931:
1820:
122:
618:
6760:
1118:
8883:
6753:
882:
1892:
5257:
1321:
6082:
6068:
2061:. The skull was also proportionally larger in the younger specimens, and the sloping of the snout's upper profile was less distinct. The ribs in the neck were fused to the vertebrae only in adults. The forelimbs appear to have become proportionally longer during growth, whereas the proportional length of the bones in the hind limbs changed very little. In 2012, palaeontologist Darla K. Zelenitsky and colleagues concluded that, since adult ornithomimosaurs had wing-like structures on their arms whereas juveniles did not (as evidenced by specimens of
1567:
767:
8335:
2122:
7548:
8321:
2399:
6061:
8328:
1202:
5263:
1306:
2005:("warm" or "cold"-blooded) metabolism, an intake which he found to be unfeasible if it was a filter feeder. He also found that ornithomimids were abundant not only in formations that represented mesic environments, but also in arid environments where there would be insufficient water to sustain a diet based on filter-feeding. In 2007, palaeontologist Espen M. Knutsen wrote that the beak shape of ornithomimids, when compared to those of modern birds, was consistent with omnivory or high-fibre herbivory.
6767:
147:
6075:
8896:
1730:
7566:
1669:
7554:
1043:
1445:
1523:
7573:
8890:
8315:
734:
1357:
1501:
1418:
7561:
2045:
1786:, extending 20 centimetres (7.9 in) below the layer with the tracks. It is possible the fossil represents an animal that died in its tracks, but the depth of the foot in the mud may be too shallow for it to have become mired. It may also have been killed by a flood, after which it was buried in a pond. However, the layers of mud and sand do not indicate flooding but probably a dry environment, and the disrupted
1545:
9018:
4685:
1791:
across the floor of a pond, breaking through the sediment layer with the tracks while it was soaked from rain or contained water. The animal may have died in this position from thirst, hunger, or another reason, and mud then deposited on the sand, thereby covering and preserving the tracks and the carcass. The foot may have become clenched and disarticulated as it decomposed, which made the
1474:
2989:
1194:(hand) was proportionally short compared to those of other ornithomimosaurs, having the smallest manus to humerus length ratio of any member of the group, but was otherwise similar in structure. It had three fingers, which were similarly developed; the first (the "thumb") was the strongest, the third was the weakest and the second was the longest. The
1849:(due to the seemingly loose connection between some of the bones at the back of the skull), a feature which allows individual bones of the skull to move in relation to each other. They also proposed that it did not use its short handed forelimbs for bringing food to the mouth, but for raking or digging in the ground to access food. The hands of
1216:(pubic bone) was long and slender, ending in a pubic boot which expanded to the front and back, a general feature of ornithomimosaurs. The hind limbs differed little from those of other ornithomimids, and were proportionally longer than in other theropods. The femur was nearly straight, long and slender, with a sideways flattened shaft. The
900:(thigh bone) was 660 millimetres (2.17 ft). It would have weighed about 400–490 kilograms (880–1,080 lb). In comparison, one juvenile specimen (ZPAL MgD-I/94) was about 2.15 metres (7.1 ft) long, 0.79 metres (2.6 ft) tall at the hip, and weighed about 26–30.2 kilograms (57–67 lb). Based on fossils of the related
1182:(upper arm bone), which had a near circular cross-section, was long and twisted. The deltopectoral crest on the upper front part of the humerus was comparatively small, and therefore provided little surface for attachment of upper arm muscles. The ulna was slender, long and weakly curved, with a nearly triangular shaft. The
1105:. The neck appears to have been proportionally longer in relation to the trunk than in other ornithomimids. The neck was divided into two distinct sections: the cervical vertebrae at the front had centra which were nearly triangular in side view and tapered towards the back, as well as low neural arches and short, broad
1265:
had a rounded beak (similar to a goose or duck), North
American ornithomimids had pointed beaks, a difference that would otherwise lead taxonomists to place modern birds in different families. In 1976, Barsbold placed Ornithomimidae in the new group Ornithomimosauria. In 2003, Kobayashi and Jun-Chang
1157:
which were about equal in length. The centra here were spool-like, flattened sideways and had fossae which appear to have continued as deep foramina in some specimens. The neural spines here were rectangular, broad, and higher than those in the dorsal vertebrae. They were higher or equal in height to
1033:
rhamphotheca (horny beak) in life. The beak may have covered a smaller area than in North
American relatives, based on the lack of nourishing foramina on the maxilla. The inner side of the beak had small, tightly packed and evenly spaced columnar structures (their exact nature is debated), which were
1660:, which was also in reference to the capsule. In 1998, Holtz instead found that troodontids were basal maniraptorans, meaning that all members of that clade would fall within Bullatosauria, which would therefore become a junior synonym of Maniraptoriformes, and the clade has since fallen out of use.
1145:, with spool-like centra that were short, but tended to become deeper and longer towards the back. Their transverse processes (processes articulating with the ribs) slightly increased in length towards the back. The two first dorsal centra had deep pneumatic foramina, while the rest only had shallow
744:
The other partially complete skeletons were juveniles; ZPAL MgD-I/1 (from
Tsaagan Khushuu) has a crushed skull with a missing tip, damaged vertebrae, fragmented ribs, pectoral girdle and forelimbs, and an incomplete left hind limb, ZPAL MgD-I/94 (from the Nemegt locality) lacks the skull, atlas, tip
524:(air-filled). The neck was proportionally long in relation to the trunk. The hands were proportionally the shortest of any ornithomimosaur and each had three digits with curved claws. The forelimbs were weak while the hindlimbs were proportionally long. The family Ornithomimidae is part of the group
1113:
that articulated between the vertebrae); the cervical vertebrae at the back had spool-like centra which became gradually higher, and long, thin zygapophyses. The pneumatic foramina here were small and oval, and the neural spines projecting outwards from the centra formed long, low and sharp ridges,
665:
in
Tsaagan Khushuu in 1964; it was preserved lying on its back, and the skull was found under its pelvis. One small specimen was also found in Tsaagan Khushuu the same year, and another small specimen was found in the Nemegt locality. A small skeleton without forelimbs was discovered in 1967 by the
1260:
was difficult because other ornithomimids known at the time were either poorly preserved or inadequately described. In 1975, Kielan-Jaworowska stated that though many dinosaurs from Asia were placed in the same families as North
American relatives, this category of classification tended to be more
942:
was very small and light compared to the vertebral column. Due to the length of its snout, the skull was long compared to other ornithomimids, and the snout had a gently convex sloping upper profile. The side profile of the snout differed from other ornithomimids in not narrowing towards its front
2035:
achieving relatively large sizes. Advantages from large body mass in herbivores include increased intake rate of food and fasting resistance, and these trends may therefore indicate that ornithomimids and deinocheirids were more herbivorous than other ornithomimosaurians. They cautioned that the
1836:
similar to that of a duck or goose, and that it would have fed on small, living prey which it swallowed whole. The mobility of the neck would have been useful in locating prey on the ground, since the eyes were positioned on the sides of the skull. They assumed that all ornithomimids had similar
1758:
had the most pneumatised skeleton among ornithomimosaurs. Pneumatisation is thought to be advantageous for flight in modern birds, but its function in non-avian dinosaurs is not known with certainty. It has been proposed that pneumatisation was used to reduce the mass of large bones, that it was
1625:
theropods, which also includes modern birds. Early ornithomimosaurs had teeth, which were lost in more derived members of the group. In 2004, Makovicky, Kobayashi, and Currie suggested that most of the early evolutionary history of ornithomimosaurs took place in Asia, where most genera have been
1790:
around the fossil indicate the animal was alive when it came to the area. The authors thus suggested that the tracks had been made over an extended amount of time and period of drying, and that probably none of them were produced by the individual that owned the foot. The animal may have walked
792:
specimens (as indicated by empty excavation pits, garbage, and scattered broken bones in the quarry). It is unusual to find tracks closely associated with body fossils; some of the tracks are consistent with ornithomimid feet, while others belong to different dinosaurs. In 2014, a slab with two
1745:
would have been more limited, which is an adaptation in some animals that improves their ability to see predators behind them. Paul considered the relatively short tails, which reduced weight, and missing halluxes of ornithomimids to be adaptations for speed. He suggested that they could have
1162:
blade and were separate, whereas in other ornithomimids they were fused together. The tail had 36–39 caudal vertebrae with the centra of those at the front being spool-shaped, while those at the back were nearly triangular, and elongated across. The neural spines here were high and flat, but
729:
youth around the neck, in reference to the bulbous capsule on the parasphenoid at the base of the dinosaur's skull. Such a feature had not been described from other reptiles at the time, and was considered unusual. The holotype consists of an almost complete skeleton with a distorted snout,
745:
of the tail, pectoral girdle and forelimbs, while the smallest specimen, IGM 100/10 (from Bugeen Tsav), lacks a pectoral girdle, forelimbs and several vertebrae and ribs. Osmólska and colleagues listed twenty-five known specimens in all, nine of which were only represented by single bones.
2008:
David J. Button and Zanno found in 2019 herbivorous dinosaurs mainly followed two distinct modes of feeding, either processing food in the gut—characterized by gracile skulls and low bite forces—or the mouth, characterized by features associated with extensive processing. Ornithomimid
3571:
2708:
Kielan-Jaworowska, Z. (1975). "Late
Cretaceous mammals and dinosaurs from the Gobi Desert: Fossils excavated by the Polish-Mongolian paleontological expeditions of 1963–71 cast new light on primitive mammals and dinosaurs and on faunal interchange between Asia and North America".
2092:
specimens. They found that the ratio was highest in young to middle aged animals, decreasing with age. In 1991, they reported that the bones of old individuals contained the highest amounts of lead and iron, while those in younger animals were lower. A study of the bone
992:
The delicate lower jaw, consisting of thin bones, was slender and shallow at the front, deepening towards the rear. The front of the mandible was shovel-like, resulting in a gap between the tips of the jaws when shut. The shovel-like shape was similar to that of the
2460:
herd. The production team became very enthusiastic as nothing similar had previously been achieved, and
Spielberg was convinced to write the scene into the script, and to also use computer graphics for other dinosaur shots in the movie instead of stop motion. The
1992:
stones) in some ornithomimids indicate the presence of a gastric mill, and therefore point towards a herbivorous diet, as these are used to grind food of animals that lack the necessary chewing apparatus. Barrett also calculated that a 440 kilograms (970 lb)
670:. The Polish-Mongolian expeditions were notable for being led by women, some of whom were among the first women to name new dinosaurs. The fossils discovered in these expeditions shed new light on the interchange of fauna between Asia and North America during the
1975:
did not appear to have been flexible bristles like those of filter-feeding birds (as there is no indication of these structures overlapping or being collapsed), but were instead more akin to the thin, regularly spaced vertical ridges in the beaks of turtles and
1876:, which were strong enough to bite off twigs, as evidenced by their gut content. He further suggested that ornithomimids were well adapted for browsing on tough plants and would have used their hands to bring branches within reach of their jaws. Palaeontologist
656:
at different growth stages from the Nemegt, Tsaagan
Khushuu, Altan Ula IV and Naran Bulak localities. Three partially complete skeletons, two with skulls, as well as many fragmentary remains, were collected. The largest skeleton (later to become the holotype of
1101:(the first vertebra that connects with the back of the skull). The atlas differed from that of other ornithomimids in that the front surface of its intercentrum was slanted downwards towards the back, instead of being concave and facing upwards to support the
2531:; these tracks were initially interpreted as representing a dinosaur stampede caused by the arrival of a theropod predator. The idea that the tracks represent a stampede has since been contested (the "theropod" may instead have been a herbivore similar to
1740:
In 1988, Paul suggested that the eyeballs of ornithomimids were flattened and had minimal mobility within the sockets, necessitating movement of the head to view objects. Since their eyes faced more sideways than in some other bird-like theropods, their
1163:
diminished backwards, where they became ridge-like. The only sign of pneumaticity in the tail were deep fossae between the neural spines and the transverse process of the two first caudal vertebrae. All the vertebrae in front of the sacrum bore
1725:
could have run at speeds of 42–56 km/h (29–34 mph). He found that ornithimimids would not have been as fast as ostriches, which can reach 70–80 km/h (43–49 mph), in part due to their arms and tails increasing their weight.
3583:
2500:. This contrasted with how dinosaurs were traditionally depicted in mass media as lumbering, tail-dragging animals, and the movie helped change the common perception of dinosaurs. This and other scenes reflected then-recent theories of
1705:) animals, although less agile than large, modern ground birds, and would have used their speed to escape predators. Russell also suggested that they had a good sense of vision and intelligence comparable to that of modern
1198:(claw bones) were strong, somewhat curved (that of the first finger was most curved) and compressed sideways with a deep groove on each side. The unguals were similarly developed, though the third was slightly smaller.
997:, and the lower beak may have had a shape similar to that of this bird. The retroarticular process at the back of the jaw (where jaw muscles attached that opened the beak) was well developed and consisted mainly of the
1831:
would have been very mobile (the hind part was more rigid), the neural arches in the vertebrae of that region being similar to chicken and other
Galliformes, indicating similar feeding habits. They found the beak of
752:
represented the most complete and best preserved ornithomimid material yet discovered, and the genus remains one of the best known members of the group. Ornithomimids were previously known mainly from North
America,
2910:
Fanti, F.; Bell, P. R.; Tighe, M.; Milan, L. A.; Dinelli, E. (2018). "Geochemical fingerprinting as a tool for repatriating poached dinosaur fossils in Mongolia: A case study for the Nemegt Locality, Gobi Desert".
2036:
correlations between herbivory and body mass were not simple, and that there was no directional trend towards increased mass seen in the clade. Furthermore, the diet of most ornithomimosaurians is poorly known.
2056:
The shape and proportions of the skull changed significantly during growth. The rear of the skull and the orbits decreased in size, whereas the snout became relatively longer; similar changes occur in modern
3677:
Naish, D.; Martill, D. M.; Frey, E. (2004). "Ecology, Systematics and biogeographical relationships of dinosaurs, including a new theropod, from the Santana Formation (?Albian, Early Cretaceous) of Brazil".
1640:
to account for the Late Cretaceous genera found there. As seen in some other dinosaur groups, ornithomimosaurs were largely restricted to Asia and North America after Europe was separated from Asia by the
977:(a bone of the braincase, at the underside of the skull's base) was thin-walled, hollow and formed a pear-shaped, bulbous structure. The structure had a shallow furrow which opened towards the front. The
3245:
Zelenitsky, D. K.; Therrien, F.; Erickson, G. M.; DeBuhr, C. L.; Kobayashi, Y.; Eberth, D. A.; Hadfield, F. (2012). "Feathered non-avian dinosaurs from North America provide insight into wing origins".
1963:
environments, while rarer in more arid environments, suggesting that they may have depended on aquatic food sources. Makovicky, Kobayashi, and Currie pointed out that if this interpretation is correct,
508:. The head was small and light with large eyes that faced to the sides. The snout was long compared to other ornithomimids, although it was broader and more rounded at the tip than in other species.
896:
Ornithomimidae. The adult holotype (specimen IGM 100/11) was about 6 metres (20 ft) long and 1.9 metres (6.2 ft) tall at the hip; its skull was 330 millimetres (1.08 ft) long and the
759:
being the only prior known member from Asia (though without a skull). Since the first discoveries, more specimens have been found by further Mongolian-led international expeditions. Three of the
1803:
mass mortality, perhaps due to a drought or famine. The fact that the animals seem to have died at the same time (the empty excavation pits were stratigraphically identical) may indicate that
4462:
2870:
2469:
was also referenced. Kielan-Jaworowska, who discovered the holotype specimen, called it a "beautiful scene". The movie's dinosaurs were one of the most widely publicised applications of
498:; adults were about 6 metres (20 ft) long, 1.9 metres (6 ft 3 in) tall at the hip and weighed about 400–490 kilograms (880–1,080 lb). As evidenced by its relative
677:. Some of the skeletons were exhibited in Warsaw in 1968, mounted in tall, semi-erect postures, which was accepted at the time, though more horizontal postures are favoured today.
869:
specimens, though this has never been published. Barsbold informally referred to a nearly complete skeleton (IGM 100/14) as "Gallimimus mongoliensis", but since it differs from
1232:
was broad when viewed end on, partly covering the adjoining two metatarsals to each side, but narrowed abruptly at mid-length, wedging between those bones and disappearing (an
1980:
dinosaurs. In these animals, such ridges are thought to be associated with herbivorous diets, used to crop tough vegetation. Barrett suggested that the ridges in the beak of
1073:, fewer than other ornithomimids. The centra (or bodies) of the vertebrae were platycoelous, with a flat front surface and a concave hind surface, except for the first six
549:) animal, using its speed to escape predators; its speed has been estimated at 42–56 km/h (29–34 mph). It may have had good vision and intelligence comparable to
2488:
s story, as they were supposed to represent the precursors to birds. The herd was shown moving as a whole, rather than individual animals running around, and the smaller
947:
at the front of the upper jaw rose upwards, instead of being horizontal. Seen from above, the snout was almost spatulate (spoon-shaped), broad and rounded at the tip (or
1240:(or dewclaw, the first toe of most other theropods). The unguals of the toes were flat on their lower sides; the outer two declined slightly outwards from their digits.
