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Foraging

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607:, foraging success increases with an increase in group size then declines once the optimal size is exceeded. A myriad number of factors affect the group sizes in different species. For example, lionesses (female lions) do not make decisions about foraging in a vacuum. They make decisions that reflect a balance between obtaining food, defending their territory and protecting their young. In fact, we see that lion foraging behavior does not maximize their energy gain. They are not behaving optimally with respect to foraging because they have to defend their territory and protect young so they hunt in small groups to reduce the risk of being caught alone. Another factor that may influence group size is the cost of hunting. To understand the behavior of wild dogs and the average group size we must incorporate the distance the dogs run. 481:, which analyzes the behavior of a forager that encounters different types of prey and must choose which to attack. This model is also known as the prey model or the attack model. In this model the predator encounters different prey items and decides whether to spend time handling or eating the prey. It predicts that foragers should ignore low profitability prey items when more profitable items are present and abundant. The objective of this model is to identify the choice that will maximize fitness. How profitable a prey item is depends on ecological variables such as the time required to find, capture, and consume the prey in addition to the energy it provides. It is likely that an individual will settle for a trade off between maximizing the intake rate while eating and minimising the search interval between prey. 341:
sex pheromones. Animals may choose to forage on their own when the resources are abundant, which can occur when the habitat is rich or when the number of conspecifics foraging are few. In these cases there may be no need for group foraging. In addition, foraging alone can result in less interaction with other foragers, which can decrease the amount of competition and dominance interactions an animal deals with. It will also ensure that a solitary forager is less conspicuous to predators. Solitary foraging strategies characterize many of the phocids (the true seals) such as the elephant and harbor seals. An example of an exclusive solitary forager is the South American species of the harvester ant,
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to come up with a new foraging strategy and introduce something it has never used before to maximize his or her fitness (survival). Forebrain size has been associated with learning behavior. Animals with larger brain sizes are expected to learn better. A higher ability to innovate has been linked to larger forebrain sizes in North American and British Isle birds according to Lefebvre et al. (1997). In this study, bird orders that contained individuals with larger forebrain sizes displayed a higher amount of foraging innovation. Examples of innovations recorded in birds include following tractors and eating frogs or other insects killed by it and using swaying trees to catch their prey.
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as an aggregation economy. The second situation occurs when a group of animals forage together but it may not be in an animal's best interest to do so known as a dispersion economy. Think of a cardinal at a bird feeder for the dispersion economy. We might see a group of birds foraging at that bird feeder but it is not in the best interest of the cardinal for any of the other birds to be there too. The amount of food the cardinal can get from that bird feeder depends on how much it can take from the bird feeder but also depends on how much the other birds take as well.
223: 546: 639:. This is the null model for thinking about what would draw animals into groups to forage and how they would behave in the process. This model predicts that animals will make an instantaneous decision about where to forage based on the quality (prey availability) of the patches available at that time and will choose the most profitable patch, the one that maximizes their energy intake. This quality depends on the starting quality of the patch and the number of predators already there consuming the prey. 600:
importantly reduction of predation threat. With regard to costs, however, group foraging results in competition for available resources by other group members. Competition for resources can be characterized by either scramble competition whereby each individual strives to get a portion of the shared resource, or by interference competition whereby the presence of competitors prevents a forager's accessibility to resources. Group foraging can thus reduce an animal's foraging payoff.
154: 43: 369: 618:(2001) defined the forager performing the activity to the optimal efficiency when the individual is having considered the balance of costs for search and pursuit of prey in considerations of prey selection. Also in selecting an area to work within the individual would have had to decide the correct time to move to another location corresponding to perception of yield remaining and potential yields of any given area available. 588: 491:
should stay at a patch or move to a new one, think of a bear in a patch of berry bushes. The longer a bear stays at the patch of berry bushes the less berries there are for that bear to eat. The bear must decide how long to stay and thus when to leave that patch and move to a new patch. Movement depends on the travel time between patches and the energy gained from one patch versus another. This is based on the
245:. Studies using quantitative trait loci (QTL) mapping have associated the following loci with the matched functions; Pln-1 and Pln-4 with onset of foraging age, Pln-1 and 2 with the size of the pollen loads collected by workers, and Pln-2 and pln-3 were shown to influence the sugar concentration of the nectar collected. 627:
activity should be dependent upon the density of juvenile fishes, and the risk of predation within the area. A balance between the growth and mortality of these juvenile fishes is reliant consequent to the duration of foraging performed by said juvenile fish. These components vary with regards to the habitat.
