
1396:
988:
922:
725:
528:the most abundant and diverse group are
4142:
590:and the presence of a quilled sternum.
388:in Spain whose strata date back to the
4496:
4227:
3740:, contemporaneous Lagerstätte of China
874:A clam shrimp belonging to the family
853:A clam shrimp belonging to the family
3615:
2935:
2875:
968:A full body impression of a juvenile
4214:
4174:
3833:
3831:
3829:
4549:Geography of the Province of Cuenca
3766:Weishampel et al., 2004, pp.556-563
704:Uncertain or tentative taxa are in
513:soft bodied animals which might be
458:
13:
4457:Sanz, J. L. & Buscalioni, A.D.
2821:
2582:
2545:
2274:
2216:
2161:
14:
4560:
4514:Lower Cretaceous Series of Europe
4475:
3940:Journal of the Geological Society
3826:
3769:
2720:Early Cretaceous stratigraphy of
2486:"Thoracic region and forelimbs."
915:
664:Taxon falsely reported as present
627:
614:Universidad Complutense de Madrid
374:Calizas de La Huérguina Formation
4032:Spanish Journal of Palaeontology
3912:10.1111/j.1475-4983.2012.01144.x
3495:
3313:
2357:
378:La Huérguina Limestone Formation
345:
338:
317:
310:
266:
43:
4244:
4196:
4124:
4115:
2361:
2165:
1998:
1755:
1640:
1393:
985:
981:
919:
722:
669:Dubious taxon or junior synonym
170:60–100 m (200–330 ft)
4060:
4019:
3978:
3927:
3878:
3751:
2714:
718:
610:Universidad Autónoma de Madrid
552:dinosaur described in Europe,
357:Show map of Castilla-La Mancha
326:La Huérguina Formation (Spain)
1:
4321:Acta Palaeontologica Polonica
3864:10.1016/j.cretres.2020.104389
3802:10.1016/j.cretres.2020.104409
3744:
2647:"Skull and partial skeleton"
1389:
596:shows the first report of an
16:Geological formation in Spain
4509:Geologic formations of Spain
4093:10.1016/j.jsames.2018.08.010
1751:
1690:
620:, in collaboration with the
431:
426:
7:
4392:The Dinosauria, 2nd edition
3720:
631:
603:
10:
4565:
3708:
3673:
3364:
3302:
2820:
1994:
1636:
1161:, originally described as
1098:, originally described as
4294:10.1017/S0016756813000095
4121:Pérez García et al., 2012
4004:10.1093/zoolinnean/zlx010
3668:
3648:
3635:
3622:
3602:
3595:
3590:
3564:
3557:
3550:
3535:
3519:
3501:
3494:
3489:
3480:
3449:
3407:
3402:
3386:
3377:
3366:
3357:
3337:
3330:
3319:
3312:
3307:
3279:
3250:
3243:
3236:
3229:
3220:
3171:
3157:
3152:
3134:
3122:
3068:
2982:
2977:
2973:
2944:
2912:
2896:
2891:
2861:
2859:
2857:
2855:
2853:
2841:
2834:
2815:
2810:
1915:
1896:
1453:
1444:
1325:
1156:
845:
770:
765:
638:
634:
304:
294:
289:
279:
261:
247:
208:
203:
193:
179:
174:
166:
156:
138:
128:
116:
26:
21:
2573:"Postcranial skeleton."
2449:"Postcranial skeleton."
829:belonging to the family
636:
467:(with many specimens of
4356:doi:10.1038/nature14905
2381:Stratigraphic position
2183:Stratigraphic position
2018:Stratigraphic position
1775:Stratigraphic position
1658:Stratigraphic position
1413:Stratigraphic position
1005:Stratigraphic position
939:Stratigraphic position
740:Stratigraphic position
561:Concavenator corcovatus
4504:La Huérguina Formation
4482:Las Hoyas site website
4045:10.7203/sjp.36.2.21966
2826:
2661:
2624:
2587:
2550:
2463:
2426:
2279:
2221:
1961:Paramacellodid lizard
1702:
1478:
712:taxa are discredited.
582:Iberomesornis romerali
555:Pelecanimimus polyodon
538:are the most abundant
438:Konservat-Lagerstätten
370:La Huérguina Formation
22:La Huérguina Formation
4544:Paleontology in Spain
4335:10.4202/app.2011.0031
3757:Martínez et al., 2017
2825:
2660:
2623:
2586:
2549:
2462:
2425:
2278:
2220:
1701:
1477:
4524:Limestone formations
4379:Weishampel, David B.
4202:Serrano et al., 2013
133:Geological formation
4534:Lacustrine deposits
4282:Geological Magazine
4139:Ortega et al., 2010
4130:Martin et al., 2015
4085:2018JSAES..88..132P
3953:10.1144/jgs2022-079
3903:2013Palgy..56...15J
3856:2020CrRes.11004389G
3844:Cretaceous Research
3794:2020CrRes.11004409B
3782:Cretaceous Research
3677:Major fossiliferous
2724:
2416:Carcharodontosaurid
771:Numerous specimens
372:(also known as the
284:South Iberian Basin
228: /
161:Tragacete Formation
143:Contreras Formation
28:Stratigraphic range
4487:2020-02-22 at the
3692:ichnofossiliferous
2827:
2719:
2662:
2651:Ornithomimosaurian
2625:
2601:M. atherfieldensis
2588:
2551:
2464:
2427:
2280:
2222:
1703:
1479:
659:Reclassified taxon
384:) is a geological
232:40.0897°N 1.8978°W
4435:
4426:(6590): 442–445.