1178:, which was large and deep from top to bottom. Overall, the forelimbs did not differ much from those of other ornithomimids, all of which were comparatively weak. The
1746:
defended themselves by pecking and kicking, but would have mainly relied on their speed for escape. In 2015, Akinobu Watanabe and colleagues found that together with
1984:
represented a natural cast of the internal surface of the beak, indicating that the animal was a herbivore that fed on material high in fibre. The discovery of many
1656:
in a clade, based on shared features such as the presence of a bulbous capsule on the parasphenoid. He named the clade Bullatosauria, based on the specific name of
1691:(or capable of grasping), and that the thumb was not opposable. They also suggested that the arms were weak compared to, for example, those of the ornithomimosaur
1857:, which may instead have used its hands for hooking and gripping, according to a 1985 article by palaeontologists Elizabeth L. Nicholls and Anthony P. Russell.
1077:(tail) vertebra—where the hind surface was also flat—and those at the end of the tail—which were amphiplatyan with both surfaces flat. Many of the centra had
2496:
says that the herd moves with "uniform direction changes, just like a flock of birds evading a predator" as he watches the movements of the fast, graceful
1236:
foot structure). The third toe was proportionally shorter in relation to the limb than in other ornithomimids. As in other ornithomimids, the foot had no
3527:
McFeeters, B.; Ryan, M.J.; Schröder-Adams, C.; Cullen, T.M. (2016). "A new ornithomimid theropod from the Dinosaur Park Formation of Alberta, Canada".
2512:
and other dinosaurs of the movie were depicted without feathers, in part because it was unknown at the time how widespread these were among the group.
1312:
1206:
696:, using the largest collected skeleton, specimen IGM 100/11 (from Tsaagan Khushuu, formerly referred to as G.I.No.DPS 100/11 and MPD 100/11), as the
2949:
Lee, H.-J.; Lee, Y.-N.; Adams, T. L.; Currie, P. J.; Kobayashi, Y.; Jacobs, L. L.; Koppelhus, E. B. (2018). "Theropod trackways associated with a
2885:
804:
concluded that the skulls of ornithomimids were more similar to each other than previously thought and moved most species into the same genus,
763:
skeletons (including the holotype) later became part of a travelling exhibit of Mongolian dinosaur fossils, which toured various countries.
2851:
565:
and other ornithomimids. The highly mobile neck may have helped locate small prey on the ground, but it may also have been an opportunistic
3894:
Pawlicki, R.; Bolechała, P. (1987). "X-ray microanalysis of fossil dinosaur bone: age differences in the calcium and phosphorus content of
3464:
Watanabe, A.; Eugenia Leone Gold, M.; Brusatte, S. L.; Benson, R. B. J.; Choiniere, J.; Davidson, A.; Norell, M. A.; Claessens, L. (2015).
784:
specimens have been looted. In 2017, Hang-Jae Lee and colleagues reported a fossil trackway discovered in 2009 associated with a clenched
3925:
Pawlicki, R.; Bolechała, P. (1991). "X-ray microanalysis of fossil dinosaur bone: age differences in lead, iron, and magnesium content".
9040:
1864:
that ate small animals and eggs as well as plants, as had previously been suggested. He pointed out that ostriches and emus are mainly
1683:
indicate that it held its neck obliquely, declining upwards at an angle of 35 degrees. Osmólska and colleagues found that the hands of
948:
9065:
788:
foot (specimen MPC-D100F/17). The rest of the skeleton appeared to have been removed previously by poachers, along with several other
2113:
dinosaurs, which were more similar to mammals. These differences may have been related to the process and rate at which bone formed.
666:
Mongolian palaeontological expedition in Bugeen Tsav outside the Nemegt Basin. The fossils were housed at the Mongolian, Polish and
8975:
3129:
Makovicky, P. J.; Kobayashi, Y.; Currie, P. J. (2004). "Ornithomimosauria". In Weishampel, D. B.; Dodson, P.; Osmolska, H. (eds.).
1636:
in Europe presents a complication in classification. The group must have dispersed once or twice from Asia to North America across
9070:
2430:
sequence with animal herds in the movie, but did not know how to achieve it, and it was initially going to be visualised through
1174:(shoulder blade) was short and curved, thin at the front end, and thick at the back. It was connected relatively weakly with the
930:
831:
in 2016. The species involved have generally been kept in separate genera by other writers. An ornithomimid vertebra from Japan
9045:
4358:
4327:
4299:
4170:
4086:
3146:
3102:
3069:
3041:
2798:
2762:
2687:
861:, but this was dismissed by Kobayashi and Barsbold, who pointed out several differences between the two. Barsbold noted some
771:
4229:
Efimov, M. B. (1983). "Peresmotr iskopayemykh krokodilov Mongolii" [Revision of the fossil crocodiles of Mongolia].
6759:
6752:
4740:
3796:
3470:(Dinosauria: Theropoda) revealed by computed tomography imaging and reappraisal of axial pneumaticity in ornithomimosauria"
2550:
8882:
4040:
2194:. Large river channels and soil deposits are evidence of a significantly more humid climate than those found in the older
1997:
would have needed between 0.07 and 3.34 kilograms (0.15 and 7.36 lb) of food per day, depending on whether it had an
4719:
573:
in water, though these structures may instead have been ridges used for feeding on tough plant material, indicative of a
1228:
of the lower leg was flat, thin and broad at the upper end narrowing towards the lower end. The lower half of the third
4607:
4512:
3389:
1734:
1959:. The authors noted that ornithomimids probably did not use their beaks to prey on large animals and were abundant in
2027:, were found to be in the former category. These researchers suggested that ornithomimid ornithomimosaurians such as
402:
4572:
2493:
1010:
1009:, a sidewards-facing opening in the lower jaw, was elongated and comparatively small. The lower jaw did not have a
873:
in some details, Yoshitsugu Kobayashi and Barsbold proposed in 2006 that it probably belongs to a different genus.
7547:
2492:
were shown in the middle of the group, as though they were being protected. During the scene, the palaeontologist
6081:
1190:(the other bone in the lower arm) was long and slender with a more expanded upper end compared to the lower. The
797:
specimens was repatriated to Mongolia along with other dinosaur skeletons, after having been smuggled to the US.
6067:
3605:
Holtz, T. R. (1994). "The Phylogenetic Position of the Tyrannosauridae: Implications for Theropod Systematics".
2080:
A 1987 study by the biologists Roman Pawlicki and P. Bolechała showed age-related differences in the content of
8320:
4624:
3165:
Chinzorig, Tsogtbaatar; Cullen, Thomas; Phillips, George; Rolke, Richard; Zanno, Lindsay E. (19 October 2022).
2516:
134:
3209:
8761:
6060:
5256:
3210:"Strength indicator values of theropod long bones, with comments on limb proportions and cursorial potential"
1819:
641:
121:
7565:
5262:
8334:
4282:
Holtz, T. R. (2004). "Tyrannosauroidea". In Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (eds.).
2505:
2267:, a possible anseriform, have been found. Herbivorous dinosaurs discovered in the Nemegt Formation include
2066:
1149:(depressions), and the neural spines were prominent being somewhat triangular or rectangular in shape. The
6766:
737:
Three skeletons, including the holotype (right) and a juvenile (middle), during a temporary exhibition in
440:(mya). Several fossils in various stages of growth were discovered by Polish-Mongolian expeditions in the
9055:
4689:
2826:
667:
617:
6074:
9008:
8327:
2470:
2443:
1117:
637:
146:
7572:
488:
dinosaur") material yet discovered, and the genus remains one of the best known members of the group.
8895:
2598:
1932:
1900:
1013:
or a supradentary bone, the lack of which is a common feature of beaked theropods (ornithomimosaurs,
3694:
3654:
2206:
deposits indicate that periodic droughts occurred. Fossil bones from the Nemegt Basin, including of
2097:
of various dinosaurs in 2000, by biologists John M. Rensberger and Mahito Watabe, revealed that the
1943:
by segregating food items from other material, and for cutting plants while grazing. They found the
4807:
918:, and that the adults bore wing-like structures as evidenced by the presence of quill-knobs on the
1837:
feeding habits, and pointed out that Russel had compared the beaks of ornithomimids with those of
1186:(the projection from the elbow) was prominent in adults, but not well developed in juveniles. The
8889:
7553:
4712:
4070:
2855:
2653:
2167:
1297:
among Ornithomimidae according to the palaeontontologist Bradley McFeeters and colleagues, 2016:
716:
662:
3952:
Rensberger, J. M.; Watabe, M. (2000). "Fine structure of bone in dinosaurs, birds and mammals".
3167:"Large-bodied ornithomimosaurs inhabited Appalachia during the Late Cretaceous of North America"
3094:
963:(the lower opening behind the orbit) was nearly triangular and smaller than that of the related
9050:
8993:
8915:
4504:
4291:
3689:
3649:
3381:
3369:
3138:
2790:
960:
4350:
4344:
4249:
3033:
3026:
2450:
skeleton in the computer and animated a test showing a herd of running skeletons, and later a
1884:
may have had an opportunistic, omnivorous diet like seagulls. He also observed that the tight
730:
incomplete lower jaw, vertebral series, pelvis, as well as some missing hand and foot bones.
9060:
8988:
7278:
7190:
2431:
2423:
2195:
1627:
1121:
1086:
711:, "mimic", in reference to the front part of the neck vertebrae which resembled those of the
521:
293:
4492:
4283:
4062:
4050:. Vol. 35. New Mexico Museum of Natural History and Science Bulletin. pp. 347–366.
3130:
2782:
2628:
2182:
of the Nemegt Formation suggest the presence of river channels, mudflats, shallow lakes and
8962:
8953:
8847:
6967:
6935:
4195:
4111:
3961:
3769:
3734:
3536:
3481:
3417:
3255:
2962:
2920:
2718:
2473:
in film, and were considered more lifelike than what had been previously accomplished with
2414:
2144:
2032:
1869:
862:
601:
317:
8314:
4162:
1320:
881:
8:
7291:
7078:
3408:
Norell, M. A.; Makovicky, P. J.; Currie, P. J. (2001). "The beaks of ostrich dinosaurs".
2654:"Geographical and geological data from the Polish-Mongolian palaeontological expeditions"
2199:
1998:
1718:
1106:
1050:
915:
505:
7560:
4199:
4115:
3965:
3773:
3738:
3540:
3485:
3421:
3259:
2966:
2924:
2722:
1637:
681:
266:
8754:
7267:
7127:
6983:
6923:
6358:
6268:
6151:
5015:
4705:
4553:
4545:
4440:
4395:
4211:
4186:
Jaworowski, Z.; Pensko, J. (1967). "Unusually radioactive fossil bones from Mongolia".
4135:
4063:
3985:
3876:
3824:"Beak morphology in extant birds with implications on beak morphology in ornithomimids"
3725:
Russell, D. A. (1972). "Ostrich dinosaurs from the Late Cretaceous of Western Canada".
3707:
3680:
3622:
3552:
3504:
3465:
3441:
3279:
3193:
3166:
3087:
2726:
2501:
2435:
2306:
2278:
2148:
2109:
and other ornithomimids were more akin to those in birds than mammals, unlike those of
1891:
1566:
1384:
1351:
1130:(A), and the cervical (B, C, D), dorsal (E), sacral (F, G) and caudal (H) vertebrae of
1094:
854:
755:
325:
141:
4528:
Kirby, D. A. (2003). "Science consultants, fictional films, and scientific practice".
3304:"A new ornithomimid dinosaur with gregarious habits from the Late Cretaceous of China"
2446:
was given a go-ahead by the movie's producers to explore possibilities. ILM created a
1971:
In 2005, palaeontologist Paul Barrett pointed out that the lamella-like structures of
770:
Specimens which were repatriated to Mongolia after having been smuggled to the US, in
444:
of Mongolia during the 1960s; a large skeleton discovered in this region was made the
8636:
8553:
8527:
8366:
8353:
8231:
7781:
7733:
6565:
5915:
4994:
4636:
4603:
4557:
4508:
4493:
4413:
Prince, S. (1996). "True lies: Perceptual realism, digital images, and film theory".
4354:
4323:
4295:
4284:
4166:
4139:
4127:
4082:
4025:
4009:
3977:
3934:
3907:
3880:
3868:
3781:
3659:
3509:
3433:
3385:
3346:
3329:
3303:
3271:
3224:
3142:
3131:
3098:
3065:
3037:
2794:
2783:
2758:
2683:
2211:
2074:
2070:
1944:
1904:
1778:
foot discovered in 2009 was associated with a trackway. The trackway is preserved in
1673:
1618:
1221:
1110:
1102:
777:
557:
may have lived in groups, based on the discovery of several specimens preserved in a
525:
233:
3711:
3556:
2121:
766:
569:. It has also been suggested that it used small columnar structures in its beak for
9022:
8740:
8497:
8407:
8196:
8120:
7983:
7902:
7874:
7834:
7715:
7685:
7203:
6956:
6820:
6677:
6577:
6426:
6333:
6291:
5858:
5168:
4537:
4430:
4422:
4387:
4378:
dinosaurs: Blockbuster schemata and cross-cultural cognition in the threat scene".
4215:
4203:
4119:
4074:
4021:
3989:
3969:
3858:
3777:
3742:
3699:
3614:
3544:
3499:
3489:
3445:
3425:
3341:
3283:
3263:
3188:
3178:
2970:
2928:
2533:
2419:
2350:
2315:
2140:
2130:
2098:
1795:
flex, and was later stepped on by heavy dinosaurs. The area may have been a single
1764:
1742:
1286:
with North American genera, which was confirmed by Kobayashi and Barsbold in 2006.
1229:
1070:
1046:
1014:
986:
956:
907:
893:
844:
832:
685:
674:
582:
433:
361:
3548:
2757:. Life of the Past. Bloomington: Indiana University Press. pp. 47–48, 50–51.
2465:
were animated by tracing frames from footage of ostriches, and footage of herding
2398:
1955:, to be the modern anseriform with structures most similar in anatomy to those of
1697:. They agreed with the interpretations of ornithomimid biology by palaeontologist
458:
mimic", referring to the similarities between its neck vertebrae and those of the
8803:
8541:
8504:
8436:
8291:
8284:
8104:
8046:
8019:
8009:
7969:
7804:
7663:
7647:
7465:
7437:
7415:
7302:
7147:
6854:
6628:
6511:
6489:
6410:
6239:
6216:
6209:
6187:
6173:
6166:
6127:
5978:
5825:
5752:
5741:
5677:
5565:
5452:
5029:
4938:
4792:
4767:
4155:
3703:
3494:
3183:
2974:
2932:
2356:
2230:
2160:
2155:
present in the fossil record indicate it was probably deposited during the early
1888:
would prevent any movement between the front and rear portions of the lower jaw.
1872:, and that the skulls of ornithomimids were most similar to those of the extinct
1846:
1649:
1630:(or "primitive") taxa, although they acknowledged that the presence of the basal
1412:
1233:
1191:
1154:
1146:
1142:
1098:
1074:
952:
809:
801:
430:
341:
30:
1713:, he suggested it would have influenced their colouration, perhaps resulting in
1201:
1082:
981:(internal openings for the nasal passage) were large and placed far back on the
8768:
8731:
8712:
8674:
8663:
8460:
8173:
8131:
8113:
8093:
7990:
7704:
7449:
7311:
7138:
7120:
6948:
6915:
6901:
6862:
6636:
6497:
6454:
6447:
6369:
6345:
6298:
6257:
6194:
6180:
5908:
5893:
5720:
5554:
5418:
5386:
5375:
5352:
5314:
5175:
5145:
5069:
5008:
4907:
2474:
2439:
2268:
2253:
2244:
2229:
fossils, as well as a variety of other aquatic animals like fish, turtles, and
2187:
2178:, mudstones, sands, as well as less frequent thin beds of sandstones. The rock
1927:
skull that preserved soft tissue structures on the beak. The inner side of the
1337:
1305:
1126:
994:
978:
922:
bone of the lower arm, bumps that indicate where feathers would have attached.
721:
606:
495:
464:
246:
4541:
3863:
3846:
3618:
9034:
8938:
8810:
8656:
8617:
8596:
8475:
8468:
8395:
8189:
8162:
8028:
7945:
7923:
7916:
7909:
7852:
7769:
7632:
7604:
7591:
7515:
7392:
7232:
7066:
6997:
6883:
6870:
6615:
6593:
6586:
6550:
6523:
6475:
6399:
6135:
5926:
5900:
5816:
5797:
5790:
5727:
5706:
5685:
5670:
5656:
5615:
5581:
5574:
5493:
5345:
5232:
5225:
5152:
5122:
5111:
5001:
4972:
4965:
4891:
4655:
4640:
4078:
3842:
3663:
3228:
2452:
2300:
2156:
2110:
2024:
1960:
1940:
1632:
1622:
1494:
1187:
1018:
970:
965:
653:
570:
72:
34:
3823:
3267:
1877:
1729:
955:(eye sockets) were large and faced sideways, as in other ornithomimids. The
8782:
8720:
8690:
8610:
8603:
8581:
8573:
8453:
8445:
8425:
8140:
8059:
7997:
7888:
7859:
7811:
7640:
7625:
7400:
7332:
7260:
7239:
7218:
7034:
6990:
6798:
6785:
6715:
6668:
6543:
6433:
6385:
6378:
6315:
6284:
6277:
6158:
5985:
5837:
5734:
5636:
5629:
5524:
5508:
5444:
5395:
5327:
5104:
4987:
4980:
4951:
4914:
4883:
4131:
4102:
Holtz, T. R. (2014). "Paleontology: Mystery of the horrible hands solved".