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willing to take, and the efficiency of their foraging patterns. For example, in environments with abundant resources, foragers may adopt less risky and energy-efficient strategies. Conversely, in resource-scarce or high-risk environments, more complex and risky foraging behaviors may evolve. For instance,
565:, the foraging process is divided between three different types of workers: nest patrollers, trail patrollers, and foragers. These workers can utilize many different methods of communicating while foraging in a group, such as guiding flights, scent paths, and "jostling runs", as seen in the eusocial bee 340:
Solitary foraging includes the variety of foraging in which animals find, capture and consume their prey alone. Individuals can manually exploit patches or they can use tools to exploit their prey. For example, Bolas spiders attack their prey by luring them with a scent identical to the female moth's
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mechanism where the loss of time and energy in avoiding food patches is traded with the decrease in risk of contracting a parasite. Adaptations in diet also help in the prevention of parasitic infection. By avoiding foods that have high potential for parasitic contamination, as well as including food
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Behavioral ecologists use economic models and categories to understand foraging; many of these models are a type of optimal model. Thus foraging theory is discussed in terms of optimizing a payoff from a foraging decision. The payoff for many of these models is the amount of energy an animal receives
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algorithms that try to follow the main concepts of group foraging by autonomous agents. An important note here is that group foraging can emerge in two types of situations. The first situation is frequently thought of and occurs when foraging in a group is beneficial and brings greater rewards known
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which describes the behavior of a forager whose prey is concentrated in small areas known as patches with a significant travel time between them. The model seeks to find out how much time an individual will spend on one patch before deciding to move to the next patch. To understand whether an animal
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Foraging behavior can also be influenced by genetics. The genes associated with foraging behavior have been widely studied in honeybees with reference to the following; onset of foraging behavior, task division between foragers and workers, and bias in foraging for either pollen or nectar. Honey bee
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repertoire, or in the environment, that had not previously been suspected. With those constraints identified, foraging behavior often does approach the optimal pattern even if it is not identical to it. In other words, we know from optimal foraging theory that animals are not foraging randomly even
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Foraging can be categorized into two main types. The first is solitary foraging, when animals forage by themselves. The second is group foraging. Group foraging includes when animals can be seen foraging together when it is beneficial for them to do so (called an aggregation economy) and when it is
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One measure of learning is 'foraging innovation'—an animal consuming new food, or using a new foraging technique in response to their dynamic living environment. Foraging innovation is considered learning because it involves behavioral plasticity on the animal's part. The animal recognizes the need
192:, young individuals learn foraging behavior from their peers and elders by watching other group members forage and by copying their behavior. Observing and learning from other members of the group ensure that the younger members of the group learn what is safe to eat and become proficient foragers. 436:
to determine the behavior that an "optimal forager" would exhibit. Such a forager has perfect knowledge of what to do to maximize usable food intake. While the behavior of real animals inevitably departs from that of the optimal forager, optimal foraging theory has proved very useful in developing
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Animals can typically be classified into two categories by their pattern of movement exhibited through foraging behaviors. These categories are "cruise" searchers and "ambush" searchers. Cruise searchers forage by continuously hunting for prey at the outer borders of the area being searched, while
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Interactions with the environment significantly influence foraging behavior by dictating the availability of resources, the competition among others, the presence of predators, and the complexity of the landscape. These factors can affect the strategies animals use to find food, the risks they're
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A quantitative model that allows for the evaluation of trade-off decisions that occur in aquatic ecosystems. 'Foraging arenas' are the areas in which a juvenile fish can forage closer to their home while also providing an easier escape from potential predators. This theory predicts that feeding
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is defined as an adaptive change or modification of a behavior based on a previous experience. Since an animal's environment is constantly changing, the ability to adjust foraging behavior is essential for maximization of fitness. Studies in social insects have shown that there is a significant
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per unit time, more specifically, the highest ratio of energetic gain to cost while foraging. Foraging theory predicts that the decisions that maximize energy per unit time and thus deliver the highest payoff will be selected for and persist. Key words used to describe foraging behavior include
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d'Ivoire also engage in foraging for meats when they can, which is achieved through group foraging. Positive correlation has been observed between the success of the hunt and the size of the foraging group. The chimps have also been observed implying rules with their foraging, where there is a
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As already mentioned, group foraging brings both costs and benefits to the members of that group. Some of the benefits of group foraging include being able to capture larger prey, being able to create aggregations of prey, being able to capture prey that are difficult or dangerous and most
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Group foraging is when animals find, capture and consume prey in the presence of other individuals. In other words, it is foraging when success depends not only on your own foraging behaviors but the behaviors of others as well. The biological behavior also inspired the development of
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strains were produced by crossing the rovers with rovers and sitters with sitters. Using the method of hybridization - crossing rovers with sitters - all of the offspring displayed the rover foraging behavior, thus demonstrating that it is an allele of complete dominance.