4387:Osmólska, Halszka
3715:
3714:
3704:
3664:
3627:
3607:
3586:
3569:
3562:
3542:
3531:
3524:
3517:
3510:
3485:
3469:
3454:
3447:
3434:
3412:
3398:
3382:
3373:
3362:
3355:
3335:
3328:
3241:
3218:
3209:
3169:
3162:
3148:
3141:
3132:
3120:
3109:
3102:
3089:
3082:
3066:
3032:
3023:
3016:
3009:
2991:
2971:
2958:
2924:
2917:
2910:
2887:
2873:
2851:
2839:
2736:Paleomap \ Basins
2712:
2711:
2355:
2354:
2159:
2158:
2117:multituberculate
1992:
1991:
1749:
1748:
1634:
1633:
1387:
1386:
979:
978:
913:
912:
876:Antronestheriidae
811:Lahuerguinagrapta
716:
715:
707:
695:
694:
691:
684:
677:
670:
665:
660:
655:
593:Eoalulavis hoyasi
366:
365:
329:Show map of Spain
256:Castile-La Mancha
4556:
4519:Cretaceous Spain
4458:
4433:10.1038/382442a0
4427:
4416:
4411:
4410:
4409:
4361:
4352:
4351:
4350:
4337:
4310:
4309:
4308:
4272:
4271:
4270:
4238:
4231:
4225:
4218:
4212:
4209:
4203:
4200:
4194:
4191:
4185:
4178:
4172:
4169:
4160:
4157:
4140:
4137:
4131:
4128:
4122:
4119:
4113:
4112:
4064:
4058:
4057:
4047:
4023:
4017:
4016:
4006:
3982:
3976:
3975:
3965:
3955:
3931:
3925:
3924:
3914:
3882:
3876:
3875:
3835:
3824:
3823:
3813:
3773:
3767:
3764:
3758:
3755:
3738:Yixian Formation
3702:
3660:
3623:
3603:
3574:
3565:
3558:
3538:
3527:
3520:
3513:
3502:
3499:
3481:
3465:
3450:
3439:
3426:
3408:
3394:
3378:
3367:
3358:
3351:
3331:
3324:
3317:
3237:
3214:
3205:
3165:
3158:
3144:
3135:
3128:
3112:
3105:
3094:
3085:
3076:
3062:
3028:
3019:
3012:
3005:
2983:
2963:
2954:
2920:
2913:
2902:
2883:
2869:
2847:
2835:
2806:
2799:
2792:
2785:
2778:
2771:
2764:
2757:
2750:
2743:
2725:
2718:
2362:
2169:Crocodyliformes
2166:
2085:multituberculate
1999:
1756:
1641:
1611:Wealdenbatrachus
1502:Gracilibatrachus
1394:
986:
920:
906:Spelaeogriphacea
756:Austropotamobius
723:
705:
687:
680:
673:
668:
663:
658:
651:
648:
632:
542:from Las Hoyas.
459:Paleoenvironment
414:enantiornithines
394:Early Cretaceous
358:
349:
348:
342:
330:
321:
320:
314:
272:
270:
269:
243:
242:
240:
239:
238:
237:40.0897; -1.8978
233:
229:
226:
225:
224:
221:
121:
112:
42:
19:
18:
4564:
4563:
4559:
4558:
4557:
4555:
4554:
4553:
4529:Marl formations
4494:
4493:
4489:Wayback Machine
4478:
4456:
4436:(HTML abstract)
4414:
4407:
4405:
4403:
4359:
4348:
4346:
4306:
4304:
4268:
4266:
4247:
4242:
4241:
4233:Listed as "cf.
4232:
4228:
4220:Listed as "cf.
4219:
4215:
4210:
4206:
4201:
4197:
4192:
4188:
4180:Listed as "cf.
4179:
4175:
4170:
4163:
4158:
4143:
4138:
4134:
4129:
4125:
4120:
4116:
4065:
4061:
4024:
4020:
3983:
3979:
3932:
3928:
3883:
3879:
3836:
3827:
3774:
3770:
3765:
3761:
3756:
3752:
3747:
3723:
3717:
3703:minor formation
3685:oofossiliferous
3679:
3654:
3642:
3582:
3370:
3201:
3138:
3079:
2950:
2941:
2802:
2795:
2788:
2781:
2774:
2767:
2760:
2753:
2746:
2739:
2717:
2577:enantiornithean
2490:enantiornithean
2453:enantiornithean
2360:
2336:Bernissartiidae
2164:
2162:Crocodyliformes
2134:S. xenarthrosus
1997:
1905:M. diazromerali
1798:B. cataphractus
1796:Indeterminate,
1754:
1709:Pleurosternidae
1639:
1533:Hylaeobatrachus
1485:C. megacephalus
1446:Albanerpetontid
1392:
1300:Gonorynchiform
984:
918:
865:Pseudestherites
817:L. multicostata
721:
703:
646:
630:
606:
550:ornithomimosaur
536:Crocodylomorphs
461:
434:
429:
400:is a Konservat-
362:
361:
360:
359:
356:
355:
352:
351:
350:
333:
332:
331:
328:
327:
324:
323:
322:
267:
265:
236:
234:
230:
227:
222:
219:
217:
215:
214:
124:
111:
110:
109:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
37:
36:
34:
17:
12:
11:
5:
4562:
4552:
4551:
4546:
4541:
4536:
4531:
4526:
4521:
4516:
4511:
4506:
4492:
4491:
4477:
4476:External links
4474:
4473:
4472:
4468:(4): 829–845.
4454:
4440:
4437:
4412:
4401:
4375:
4357:
4353:
4328:(2): 285–298,
4311:
4288:(3): 572–576,
4273:
4246:
4243:
4240:
4239:
4235:Ricardoestesia
4226:
4213:
4204:
4195:
4186:
4173:
4161:
4141:
4132:
4123:
4114:
4059:
4038:(2): 191–204.
4018:
3997:(3): 604–637.