4041:"A taxonomic review of the Pachycephalosauridae (Dinosauria: Ornithischia)"
3981:
3872:
3847:"Repeated evolution of divergent modes of herbivory in non-avian dinosaurs"
3513:
3437:
3275:
2282:
1952:
1936:
1710:
1698:
1693:
1561:
1279:
1271:
1213:
1159:
1055:
998:
974:
951:), whereas it was acute (or V-shaped) in North American ornithomimids. The
902:
840:
726:
649:
591:
500:
481:
469:
335:
3938:
3911:
2629:"Narrative of the Polish-Mongolian palaeontological expeditions 1963–1965"
1811:(lived in groups), which has also been suggested for other ornithomimids.
1774:
circumstances (changes during decay and fossilisation) to explain how the
1668:
1326:
Reconstructed cast of the holotype skeleton in a semi-erect posture, MEPAS
1256:
as the closest relative, while lamenting the fact that comparison between
780:
has become a serious problem in Mongolia in the 21st century, and several
609:, and in shaping the common conception of dinosaurs as bird-like animals.
8947:
8836:
8817:
8747:
8697:
8683:
8625:
8566:
8511:
8388:
8182:
8066:
8035:
7952:
7938:
7670:
7618:
7486:
7408:
7371:
7253:
7225:
7178:
7098:
6976:
6833:
6684:
6657:
6557:
6504:
6440:
6419:
6392:
6143:
5956:
5949:
5935:
5761:
5608:
5599:
5501:
5472:
5402:
5367:
5360:
5160:
5138:
5131:
5090:
5082:
5058:
5043:
5022:
4930:
4922:
4231:
Sovmestnaya Sovetsko-Mongol'skaya Paleontologicheskaya Ekspeditsiya Trudy
2523:, Australia, served as inspiration and "scientific underpinning" for the
2508:, ideas which were still contentious at the time. Despite such theories,
2368:
2344:
2320:
2288:
2235:
2218:
accumulated in the bones, transported there by percolating ground water.
2019:
2014:
1985:
1977:
1916:
1908:
1838:
1701:
from earlier in 1972, including that they would have been very fleet (or
1653:
1539:
1444:
1042:
969:. It had deep muscle scars at the back part of the skull roof, along the
849:
712:
645:
530:
459:
441:
47:
4123:
3463:
2730:
2243:
fossils are rare in the Nemegt Formation, but many birds, including the
1522:
1224:(rounded end of a bone) on the upper end and a flattened lower end. The
1005:
was the largest bone of the lower jaw, which is usual in theropods. The
8980:
8704:
8649:
8270:
8077:
7895:
7881:
7866:
7818:
7752:
7678:
7500:
7493:
7479:
7385:
7318:
7246:
7210:
7163:
7105:
7091:
7058:
7050:
7043:
7024:
6847:
6812:
6600:
6201:
5992:
5942:
5879:
5850:
5805:
5783:
5776:
5768:
5692:
5588:
5547:
5540:
5532:
5516:
5436:
5307:
5183:
5036:
4899:
4758:
4600:
Dinosaurs Without Bones: Dinosaur Lives Revealed by their Trace Fossils
4549:
4444:
4435:
4399:
3626:
2603:
n. gen., n. sp. (Ornithomimidae) from the Upper Cretaceous of Mongolia"
2520:
2362:
2332:
2294:
2257:
2183:
2085:
1827:
Osmólska and colleagues pointed out that the front part of the neck of
1760:
1714:
1688:
1642:
1026:
1006:
1002:
944:
738:
671:
648:
of Mongolia. Among the dinosaur remains discovered in sand beds of the
621:
513:
472:
youth, in reference to a bulbous structure at the base of the skull of
437:
204:
92:
57:
2354:. Herbivorous or omnivorous theropods include therizinosaurs, such as
534:, also from Mongolia, is thought to have been the closest relative of
8796:
8789:
8489:
8482:
8155:
8147:
8086:
7740:
7458:
7004:
6908:
6643:
6228:
6113:
6100:
6027:
5963:
5886:
5865:
5622:
5485:
5293:
5280:
5198:
5050:
4869:
4856:
4819:
4728:
4500:
4207:
3973:
3640:
Holtz, T. R. (1998). "A new phylogeny of the carnivorous dinosaurs".
3429:
2374:
2338:
2326:
2263:
2248:
2175:
2094:
2058:
2002:
1968:
would have been one of the largest known terrestrial filter feeders.
1808:
1779:
1771:
1702:
1517:
1468:
1356:
1311:
Adult holotype skeleton IGM 100/11, during a temporary exhibition in
1290:
1261:
inclusive than was used for modern birds. She highlighted that while
1183:
733:
574:
546:
419:
217:
158:
97:
41:
8909:
4426:
4391:
3746:
2871:"Dinosaurer, æg og unger – en særudstilling om en forsvunden verden"
835:"Sanchusaurus" was reported in a 1988 magazine, but was assigned to
8932:
8380:
7797:
7790:
7762:
7655:
7155:
6536:
5872:
5699:
5460:
5097:
4776:
4752:
2427:
2226:
2191:
2102:
1948:
1939:
birds, which use these for manipulating food, straining sediments,
1931:
beak had columnar structures that the authors found similar to the
1885:
1861:
1799:(based on the possible number of poached specimens) representing a
1796:
1787:
1783:
1500:
1175:
1090:
1078:
1066:
697:
644:
organised the Polish-Mongolian palaeontological expeditions to the
625:
566:
558:
517:
445:
426:
422:
191:
178:
130:
87:
82:
67:
62:
52:
3572:"An overview on non-avian theropod discoveries and classification"
3064:(2 ed.). Princeton: Princeton University Press. p. 131.
2519:(one of the world's largest concentrations of dinosaur tracks) in
2504:
encouraged by the movie's scientific advisor, the palaeontologist
2434:. At the time, there was little faith in creating animals through
1417:
7930:
7692:
7422:
7325:
6650:
5971:
4250:"Giant theropod dinosaurs from Asia and North America: Skulls of
3526:
3137:(2 ed.). Berkeley: University of California Press. pp.
2466:
2405:
was formerly depicted without feathers, like this model in MEPAS.
2273:
2215:
2203:
2081:
2044:
1989:
1865:
1544:
1179:
1171:
1030:
911:
485:
455:
102:
77:
8967:
4290:(2 ed.). Berkeley: University of California Press. p.
3244:
2069:, which could have been used for reproductive behaviour such as
1021:
and birds), but unusual among theropods in general. The jaws of
8589:
8277:
5713:
4746:
4684:
3797:"Structure and function of the pectoral girdle and forelimb of
2240:
2179:
1792:
1706:
1237:
1225:
1195:
1150:
982:
550:
168:
4010:"Late Mesozoic stratigraphy and vertebrates of the Gobi Basin"
3093:(1 ed.). Princeton: Princeton University Press. pp.
1252:
to the family Ornithomimidae in 1972, with the North American
7429:
7170:
7112:
3330:"The diet of ostrich dinosaurs (Theropoda: Ornihomimosauria)"
2152:
1283:
1257:
1217:
1153:(fused vertebrae between the pelvic bones) consisted of five
1029:(toothless), and the front part would have been covered in a
897:
853:, which is only known from one skeleton from Mongolia, was a
701:
689:
415:
4697:
3164:
2820:
2818:
2816:
2814:
2812:
2810:
1473:
561:. Various theories have been proposed regarding the diet of
8853:
7472:
3376:. In Tanke, D. H.; Carpenter, K.; Skrepnick, M. W. (eds.).
2651:
2541:
has denied the tracks served as inspiration for the movie.
2171:
2052:, the juvenile specimen IGM 100/10, exhibited in CosmoCaixa
959:
at the side of the skull behind the eyes was deep, and the
919:
391:
385:
4003:
4001:
3999:
2225:
lived, were home to a wide array of organisms. Occasional
3760:
Thulborn, R. A. (1982). "Speeds and gaits of dinosaurs".
2807:
2596:
1880:
suggested in 2001 that due to its similar jaw structure,
1873:
1164:
376:
370:
367:
3128:
2626:
516:(horny) beak, and had a delicate lower jaw. Many of the
4495:
Lab Coats in Hollywood: Science, Scientists, and Cinema
3996:
2031:
and deinocheirids had invaded these niches separately,
1770:
In 2017, Lee and colleagues suggested various possible
3569:
2390:
is the most common member of the group in the Nemegt.
2214:
than fossils from surrounding areas, possibly because
2129:(right foreground) with contemporary dinosaurs of the
1034:
longest at the front and shortening towards the back.
9006:
4048:
Late Cretaceous vertebrates from the Western Interior
3407:
2827:"Ornithomimids from the Nemegt Formation of Mongolia"
1899:
contained structures which have been compared to the
843:
and colleagues in 1990. In 2000, the palaeontologist
403:
394:
364:
4046:. In Lucas, Spencer G.; Sullivan, Robert M. (eds.).
3828:
Det Matematisk-Naturvitenskapelige Fakultet – Thesis
2953:
foot skeleton from the Nemegt Formation, Mongolia".
2682:. Princeton: Princeton University Press. p. 9.
2652:
Gradzihski, R.; Kazmierczak, J.; Lefeld, J. (1969).
2620:
2480:
Emphasising the bird-like flocking behaviour of the
2426:
instead featured hadrosaurs. Spielberg had wanted a
373:
8262:
4061:Gradstein, F. M.; Ogg, J. G.; Smith, A. G. (2005).
4007:
2909:
2592:
2590:
2588:
2586:
2088:(important components in the formation of bone) of
1167:except for the atlas and the last dorsal vertebra.
388:
382:
379:
4322:. New York: Boxtree Ltd. pp. 48–52, 135–136.
4154:
4069:. Cambridge: Cambridge University Press. pp.
3834:
3570:Hendrickx, C.; Hartman, S. A.; Mateus, O. (2015).
3380:. Bloomington: Indiana University Press. pp.
3297:
3295:
3293:
3086:
3025:
2597:Osmolska, H.; Roniewicz, E.; Barsbold, R. (1972).
2584:
2582:
2580:
2578:
2576:
2574:
2572:
2570:
2568:
2566:
1733:Skull cast of the juvenile specimen ZPAL MgD-I/1,
839:sp. (of uncertain species) by the palaeontologist
605:, in a scene that was important to the history of
581:is the most commonly found ornithomimosaur in the
16:Genus of dinosaurs from the Late Cretaceous Period
4060:
4054:
3924:
3893:
3762:Palaeogeography, Palaeoclimatology, Palaeoecology
3403:
3401:
3124:
3122:
3120:
3118:
3116:
3114:
2990:"Stolen 'Nest of Dinosaurs' Returned to Mongolia"
2955:Palaeogeography, Palaeoclimatology, Palaeoecology
2948:
2913:Palaeogeography, Palaeoclimatology, Palaeoecology
2824:
2752:
2707:
2163:about 70 million years ago. The sediments of the
1207:Museum of Evolution of Polish Academy of Sciences
480:represented the most complete and best preserved
9032:
8364:
8230:
4185:
3951:
3676:
7713:
4241:
3794:
3466:"Vertebral pneumaticity in the ornithomimosaur
3459:
3457:
3455:
3290:
2834:Journal of the Paleontological Society of Korea
2563:
1860:In 1988 Paul disagreed that ornithomimids were
1853:may have been weaker than for example those of
1823:Reconstructed skull and neck in front view, NHM
1114:except for in the hindmost cervical vertebrae.
1097:, which were all long and wide, except for the
6237:
4791:
3398:
3323:
3321:
3111:
2868:
2143:in the Gobi Desert of southern Mongolia. This
1841:birds. Osmólska and colleagues suggested that
4713:
4656:"The great dinosaur stampede that never was?"
3576:PalArch's Journal of Vertebrate Palaeontology
3240:
3238:
2703:
2701:
2699:
827:", but returned to using only the genus name
7602:
4570:
4032:
3452:
3363:
3361:
3359:
3357:
3207:
1907:, or the ridges in the beaks of turtles and
1220:was straight, long, with two well developed
6796:
6714:
5836:
4313:
4311:
4247:
4179:
3840:
3318:
3301:
3055:
3053:
3032:. New York: Simon & Schuster. pp.
2944:
2942:
2903:
2627:Kielan-Jaworowska, Z.; Dovchin, N. (1968).
2310:. Predatory theropods that lived alongside
476:. At the time it was named, the fossils of
129:Reconstructed skeleton (based on the adult
4720:
4706:
4463:"What we've learned about dinosaurs since
4317:
4222:
3235:
2869:Oest, S. G.; Hansen, S.; Oest, T. (1997).
2789:. Jefferson: McFarland & Co. pp.
2696:
1709:birds. Since their predators may have had
120:
4625:"Dinosaur stampede stopped in its tracks"
4602:. London: Pegasus Books. pp. 66–67.
4460:
4456:
4454:
4434:
4349:. London: Random Century Group. pp.
4275:
4095:
4008:Jerzykiewicz, T.; Russell, D. A. (1991).
3862:
3693:
3653:
3563:
3503:
3493:
3354:
3345:
3192:
3182:
3019:
3017:
3015:
3013:
3011:
2748:
2746:
2744:
2742:
2740:
2647:
2645:
2360:, as well as oviraptorosaurians, such as
1763:, balance during locomotion, or used for
748:At the time it was named, the fossils of
585:, where it lived alongside its relatives
6111:
6026:
5291:
4867:
4342:
4308:
4038:
3795:Nicholls, E. L.; Russell, A. P. (1985).
3788:
3759:
3078:
3050:
2981:
2939:
2776:
2774:
2397:
2393:
2120:
2101:(channels which connect bone cells) and
2043:
1890:
1818:
1728:
1667:
1200:
1116:
1053:, based on those known from the related
1041:
943:half, and the lower front margin of the
929:
880:
765:
732:
616:
4653:
4622:
4152:
3724:
3327:
2680:The Horned Dinosaurs: A Natural History
2671:
2239:, have been discovered in this region.
1617:Ornithomimosaurs belonged to the clade
1089:(with their hollow chambers invaded by
1081:(openings which have also been called "
1037:
700:. The generic name is derived from the
612:
9033:
4597:
4451:
4412:
4336:
4228:
3821:
3753:
3160:
3158:
3089:The Princeton Field Guide to Dinosaurs
3062:The Princeton Field Guide to Dinosaurs
3008:
2987:
2737:
2677:
2642:
1124:structures in the caudal vertebrae of
985:, due to the presence of an extensive
448:specimen of the new genus and species
8914:
8913:
8352:
8351:
7589:
6783:
6098:
5278:
4854:
4739:
4701:
4527:
4490:
4373:
4281:
4101:
3639:
3604:
3598:
3367:
2771:
989:, which was common to ornithomimids.
772:Central Museum of Mongolian Dinosaurs
520:had openings that indicate they were
3084:
3059:
3023:
2844:
2825:Kobayashi, Y.; Barsbold, R. (2006).
2780:
2551:Timeline of ornithomimosaur research
2378:. Other ornithomimosaurs, including
2116:
2065:), these structures were originally
815:. In 2010, he instead listed it as "
661:) was discovered by palaeontologist
462:. The specific name is derived from
4146:
3927:Folia Histochemica et Cytobiologica
3900:Folia Histochemica et Cytobiologica
3670:
3155:
1947:, which feeds on plants, molluscs,
1919:, Makovicky, and Currie reported a
1814:
892:is the largest known member of the
13:
3727:Canadian Journal of Earth Sciences
3529:Journal of Vertebrate Paleontology
2422:; a similar scene in the original
1735:National Museum of Natural History
14:
9082:
9041:Late Cretaceous dinosaurs of Asia
7590:
4677:
4248:Hurum, J. H.; Sabath, K. (2003).
3302:Kobayashi, Y.; Lü, J.-C. (2003).