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Behavioral ecologists first tackled this topic in the 1960s and 1970s. Their goal was to quantify and formalize a set of models to test their null hypothesis that animals forage randomly. Important contributions to foraging theory have been made by:
254:), there were two types of foraging strategies: rovers and sitters. Rovers used the strategy of moving across multiple patches in search for food, while sitters remained in one patch with no inclination to go searching. Both of these strategies are 1414:
Torres-Contreras, Hugo; Ruby Olivares-Donoso; Hermann M. Niemeyer (2007). "Solitary Foraging in the Ancestral South American Ant, Pogonomyrmex vermiculatus. Is it Due to Constraints in the Production or Perception of Trail Pheromones?".
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can affect the way in which animals forage. For an organism to counteract the procurement of a parasite, they may display avoidance towards certain areas where parasites have previously been discovered. This avoidance behavior is a
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Hrncir, Michael; Jarau, Stefan; Zucchi, Ronaldo; Barth, Friedrich G. (January 2000). "Recruitment behavior in stingless bees, Melipona scutellaris and M. quadrifasciata . II. Possible mechanisms of communication".
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Hutchings, Michael R.; Judge, Johanna; Gordon, Iain J.; Athanasiadou, Spiridoula; Kyriazakis, Ilias (January 2006). "Use of trade-off theory to advance understanding of herbivore–parasite interactions".
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There are many versions of optimal foraging theory that are relevant to different foraging situations. These models generally possess the following components according to Stephens et al. 2007;
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Roch, S.; von Ammon, L.; Geist, J.; Brinker, A. (2018). "Foraging habits of invasive three-spined sticklebacks ( Gasterosteus aculeatus ) – impacts on fisheries yield in Upper Lake Constance".
432:, and by J. Merritt Emlen. This theory argues that because of the key importance of successful foraging to an individual's survival, it should be possible to predict foraging behavior by using 501:
theory is a version of the patch model. This model describes the behavior of a forager that must return to a particular place to consume food, or perhaps to hoard food or feed it to a
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Tupper, Mark; Juanes, Francis (February 2017). "Testing foraging arena theory: The effects of conspecific density and habitat type on time and energy budgets of juvenile cunner".
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le Roux, Aliza; Michael I. Cherry; Lorenz Gygax (5 May 2009). "Vigilance behaviour and fitness consequences: comparing a solitary foraging and an obligate group-foraging mammal".
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traits that naturally occur within the larval stages of fruit flies. The gene responsible for major effects on foraging behavior in Drosophila melanogaster larvae is the chaser (
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individuals learned the locations and times of feeding events, and arrived to those locations up to thirty minutes before the feeding event in anticipation of the food reward.
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ambush searchers forage by sitting and waiting. They remain motionless for long durations as they wait on the prey to pass by, therefore initiating the ambusher to attack.
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with their needs, thus deviating from the foraging behaviour that would be expected in the absence of predators. An example of this balanced risk can be observed in the
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differ in their behavior based on the food resources available in their environment. They will take on a more solitary or active role depending on their environment.