3977:
3926:
3877:
3825:
3768:
3759:
3749:
3748:
3746:
3743:
3742:
3741:
3735:
3732:Paja Formation
3729:
3722:
3719:
3713:
3712:
3706:
3705:
3675:
3672:
3666:
3665:
3657:
3656:
3650:
3646:
3645:
3638:
3634:
3621:
3614:
3601:
3594:
3588:
3587:
3571:
3570:
3563:
3556:
3549:
3536:
3533:
3532:
3525:
3518:
3511:
3500:
3493:
3487:
3486:
3479:
3470:
3462:
3461:
3448:
3436:
3435:
3424:
3413:
3406:
3400:
3399:
3392:
3384:
3383:
3375:
3374:
3365:
3363:
3356:
3345:
3336:
3329:
3318:
3311:
3306:
3300:
3299:
3294:
3285:
3278:
3267:
3256:
3248:
3247:
3242:
3235:
3228:
3219:
3211:
3210:
3203:
3197:
3194:
3189:
3182:
3177:
3170:
3163:
3156:
3150:
3149:
3142:
3133:
3126:
3121:
3110:
3103:
3091:
3090:
3083:
3074:
3067:
3060:
3055:
3044:
3033:
3025:
3024:
3017:
3010:
3003:
2992:
2981:
2975:
2974:
2972:
2960:
2959:
2952:
2946:
2943:
2937:
2934:
2926:
2925:
2918:
2911:
2900:
2895:
2889:
2888:
2880:
2879:
2874:
2866:
2865:
2860:
2858:
2856:
2854:
2852:
2845:
2840:
2833:
2828:
2819:
2814:
2808:
2807:
2800:
2793:
2786:
2779:
2772:
2765:
2758:
2751:
2744:
2737:
2734:
2731:
2716:
2713:
2710:
2709:
2707:
2705:
2703:
2701:
2698:
2697:Indeterminate
2695:
2692:Ricardoestesia
2687:
2686:
2684:
2682:
2680:
2678:
2675:
2674:Indeterminate
2672:
2664:
2663:
2654:
2648:
2645:
2643:
2640:
2635:
2627:
2626:
2617:
2611:
2608:
2606:
2603:
2598:
2595:Mantellisaurus
2590:
2589:
2580:
2574:
2571:
2569:
2566:
2561:
2553:
2552:
2543:
2537:
2535:
2533:
2530:
2528:E. olcadesorum
2525:
2517:
2516:
2514:
2512:
2510:
2508:
2505:
2504:Indeterminate
2502:
2494:
2493:
2487:
2484:
2482:
2479:
2474:
2466:
2465:
2456:
2450:
2447:
2445:
2442:
2437:
2429:
2428:
2419:
2413:
2410:
2408:
2405:
2400:
2392:
2391:
2388:
2385:
2382:
2379:
2376:
2373:
2369:
2368:
2359:
2356:
2353:
2352:
2350:
2348:
2346:
2344:
2341:
2340:Indeterminate
2338:
2332:
2331:
2329:
2327:
2325:
2323:
2320:
2315:
2307:
2306:
2304:
2302:
2300:
2298:
2295:
2290:
2282:
2281:
2272:
2270:
2268:
2266:
2263:
2262:Indeterminate
2260:
2257:Montsecosuchus
2252:
2251:
2249:
2244:
2242:
2240:
2237:
2232:
2224:
2223:
2214:
2212:
2210:
2208:
2205:
2204:Indeterminate
2202:
2194:
2193:
2190:
2187:
2184:
2181:
2178:
2175:
2171:
2170:
2163:
2160:
2157:
2156:
2154:
2152:
2144:
2141:
2139:
2136:
2131:
2123:
2122:
2120:
2118:
2115:Paulchoffatiid
2112:
2110:
2108:
2105:
2100:
2092:
2091:
2089:
2087:
2079:
2077:
2075:
2072:
2067:
2059:
2058:
2056:
2054:
2049:
2047:
2045:
2042:
2037:
2029:
2028:
2025:
2022:
2019:
2016:
2013:
2010:
2006:
2005:
1996:
1993:
1990:
1989:
1987:
1985:
1983:
1981:
1978:
1973:
1965:
1964:
1962:
1959:
1957:
1955:
1952:
1947:
1944:Paramacellodus
1939:
1938:
1936:
1934:
1932:
1929:
1923:
1922:
1920:
1914:
1912:
1910:
1907:
1902:
1894:
1893:
1891:
1886:
1884:
1882:
1879:
1874:
1866:
1865:
1863:
1861:
1859:
1857:
1854:
1849:
1841:
1840:
1838:
1836:
1834:
1832:
1829:
1824:
1816:
1815:
1813:
1810:Paramacellodid
1807:
1805:
1803:
1800:
1794:
1786:
1785:
1782:
1779:
1776:
1773:
1770:
1767:
1763:
1762:
1753:
1750:
1747:
1746:
1744:
1742:
1740:
1738:
1735:
1734:Indeterminate
1732:
1726:
1725:
1723:
1721:
1719:
1717:
1714:
1713:Indeterminate
1711:
1705:
1704:
1695:
1689:
1687:
1685:
1682:
1677:
1669:
1668:
1665:
1662:
1659:
1656:
1653:
1650:
1646:
1645:
1638:
1635:
1632:
1631:
1629:
1626:
1624:
1622:
1619:
1614:
1606:
1605:
1603:
1600:
1598:
1596:
1593:
1588:
1580:
1579:
1577:
1574:
1572:
1570:
1567:
1562:
1559:Iberobatrachus
1554:
1553:
1551:
1546:
1544:
1542:
1539:
1538:Indeterminate
1536:
1528:
1527:
1525:
1517:
1515:
1513:
1510:
1505:
1497:
1496:
1494:
1492:
1490:
1487:
1481:
1480:
1471:
1469:
1467:
1464:
1459:
1451:
1450:
1448:
1443:
1440:
1438:
1435:
1434:Indeterminate
1432:
1424:
1423:
1420:
1417:
1414:
1411:
1408:
1405:
1401:
1400:
1391:
1388:
1385:
1384:
1382:
1379:
1377:
1375:
1372:
1363:
1355:
1354:
1352:
1350:
1348:
1345:
1340:
1332:
1331:
1329:
1324:
1322:
1320:
1317:
1312:
1304:
1303:
1301:
1298:
1296:
1294:
1291:
1286:
1278:
1277:
1275:
1272:
1270:
1268:
1265:
1259:
1251:
1250:
1248:
1246:
1244:
1242:
1239:
1233:
1225:
1224:
1222:
1217:
1215:
1213:
1210:
1201:
1193:
1192:
1190:
1188:
1186:
1183:
1178:
1170:
1169:
1167:
1155:
1153:
1151:
1148:
1143:
1135:
1134:
1132:
1127:
1125:
1123:
1120:
1119:Indeterminate
1117:
1107:
1106:
1104:
1093:
1091:
1089:
1086:
1081:
1073:
1072:
1070:
1068:Gonorynchiform
1065:
1063:
1061:
1058:
1053:
1045:
1044:
1042:
1037:
1035:
1033:
1030:
1024:
1016:
1015:
1012:
1009:
1006:
1003:
1000:
997:
993:
992:
983:
980:
977:
976:
974:
969:
966:
964:
961:
960:Indeterminate
958:
950:
949:
946:
943:
940:
937:
934:
931:
927:
926:
924:Chondrichthyes
917:
916:Chondrichthyes
914:
911:
910:
908:
898:
896:
894:
889:
881:
880:
878:
872:
870:
868:
860:
859:
857:
851:
849:
847:
846:Indeterminate
844:
836:
835:
833:
823:
821:
819:
814:
806:
805:
803:
796:
794:
789:
781:
780:
778:
772:
769:
767:
764:
759:
751:
750:
747:
744:
741:
738:
735:
732:
728:
727:
720:
717:
714:
713:
698:
696:
693:
692:
685:
678:
671:
666:
661:
656:
640:
639:
637:
635:
629:
628:Fossil content
626:
605:
602:
577:enantiornithes
489:) and diverse
460:
457:
433:
430:
428:
425:
364:
363:
353:
344:
343:
337:
336:
335:
334:
325:
316:
315:
309:
308:
307:
306:
305:
302:
301:
296:
292:
291:
287:
286:
281:
277:
276:
263:
259:
258:
249:
245:
244:
212:
206:
205:
201:
200:
195:
191:
190:
181:
177:
176:
172:
171:
168:
164:
163:
158:
154:
153:
151:Utrillas Group
140:
136:
135:
130:
126:
125:
122:
114:
113:
106:
105:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
24:
23:
15:
9:
6:
4:
3:
2:
4561:
4550:
4547:
4545:
4542:
4540:
4537:
4535:
4532:
4530:
4527:
4525:
4522:
4520:
4517:
4515:
4512:
4510:
4507:
4505:
4502:
4501:
4499:
4490:
4486:
4483:
4480:
4479:
4471:
4467:
4464:
4463:
4462:Palaeontology
4455:
4452:
4449:
4445:
4441:
4438:
4434:
4430:
4425:
4422:
4421:
4413:
4404:
4402:0-520-24209-2
4398:
4394:
4393:
4388:
4384:
4383:Dodson, Peter
4380:
4376:
4374:
4373:HTML abstract
4370:
4367:
4366:
4358:
4354:
4345:
4341:
4336:
4331:
4327:
4323:
4322:
4317:
4312:
4303:
4299:
4295:
4291:
4287:
4283:
4279:
4274:
4264:
4260:
4259:
4254:
4249:
4248:
4236:
4230:
4223:
4217:
4208:
4199:
4190:
4183:
4177:
4168:
4166:
4156:
4154:
4152:
4150:
4148:
4146:
4136:
4127:
4118:
4110:
4106:
4102:
4098:
4094:
4090:
4086:
4082:
4078:
4074:
4070:
4063:
4055:
4051:
4046:
4041:
4037:
4033:
4029:
4022:
4014:
4010:
4005:
4000:
3996:
3992:
3988:
3981:
3973:
3969:
3964:
3959:
3954:
3949:
3945:
3941:
3937:
3930:
3922:
3918:
3913:
3908:
3904:
3900:
3896:
3892:
3891:Palaeontology
3888:
3881:
3873:
3869:
3865:
3861:
3857:
3853:
3849:
3845:
3841:
3834:
3832:
3830:
3821:
3817:
3812:
3807:
3803:
3799:
3795:
3791:
3787:
3783:
3779:
3772:
3763:
3754:
3750:
3739:
3736:
3733:
3730:
3728:
3725:
3724:
3718:
3711:
3707:
3700:
3699:
3698:coproliferous
3694:
3693:
3688:
3687:
3686:
3680:
3678:
3671:
3667:
3663:
3659:
3658:
3655:
3653:
3647:
3644:
3643:
3641:
3633:
3632:
3626:
3620:
3619:
3613:
3612:
3606:
3600:
3599:
3593:
3589:
3585:
3584:Sierra Matute
3581:
3577:
3573:
3572:
3568:
3561:
3555:
3554:
3548:
3547:
3541:
3537:
3534:
3530:
3526:
3523:
3516:
3512:
3509:
3505:
3498:
3492:
3488:
3484:
3478:
3477:
3476:
3471:
3468:
3464:
3463:
3460:
3459:
3453:
3446:
3442:
3438:
3437:
3433:
3429:
3425:
3423:
3422:
3417:
3414:
3411:
3405:
3401:
3397:
3393:
3391:
3390:
3385:
3381:
3376:
3372:
3361:
3354:
3349:
3346:
3344:
3343:
3342:
3334:
3327:
3322:
3316:
3310:
3305:
3301:
3298:
3295:
3293:
3292:
3291:
3286:
3284:
3283:
3277:
3276:
3272:
3268:
3266:
3265:
3260:
3257:
3255:
3254:
3249:
3246:
3240:
3234:
3233:
3227:
3226:
3225:
3217:
3213:
3212:
3208:
3204:
3202:
3200:
3195:
3193:
3190:
3188:
3187:
3183:
3181:
3178:
3176:
3175:
3168:
3164:
3161:
3155:
3151:
3147:
3143:
3140:
3131:
3127:
3125:
3119:
3115:
3111:
3108:
3104:
3101:
3097:
3093:
3092:
3088:
3084:
3081:
3075:
3073:
3072:
3065:
3061:
3059:
3056:
3054:
3050:
3049:
3045:
3043:
3042:
3037:
3034:
3031:
3027:
3026:
3022:
3018:
3015:
3011:
3008:
3004:
3002:
3001:
2996:
2993:
2990:
2986:
2980:
2976:
2970:
2966:
2962:
2961:
2957:
2953:
2951:
2949:
2942:
2940:
2933:
2932:
2928:
2927:
2923:
2919:
2916:
2909:
2905:
2901:
2899:
2894:
2890:
2886:
2882:
2881:
2878:
2872:
2868:
2867:
2864:
2863:Caranguejeira
2850:
2846:
2844:
2838:
2832:
2829:
2824:
2818:
2813:
2809:
2805:
2801:
2798:
2794:
2791:
2790:South Iberian
2787:
2784:
2780:
2777:
2773:
2770:
2766:
2763:
2759:
2756:
2752:
2749:
2745:
2742:
2738:
2735:
2732:
2730:
2727:
2726:
2723:
2708:
2706:
2704:
2702:
2699:
2696:
2694:
2693:
2689:
2688:
2685:
2683:
2681:
2679:
2676:
2673:
2671:
2670:
2666:
2665:
2659:
2655:
2652:
2649:
2646:
2644:
2641:
2639:
2636:
2634:
2633:
2632:Pelecanimimus
2629:
2628:
2622:
2618:
2615:
2614:Iguanodontian
2612:
2609:
2607:
2604:
2602:
2599:
2597:
2596:
2592:
2591:
2585:
2581:
2578:
2575:
2572:
2570:
2567:
2565:
2562:
2560:
2559:
2558:Iberomesornis
2555:
2554:
2548:
2544:
2541:
2538:
2536:
2534:
2531:
2529:
2526:
2524:
2523:
2519:
2518:
2515:
2513:
2511:
2509:
2506:
2503:
2501:
2500:
2496:
2495:
2491:
2488:
2485:
2483:
2480:
2478:
2475:
2473:
2472:
2468:
2467:
2461:
2457:
2454:
2451:
2448:
2446:
2443:
2441:
2438:
2436:
2435:
2431:
2430:
2424:
2420:
2417:
2414:
2411:
2409:
2406:
2404:
2403:C. corcovatus
2401:
2399:
2398:
2394:
2393:
2389:
2386:
2383:
2380:
2377:
2374:
2371:
2370:
2367:
2366:Ornithodirans
2363:
2358:Ornithodirans
2351:
2349:
2347:
2345:
2342:
2339:
2337:
2334:
2333:
2330:
2328:
2326:
2324:
2321:
2319:
2316:
2314:
2313:
2309:
2308:
2305:
2303:
2301:
2299:
2296:
2294:
2291:
2289:
2288:
2284:
2283:
2277:
2273:
2271:
2269:
2267:
2264:
2261:
2259:
2258:
2254:
2253:
2250:
2248:
2245:
2243:
2241:
2238:
2236:
2233:
2231:
2230:
2226:
2225:
2219:
2215:
2213:
2211:
2209:
2206:
2203:
2201:
2200:
2196:
2195:
2191:
2188:
2185:
2182:
2179:
2176:
2173:
2172:
2167:
2155:
2153:
2151:
2150:Eutriconodont
2148:
2145:
2142:
2140:
2137:
2135:
2132:
2130:
2129:
2125:
2124:
2121:
2119:
2116:
2113:
2111:
2109:
2106:
2104:
2101:
2099:
2098:
2094:
2093:
2090:
2088:
2086:
2083:
2080:
2078:
2076:
2073:
2071:
2070:E. hispanicus
2068:
2066:
2065:
2061:
2060:
2057:
2055:
2053:
2050:
2048:
2046:
2043:
2041:
2038:
2036:
2035:
2031:
2030:
2026:
2023:
2020:
2017:
2014:
2011:
2008:
2007:
2004:
2000:
1988:
1986:
1984:
1982:
1979:
1977:
1976:S. ciervensis
1974:
1972:
1971:
1967:
1966:
1963:
1960:
1958:
1956:
1953:
1951:
1948:
1946:
1945:
1941:
1940:
1937:
1935:
1933:
1930:
1928:
1925:
1924:
1921:
1918:
1913:
1911:
1908:
1906:
1903:
1901:
1900:
1895:
1892:
1890:
1887:
1885:
1883:
1880:
1878:
1875:
1873:
1872:
1868:
1867:
1864:
1862:
1860:
1858:
1855:
1853:
1850:
1848:
1847:
1843:
1842:
1839:
1837:
1835:
1833:
1830:
1828:
1825:
1823:
1822:
1818:
1817:
1814:
1811:
1808:
1806:
1804:
1801:
1799:
1795:
1793:
1792:
1788:
1787:
1783:
1780:
1777:
1774:
1771:
1768:
1765:
1764:
1761:
1757:
1745:
1743:
1741:
1739:
1736:
1733:
1731:
1730:Pelomedusidae
1728:
1727:
1724:
1722:
1720:
1718:
1715:
1712:
1710:
1707:
1706:
1700:
1696:
1694:
1688:
1686:
1683:
1681:
1678:
1676:
1675:
1671:
1670:
1666:
1663:
1660:
1657:
1654:
1651:
1648:
1647:
1642:
1630:
1627:
1625:
1623:
1620:
1618:
1617:W. jucarensis
1615:
1613:
1612:
1608:
1607:
1604:
1601:
1599:
1597:
1594:
1592:
1589:
1587:
1586:
1582:
1581:
1578:
1575:
1573:
1571:
1568:
1566:
1563:
1561:
1560:
1556:
1555:
1552:
1550:
1547:
1545:
1543:
1540:
1537:
1535:
1534:
1530:
1529:
1526:
1524:
1521:
1518:
1516:
1514:
1511:
1509:
1506:
1504:
1503:
1499:
1498:
1495:
1493:
1491:
1488:
1486:
1483:
1482:
1476:
1472:
1470:
1468:
1465:
1463:
1460:
1458:
1457:
1452:
1449:
1447:
1441:
1439:
1436:
1433:
1431:
1430:
1426:
1425:
1421:
1418:
1415:
1412:
1409:
1406:
1403:
1402:
1399:
1395:
1383:
1380:
1378:
1376:
1373:
1371:
1367:
1364:
1362:
1361:
1357:
1356:
1353:
1351:
1349:
1346:
1344:
1343:T. praeclarus
1341:
1339:
1338:
1334:
1333:
1330:
1328:
1323:
1321:
1318:
1316:
1313:
1311:
1310:
1306:
1305:
1302:
1299:
1297:
1295:
1292:
1290:
1287:
1285:
1284:
1283:Rubiesichthys
1280:
1279:
1276:
1273:
1271:
1269:
1266:
1263:
1260:
1258:
1257:
1253:
1252:
1249:
1247:
1245:
1243:
1240:
1237:
1234:
1232:
1231:
1227:
1226:
1223:
1221:
1218:
1216:
1214:
1211:
1209:
1205:
1202:
1200:
1199:
1195:
1194:
1191:
1189:
1187:
1184:
1182:
1179:
1177:
1176:
1172:
1171:
1168:
1166:
1165:
1160:
1154:
1152:
1149:
1147:
1144:
1142:
1141:
1137:
1136:
1133:
1131:
1128:
1126:
1124:
1121:
1118:
1115:
1114:
1109:
1108:
1105:
1103:
1102:
1097:
1094:
1092:
1090:
1087:
1085:
1082:
1080:
1079:
1075:
1074:
1071:
1069:
1066:
1064:
1062:
1059:
1057:
1056:G. conquensis
1054:
1052:
1051:
1047:
1046:
1043:
1041:
1038:
1036:
1034:
1031:
1028:
1025:
1023:
1022:
1018:
1017:
1013:
1010:
1007:
1004:
1001:
998:
995:
994:
991:
987:
975:
973:
970:
967:
965:
962:
959:
957:
956:
952:
951:
947:
944:
941:
938:
935:
932:
929:
928:
925:
921:
909:
907:
904:belonging to
903:
899:
897:
895:
893:
890:
888:
887:
883:
882:
879:
877:
873:
871:
869:
867:
866:
862:
861:
858:
856:
855:Polygraptidae
852:
850:
848:
843:
842:
838:
837:
834:
832:
831:Afrograptidae
828:
824:
822:
820:
818:
815:
813:
812:
808:
807:
804:
801:
797:
795:
793:
792:D. martinelli
790:
788:
787:
783:
782:
779:
777:
773:
768:
763:
760:
758:
757:
753:
752:
748:
745:
742:
739:
736:
733:
730:
729:
724:
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702:
699:
697:
690:
686:
683:
679:
676:
672:
667:
662:
657:
654:
650:
649:
645:
642:
641:
633:
625:
623:
619:
615:
611:
601:
599:
595:
594:
589:
584:
583:
578:
574:
569:
567:
563:
562:
557:
556:
551:
547:
543:
541:
537:
533:
531:
527:
522:
520:
516:
512:
508:
504:
500:
496:
492:
488:
487:
482:
481:
476:
475:
470:
466:
456:
454:
448:
446:
445:Pelecanimimus
441:
439:
424:
422:
419:
415:
412:, especially
411:
407:
406:Cuenca, Spain
403:
399:
395:
392:stage of the
391:
387:
383:
382:Una Formation
379:
375:
371:
341:
313:
303:
300:
297:
293:
288:
285:
282:
278:
275:
264:
260:
257:
253:
250:
246:
241:
213:
211:
207:
202:
199:
196:
192:
189:
185:
182:
178:
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155:
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144:
141:
137:
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127:
120:
115:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
41:
33:
29:
25:
20:
4470:PDF fulltext
4465:
4460:
4450:
4447:
4443:
4423:
4418:
4406:, retrieved
4391:
4368:
4363:
4347:, retrieved
4325:
4319:
4305:, retrieved
4285:
4281:
4267:, retrieved
4262:
4256:
4245:Bibliography
4234:
4229:
4222:Paranychodon
4221:
4216:
4207:
4198:
4189:
4182:Euronychodon
4181:
4176:
4135:
4126:
4117:
4076:
4072:
4062:
4035:
4031:
4021:
3994:
3990:
3980:
3963:10486/710760
3943:
3939:
3929:
3897:(1): 15–28.
3894:
3890:
3880:
3847:
3843:
3811:10486/695365
3785:
3781:
3771:
3762:
3753:
3716:
3696:
3690:
3683:
3682:
3674:
3669:
3649:
3637:
3636:
3629:
3616:
3609:
3596:
3559:
3551:
3544:
3521:
3473:
3472:
3456:
3419:
3387:
3379:
3339:
3338:
3303:
3288:
3287:
3280:
3269:
3262:
3251:
3239:Moutonianum
3230:
3222:
3221:
3198:
3196:
3184:
3172:
3069:
3046:
3039:
2998:
2945:
2936:
2931:Monte Grande
2929:
2892:
2811:
2690:
2669:Paronychodon
2667:
2637:
2630:
2600:
2593:
2563:
2556:
2527:
2520:
2499:Euronychodon
2497:
2476:
2469:
2440:C. lacustris
2439:
2432:
2402:
2397:Concavenator
2395:
2317:
2310:
2292:
2285:
2255:
2235:C. sanziuami
2234:
2229:Cassissuchus
2227:
2199:Bernissartia
2197:
2147:Gobiconodont
2133:
2126:
2103:G. nannothus
2102:
2095:
2069:
2062:
2039:
2032:
1975:
1968:
1949:
1942:
1926:
1904:
1897:
1889:Scleroglossa
1877:J. grandipes
1876:
1869:
1851:
1844:
1826:
1821:Cuencasaurus
1819:
1797:
1789:
1693:eucryptodire
1679:
1672:
1616:
1609:
1590:
1583:
1564:
1557:
1531:
1507:
1500:
1484:
1461:
1454:
1429:Albanerpeton
1427:
1369:
1365:
1358:
1342:
1337:Turbomesodon
1335:
1314:
1307:
1288:
1281:
1274:Macrosemiid
1261:
1254:
1235:
1230:Pleuropholis
1228:
1207:
1203:
1196:
1181:L. microrhis
1180:
1173:
1162:
1145:
1138:
1111:
1099:
1083:
1076:
1055:
1048:
1026:
1019:
990:Osteichthyes
982:Osteichthyes
953:
900:A member of
891:
886:Spinogriphus
884:
863:
839:
816:
809:
791:
784:
761:
754:
726:Crustaceans
709:
700:
643:
607:
591:
588:neornithines
580:
570:
559:
553:
544:
534:
523:
484:
478:
472:
462:
452:
449:
444:
442:
435:
397:
381:
377:
373:
369:
367:
290:Type section
198:Conglomerate
147:Unconformity
4371:: 203–206.