2186:in an environment similar to the
1248:Osmólska and colleagues assigned
1243:
934:Reconstructed skull and neck, NHM
454:in 1972. The generic name means "
9016:
8894:
8888:
8881:
8333:
8326:
8319:
8313:
7571:
7564:
7559:
7552:
7546:
6784:
6765:
6758:
6751:
6080:
6073:
6066:
6059:
5261:
5255:
4683:
3347:10.1111/j.1475-4983.2005.00448.x
3028:Predatory Dinosaurs of the World
2852:"Cretaceous Mongolian Dinosaurs"
2067:secondary sexual characteristics
1663:
1565:
1543:
1521:
1499:
1472:
1443:
1416:
1355:
1319:
1304:
1205:Hindlimb bones of ZPAL MgD-I/8,
1085:"), and were therefore probably
632:fossils were collected in area A
360:
145:
45:
4647:
4616:
4591:
4564:
4521:
4484:
4406:
4374:Baird, R. (1998). "Animalizing
4367:
4157:Dinosaurs of the Flaming Cliffs
3945:
3918:
3887:
3815:
3718:
3633:
3520:
3201:
2862:
2854:. DinoCasts.com. Archived from
2412:was featured in the 1993 movie
1782:while the foot is preserved in
1626:discovered, including the most
628:fossil localities of Mongolia;
9071:Taxa named by Rinchen Barsbold
9066:Taxa named by Halszka Osmólska
4577:dinosaurs looked in real life"
2753:Kielan-Jaworowska, Z. (2013).
2039:
876:
1:
9046:Fossil taxa described in 1972
6099:
5279:
4855:
4727:
4318:Shay, D.; Duncan, J. (1993).
4265:Acta Palaeontologica Polonica
3549:10.1080/02724634.2016.1221415
3308:Acta Palaeontologica Polonica
2556:
2515:It has been claimed that the
2286:, large hadrosaurids such as
1652:grouped ornithomimosaurs and
1648:In 1994, the palaeontologist
800:In 1988, the palaeontologist
642:Mongolian Academy of Sciences
4161:. New York: Anchor. p.
4026:10.1016/0195-6671(91)90015-5
3801:(Theropoda: Ornithomimidae)"
3782:10.1016/0031-0182(82)90005-0
3704:10.1080/08912960410001674200
3495:10.1371/journal.pone.0145168
3184:10.1371/journal.pone.0266648
2975:10.1016/j.palaeo.2017.10.020
2933:10.1016/j.palaeo.2017.10.032
1923:skull (IGM 100/1133) and an
1093:). The neck consisted of 10
545:would have been a fleet (or
7:
4320:The Making of Jurassic Park
2785:Dinosaurs: The Encyclopedia
2755:In Pursuit of Early Mammals
2544:
2170:Tsaagan Khushuu consist of
1717:. In 1982, palaeontologist
707:, "chicken", and the Greek
636:Between 1963 and 1965, the
468:, a golden capsule worn by
10:
9087:
4065:A Geologic Time Scale 2004
3208:Christiansen, Per (1998).
2471:computer-generated imagery
2444:Industrial Light and Magic
2336:, and troodontids such as
2017:and titanosaur sauropods,
1915:In 2001, palaeontologists
1679:The cervical vertebrae of
719:is derived from the Latin
680:In 1972, palaeontologists
638:Polish Academy of Sciences
599:was featured in the movie
425:that lived in what is now
133:and a juvenile specimen),
8922:
8878:
8835:
8730:
8673:
8644:
8635:
8561:
8552:
8536:
8525:
8435:
8420:
8416:
8405:
8375:
8360:
8347:
8311:
8258:
8226:
8172:
8130:
8103:
8076:
8054:
8045:
8018:
8007:
7978:
7967:
7847:
7843:
7832:
7780:
7751:
7728:
7724:
7702:
7613:
7598:
7585:
7544:
7514:
7448:
7380:
7370:
7366:
7301:
7286:
7277:
7198:
7189:
7137:
7086:
7077:
7033:
7022:
6966:
6943:
6934:
6896:
6892:
6881:
6842:
6831:
6807:
6792:
6779:
6748:
6710:
6667:
6623:
6614:
6576:
6531:
6522:
6484:
6473:
6409:
6368:
6353:
6344:
6328:
6324:
6313:
6267:
6252:
6248:
6226:
6122:
6107:
6094:
6056:
6022:
5925:
5845:
5815:
5751:
5665:
5654:
5598:
5564:
5480:
5471:
5431:
5427:
5416:
5385:
5340:
5336:
5325:
5302:
5287:
5274:
5252:
5217:
5197:
5121:
5077:
5068:
4960:
4949:
4878:
4863:
4850:
4818:
4800:
4787:
4735:
4542:10.1177/03063127030332015
4530:Social Studies of Science
3864:10.1016/j.cub.2019.10.050
3619:10.1017/S0022336000026706
3582:(1): 1–73. Archived from
2456:chasing a fully rendered
2221:The Nemegt rivers, where
2159:stage, at the end of the
1558:
1536:
1514:
1492:
1465:
1458:
1436:
1409:
1402:
1382:
1375:
1348:
1341:
725:, a gold capsule worn by
331:
324:
299:
292:
142:Scientific classification
140:
128:
119:
23:
4417:(Submitted manuscript).
4079:10.1017/CBO9780511536045
4039:Sullivan, R. M. (2006).
3378:Mesozoic Vertebrate Life
2661:Palaeontologica Polonica
2636:Palaeontologica Polonica
2298:, and sauropods such as
1158:the upper margin of the
925:
885:Size compared to a human
668:USSR Academy of Sciences
4654:Shapiro, A. D. (2013).
3607:Journal of Paleontology
3328:Barrett, P. M. (2005).
3268:10.1126/science.1225376
2884:: 24–25. Archived from
2610:Palaeontologia Polonica
2537:), and a consultant to
1293:shows the placement of
663:Zofia Kielan-Jaworowska
4598:Martin, A. J. (2014).
4461:Montanari, S. (2015).
3822:Madsen, E. K. (2007).
2406:
2386:, are also found, but
2133:
2053:
1912:
1824:
1737:
1676:
1209:
1134:
1059:
961:infratemporal fenestra
935:
886:
865:variation among newer
774:
741:
633:
135:Natural History Museum
8989:Paleobiology Database
7279:Carcharodontosaurinae
7191:Carcharodontosauridae
4571:Phillips, I. (2015).
4491:Kirby, D. A. (2011).
4343:Crichton, M. (1990).
2432:stop-motion animation
2401:
2394:Cultural significance
2202:formations, although
2149:dated radiometrically
2124:
2047:
1903:of, for example, the
1894:
1886:intramandibular joint
1822:
1732:
1671:
1385:Bissekty ornithomimid
1204:
1120:
1045:
933:
884:
769:
736:
620:
512:was toothless with a
494:is the largest known
8263:Dubious coelurosaurs
6968:Metriacanthosaurinae
6936:Metriacanthosauridae
5218:Dubious neotheropods
4694:at Wikimedia Commons
4629:Australasian Science
4623:Romilio, A. (2015).
4153:Novacek, M. (1996).
3085:Paul, G. S. (2010).
3060:Paul, G. S. (2016).
3024:Paul, G. S. (1988).
2781:Glut, D. F. (1997).
2484:herd was a point in
1038:Postcranial skeleton
684:, Ewa Roniewicz and
613:History of discovery
541:As an ornithomimid,
7292:Carcharodontosaurus
7079:Carcharodontosauria
4200:1967Natur.214..161J
4124:10.1038/nature13930
4116:2014Natur.515..203H
4014:Cretaceous Research
3966:2000Natur.406..619R
3896:Gallimimus bullatus
3799:Struthiomimus altus
3774:1982PPP....38..227T
3739:1972CaJES...9..375R
3541:2016JVPal..36E1415M
3486:2015PLoSO..1045168W
3422:2001Natur.412..873N
3372:Gallimimus bullatus
3260:2012Sci...338..510Z
2988:Gannon, M. (2014).
2967:2018PPP...494..160L
2925:2018PPP...494...51F
2858:on 2 February 2015.
2723:1975AmSci..63..150K
2678:Dodson, P. (1998).
2601:Gallimimus bullatus
2279:pachycephalosaurian
1719:Richard A. Thulborn
1658:Gallimimus bullatus
1007:mandibular fenestra
906:, it is known that
808:, resulting in the
694:Gallimimus bullatus
659:Gallimimus bullatus
451:Gallimimus bullatus
306:Gallimimus bullatus
9056:Maastrichtian life
8755:Archaeornithomimus
7268:Veterupristisaurus
7128:Veterupristisaurus
6984:Metriacanthosaurus
6924:Piatnitzkysauridae
6359:Eustreptospondylus
6269:Piatnitzkysauridae
6152:Chuandongocoelurus
5016:Notatesseraeraptor
4579:. Business Insider
4252:Tarbosaurus bataar
3681:Historical Biology
3468:Archaeornithomimus
3368:Hurum, J. (2001).
2711:American Scientist
2527:stampede scene in
2517:Lark Quarry tracks
2436:computer animation
2407:
2307:Opisthocoelicaudia
2145:geologic formation
2139:is known from the
2134:
2054:
2009:ornithomimosaurs,
1945:Northern shoveller
1913:
1905:Northern shoveller
1825:
1752:Archaeornithomimus
1738:
1677:
1352:Archaeornithomimus
1234:arctometatarsalian
1210:
1135:
1095:cervical vertebrae
1060:
936:
887:
775:
756:Archaeornithomimus
742:
634:
9004:
9003:
8916:Taxon identifiers
8907:
8906:
8903:
8902:
8876:
8875:
8872:
8871:
8868:
8867:
8864:
8863:
8831:
8830:
8827:
8826:
8637:Macrocheiriformes
8554:Ornithomimosauria
8528:Maniraptoriformes
8521:
8520:
8367:Maniraptoromorpha
8354:Maniraptoromorpha
8343:
8342:
8309:
8308:
8305:
8304:
8301:
8300:
8254:
8253:
8250:
8249:
8232:Maniraptoromorpha
8222:
8221:
8218:
8217:
8214:
8213:
8210:
8209:
8206:
8205:
7963:
7962:
7828:
7827:
7782:Proceratosauridae
7734:Chingkankousaurus
7581:
7580:
7542:
7541:
7538:
7537:
7534:
7533:
7510:
7509:
7362:
7361:
7358:
7357:
7354:
7353:
7350:
7349:
7346:
7345:
7342:
7341:
7018:
7017:
7014:
7013:
6775:
6774:
6746:
6745:
6742:
6741:
6738:
6737:
6734:
6733:
6706:
6705:
6702:
6701:
6698:
6697:
6694:
6693:
6610:
6609:
6566:Vallibonavenatrix
6469:
6468:
6465:
6464:
6309:
6308:
6090:
6089:
6054:
6053:
6050:
6049:
6046:
6045:
6018:
6017:
6014:
6013:
6010:
6009:
6006:
6005:
6002:
6001:
5650:
5649:
5646:
5645:
5412:
5411:
5270:
5269:
5250:
5249:
5246:
5245:
5242:
5241:
5193:
5192:
4846:
4845:
4842:
4841:
4838:
4837:
4688:Media related to
4499:. Massachusetts:
4360:978-0-394-58816-2
4329:978-1-85283-774-7
4301:978-0-520-24209-8
4256:Tyrannosaurus rex
4194:(5084): 161–163.
4172:978-0-385-47775-8
4110:(7526): 203–205.
4088:978-0-521-78142-8
3960:(6796): 619–622.
3857:(1): 158–168.e4.
3416:(6850): 873–874.
3254:(6106): 510–514.
3148:978-0-520-24209-8
3104:978-0-691-13720-9
3071:978-0-691-16766-4
3043:978-0-671-61946-6
2996:. LiveScience.com
2994:uk.news.yahoo.com
2800:978-0-89950-917-4
2764:978-0-253-00824-4
2689:978-0-691-05900-6
2599:"A new dinosaur,
2254:hesperornithiform
2117:Palaeoenvironment
2105:fibre bundles of
1674:Dinosauria Museum
1619:Maniraptoriformes
1614:
1613:
1605:
1604:
1596:
1595:
1587:
1586:
1578:
1577:
1278:, both forming a
1103:occipital condyle
1049:showing an adult
914:dinosaurs") were
526:Ornithomimosauria
438:million years ago
351:
350:
345:
273:
234:Ornithomimosauria
9078:
9021:
9020:
9019:
9012:
8997:
8996:
8984:
8983:
8971:
8970:
8958:
8957:
8956:
8943:
8942:
8941:
8911:
8910:
8898:
8892:
8885:
8741:Aepyornithomimus
8642:
8641:
8559:
8558:
8534:
8533:
8498:Sinocalliopteryx
8418:
8417:
8414:
8413:
8408:Neocoelurosauria
8373:
8372:
8362:
8361:
8349:
8348:
8337:
8330:
8323:
8317:
8260:
8259:
8228:
8227:
8197:Zhuchengtyrannus
8121:Thanatotheristes
8052:
8051:
8016:
8015:
7984:Appalachiosaurus
7976:
7975:
7845:
7844:
7841:
7840:
7835:Pantyrannosauria
7726:
7725:
7722:
7721:
7716:Tyrannosauroidea
7711:
7710:
7686:Xinjiangovenator
7611:
7610:
7600:
7599:
7587:
7586:
7575:
7568:
7563:
7556:
7550:
7378:
7377:
7368:
7367:
7284:
7283:
7204:Acrocanthosaurus
7196:
7195:
7084:
7083:
7031:
7030:
6957:Yangchuanosaurus
6941:
6940:
6894:
6893:
6890:
6889:
6840:
6839:
6821:Lourinhanosaurus
6805:
6804:
6794:
6793:
6781:
6780:
6769:
6762:
6755:
6712:
6711:
6678:Sigilmassasaurus
6621:
6620:
6578:Ceratosuchopsini
6529:
6528:
6482:
6481:
6427:Dubreuillosaurus
6351:
6350:
6334:Streptospondylus
6326:
6325:
6322:
6321:
6292:Piatnitzkysaurus
6250:
6249:
6246:
6245:
6235:
6234:
6120:
6119:
6109:
6108:
6096:
6095:
6084:
6077:
6070:
6063:
6024:
6023:
5859:Ekrixinatosaurus
5843:
5842:
5834:
5833:
5663:
5662:
5478:
5477:
5429:
5428:
5425:
5424:
5338:
5337:
5334:
5333:
5300:
5299:
5289:
5288:
5276:
5275:
5265:
5259:
5169:Procompsognathus
5075:
5074:
4958:
4957:
4876:
4875:
4865:
4864:
4852:
4851:
4798:
4797:
4789:
4788:
4782:
4781:
4737:
4736:
4722:
4715:
4708:
4699:
4698:
4687:
4671:
4670:
4668:
4666:
4651:
4645:
4644:
4620:
4614:
4613:
4595:
4589:
4588:
4586:
4584:
4568:
4562:
4561:
4525:
4519:
4518:
4498:
4488:
4482:
4481:
4479:
4477:
4458:
4449:
4448:
4438:
4410:
4404:
4403:
4371:
4365:
4364:
4340:
4334:
4333:
4315:
4306:
4305:
4289:
4279:
4273:
4272:
4262:
4245:
4239:
4238:
4226:
4220:
4219:
4208:10.1038/214161a0
4183:
4177:
4176:
4160:
4150:
4144:
4143:
4099:
4093:
4092:
4068:
4058:
4052:
4051:
4045:
4036:
4030:
4029:
4005:
3994:
3993:
3974:10.1038/35020550
3949:
3943:
3942:
3922:
3916:
3915:
3906:(3–4): 241–244.
3891:
3885:
3884:
3866:
3838:
3832:
3831:
3819:
3813:
3812:
3792:
3786:
3785:
3768:(3–4): 227–256.
3757:
3751:
3750:
3722:
3716:
3715:
3697:
3674:
3668:
3667:
3657:
3637:
3631:
3630:
3613:(5): 1100–1117.
3602:
3596:
3595:
3593:
3591:
3567:
3561:
3560:
3524:
3518:
3517:
3507:
3497:
3480:(12): e0145168.
3461:
3450:
3449:
3430:10.1038/35091139
3405:
3396:
3395:
3365:
3352:
3351:
3349:
3325:
3316:
3315:
3299:
3288:
3287:
3242:
3233:
3232:
3214:
3205:
3199:
3198:
3196:
3186:
3177:(10). e0266648.
3162:
3153:
3152:
3136:
3126:
3109:
3108:
3092:
3082:
3076:
3075:
3057:
3048:
3047:
3034:384–386, 393–394
3031:
3021:
3006:
3005:
3003:
3001:
2985:
2979:
2978:
2946:
2937:
2936:
2907:
2901:
2900:
2898:
2896:
2890:
2875:
2866:
2860:
2859:
2848:
2842:
2841:
2831:
2822:
2805:
2804:
2788:
2778:
2769:
2768:
2750:
2735:
2734:
2705:
2694:
2693:
2675:
2669:
2668:
2658:
2649:
2640:
2639:
2633:
2624:
2618:
2617:
2607:
2594:
2534:Muttaburrasaurus
2420:Steven Spielberg
2141:Nemegt Formation
2131:Nemegt Formation
2077:, and brooding.