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Murphy, Christina M.; Breed, Michael D. (April 2008). "Time-Place Learning in a Neotropical Stingless Bee, Trigona fulviventris Guérin (Hymenoptera: Apidae)".
509:. Chipmunks are a good example of this model. As travel time between the patch and their hiding place increased, the chipmunks stayed longer at the patch. 239:
foraging activity occurs both inside and outside the hive for either pollen or nectar. Similar behavior is seen in many social wasps, such as the species
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Cruz-Rivera, Edwin; Hay, Mark E. (January 2000). "Can quantity replace quality? food choice, compensatory feeding, and fitness of marine mesograzers".
557:. Birds feeding at a bird feeder is an example of a dispersion economy. This is when it may not be in an animal's best interest to forage in a group. 437:
hypotheses for describing real foraging behavior. Departures from optimality often help to identify constraints either in the animal's behavioral or
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Rapaport, L.G.; Brown, G.R. (2008). "Social influences on foraging behavior in young nonhuman primates:learning what, where and how to eat".
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make foraging decisions and more specifically decisions about hunting group size with protection of their cubs and territory defense in mind.
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Lefebvre, Louis; Whittle, Patrick; Lascaris, Evan; Finkelstein, Adam (March 1997). "Feeding innovations and forebrain size in birds".
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The emergence and stability of cooperative fishing on Ifaluk Atoll, for Human Behavior and Adaptation: an Anthropological Perspective
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The presence of predators while a (prey) animal is foraging affects its behaviour. In general, foragers balance the risk of
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Benoit-Bird, Kelly J.; Au, Whitlow W. L. (2009). "Cooperative prey herding by the pelagic dolphin, Stenella longirostris".
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Creel, Scott; Creel, Nancy Marusha (1995). "Communal hunting and pack size in African wild dogs, Lycaon pictus".
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items that contain anti-parasitic properties in the diet. These anti-parasitic properties can be used in a
3557: 2448: 2278: 1530:"The Ecological Conditions That Favor Tool Use and Innovation in Wild Bottlenose Dolphins (Tursiops sp.)" 457:: an objective function, what we want to maximize, in this case energy over time as a currency of fitness 3622: 3342: 2829: 2546: 2443: 516:. Although this is controversial, coming under some of the same kinds of attack as the application of 424:. Optimal foraging theory (OFT) was first proposed in 1966, in two papers published independently, by 3898: 3803: 3415: 3307: 3165: 3150: 3145: 2824: 2536: 1008:"Genetic localization of foraging (for): a major gene for larval behavior in Drosophila melanogaster" 648: 141: 80:
that studies the foraging behavior of animals in response to the environment where the animal lives.
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In recent decades, optimal foraging theory has often been applied to the foraging behavior of human
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Rutz, C.; et al. (2010). "The ecological significance of tool use in New Caledonian Crows".
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Hutchings, Michael R.; Athanasiadou, Spiridoula; Kyriazakis, Ilias; J. Gordon, Iain (May 2003).
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Ahrens, Robert N M; Walters, Carl J; Christensen, Villy (March 2012). "Foraging arena theory".
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1 Gomes 2 Boesch, 1 CM 2 C (2011). "Reciprocity and trades in wild west African chimpanzees".
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benefit to becoming involved through allowing successful hunters first access to their kills.
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South West Outdoor Travelers- Wild Edibles, Medicinals, Foraging, Primitive Skills & More
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The Association of Foragers: An international association for teachers of foraging skills.
463:: set of choices under the organism's control, or the decisions that the organism exhibits 8: 3863: 3838: 3702: 3672: 3617: 3530: 3420: 3405: 3352: 3185: 3120: 3002: 2932: 2864: 2463: 2023:
Packer, C.; Scheel, D.; Pusey, A.E. (1990). "Why lions form groups: food is not enough".
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Group foraging may be influenced by the size of a group. In some species like lions and
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Several factors affect an animal's ability to forage and acquire profitable resources.
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because it plays an important role in an animal's ability to survive and reproduce.
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Some behaviors are more dominant than others. In a study using fruit fly larvae (
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MacArthur RH, Pianka ER (1966), "On the optimal use of a patchy environment.",
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Pereira, H S; MacDonald, D E; Hilliker, A J; Sokolowski, M B (September 1995).