4079:: 132–143.
3404:Valanginian
3309:Hauterivian
3160:Vega de Pas
2715:Correlation
2638:P. polyodon
2564:I. romerali
2287:Sabresuchus
2247:Gobiosuchid
2128:Spinolestes
2040:C. cuencana
2034:Crusafontia
1950:P. sinuosus
1927:M. unaensis
1846:Hoyalacerta
1680:H. jimenezi
1602:Salamander
1591:V. gracilis
1585:Valdotriton
1462:C. ibericus
1289:R. gregalis
1220:Macrosemiid
1159:Lepidotidae
1146:H. tanyrhis
1084:H. newbreyi
1050:Gordichthys
892:S. ibericus
841:Skyestheria
827:clam shrimp
719:Crustaceans
710:crossed out
689:Morphotaxon
566:quill knobs
526:vertebrates
486:Frenelopsis
469:Charophytes
402:Lagerstätte
235: /
210:Coordinates
4498:Categories
4408:2019-02-21
4349:2019-10-12
4307:2019-10-12
4269:2019-10-12
3850:: 104389.
3788:: 104409.
3745:References
3618:Higuereles
3515:Valdeprado
3491:Berriasian
3458:Piedrahita
3445:Hortigüela
3432:Hortigüela
3290:Camarillas
3096:Patrocinio
2989:Tellamendi
2849:Mosquerela
2817:Cenomanian
2804:Lusitanian
2762:Maestrazgo
2741:Cantabrian
2700:Las Hoyas
2677:Las Hoyas
2642:Las Hoyas
2605:Las Hoyas
2568:Las Hoyas
2542:pterosaur
2532:Las Hoyas
2522:Europejara
2507:Las Hoyas
2481:Las Hoyas
2471:Eoalulavis
2444:Las Hoyas
2407:Las Hoyas
2343:Las Hoyas
2318:U. reginae
2293:S.ibericus
2265:Las Hoyas
2239:Las Hoyas
2138:Las Hoyas
2082:Eobaatarid
2052:Dryolestid
1980:Las Hoyas
1970:Scandensia
1909:Las Hoyas
1899:Meyasaurus
1881:Las Hoyas
1871:Jucaraseps
1856:Las Hoyas
1791:Becklesius
1684:Las Hoyas
1595:Las Hoyas
1569:Las Hoyas
1565:I. angelae
1549:Salamander
1541:Las Hoyas
1512:Las Hoyas
1508:G. avallei
1466:Las Hoyas
1398:Amphibians
1390:Amphibians
1374:Las Hoyas
1370:catalunica
1347:Las Hoyas
1319:Las Hoyas
1293:Las Hoyas
1267:Las Hoyas
1241:Las Hoyas
1212:Las Hoyas
1208:N. ferreri
1185:Las Hoyas
1175:Lepidohyas
1157:Member of
1150:Las Hoyas
1130:Coelacanth
1122:Las Hoyas
1113:Holophagus
1088:Las Hoyas
1078:Hispanamia
1060:Las Hoyas
1040:Amiiformes
1032:Las Hoyas
963:Las Hoyas
955:Lonchidion
902:Peracarida
766:Las Hoyas
762:A. llopisi
706:small text
675:Ichnotaxon
571:Las Hoyas
515:Nemertines
499:arthropods
480:Weischelia
474:Montsechia
418:lacustrine
380:or as the
220:40°05′23″N
4344:0567-7920
4302:0016-7568
4265:: 113–142
4109:134986658
4101:0895-9811
4054:2660-9568
4013:0024-4082
3972:0016-7649
3921:0031-0239
3872:214213415
3820:213738761
3652:Arzobispo
3592:Tithonian
3580:Huérteles
3553:Arzobispo
3483:Miravetes
3341:Castellar
3245:Papo Seco
3199:Huérguina
3154:Barremian
3139:Bedoulian
3130:Villaroya
3064:Villaroya
3000:Castrillo
2985:Valmaseda
2922:Traiguera
2908:Balmaseda
2831:La Cabana
2797:Pre-betic
2653:dinosaur
2616:dinosaur
2540:Tapejarid
2477:E. hoyasi
2434:Concornis
2418:dinosaur
2384:Material
2378:Location
2312:Unasuchus
2186:Material
2180:Location
2097:Galveodon
2021:Material
2015:Location
1827:C. estesi
1778:Material
1772:Location
1760:Squamates
1752:Squamates
1674:Hoyasemys
1661:Material
1655:Location
1520:Pipimorph
1456:Celtedens
1442:Frontals
1416:Material
1410:Location
1360:Vidalamia
1327:Pycnodont
1309:Stenamara
1256:Propterus
1198:Notagogus
1164:Lepidotes
1140:Hoyasotes
1008:Material
1002:Location
942:Material
936:Location
786:Delclosia
743:Material
737:Location
644:Color key
546:Dinosaurs
511:vermiform
509:and many
453:Concornis
432:Taphonomy
427:Las Hoyas
421:limestone
410:dinosaurs
398:Las Hoyas
390:Barremian
386:formation
299:Huérguina
295:Named for
223:1°53′52″W
184:Limestone
175:Lithology
167:Thickness
149:with the
139:Underlies
38:125
32:Barremian
4485:Archived
4453:: 38–49.
4389:(2004),
3721:See also
3640:Lourinhã
3625:Tera Gp.
3605:Tera Gp.
3529:Alfambra
3467:Peñacoba
3369:Porto da
3360:Villares
3348:Tera Gp.