1935:in the beaks of
1815:Feeding and diet
1765:thermoregulation
1759:related to high
1743:binocular vision
1569:
1547:
1525:
1503:
1489:
1476:
1461:
1460:
1447:
1433:
1420:
1405:
1404:
1399:
1378:
1377:
1372:
1359:
1344:
1343:
1334:
1333:
1323:
1308:
1282:(or "advanced")
1155:sacral vertebrae
1143:dorsal vertebrae
1047:Life restoration
1011:coronoid process
987:secondary palate
908:ornithomimosaurs
845:Philip J. Currie
833:informally named
686:Rinchen Barsbold
682:Halszka Osmólska
583:Nemegt Formation
504:, it would have
436:, about seventy
410:
406:
401:
400:
397:
396:
393:
390:
387:
384:
381:
378:
375:
372:
369:
366:
340:
308:
304:
285:G. bullatus
265:
258:
245:
232:
150:
149:
124:
114:
44:
29:Temporal range:
21:
20:
9086:
9085:
9081:
9080:
9079:
9077:
9076:
9075:
9031:
9030:
9027:
9017:
9015:
9007:
9005:
9000:
8992:
8987:
8979:
8974:
8966:
8961:
8952:
8951:
8946:
8937:
8936:
8931:
8918:
8908:
8899:
8886:
8860:
8823:
8804:Sinornithomimus
8726:
8669:
8631:
8548:
8542:Compsognathidae
8530:
8517:
8505:Sinosauropteryx
8437:Compsognathidae
8431:
8410:
8401:
8369:
8356:
8339:
8338:
8324:
8297:
8292:Shanyangosaurus
8285:Phaedrolosaurus
8246:
8202:
8168:
8126:
8105:Daspletosaurini
8099:
8072:
8047:Tyrannosaurinae
8041:
8020:Albertosaurinae
8012:
8010:Tyrannosauridae
8003:
7972:
7970:Eutyrannosauria
7959:
7837:
7824:
7805:Proceratosaurus
7776:
7747:
7718:
7707:
7698:
7664:Richardoestesia
7648:Chilantaisaurus
7607:
7594:
7577:
7576:
7557:
7530:
7506:
7466:Australovenator
7444:
7438:Chilantaisaurus
7416:Phuwiangvenator
7338:
7303:Giganotosaurini
7297:
7273:
7185:
7148:Chilantaisaurus
7133:
7073:
7027:
7010:
6962:
6930:
6886:
6877:
6855:Monolophosaurus
6836:
6827:
6801:
6788:
6771:
6770:
6756:
6730:
6690:
6663:
6629:Camarillasaurus
6606:
6572:
6518:
6512:Ostafrikasaurus
6490:Camarillasaurus
6478:
6461:
6411:Afrovenatorinae
6405:
6364:
6340:
6318:
6305:
6263:
6242:
6240:Megalosauroidea
6231:
6222:
6217:Vectaerovenator
6210:Szechuanosaurus
6188:Monolophosaurus
6174:Kaijiangosaurus
6167:Cryolophosaurus
6128:Calamospondylus
6116:
6103:
6086:
6085:
6078:
6071:
6064:
6042:
5998:
5979:Pycnonemosaurus
5921:
5832:
5826:Majungasaurinae
5811:
5753:Majungasaurinae
5747:
5742:Xenotarsosaurus
5678:Dryptosauroides
5659:
5642:
5594:
5566:Elaphrosaurinae
5560:
5467:
5453:Ornithomimoides
5421:
5408:
5381:
5330:
5321:
5296:
5283:
5266:
5260:
5238:
5213:
5189:
5117:
5064:
5030:Shuangbaisaurus
4954:
4945:
4872:
4859:
4834:
4814:
4808:Avemetatarsalia
4793:Avemetatarsalia
4783:
4768:Avemetatarsalia
4742:
4741:
4731:
4726:
4680:
4675:
4674:
4664:
4662:
4652:
4648:
4621:
4617:
4610:
4596:
4592:
4582:
4580:
4569:
4565:
4526:
4522:
4515:
4489:
4485:
4475:
4473:
4459:
4452:
4427:10.2307/1213468
4411:
4407:
4392:10.2307/1225728
4376:Jurassic Park's
4372:
4368:
4361:
4341:
4337:
4330:
4316:
4309:
4302:
4280:
4276:
4260:
4246:
4242:
4227:
4223:
4184:
4180:
4173:
4151:
4147:
4100:
4096:
4089:
4059:
4055:
4043:
4037:
4033:
4006:
3997:
3950:
3946:
3923:
3919:
3892:
3888:
3851:Current Biology
3841:Button, D. J.;
3839:
3835:
3820:
3816:
3793:
3789:
3758:
3754:
3747:10.1139/e72-031
3723:
3719:
3695:10.1.1.394.9219
3675:
3671:
3655:10.1.1.211.3044
3638:
3634:
3603:
3599:
3589:
3587:
3586:on 22 June 2018
3568:
3564:
3535:(6): e1221415.
3525:
3521:
3462:
3453:
3406:
3399:
3392:
3366:
3355:
3326:
3319:
3300:
3291:
3243:
3236:
3212:
3206:
3202:
3163:
3156:
3149:
3127:
3112:
3105:
3083:
3079:
3072:
3058:
3051:
3044:
3022:
3009:
2999:
2997:
2986:
2982:
2947:
2940:
2908:
2904:
2894:
2892:
2888:
2873:
2867:
2863:
2850:
2849:
2845:
2829:
2823:
2808:
2801:
2779:
2772:
2765:
2751:
2738:
2706:
2697:
2690:
2676:
2672:
2656:
2650:
2643:
2631:
2625:
2621:
2605:
2595:
2564:
2559:
2547:
2475:special effects
2396:
2357:Therizinosaurus
2316:tyrannosauroids
2245:enantiornithine
2231:crocodylomorphs
2190:of present-day
2161:Late Cretaceous
2147:has never been
2119:
2048:Smallest known
2042:
1847:cranial kinesis
1845:was capable of
1817:
1721:estimated that
1666:
1650:Thomas R. Holtz
1615:
1606:
1597:
1588:
1579:
1504:
1490:
1487:
1434:
1431:
1413:Sinornithomimus
1400:
1397:
1373:
1370:
1331:
1330:
1329:
1328:
1327:
1324:
1316:
1315:
1313:Experimentarium
1309:
1246:
1040:
1015:oviraptorosaurs
957:temporal region
928:
879:
810:new combination
802:Gregory S. Paul
778:Fossil poaching
615:
607:special effects
431:Late Cretaceous
408:
404:
363:
359:
320:
310:
302:
301:
288:
264:
256:
243:
230:
144:
115:
113:
112:
111:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
39:
38:
31:Late Cretaceous
27:
17:
12:
11:
5:
9084:
9074:
9073:
9068:
9063:
9058:
9053:
9048:
9043:
9026:
9025:
9002:
9001:
8999:
8998:
8985:
8972:
8959:
8944:
8928:
8926:
8920:
8919:
8905:
8904:
8901:
8900:
8880:
8879:
8877:
8874:
8873:
8870:
8869:
8866:
8865:
8862:
8861:
8859:
8858:
8857:
8856:
8850:
8841:
8839:
8833:
8832:
8829:
8828:
8825:
8824:
8822:
8821:
8814:
8807:
8800:
8793:
8786:
8779:
8772:
8769:Dromiceiomimus
8765:
8758:
8751:
8744:
8736:
8734:
8732:Ornithomimidae
8728:
8727:
8725:
8724:
8717:
8713:Paraxenisaurus
8709:
8701:
8694:
8687:
8679:
8677:
8675:Deinocheiridae
8671:
8670:
8668:
8667:
8664:Shenzhousaurus
8660:
8653:
8645:
8639:
8633:
8632:
8630:
8629:
8622:
8614:
8607:
8600:
8593:
8586:
8578:
8570:
8562:
8556:
8550:
8549:
8547:
8546:
8537:
8531:
8526:
8523:
8522:
8519:
8518:
8516:
8515:
8508:
8501:
8494:
8486:
8479:
8472:
8465:
8461:Beipiaognathus
8457:
8450:
8441:
8439:
8433:
8432:
8430:
8429:
8421:
8411:
8406:
8403:
8402:
8400:
8399:
8392:
8385:
8376:
8370:
8365:
8358:
8357:
8345:
8344:
8341:
8340:
8332:
8318:
8312:
8310:
8307:
8306:
8303:
8302:
8299:
8298:
8296:
8295:
8288:
8281:
8274:
8266:
8264:
8256:
8255:
8252:
8251:
8248:
8247:
8245:
8244:
8243:
8242:
8236:
8234:
8224:
8223:
8220:
8219:
8216:
8215:
8212:
8211:
8208:
8207:
8204:
8203:
8201:
8200:
8193:
8186:
8178:
8176:
8174:Tyrannosaurini
8170:
8169:
8167:
8166:
8159:
8152:
8144:
8136:
8134:
8132:Teratophoneini
8128:
8127:
8125:
8124:
8117:
8114:Daspletosaurus
8109:
8107:
8101:
8100:
8098:
8097:
8094:Qianzhousaurus
8090:
8082:
8080:
8074:
8073:
8071:
8070:
8063:
8055:
8049:
8043:
8042:
8040:
8039:
8032:
8024:
8022:
8013:
8008:
8005:
8004:
8002:
8001:
7994:
7991:Bistahieversor
7987:
7979:
7973:
7968:
7965:
7964:
7961:
7960:
7958:
7957:
7949:
7942:
7935:
7927:
7920:
7913:
7906:
7899:
7892:
7885:
7878:
7871:
7863:
7856:
7848:
7838:
7833:
7830:
7829:
7826:
7825:
7823:
7822:
7815:
7808:
7801:
7794:
7786:
7784:
7778:
7777:
7775:
7774:
7766:
7758:
7756:
7749:
7748:
7746:
7745:
7737:
7729:
7719:
7714:
7708:
7705:Tyrannoraptora
7703:
7700:
7699:
7697:
7696:
7689:
7682:
7675:
7667:
7660:
7652:
7644:
7637:
7629:
7622:
7614:
7608:
7603:
7596:
7595:
7583:
7582:
7579:
7578:
7570:
7551:
7545:
7543:
7540:
7539:
7536:
7535:
7532:
7531:
7529:
7528:
7527:
7526:
7520:
7518:
7512:
7511:
7508:
7507:
7505:
7504:
7497:
7490:
7483:
7476:
7469:
7462:
7454:
7452:
7450:Megaraptoridae
7446:
7445:
7443:
7442:
7434:
7426:
7419:
7412:
7405:
7397:
7389:
7381:
7375:
7364:
7363:
7360:
7359:
7356:
7355:
7352:
7351:
7348:
7347:
7344:
7343:
7340:
7339:
7337:
7336:
7329:
7322:
7315:
7312:Giganotosaurus
7307:
7305:
7299:
7298:
7296:
7295:
7287:
7281:
7275:
7274:
7272:
7271:
7264:
7257:
7250:
7243:
7236:
7229:
7222:
7215:
7207:
7199:
7193:
7187:
7186:
7184:
7183:
7175:
7167:
7160:
7152:
7143:
7141:
7139:Neovenatoridae
7135:
7134:
7132:
7131:
7124:
7121:Ulughbegsaurus
7117:
7109:
7102:
7095:
7087:
7081:
7075:
7074:
7072:
7071:
7063:
7055:
7047:
7039:
7037:
7028:
7023:
7020:
7019:
7016:
7015:
7012:
7011:
7009:
7008:
7001:
6994:
6987:
6980:
6972:
6970:
6964:
6963:
6961:
6960:
6953:
6949:Xuanhanosaurus
6944:
6938:
6932:
6931:
6929:
6928:
6920:
6916:Xuanhanosaurus
6912:
6905:
6902:Asfaltovenator
6897:
6887:
6882:
6879:
6878:
6876:
6875:
6867:
6863:Megalosauridae
6859:
6851:
6843:
6837:
6832:
6829:
6828:
6826:
6825:
6817:
6808:
6802:
6797:
6790:
6789:
6777:
6776:
6773:
6772:
6764:
6750:
6749:
6747:
6744:
6743:
6740:
6739:
6736:
6735:
6732:
6731:
6729:
6728:
6727:
6726:
6720:
6718:
6708:
6707:
6704:
6703:
6700:
6699:
6696:
6695:
6692:
6691:
6689:
6688:
6681:
6673:
6671:
6665:
6664:
6662:
6661:
6654:
6647:
6640:
6637:Ichthyovenator
6633:
6624:
6618:
6612:
6611:
6608:
6607:
6605:
6604:
6597:
6590:
6582:
6580:
6574:
6573:
6571:
6570:
6562:
6554:
6547:
6540:
6532:
6526:
6520:
6519:
6517:
6516:
6508:
6501:
6498:Cristatusaurus
6494:
6485:
6479:
6474:
6471:
6470:
6467:
6466:
6463:
6462:
6460:
6459:
6455:Poekilopleuron
6451:
6448:Piveteausaurus
6444:
6437:
6430:
6423:
6415:
6413:
6407:
6406:
6404:
6403:
6396:
6389:
6382:
6374:
6372:
6370:Megalosaurinae
6366:
6365:
6363:
6362:
6354:
6348:
6346:Megalosauridae
6342:
6341:
6339:
6338:
6329:
6319:
6314:
6311:
6310:
6307:
6306:
6304:
6303:
6299:Xuanhanosaurus
6295:
6288:
6281:
6273:
6271:
6265:
6264:
6262:
6261:
6258:Yunyangosaurus
6253:
6243:
6238:
6232:
6227:
6224:
6223:
6221:
6220:
6213:
6206:
6198:
6195:Pandoravenator
6191:
6184:
6181:Kayentavenator
6177:
6170:
6163:
6155:
6148:
6140:
6132:
6123:
6117:
6112:
6105:
6104:
6092:
6091:
6088:
6087:
6079:
6072:
6065:
6058:
6057:
6055:
6052:
6051:
6048:
6047:
6044:
6043:
6041:
6040:
6039:
6038:
6032:
6030:
6020:
6019:
6016:
6015:
6012:
6011:
6008:
6007:
6004:
6003:
6000:
5999:
5997:
5996:
5989:
5982:
5975:
5968:
5960:
5953:
5946:
5939:
5931:
5929:
5923:
5922:
5920:
5919:
5912:
5909:Skorpiovenator
5905:
5897:
5890:
5883:
5876:
5869:
5862:
5855:
5846:
5840:
5831:
5830:
5821:
5819:
5813:
5812:
5810:
5809:
5802:
5794:
5787:
5780:
5773:
5765:
5757:
5755:
5749:
5748:
5746:
5745:
5738:
5731:
5724:
5721:Spectrovenator
5717:
5710:
5703:
5696:
5689:
5682:
5674:
5666:
5660:
5655:
5652:
5651:
5648:
5647:
5644:
5643:
5641:
5640:
5633:
5626:
5619:
5612:
5604:
5602:
5596:
5595:
5593:
5592:
5585:
5578:
5570:
5568:
5562:
5561:
5559:
5558:
5555:Spinostropheus
5551:
5544:
5537:
5529:
5521:
5513:
5505:
5498:
5490:
5481:
5475:
5469:
5468:
5466:
5465:
5457:
5449:
5441:
5432:
5422:
5419:Abelisauroidea
5417:
5414:
5413:
5410:
5409:
5407:
5406:
5399:
5391:
5389:
5387:Ceratosauridae
5383:
5382:
5380:
5379:
5376:Saltriovenator
5372:
5364:
5357:
5353:Fosterovenator
5349:
5341:
5331:
5326:
5323:
5322:
5320:
5319:
5315:Lophostropheus
5311:
5303:
5297:
5292:
5285:
5284:
5272:
5271:
5268:
5267:
5254:
5253:
5251:
5248:
5247:
5244:
5243:
5240:
5239:
5237:
5236:
5229:
5221:
5219:
5215:
5214:
5212:
5211:
5210:
5209:
5203:
5201:
5195:
5194:
5191:
5190:
5188:
5187:
5180:
5176:Pterospondylus
5172:
5165:
5157:
5149:
5146:Lucianovenator
5142:
5135:
5127:
5125:
5119:
5118:
5116:
5115:
5108:
5101:
5094:
5087:
5078:
5072:
5070:Coelophysoidea
5066:
5065:
5063:
5062:
5055:
5047:
5040:
5033:
5026:
5019:
5012:
5009:Lophostropheus
5005:
4998:
4991:
4984:
4977:
4969:
4961:
4955:
4950:
4947:
4946:
4944:
4943:
4935:
4927:
4919:
4911:
4908:Erythrovenator
4904:
4896:
4888:
4879:
4873:
4868:
4861:
4860:
4848:
4847:
4844:
4843:
4840:
4839:
4836:
4835:
4833:
4832:
4831:
4830:
4824:
4822:
4816:
4815:
4813:
4812:
4811:
4810:
4801:
4795:
4785:
4784:
4780:
4779:
4770:
4761:
4755:
4749:
4733:
4732:
4725:
4724:
4717:
4710:
4702:
4696:
4695:
4679:
4678:External links
4676:
4673:
4672:
4646:
4615:
4609:978-1605984995
4608:
4590:
4575:Jurassic World
4563:
4536:(2): 231–268.
4520:
4514:978-0262294867
4513:
4483:
4450:
4415:Film Quarterly
4405:
4380:Cinema Journal
4366:
4359:
4335:
4328:
4307:
4300:
4286:The Dinosauria
4274:
4240:
4233:(in Russian).