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ISBN The pinnipeds: seals, sea lions, and walruses By Marianne Riedman 1990.
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theory to human behavior, it does represent a convergence of ideas from
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Emlen, J. M. (1966), "The role of time and energy in food preference",
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The theory scientists use to understand group foraging is called the
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Another measure of learning is spatio-temporal learning (also called
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if their behavior doesn't perfectly match what is predicted by OFT.
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The theory scientists use to understand solitary foraging is called
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Institute for the Study of Edible Wild Plants and Other Foragables
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de Belle, J S; Hilliker, A J; Sokolowski, M B (September 1989).
415: 659: 502: 401: 373: 231: 58: 36: 140:, and then proceeded to an extensive study of foraging in the 1898:"Wild chimpanzees exchange meat for sex on a long term basis" 1861:
Boesch, C (1994). "Cooperative hunting in wild Chimpanzees".
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detrimental for them to do so (called a dispersion economy).
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is searching for wild food resources. It affects an animal's
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10.1890/0012-9658(2000)081[0201:CQRQFC]2.0.CO;2
948:"Behavioral genomics of honeybee foraging and nest defense" 592: 1005: 35:. For the material that is eaten by foraging animals, see 1810:
Search and Foraging: Individual Motion and Swarm Dynamics
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Stephens, D.W.; Brown, J.S. & Ydenberg, R.C. (2007).
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Proceedings of the Royal Society B: Biological Sciences
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correlation between learning and foraging performance.
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to feed on fish that bury themselves in the sediment,
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Liang, Tong; Brinkman, Braden A. W. (14 March 2022).
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Danchin, E.; Giraldeau, L. & Cezilly, F. (2008).
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Evolutionary Anthropology: Issues, News, and Reviews
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Berkeley: University of California Press. 2051: 2003:: CS1 maint: numeric names: authors list ( 1954:: CS1 maint: numeric names: authors list ( 918:Journal of the Kansas Entomological Society 915: 3303:Latitudinal gradients in species diversity 3089: 3075: 2342: 2328: 2294:The Big Green Idea Wild Foraging Factsheet 2100: 945: 854: 852: 850: 848: 610:Theorizing on hominid foraging during the 528:that has proved fruitful and interesting. 1931: 1921: 1729: 1563: 1553: 1521: 1299: 1289: 1204: 1168: 1080: 1031: 979: 793: 725: 702: 700: 698: 696: 694: 3201:Predator–prey (Lotka–Volterra) equations 2840:Tritrophic interactions in plant defense 2237: 1896:1. Gomes 2. Boesch, 1. C M 2. C (2009). 1788: 1786: 1784: 1782: 1780: 1778: 1776: 1774: 1586: 1324:"Foraging Strategies | Encyclopedia.com" 858: 734:Behavioural Mechanisms of Food Selection 621: 586: 544: 367: 270: 221: 152: 41: 23:. For a prominent American forager, see 3233:Random generalized Lotka–Volterra model 1797:. Chicago: University of Chicago Press. 1637: 1631: 1496:"Search Strategies of Foraging Animals" 1372: 1346: 1340: 877: 845: 763: 761: 3886: 3041:Herbivore adaptations to plant defense 1860: 1407: 1182: 1180: 941: 939: 731: 691: 3070: 2323: 1808:Kagan, Eugene; Ben-Gal, Irad (2015). 1771: 1489: 1487: 1001: 999: 711:. New York: Oxford University Press. 