3297:Mirambel
3259:Tera Gp.
3146:Figueira
3087:Almargem
3014:Benassal
3007:Benassal
2877:Utrillas
2871:Altamira
2843:Utrillas
2375:Species
2177:Species
2064:Eobaatar
2012:Species
1852:H. sanzi
1769:Species
1652:Species
1644:Turtles
1407:Species
1101:Amiopsis
999:Species
972:Hybodont
933:Species
776:crayfish
734:Species
616:and the
604:Research
540:amniotes
519:annelids
507:Chordata
503:molluscs
204:Location
157:Overlies
4081:Bibcode
3899:Bibcode
3852:Bibcode
3790:Bibcode
3710:Sources
3598:Lastres
3540:TdL Gp.
3504:Cab Gp.
3452:Ped Gp.
3441:Ped Gp.
3428:Ped Gp.
3421:Larriba
3410:Villaro
3353:Pinilla
3333:Llacova
3326:Pinilla
3264:Golmayo
3232:Collado
3224:Artoles
3207:Assises
3124:Forcall
3071:Morella
3048:Morella
2948:Escucha
2939:Escucha
2837:Sopeira
2783:Morella
2755:Cameros
2390:Images
2192:Images
2027:Images
2003:Mammals
1995:Mammals
1919:lizard
1917:teiioid
1812:lizard
1784:Images
1667:Images
1637:Turtles
1422:Images
1027:Caturus
1021:Caturus
1014:Images
948:Images
802:shrimp
749:Images
682:Ootaxon
262:Country
180:Primary
107:↓
4420:Nature
4399:
4365:Nature
4342:
4300:
4107:
4099:
4052:
4011:
3970:
3919:
3870:
3818:
3670:Legend
3662:Ágreda
3631:Magaña
3611:Magaña
3576:On Gp.
3560:hiatus
3546:Rupelo
3522:hiatus
3508:Arcera
3416:Ur Gp.
3389:Huerva
3380:hiatus
3371:Calada
3321:Ur Gp.
3271:Alacón
3253:Rúbies
3186:Alacón
3174:Abejar
3118:Jubela
3114:En Gp.
3107:Senyús
3100:Ernaga
3058:Oliete
3053:Oliete
3036:En Gp.
2995:Ol Gp.
2979:Aptian
2965:Itxina
2956:Jijona
2904:Ullaga
2898:Albian
2885:Eguino
2769:Oliete
2748:Olanyà
2722:Iberia
2387:Notes
2372:Genus
2189:Notes
2174:Genus
2024:Notes
2009:Genus
1781:Notes
1766:Genus
1664:Notes
1649:Genus
1419:Notes
1404:Genus
1381:Amiid
1315:S. mia
1011:Notes
996:Genus
945:Notes
930:Genus
746:Notes
731:Genus
612:, the
524:Among
280:Extent
271:
252:Cuenca
248:Region
4105:S2CID
3946:(3).
3868:S2CID
3816:S2CID
3567:Tollo
3475:Galve
3396:Gaita
3282:Blesa
3275:Blesa
3216:Prada
3137:Upper
3080:Rojas
3078:Capas
3021:Olhos
2969:Miono
2915:Lluçà
2776:Galve
2579:bird
2492:bird
2455:bird
1628:Frog
1576:Frog
1206:aff.
1110:cf. "
1096:Amiid
800:atyid
701:Notes
653:Taxon
598:alula
573:birds
495:Phyla
491:fauna
465:flora
436:As a
274:Spain
194:Other
4397:ISBN
4340:ISSN
4298:ISSN
4097:ISSN
4050:ISSN
4009:ISSN
3968:ISSN
3917:ISSN
3192:Xert
3180:Xert
3167:Cabó
3041:Leza
3030:Font
2322:Uña
2297:Uña
2207:Uña
2107:Uña
2074:Uña
2044:Uña
1954:Uña
1931:Uña
1831:Uña
1802:Uña
1737:Uña
1716:Uña
1621:Uña
1523:frog
1489:Uña
1437:Uña
1368:cf.
1264:sp.
1238:sp.
1029:sp.
575:are
530:fish
368:The
188:marl
129:Type
46:PreꞒ
4429:doi
4424:382
4369:467
4330:doi
4290:doi
4286:150
4089:doi
4040:doi
3999:doi
3995:181
3958:hdl
3948:doi
3944:180
3907:doi
3860:doi
3848:110
3806:hdl
3798:doi
3786:110
3304:150
2906:-
2893:125
2812:100
2733:Age
798:An
517:or
483:or
145:or
4500::
4466:35
4451:36
4444:In
4385:;
4381:;
4338:,
4326:57
4324:,
4318:,
4296:,
4284:,
4280:,
4263:30
4261:,
4255:,
4164:^
4144:^
4103:.
4095:.
4087:.
4077:88
4075:.
4071:.
4048:.
4036:36
4034:.
4030:.
4007:.
3993:.
3989:.
3966:.
3956:.
3942:.
3938:.
3915:.
3905:.
3895:56
3893:.
3889:.
3866:.
3858:.
3846:.
3842:.
3828:^
3814:.
3804:.
3796:.
3784:.
3780:.
3701:,
3695:,
3689:,
3681:,
3628:-
3608:-
3578:-
3543:-
3506:-
3455:-
3443:-
3430:-
3418:-
3350:-
3323:-
3261:-
3116:-
3098:-
3038:-
2997:-
2987:-
2967:-
2729:Ma
1691:A
1366:V.
1262:P.
1236:P.
1204:N.
1116:"
825:A
774:A
708:;
521:.
505:,
501:,
497::
477:,
471:,
396:.
376:,
254:,
186:,
96:Pg
40:Ma
30::
4431::
4332::
4292::
4111:.
4091::
4083::
4056:.
4042::
4015:.
4001::
3974:.
3960::
3950::
3923:.
3909::
3901::
3874:.
3862::
3854::
3822:.
3808::
3800::
3792::
3273:/
3051:/
101:N
91:K
86:J
81:T
76:P
71:C
66:D
61:S
56:O
51:Ꞓ
35:~
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