4221:
4178:
4171:
4145:
4094:
4087:
4053:
4031:
4020:(4): 345–377.
3995:
3944:
3917:
3886:
3833:
3814:
3787:
3752:
3733:(4): 375–402.
3717:
3688:(2–4): 57–70.
3669:
3632:
3597:
3562:
3519:
3451:
3397:
3391:978-0253339072
3390:
3370:"Lower jaw of
3353:
3340:(2): 347–358.
3317:
3289:
3234:
3200:
3154:
3147:
3133:The Dinosauria
3110:
3103:
3077:
3070:
3049:
3042:
3007:
2980:
2938:
2902:
2891:on 14 May 2018
2861:
2843:
2806:
2799:
2770:
2763:
2736:
2717:(2): 150–159.
2695:
2688:
2670:
2641:
2619:
2561:
2560:
2558:
2555:
2554:
2553:
2546:
2543:
2506:John R. Horner
2502:bird evolution
2486:Jurassic Park'
2440:visual effects
2395:
2392:
2188:Okavango Delta
2118:
2115:
2041:
2038:
1953:foraminiferans
1941:filter-feeding
1917:Mark A. Norell
1816:
1813:
1665:
1662:
1623:coelurosaurian
1612:
1611:
1608:
1607:
1603:
1602:
1599:
1598:
1594:
1593:
1590:
1589:
1585:
1584:
1581:
1580:
1576:
1575:
1572:
1571:
1557:
1554:
1553:
1550:
1549:
1535:
1532:
1531:
1528:
1527:
1513:
1510:
1509:
1506:
1505:
1498:
1491:
1486:
1483:
1482:
1479:
1478:
1464:
1459:
1457:
1454:
1453:
1450:
1449:
1435:
1430:
1427:
1426:
1423:
1422:
1408:
1403:
1401:
1396:
1393:
1392:
1389:
1388:
1381:
1376:
1374:
1369:
1366:
1365:
1362:
1361:
1347:
1342:
1340:
1338:Ornithomimidae
1332:
1325:
1318:
1317:
1310:
1303:
1302:
1301:
1300:
1299:
1289:The following
1266:Lü found that
1245:
1244:Classification
1242:
1127:Shenzhousaurus
1039:
1036:
1019:therizinosaurs
995:common seagull
979:internal nares
927:
924:
878:
875:
855:junior synonym
847:proposed that
688:named the new
652:were numerous
614:
611:
571:filter-feeding
349:
348:
347:
346:
329:
328:
322:
321:
311:
297:
296:
290:
289:
281:
279:
275:
274:
254:
250:
249:
247:Ornithomimidae
241:
237:
236:
228:
221:
220:
215:
208:
207:
202:
195:
194:
189:
182:
181:
176:
172:
171:
166:
162:
161:
156:
152:
151:
138:
137:
126:
125:
117:
116:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
28:
15:
9:
6:
4:
3:
2:
9083:
9072:
9069:
9067:
9064:
9062:
9059:
9057:
9054:
9052:
9051:Ornithomimids
9049:
9047:
9044:
9042:
9039:
9038:
9036:
9029:
9024:
9014:
9013:
9010:
8995:
8990:
8986:
8982:
8977:
8973:
8969:
8964:
8960:
8955:
8949:
8945:
8940:
8934:
8930:
8929:
8927:
8925:
8921:
8917:
8912:
8897:
8893:
8891:
8884:
8855:
8851:
8849:
8845:
8844:
8843:
8842:
8840:
8838:
8834:
8820:
8819:
8815:
8813:
8812:
8811:Struthiomimus
8808:
8806:
8805:
8801:
8799:
8798:
8794:
8792:
8791:
8787:
8785:
8784:
8780:
8778:
8777:
8773:
8771:
8770:
8766:
8764:
8763:
8762:"Coelosaurus"
8759:
8757:
8756:
8752:
8750:
8749:
8745:
8743:
8742:
8738:
8737:
8735:
8733:
8729:
8723:
8722:
8718:
8715:
8714:
8710:
8707:
8706:
8702:
8700:
8699:
8695:
8693:
8692:
8688:
8686:
8685:
8681:
8680:
8678:
8676:
8672:
8666:
8665:
8661:
8659:
8658:
8657:Pelecanimimus
8654:
8652:
8651:
8647:
8646:
8643:
8640:
8638:
8634:
8628:
8627:
8623:
8620:
8619:
8618:Thecocoelurus
8615:
8613:
8612:
8608:
8606:
8605:
8601:
8599:
8598:
8597:Kinnareemimus
8594:
8592:
8591:
8587:
8584:
8583:
8579:
8576:
8575:
8571:
8569:
8568:
8564:
8563:
8560:
8557:
8555:
8551:
8544:
8543:
8539:
8538:
8535:
8532:
8529:
8524:
8514:
8513:
8509:
8507:
8506:
8502:
8500:
8499:
8495:
8492:
8491:
8487:
8485:
8484:
8480:
8478:
8477:
8476:Huaxiagnathus
8473:
8471:
8470:
8469:Compsognathus
8466:
8463:
8462:
8458:
8456:
8455:
8451:
8448:
8447:
8443:
8442:
8440:
8438:
8434:
8428:
8427:
8423:
8422:
8419:
8415:
8412:
8409:
8404:
8398:
8397:
8396:Ornitholestes
8393:
8391:
8390:
8386:
8383:
8382:
8378:
8377:
8374:
8371:
8368:
8363:
8359:
8355:
8350:
8346:
8336:
8331:
8329:
8322:
8316:
8294:
8293:
8289:
8287:
8286:
8282:
8280:
8279:
8275:
8273:
8272:
8268:
8267:
8265:
8261:
8257:
8240:
8239:
8238:
8237:
8235:
8233:
8229:
8225:
8199:
8198:
8194:
8192:
8191:
8190:Tyrannosaurus
8187:
8185:
8184:
8180:
8179:
8177:
8175:
8171:
8165:
8164:
8163:Teratophoneus
8160:
8158:
8157:
8153:
8150:
8149:
8145:
8143:
8142:
8138:
8137:
8135:
8133:
8129:
8123:
8122:
8118:
8116:
8115:
8111:
8110:
8108:
8106:
8102:
8096:
8095:
8091:
8089:
8088:
8084:
8083:
8081:
8079:
8075:
8069:
8068:
8064:
8062:
8061:
8057:
8056:
8053:
8050:
8048:
8044:
8038:
8037:
8033:
8031:
8030:
8029:Albertosaurus
8026:
8025:
8023:
8021:
8017:
8014:
8011:
8006:
8000:
7999:
7995:
7993:
7992:
7988:
7986:
7985:
7981:
7980:
7977:
7974:
7971:
7966:
7955:
7954:
7950:
7948:
7947:
7946:Xiongguanlong
7943:
7941:
7940:
7936:
7933:
7932:
7928:
7926:
7925:
7924:Suskityrannus
7921:
7919:
7918:
7917:Stokesosaurus
7914:
7912:
7911:
7910:Santanaraptor
7907:
7905:
7904:
7900:
7898:
7897:
7893:
7891:
7890:
7886:
7884:
7883:
7879:
7877:
7876:
7872:
7869:
7868:
7864:
7862:
7861:
7857:
7855:
7854:
7853:Alectrosaurus
7850:
7849:
7846:
7842:
7839:
7836:
7831:
7821:
7820:
7816:
7814:
7813:
7809:
7807:
7806:
7802:
7800:
7799:
7795:
7793:
7792:
7788:
7787:
7785:
7783:
7779:
7772:
7771:
7770:Tanycolagreus
7767:
7765:
7764:
7760:
7759:
7757:
7754:
7750:
7743:
7742:
7738:
7736:
7735:
7731:
7730:
7727:
7723:
7720:
7717:
7712:
7709:
7706:
7701:
7695:
7694:
7690:
7688:
7687:
7683:
7681:
7680:
7676:
7673:
7672:
7668:
7666:
7665:
7661:
7658:
7657:
7653:
7650:
7649:
7645:
7643:
7642:
7638:
7635:
7634:
7633:Bahariasaurus
7630:
7628:
7627:
7623:
7621:
7620:
7616:
7615:
7612:
7609:
7606:
7605:Coelurosauria
7601:
7597:
7593:
7592:Coelurosauria
7588:
7584:
7574:
7569:
7567:
7562:
7555:
7549:
7524:
7523:
7522:
7521:
7519:
7517:
7516:Coelurosauria
7513:
7503:
7502:
7498:
7496:
7495:
7491:
7489:
7488:
7484:
7482:
7481:
7477:
7475:
7474:
7470:
7468:
7467:
7463:
7461:
7460:
7456:
7455:
7453:
7451:
7447:
7441:
7439:
7435:
7433:
7431:
7427:
7425:
7424:
7420:
7418:
7417:
7413:
7411:
7410:
7406:
7403:
7402:
7398:
7395:
7394:
7393:Bahariasaurus
7390:
7388:
7387:
7383:
7382:
7379:
7376:
7373:
7369:
7365:
7335:
7334:
7330:
7328:
7327:
7323:
7321:
7320:
7316:
7314:
7313:
7309:
7308:
7306:
7304:
7300:
7294:
7293:
7289:
7288:
7285:
7282:
7280:
7276:
7270:
7269:
7265:
7263:
7262:
7258:
7256:
7255:
7251:
7249:
7248:
7244:
7242:
7241:
7237:
7235:
7234:
7233:Kelmayisaurus
7230:
7228:
7227:
7223:
7221:
7220:
7216:
7213:
7212:
7208:
7206:
7205:
7201:
7200:
7197:
7194:
7192:
7188:
7181:
7180:
7176:
7173:
7172:
7168:
7166:
7165:
7161:
7158:
7157:
7153:
7150:
7149:
7145:
7144:
7142:
7140:
7136:
7130:
7129:
7125:
7123:
7122:
7118:
7115:
7114:
7110:
7108:
7107:
7103:
7101:
7100:
7096:
7094:
7093:
7089:
7088:
7085:
7082:
7080:
7076:
7069:
7068:
7067:Saurophaganax
7064:
7061:
7060:
7056:
7053:
7052:
7048:
7046:
7045:
7041:
7040:
7038:
7036:
7032:
7029:
7026:
7021:
7007:
7006:
7002:
7000:
6999:
6998:Siamotyrannus
6995:
6993:
6992:
6988:
6986:
6985:
6981:
6979:
6978:
6974:
6973:
6971:
6969:
6965:
6959:
6958:
6954:
6951:
6950:
6946:
6945:
6942:
6939:
6937:
6933:
6926:
6925:
6921:
6918:
6917:
6913:
6911:
6910:
6906:
6904:
6903:
6899:
6898:
6895:
6891:
6888:
6885:
6884:Allosauroidea
6880:
6873:
6872:
6871:Spinosauridae
6868:
6865:
6864:
6860:
6857:
6856:
6852:
6850:
6849:
6845:
6844:
6841:
6838:
6835:
6830:
6823:
6822:
6818:
6815:
6814:
6810:
6809:
6806:
6803:
6800:
6795:
6791:
6787:
6782:
6778:
6768:
6763:
6761:
6754:
6724:
6723:
6722:
6721:
6719:
6717:
6713:
6709:
6687:
6686:
6682:
6680:
6679:
6675:
6674:
6672:
6670:
6666:
6660:
6659:
6655:
6653:
6652:
6648:
6646:
6645:
6641:
6639:
6638:
6634:
6631:
6630:
6626:
6625:
6622:
6619:
6617:
6616:Spinosaurinae
6613:
6603:
6602:
6598:
6596:
6595:
6594:Riparovenator
6591:
6589:
6588:
6587:Ceratosuchops
6584:
6583:
6581:
6579:
6575:
6568:
6567:
6563:
6560:
6559:
6555:
6553:
6552:
6551:Riojavenatrix
6548:
6546:
6545:
6541:
6539:
6538:
6534:
6533:
6530:
6527:
6525:
6524:Baryonychinae
6521:
6514:
6513:
6509:
6507:
6506:
6502:
6500:
6499:
6495:
6492:
6491:
6487:
6486:
6483:
6480:
6477:
6476:Spinosauridae
6472:
6457:
6456:
6452:
6450:
6449:
6445:
6443:
6442:
6438:
6436:
6435:
6431:
6429:
6428:
6424:
6422:
6421:
6417:
6416:
6414:
6412:
6408:
6402:
6401:
6400:Wiehenvenator
6397:
6395:
6394:
6390:
6388:
6387:
6383:
6381:
6380:
6376:
6375:
6373:
6371:
6367:
6361:
6360:
6356:
6355:
6352:
6349:
6347:
6343:
6336:
6335:
6331:
6330:
6327:
6323:
6320:
6317:
6312:
6301:
6300:
6296:
6294:
6293:
6289:
6287:
6286:
6282:
6280:
6279:
6275:
6274:
6272:
6270:
6266:
6260:
6259:
6255:
6254:
6251:
6247:
6244:
6241:
6236:
6233:
6230:
6225:
6219:
6218:
6214:
6212:
6211:
6207:
6204:
6203:
6199:
6197:
6196:
6192:
6190:
6189:
6185:
6183:
6182:
6178:
6176:
6175:
6171:
6169:
6168:
6164:
6161:
6160:
6156:
6154:
6153:
6149:
6146:
6145:
6141:
6138:
6137:
6136:Chienkosaurus
6133:
6130:
6129:
6125:
6124:
6121:
6118:
6115:
6110:
6106:
6102:
6097:
6093:
6083:
6076:
6069:
6062:
6036:
6035:
6034:
6033:
6031:
6029:
6025:
6021:
5995:
5994:
5990:
5988:
5987:
5983:
5981:
5980:
5976:
5974:
5973:
5969:
5966:
5965:
5961:
5959:
5958:
5954:
5952:
5951:
5947:
5945:
5944:
5940:
5938:
5937:
5933:
5932:
5930:
5928:
5927:Furileusauria
5924:
5918:
5917:
5913:
5911:
5910:
5906:
5903:
5902:
5901:Rahiolisaurus
5898:
5896:
5895:
5891:
5889:
5888:
5884:
5882:
5881:
5877:
5875:
5874:
5870:
5868:
5867:
5863:
5861:
5860:
5856:
5853:
5852:
5848:
5847:
5844:
5841:
5839:
5835:
5828:
5827:
5823:
5822:
5820:
5818:
5817:Carnotaurinae
5814:
5808:
5807:
5803:
5800:
5799:
5798:Rahiolisaurus
5795:
5793:
5792:
5791:Majungasaurus
5788:
5786:
5785:
5781:
5779:
5778:
5774:
5771:
5770:
5766:
5764:
5763:
5759:
5758:
5756:
5754:
5750:
5744:
5743:
5739:
5737:
5736:
5732:
5730:
5729:
5728:Tarascosaurus
5725:
5723:
5722:
5718:
5716:
5715:
5711:
5709:
5708:
5704:
5702:
5701:
5697:
5695:
5694:
5690:
5688:
5687:
5686:Eoabelisaurus
5683:
5680:
5679:
5675:
5673:
5672:
5671:Chenanisaurus
5668:
5667:
5664:
5661:
5658:
5657:Abelisauridae
5653:
5639:
5638:
5634:
5632:
5631:
5627:
5625:
5624:
5620:
5618:
5617:
5616:Masiakasaurus
5613:
5611:
5610:
5606:
5605:
5603:
5601:
5597:
5591:
5590:
5586:
5584:
5583:
5582:Huinculsaurus
5579:
5577:
5576:
5575:Elaphrosaurus
5572:
5571:
5569:
5567:
5563:
5557:
5556:
5552:
5550:
5549:
5545:
5543:
5542:
5538:
5535:
5534:
5530:
5527:
5526:
5522:
5519:
5518:
5514:
5511:
5510:
5506:
5504:
5503:
5499:
5496:
5495:
5494:Austrocheirus
5491:
5488:
5487:
5483:
5482:
5479:
5476:
5474:
5470:
5463:
5462:
5458:
5455:
5454:
5450:
5447:
5446:
5442:
5439:
5438:
5434:
5433:
5430:
5426:
5423:
5420:
5415:
5405:
5404:
5400:
5398:
5397:
5393:
5392:
5390:
5388:
5384:
5378:
5377:
5373:
5370:
5369:
5365:
5363:
5362:
5358:
5355:
5354:
5350:
5348:
5347:
5346:Berberosaurus
5343:
5342:
5339:
5335:
5332:
5329:
5324:
5317:
5316:
5312:
5310:
5309:
5305:
5304:
5301:
5298:
5295:
5290:
5286:
5282:
5277:
5273:
5264:
5258:
5235:
5234:
5233:Halticosaurus
5230:
5228:
5227:
5226:Teinurosaurus
5223:
5222:
5220:
5216:
5207:
5206:
5205:
5204:
5202:
5200:
5196:
5186:
5185:
5181:
5178:
5177:
5173:
5171:
5170:
5166:
5163:
5162:
5158:
5155:
5154:
5153:Megapnosaurus
5150:
5148:
5147:
5143:
5141:
5140:
5136:
5134:
5133:
5129:
5128:
5126:
5124:
5123:Coelophysidae
5120:
5114:
5113:
5112:Powellvenator
5109:
5107:
5106:
5102:
5100:
5099:
5095:
5093:
5092:
5088:
5085:
5084:
5080:
5079:
5076:
5073:
5071:
5067:
5061:
5060:
5056:
5053:
5052:
5048:
5046:
5045:
5041:
5039:
5038:
5034:
5032:
5031:
5027:
5025:
5024:
5020:
5018:
5017:
5013:
5011:
5010:
5006:
5004:
5003:
5002:Liliensternus
4999:
4997:
4996:
4992:
4990:
4989:
4985:
4983:
4982:
4978:
4975:
4974:
4973:Dolichosuchus
4970:
4968:
4967:
4966:Dilophosaurus
4963:
4962:
4959:
4956:
4953:
4948:
4941:
4940:
4939:Herrerasauria
4936:
4933:
4932:
4928:
4925:
4924:
4920:
4917:
4916:
4912:
4910:
4909:
4905:
4902:
4901:
4897:
4894:
4893:
4892:Daemonosaurus
4889:
4886:
4885:
4881:
4880:
4877:
4874:
4871:
4866:
4862:
4858:
4853:
4849:
4828:
4827:
4826:
4825:
4823:
4821:
4817:
4809:
4805:
4804:
4803:
4802:
4799:
4796:
4794:
4790:
4786:
4778:
4774:
4771:
4769:
4765:
4762:
4760:
4756:
4754:
4750:
4748:
4744:
4743:
4738:
4734:
4730:
4723:
4718:
4716:
4711:
4709:
4704:
4703:
4700:
4693:
4692:
4686:
4682:
4681:
4661:
4657:
4650:
4642:
4638:
4634:
4630:
4626:
4619:
4611:
4605:
4601:
4594:
4578:
4576:
4567:
4559:
4555:
4551:
4547:
4543:
4539:
4535:
4531:
4524:
4516:
4510:
4506:
4502:
4497:
4496:
4487:
4472:
4468:
4466:
4465:Jurassic Park
4457:
4455:
4446:
4442:
4437:
4432:
4428:
4424:
4420:
4416:
4409:
4401:
4397:
4393:
4389:
4386:(4): 82–103.
4385:
4381:
4377:
4370:
4362:
4356:
4352:
4348:
4347:
4346:Jurassic Park
4339:
4331:
4325:
4321:
4314:
4312:
4303:
4297:
4293:
4288:
4287:
4278:
4270:
4266:
4259:
4257:
4253:
4244:
4236:
4232:
4225:
4217:
4213:
4209:
4205:
4201:
4197:
4193:
4189:
4182:
4174:
4168:
4164:
4159:
4158:
4149:
4141:
4137:
4133:
4129:
4125:
4121:
4117:
4113:
4109:
4105:
4098:
4090:
4084:
4080:
4076:
4072:
4067:
4066:
4057:
4049:
4042:
4035:
4027:
4023:
4019:
4015:
4011:
4004:
4002:
4000:
3991:
3987:
3983:
3979:
3975:
3971:
3967:
3963:
3959:
3955:
3948:
3940:
3936:
3932:
3928:
3921:
3913:
3909:
3905:
3901:
3897:
3890:
3882:
3878:
3874:
3870:
3865:
3860:
3856:
3852:
3848:
3844:
3837:
3829:
3825:
3818:
3810:
3806:
3805:Palaeontology
3802:
3800:
3791:
3783:
3779:
3775:
3771:
3767:
3763:
3756:
3748:
3744:
3740:
3736:
3732:
3728:
3721:
3713:
3709:
3705:
3701:
3696:
3691:
3687:
3683:
3682:
3673:
3665:
3661:
3656:
3651:
3647:
3643:
3636:
3628:
3624:
3620:
3616:
3612:
3608:
3601:
3585:
3581:
3577:
3573:
3566:
3558:
3554:
3550:
3546:
3542:
3538:
3534:
3530:
3523:
3515:
3511:
3506:
3501:
3496:
3491:
3487:
3483:
3479:
3475:
3471:
3469:
3460:
3458:
3456:
3447:
3443:
3439:
3435:
3431:
3427:
3423:
3419:
3415:
3411:
3404:
3402:
3393:
3387:
3383:
3379:
3375:
3373:
3364:
3362:
3360:
3358:
3348:
3343:
3339:
3335:
3334:Palaeontology
3331:
3324:
3322:
3314:(2): 235–259.
3313:
3309:
3305:
3298:
3296:
3294:
3285:
3281:
3277:
3273:
3269:
3265:
3261:
3257:
3253:
3249:
3241:
3239:
3230:
3226:
3222:
3218:
3211:
3204:
3195:
3190:
3185:
3180:
3176:
3172:
3168:
3161:
3159:
3150:
3144:
3140:
3135:
3134:
3125:
3123:
3121:
3119:
3117:
3115:
3106:
3100:
3096:
3091:
3090:
3081:
3073:
3067:
3063:
3056:
3054:
3045:
3039:
3035:
3030:
3029:
3020:
3018:
3016:
3014:
3012:
2995:
2991:
2984:
2976:
2972:
2968:
2964:
2960:
2956:
2952:
2945:
2943:
2934:
2930:
2926:
2922:
2918:
2914:
2906:
2887:
2883:
2880:(in Danish).
2879:
2872:
2865:
2857:
2853:
2847:
2840:(1): 195–207.
2839:
2835:
2828:
2821:
2819:
2817:
2815:
2813:
2811:
2802:
2796:
2792:
2787:
2786:
2777:
2775:
2766:
2760:
2756:
2749:
2747:
2745:
2743:
2741:
2732:
2728:
2724:
2720:
2716:
2712:
2704:
2702:
2700:
2691:
2685:
2681:
2674:
2666:
2662:
2655:
2648:
2646:
2637:
2630:
2623:
2615:
2611:
2604:
2602:
2593:
2591:
2589:
2587:
2585:
2583:
2581:
2579:
2577:
2575:
2573:
2571:
2569:
2567:
2562:
2552:
2549:
2548:
2542:
2540:
2539:Jurassic Park
2536:
2535:
2530:
2529:Jurassic Park
2526:
2522:
2518:
2513:
2511:
2507:
2503:
2499:
2495:
2491:
2487:
2483:
2478:
2476:
2472:
2468:
2464:
2459:
2455:
2454:
2453:Tyrannosaurus
2449:
2445:
2441:
2437:
2433:
2429:
2425:
2421:
2417:
2416:
2415:Jurassic Park
2411:
2404:
2400:
2391:
2389:
2385:
2381:
2377:
2376:
2371:
2370:
2365:
2364:
2359:
2358:
2353:
2352:
2347:
2346:
2341:
2340:
2335:
2334:
2329:
2328:
2323:
2322:
2317:
2313:
2309:
2308:
2303:
2302:
2301:Nemegtosaurus
2297:
2296:
2291:
2290:
2285:
2284:
2280:
2276:
2275:
2270:
2269:ankylosaurids
2266:
2265:
2261:, as well as
2260:
2259:
2255:
2251:
2250:
2246:
2242:
2238:
2237:
2232:
2228:
2224:
2219:
2217:
2213:
2209:
2205:
2201:
2197:
2193:
2189:
2185:
2181:
2177:
2173:
2169:
2168:type locality
2166:
2162:
2158:
2157:Maastrichtian
2154:
2150:
2146:
2142:
2138:
2132:
2128:
2123:
2114:
2112:
2111:ornithischian
2108:
2104:
2100:
2096:
2091:
2087:
2083:
2078:
2076:
2072:
2068:
2064:
2060:
2051:
2046:
2037:
2034:
2030:
2026:
2025:caenagnathids
2022:
2021:
2016:
2012:
2006:
2004:
2000:
1996:
1991:
1987:
1983:
1979:
1974:
1969:
1967:
1962:
1958:
1954:
1950:
1946:
1942:
1938:
1934:
1930:
1926:
1922:
1918:
1910:
1906:
1902:
1898:
1893:
1889:
1887:
1883:
1879:
1875:
1871:
1867:
1863:
1858:
1856:
1855:Struthiomimus
1852:
1848:
1844:
1840:
1839:insectivorous
1835:
1830:
1821:
1812:
1810:
1806:
1802:
1798:
1794:
1789:
1785:
1781:
1777:
1773:
1768:
1766:
1762:
1757:
1753:
1749:
1744:
1736:
1731:
1727:
1724:
1720:
1716:
1712:
1711:colour vision
1708:
1704:
1700:
1696:
1695:
1690:
1686:
1682:
1675:
1670:
1664:Palaeobiology
1661:
1659:
1655:
1651:
1646:
1644:
1643:Turgai Strait
1639:
1635:
1634:
1633:Pelecanimimus
1629:
1624:
1620:
1610:
1609:
1601:
1600:
1592:
1591:
1583:
1582:
1574:
1573:
1570:
1568:
1564:
1563:
1556:
1555:
1552:
1551:
1548:
1546:
1542:
1541:
1534:
1533:
1530:
1529:
1526:
1524:
1520:
1519:
1512:
1511:
1508:
1507:
1502:
1497:
1496:
1495:Struthiomimus
1485:
1484:
1481:
1480:
1477:
1475:
1471:
1470:
1463:
1462:
1456:
1455:
1452:
1451:
1448:
1446:
1442:
1441:
1429:
1428:
1425:
1424:
1421:
1419:
1415:
1414:
1407:
1406:
1395:
1394:
1391:
1390:
1387:
1386:
1380:
1379:
1368:
1367:
1364:
1363:
1360:
1358:
1354:
1353:
1346:
1345:
1339:
1336:
1335:
1322:
1314:
1307:
1298:
1296:
1292:
1287:
1285:
1281:
1277:
1273:
1269:
1264:
1259:
1255:
1254:Struthiomimus
1251:
1241:
1239:
1235:
1231:
1227:
1223:
1219:
1215:
1208:
1203:
1199:
1197:
1193:
1189:
1185:
1181:
1177:
1173:
1168:
1166:
1161:
1156:
1152:
1148:
1144:
1140:
1133:
1129:
1128:
1123:
1119:
1115:
1112:
1108:
1104:
1100:
1096:
1092:
1088:
1084:
1080:
1076:
1072:
1068:
1064:
1058:
1057:
1052:
1051:with feathers
1048:
1044:
1035:
1032:
1028:
1024:
1020:
1016:
1012:
1008:
1004:
1000:
996:
990:
988:
984:
980:
976:
972:
971:parietal bone
968:
967:
966:Struthiomimus
962:
958:
954:
950:
946:
941:
932:
923:
921:
917:
913:
909:
905:
904:
899:
895:
891:
883:
874:
872:
868:
864:
863:morphological
860:
856:
852:
851:
846:
842:
838:
834:
830:
826:
822:
821:Struthiomimus
818:
814:
811:
807:
803:
798:
796:
791:
787:
783:
779:
773:
768:
764:
762:
758:
757:
751:
746:
740:
735:
731:
728:
724:
723:
718:
717:specific name
714:
710:
706:
703:
699:
695:
691:
687:
683:
678:
676:
673:
669:
664:
660:
655:
654:ornithomimids
651:
647:
643:
639:
631:
627:
623:
619:
610:
608:
604:
603:
602:Jurassic Park
598:
594:
593:
588:
584:
580:
576:
572:
568:
564:
560:
556:
552:
548:
544:
539:
537:
533:
532:
527:
523:
519:
515:
511:
507:
503:
502:
497:
493:
489:
487:
483:
479:
475:
471:
467:
466:
461:
457:
453:
452:
447:
443:
439:
435:
432:
428:
424:
421:
417:
413:
412:
399:
357:
356:
343:
339:
337:
333:
332:
330:
327:
323:
319:
315:
309:
307:
298:
295:
294:Binomial name
291:
287:
286:
280:
277:
276:
271:
268:
263:
262:
255:
252:
251:
248:
242:
239:
238:
235:
229:
226:
223:
222:
219:
216:
213:
210:
209:
206:
203:
200:
197:
196:
193:
190:
187:
184:
183:
180:
177:
174:
173:
170:
167:
164:
163:
160:
157:
154:
153:
148:
143:
139:
136:
132:
127:
123:
118:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
43:
36:
35:Maastrichtian
32:
26:
22:
19:
9061:Nemegt fauna
9028:
8923:
8887:
8816:
8809:
8802:
8795:
8788:
8783:Ornithomimus
8781:
8775:
8774:
8767:
8760:
8753:
8746:
8739:
8721:Tyrannomimus
8719:
8711:
8703:
8696:
8691:Deinocheirus
8689:
8682:
8662:
8655:
8648:
8624:
8616:
8611:Nqwebasaurus
8609:
8604:Nedcolbertia
8602:
8595:
8588:
8582:Calamosaurus
8580:
8574:Aviatyrannis
8572:
8565:
8540:
8510:
8503:
8496:
8488:
8481:
8474:
8467:
8459:
8454:Aristosuchus
8452:
8446:Aniksosaurus
8444:
8426:Aniksosaurus
8424:
8394:
8387:
8379:
8325:
8290:
8283:
8276:
8269:
8195:
8188:
8181:
8161:
8154:
8146:
8141:Dynamoterror
8139:
8119:
8112:
8092:
8085:
8065:
8060:Asiatyrannus
8058:
8034:
8027:
7998:Dryptosaurus
7996:
7989:
7982:
7951:
7944:
7937:
7929:
7922:
7915:
7908:
7901:
7894:
7889:Jinbeisaurus
7887:
7880:
7873:
7865:
7860:Aviatyrannis
7858:
7851:
7817:
7812:Sinotyrannus
7810:
7803:
7796:
7789:
7768:
7761:
7739:
7732:
7691:
7684:
7677:
7669:
7662:
7654:
7646:
7641:Bicentenaria
7639:
7631:
7626:Asiamericana
7624:
7617:
7558:
7499:
7492:
7485:
7478:
7471:
7464:
7457:
7436:
7428:
7421:
7414:
7407:
7401:Deltadromeus
7399:
7391:
7384:
7333:Tyrannotitan
7331:
7324:
7317:
7310:
7290:
7266:
7261:Taurovenator
7259:
7252:
7245:
7240:Lajasvenator
7238:
7231:
7224:
7219:Concavenator
7217:
7209:
7202:
7177:
7169:
7162:
7154:
7146:
7126:
7119:
7111:
7104:
7097:
7090:
7065:
7057:
7049:
7042:
7035:Allosauridae
7003:
6996:
6991:Shidaisaurus
6989:
6982:
6975:
6955:
6947:
6922:
6914:
6907:
6900:
6869:
6861:
6853:
6846:
6819:
6811:
6799:Avetheropoda
6786:Avetheropoda
6757:
6716:Avetheropoda
6683:
6676:
6669:Spinosaurini
6656:
6649:
6642:
6635:
6627:
6599:
6592:
6585:
6564:
6556:
6549:
6544:Protathlitis
6542:
6535:
6510:
6503:
6496:
6488:
6453:
6446:
6439:
6434:Leshansaurus
6432:
6425:
6418:
6398:
6391:
6386:Megalosaurus
6384:
6379:Duriavenator
6377:
6357:
6332:
6316:Megalosauria
6297:
6290:
6285:Marshosaurus
6283:
6278:Condorraptor
6276:
6256:
6215:
6208:
6200:
6193:
6186:
6179:
6172:
6165:
6159:Cruxicheiros
6157:
6150:
6142:
6134:
6126:
5991:
5986:Quilmesaurus
5984:
5977:
5970:
5962:
5955:
5948:
5941:
5934:
5914:
5907:
5899:
5892:
5885:
5878:
5871:
5864:
5857:
5849:
5838:Brachyrostra
5824:
5804:
5796:
5789:
5782:
5775:
5767:
5760:
5740:
5735:Tralkasaurus
5733:
5726:
5719:
5712:
5705:
5698:
5691:
5684:
5676:
5669:
5637:Vespersaurus
5635:
5630:Velocisaurus
5628:
5621:
5614:
5607:
5587:
5580:
5573:
5553:
5546:
5539:
5531:
5525:Deltadromeus
5523:
5515:
5509:Compsosuchus
5507:
5500:
5492:
5484:
5459:
5451:
5445:Lametasaurus
5443:
5435:
5401:
5396:Ceratosaurus
5394:
5374:
5366:
5359:
5351:
5344:
5328:Ceratosauria
5313:
5306:
5231:
5224:
5182:
5174:
5167:
5159:
5151:
5144:
5137:
5130:
5110:
5105:Podokesaurus
5103:
5096:
5089:
5081:
5057:
5049:
5042:
5035:
5028:
5021:
5014:
5007:
5000:
4993:
4988:Gojirasaurus
4986:
4981:Dracovenator
4979:
4971:
4964:
4952:Neotheropoda
4937:
4929:
4921:
4915:Guaibasaurus
4913:
4906:
4898:
4890:
4884:Chindesaurus
4882:
4772:
4763:
4690:
4663:. Retrieved
4659:
4649:
4635:(2): 24–27.
4632:
4628:
4618:
4599:
4593:
4581:. Retrieved
4574:
4566:
4533:
4529:
4523:
4494:
4486:
4474:. Retrieved
4470:
4464:
4421:(3): 27–37.
4418:
4414:
4408:
4383:
4379:
4375:
4369:
4345:
4338:
4319:
4285:
4277:
4268:
4264:
4255:
4251:
4243:
4234:
4230:
4224:
4191:
4187:
4181:
4156:
4148:
4107:
4103:
4097:
4064:
4056:
4047:
4034:
4017:
4013:
3957:
3953:
3947:
3933:(2): 81–83.
3930:
3926:
3920:
3903:
3899:
3895:
3889:
3854:
3850:
3843:Zanno, L. E.
3836:
3827:
3817:
3811:(4): 64–677.
3808:
3804:
3798:
3790:
3765:
3761:
3755:
3730:
3726:
3720:
3685:
3679:
3672:
3645:
3641:
3635:
3610:
3606:
3600:
3588:. Retrieved
3584:the original
3579:
3575:
3565:
3532:
3528:
3522:
3477:
3473:
3467:
3413:
3409:
3377:
3371:
3337:
3333:
3311:
3307:
3251:
3247:
3220:
3216:
3203:
3174:
3170:
3132:
3088:
3080:
3061:
3027:
2998:. Retrieved
2993:
2983:
2958:
2954:
2950:
2916:
2912:
2905:
2893:. Retrieved
2886:the original
2881:
2878:GeologiskNyt
2877:
2864:
2856:the original
2846:
2837:
2833:
2784:
2754:
2714:
2710:
2679:
2673:
2664:
2660:
2635:
2622:
2613:
2609:
2600:
2538:
2532:
2528:
2524:
2514:
2509:
2497:
2489:
2485:
2481:
2479:
2462:
2457:
2451:
2447:
2418:by director
2413:
2409:
2408:
2402:
2387:
2384:Deinocheirus
2383:
2379:
2373:
2367:
2361:
2355:
2349:
2343:
2337:
2331:
2325:
2319:
2311:
2305:
2299:
2293:
2287:
2283:Prenocephale
2281:
2272:
2262:
2256:
2247:
2234:
2233:, including
2222:
2220:
2207:
2164:
2136:
2135:
2126:
2106:
2089:
2079:
2063:Ornithomimus
2062:
2055:
2049:
2033:convergently
2028:
2018:
2011:Deinocheirus
2010:
2007:
1994:
1981:
1972:
1970:
1965:
1956:
1928:
1925:Ornithomimus
1924:
1920:
1914:
1909:hadrosaurids
1896:
1895:The beak of
1881:
1859:
1854:
1850:
1842:
1833:
1828:
1826:
1804:
1800:
1775:
1769:
1755:
1751:
1748:Deinocheirus
1747:
1739:
1722:
1699:Dale Russell
1694:Deinocheirus
1692:
1684:
1680:
1678:
1657:
1647:
1631:
1616:
1562:Ornithomimus
1560:
1559:
1538:
1537:
1516:
1515:
1493:
1467:
1466:
1439:
1438:
1437:
1411:
1410:
1383:
1350:
1349:
1294:
1288:
1275:
1272:sister taxon
1267:
1262:
1253:
1249:
1247:
1211:
1169:
1138:
1137:The back of
1136:
1131:
1125:
1107:zygapophyses
1062:
1061:
1056:Ornithomimus
1054:
1022:
999:angular bone
991:
975:parasphenoid
964:
939:
938:The head of
937:
903:Ornithomimus
901:
889:
888:
870:
866:
858:
848:
841:Dong Zhiming
836:
828:
824:
820:
816:
812:
806:Ornithomimus
805:
799:
794:
789:
785:
781:
776:
760:
754:
749:
747:
743:
720:
708:
704:
693:
692:and species
679:
658:
650:Nemegt Basin
635:
629:
600:
596:
592:Deinocheirus
590:
586:
578:
562:
554:
542:
540:
535:
529:
509:
506:had feathers
501:Ornithomimus
499:
496:ornithomimid
491:
490:
482:ornithomimid
477:
473:
463:
450:
449:
354:
353:
352:
336:Ornithomimus
334:
313:
305:
300:
284:
283:
269:
260:
259:
224:
211:
198:
185:
24:
18:
8948:Wikispecies
8848:Maniraptora
8837:Maniraptora
8818:Tototlmimus
8748:Anserimimus
8698:Garudimimus
8684:Beishanlong
8626:Valdoraptor
8567:Arkansaurus
8512:Xunmenglong
8389:Juravenator
8183:Tarbosaurus
8067:Nanuqsaurus
8036:Gorgosaurus
7953:Megaraptora
7939:Timurlengia
7671:Sciurumimus
7619:Aratasaurus
7487:Murusraptor
7409:Fukuiraptor
7372:Megaraptora
7254:Shaochilong
7226:Eocarcharia
7179:Megaraptora
7099:Lusovenator
6977:Alpkarakush
6834:Carnosauria
6685:Spinosaurus
6658:Siamosaurus
6558:Suchosaurus
6505:Iberospinus
6441:Magnosaurus
6420:Afrovenator
6393:Torvosaurus
6144:Chilesaurus
5957:Carnotaurus
5950:Caletodraco
5936:Abelisaurus
5762:Arcovenator
5609:Laevisuchus
5600:Noasaurinae
5502:Berthasaura
5473:Noasauridae
5403:Genyodectes
5368:Lukousaurus
5361:Jubbulpuria
5161:Panguraptor
5139:Coelophysis
5132:Camposaurus
5091:Panguraptor
5083:Dracoraptor
5059:Zupaysaurus
5044:Tachiraptor
5023:Sarcosaurus
4923:Nhandumirim
4503:. pp.
4436:10919/25811
3223:: 241–255.
2961:: 160–167.
2380:Anserimimus
2369:Nemegtomaia
2345:Tochisaurus
2321:Tarbosaurus
2289:Saurolophus
2236:Shamosuchus
2212:radioactive
2210:, are more
2196:Barun Goyot
2184:floodplains
2040:Development
2020:Segnosaurus
2015:diplodocoid
2003:ectothermic
1999:endothermic
1986:gastroliths
1978:hadrosaurid
1654:troodontids
1540:Anserimimus
1268:Anserimimus
1083:pleurocoels
877:Description
850:Anserimimus
813:O. bullatus
713:Galliformes
646:Gobi Desert
587:Anserimimus
575:herbivorous
531:Anserimimus
460:Galliformes
442:Gobi Desert
429:during the
9035:Categories
8954:Gallimimus
8924:Gallimimus
8776:Gallimimus
8705:Harpymimus
8650:Harpymimus
8271:Iliosuchus
8241:see below↓
8078:Alioramini
7896:Juratyrant
7882:Eotyrannus
7867:Bagaraatan
7819:Yutyrannus
7753:Coeluridae
7679:Vayuraptor
7525:see below↓
7501:Tratayenia
7494:Orkoraptor
7480:Megaraptor
7386:Aoniraptor
7319:Mapusaurus
7247:Sauroniops
7211:Altispinax
7164:Neovenator
7106:Siamraptor
7092:Datanglong
7059:Epanterias
7051:Antrodemus
7044:Allosaurus
7025:Allosauria
6848:Altispinax
6813:Gasosaurus
6725:see below↓
6601:Suchomimus
6202:Sinosaurus
6037:see below↓
5993:Viavenator
5943:Aucasaurus
5880:Ilokelesia
5851:Dahalokely
5806:Rajasaurus
5784:Indosaurus
5777:Genusaurus
5769:Dahalokely
5693:Indosuchus
5589:Limusaurus
5548:Ligabueino
5541:Kiyacursor
5533:Genusaurus
5517:Dahalokely
5437:Betasuchus
5308:Dornraptor
5208:see below↓
5184:Segisaurus
5037:Sinosaurus
4900:Eodromaeus
4829:see below↓
4777:Dinosauria
4759:Sauropsida
4691:Gallimimus
4583:3 December
2951:Gallimimus
2616:: 103–143.
2557:References
2525:Gallimimus
2521:Queensland
2510:Gallimimus
2498:Gallimimus
2494:Alan Grant
2490:Gallimimus
2482:Gallimimus
2463:Gallimimus
2458:Gallimimus
2448:Gallimimus
2438:, but the
2424:1990 novel
2410:Gallimimus
2403:Gallimimus
2388:Gallimimus
2363:Elmisaurus
2333:Bagaraatan
2312:Gallimimus
2295:Barsboldia
2258:Judinornis
2223:Gallimimus
2208:Gallimimus
2176:siltstones
2165:Gallimimus
2151:, but the
2137:Gallimimus
2127:Gallimimus
2107:Gallimimus
2099:canaliculi
2090:Gallimimus
2086:phosphorus
2059:crocodiles
2050:Gallimimus
2029:Gallimimus
1995:Gallimimus
1982:Gallimimus
1973:Gallimimus
1966:Gallimimus
1957:Gallimimus
1937:anseriform
1929:Gallimimus
1921:Gallimimus
1897:Gallimimus
1882:Gallimimus
1878:Jørn Hurum
1851:Gallimimus
1843:Gallimimus
1834:Gallimimus
1829:Gallimimus
1809:gregarious
1805:Gallimimus
1801:Gallimimus
1776:Gallimimus
1772:taphonomic
1761:metabolism
1756:Gallimimus
1723:Gallimimus
1715:camouflage
1689:prehensile
1685:Gallimimus
1681:Gallimimus
1440:Gallimimus
1295:Gallimimus
1276:Gallimimus
1263:Gallimimus
1250:Gallimimus
1230:metatarsal
1139:Gallimimus
1132:Gallimimus
1099:atlas bone
1065:had 64–66
1063:Gallimimus
1031:keratinous
1027:edentulous
1023:Gallimimus
1003:surangular
945:premaxilla
940:Gallimimus
890:Gallimimus
871:Gallimimus
867:Gallimimus
859:Gallimimus
837:Gallimimus
829:Gallimimus
817:Gallimimus
795:Gallimimus
790:Gallimimus
786:Gallimimus
782:Gallimimus
761:Gallimimus
750:Gallimimus
739:CosmoCaixa
672:Cretaceous
630:Gallimimus
622:Cretaceous
597:Gallimimus
579:Gallimimus
563:Gallimimus
555:Gallimimus
543:Gallimimus
536:Gallimimus
514:keratinous
510:Gallimimus
492:Gallimimus
478:Gallimimus
474:Gallimimus
355:Gallimimus
261:Gallimimus
205:Saurischia
192:Dinosauria
25:Gallimimus
9023:Dinosaurs
8852:includes
8797:Rativates
8790:Qiupalong
8490:Scipionyx
8483:Mirischia
8156:Lythronax
8148:Labocania
8087:Alioramus
7741:Labocania
7459:Aerosteon
7005:Sinraptor
6909:Erectopus
6644:Irritator
6229:Orionides
6114:Tetanurae
6101:Tetanurae
6028:Tetanurae
5964:Elemgasem
5887:Llukalkan
5866:Elemgasem
5623:Noasaurus
5486:Afromimus
5294:Averostra
5281:Averostra
5199:Averostra
5051:Velocipes
4870:Theropoda
4857:Theropoda
4820:Theropoda
4745:Kingdom:
4729:Theropoda
4641:1442-679X
4558:111227921
4501:MIT Press
4467:came out"
4271:(2): 188.
4258:compared"
4140:205241353
3881:208652510
3690:CiteSeerX
3664:0871-5424
3650:CiteSeerX
3229:0871-5424
2919:: 51–64.
2375:Rinchenia
2351:Zanabazar
2339:Borogovia
2327:Alioramus
2264:Teviornis
2249:Gurilynia
2200:Djadochta
2095:histology
2071:courtship
1949:ostracods
1862:omnivores
1788:sediments
1780:sandstone
1703:cursorial
1687:were not
1672:Skull in
1518:Qiupalong
1469:Rativates
1291:cladogram
1184:olecranon
1122:Pneumatic
1111:processes
1087:pneumatic
1067:vertebrae
916:feathered
547:cursorial
522:pneumatic
518:vertebrae
312:Osmólska
278:Species:
218:Theropoda
165:Kingdom:
159:Eukaryota
8933:Wikidata
8381:Coelurus
7798:Kileskus
7791:Guanlong
7763:Coelurus
7656:Gualicho
7156:Gualicho
6537:Baryonyx
5873:Guemesia
5700:Kryptops
5461:Ozraptor
5098:Pendraig
4753:Chordata
4751:Phylum:
4747:Animalia
4660:BBC News
4237:: 76–96.
4132:25337885
3982:10949300
3898:bones".
3873:31813611
3845:(2019).
3712:18592288
3648:: 5–61.
3590:16 April
3557:89242374
3514:26682888
3474:PLOS ONE
3438:11528466
3276:23112330
3171:PLOS One
2731:27845359
2667:: 33–82.
2545:See also
2467:gazelles
2442:company
2428:stampede
2318:such as
2314:include
2271:such as
2192:Botswana
2103:collagen
1933:lamellae
1901:lamellae
1870:browsers
1797:bone bed
1784:mudstone
1638:Beringia
1270:was the
1222:condyles
1176:coracoid
1091:air sacs
1079:foramina
949:U-shaped
825:bullatus
698:holotype
640:and the
626:dinosaur
567:omnivore
559:bone bed
446:holotype
427:Mongolia
423:dinosaur
420:theropod
338:bullatus
326:Synonyms
267:Osmólska
240:Family:
179:Chordata
175:Phylum:
169:Animalia
155:Domain:
131:holotype
40:70
8981:4822903
8968:4433732
8939:Q309467
7931:Timimus
7693:Zuolong
7423:Rapator
7326:Meraxes
6651:Oxalaia
5972:Koleken
4995:Lepidus
4757:Class:
4550:3183078
4445:1213468
4400:1225728
4351:137–138
4216:4148855
4196:Bibcode
4112:Bibcode
3990:4384549
3962:Bibcode
3939:1804726
3912:3450541
3830:: 1–21.
3770:Bibcode
3735:Bibcode
3627:1306180
3537:Bibcode
3505:4684312
3482:Bibcode
3446:4313779
3418:Bibcode
3284:2057698
3256:Bibcode
3248:Science
3194:9581415
3095:112–114
2963:Bibcode
2921:Bibcode
2719:Bibcode
2638:: 7–30.
2274:Tarchia
2227:mollusc
2216:uranium
2204:caliche
2082:calcium
2075:display
1990:gizzard
1866:grazers
1793:tendons
1488:unnamed
1432:unnamed
1398:unnamed
1371:unnamed
1280:derived
1196:unguals
1180:humerus
1172:scapula
1141:had 13
1069:in its
912:ostrich
553:birds.
486:ostrich
456:chicken
414:) is a
303:†
282:†
253:Genus:
109:↓
9009:Portal
8590:Hexing
8278:Kakuru
7875:Dilong
5916:Thanos
5894:Niebla
5714:Rugops
5707:Kurupi
4639:
4606:
4556:
4548:
4511:
4507:–129.
4471:Forbes
4443:
4398:
4357:
4326:
4298:
4214:
4188:Nature
4169:
4138:
4130:
4104:Nature
4085:
4073:–371.
3988:
3980:
3954:Nature
3937:
3910:
3879:
3871:
3710:
3692:
3662:
3652:
3625:
3555:
3512:
3502:
3444:
3436:
3410:Nature
3388:
3282:
3274:
3227:
3191:
3145:
3141:–150.
3101:
3068:
3040:
3000:21 May
2895:13 May
2797:
2793:–432.
2761:
2729:
2686:
2372:, and
2277:, the
2252:, the
2241:Mammal
2180:facies
2023:, and
2001:or an
1951:, and
1707:ratite
1238:hallux
1226:fibula
1188:radius
1151:sacrum
1147:fossae
1075:caudal
1001:. The
983:palate
973:. The
953:orbits
894:family
715:. The
705:gallus
675:period
624:-aged
577:diet.
551:ratite
434:period
344:, 1988
314:et al.
272:, 1972
270:et al.
8994:38547
8854:birds
7903:Moros
7430:Siats
7171:Siats
7113:Siats
4773:Clade
4764:Clade
4665:7 May
4573:"How
4554:S2CID
4546:JSTOR
4476:7 May
4441:JSTOR
4396:JSTOR
4261:(PDF)
4212:S2CID
4136:S2CID
4044:(PDF)
3986:S2CID
3877:S2CID
3708:S2CID
3623:JSTOR
3553:S2CID
3442:S2CID
3382:34–41
3280:S2CID
3213:(PDF)
2889:(PDF)
2874:(PDF)
2830:(PDF)
2727:JSTOR
2657:(PDF)
2632:(PDF)
2606:(PDF)
2172:silts
2153:fauna
1961:mesic
1628:basal
1284:clade
1218:tibia
1214:pubis
1192:manus
1160:iliac
1109:(the
1071:spine
1025:were
926:Skull
898:femur
727:Roman
722:bulla
709:mimos
702:Latin
690:genus
470:Roman
465:bulla
416:genus
225:Clade
212:Clade
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