149:Factors influencing foraging behavior 92:, any organism that consumes others, 3056:Predator avoidance in schooling fish 1580: 1193:Proceedings of the Nutrition Society 911: 909: 907: 758: 335: 326: 3506:Intermediate disturbance hypothesis 1971:Behavioral Ecology and Sociobiology 1381:Behavioral Ecology and Sociobiology 1177: 936: 583:Cost and benefits of group foraging 13: 3259:Ecological effects of biodiversity 1484: 996: 445: 348: 14: 3910: 2595:Generalist and specialist species 2267: 904: 768:Raine, N.E.; Chittka, L. (2008). 531: 16:Searching for wild food resources 3318:Occupancy–abundance relationship 2248:10.1016/B978-008045405-4.00026-4 2190:10.1111/j.1467-2979.2011.00432.x 1251:10.1111/j.1365-2907.2006.00080.x 946:Hunt, G.J.; et al. (2007). 473:Some of these versions include: 3338:Relative abundance distribution 3051:Plant defense against herbivory 2918:Competitive exclusion principle 2630:Mesopredator release hypothesis 2231: 2196: 2161: 2129: 1962: 1889: 1854: 1826: 1801: 1459: 1316: 1257: 1229: 1140: 1097: 1048: 675:LĂ©vy flight foraging hypothesis 100:, concentrations of resources. 2923:Consumer–resource interactions 1795:Foraging: Behavior and Ecology 732:Hughes, Roger N., ed. (1990). 53:) mother and cubs foraging in 1: 3769:Biological data visualization 3596:Environmental niche modelling 3323:Population viability analysis 1126:10.1016/j.fishres.2018.02.014 861:Principles of Animal Behavior 685: 380:with a tool, a prepared stick 358:Tool use in solitary foraging 312:Interactions with environment 290: 3254:Density-dependent inhibition 2279:Forager's Buddy GPS Foraging 2115:10.1016/0003-3472(95)80048-4 1923:10.1371/journal.pone.0005116 1555:10.1371/journal.pone.0022243 1291:10.1371/journal.pcbi.1009934 19:For foraging by humans, see 7: 3723:Liebig's law of the minimum 3558:Resource selection function 2449:Metabolic theory of ecology 2225:10.1016/j.jembe.2016.12.001 1417:Journal of Chemical Ecology 859:Dugatkin, Lee Alan (2004). 642: 217: 176: 10: 3915: 3623:Niche apportionment models 3343:Relative species abundance 2547:Primary nutritional groups 2444:List of feeding behaviours 2137:Aurignacian Lithic Economy 1347:Riedman, Marianne (1990). 1270:PLOS Computational Biology 1073:10.1093/genetics/141.1.263 1024:10.1093/genetics/123.1.157 361: 343:Pogonomyrmex vermiculatus 262:) gene. During the study, 18: 3872: 3804:Ecosystem based fisheries 3746: 3646: 3571: 3444: 3416:Interspecific competition 3381: 3308:Minimum viable population 3241: 3166:Maximum sustainable yield 3151:Intraspecific competition 3146:Effective population size 3109: 3026:Anti-predator adaptations 3011: 2890: 2817: 2774: 2696: 2663: 2560: 2537:Photosynthetic efficiency 2472: 2366: 2305:Caress, Badiday. (2000), 1983:10.1007/s00265-011-1227-x 1437:10.1007/s10886-006-9240-7 1393:10.1007/s00265-009-0762-1 972:10.1007/s00114-006-0183-1 742:10.1007/978-3-642-75118-9 649:List of forageable plants 142:common pied oystercatcher 3794:Ecological stoichiometry 3759:Alternative stable state 3638:Ontogenetic niche shift 3501:Ideal free distribution 3411:Ecological facilitation 3161:Malthusian growth model 3131:Consumer-resource model 2988:Paradox of the plankton 2953:Energy systems language 2673:Chemoorganoheterotrophy 2640:Optimal foraging theory 2615:Heterotrophic nutrition 2240:Encyclopedia of Ecology 1738:The American Naturalist 1609:10.1126/science.1192053 637:Ideal free distribution 539:Artificial Intelligence 422:optimal foraging theory 400:that use sticks to get 251:Drosophila melanogaster 51:Ursus arctos horribilis 3784:Ecological forecasting 3728:Marginal value theorem 3526:Landscape epidemiology 3461:Cross-boundary subsidy 3396:Biological interaction 2746:Microbial intelligence 2434:Green world hypothesis 2242:. pp. 2561–2566. 1875:10.1006/anbe.1994.1285 1467:"About Forest to Food" 898:10.1006/anbe.1996.0330 786:10.1098/rspb.2007.1652 596: 558: 499:Central place foraging 493:marginal value theorem 486:Patch selection theory 478:The optimal diet model 381: 235: 170: 113:marginal value theorem 62: 3789:Ecological humanities 3688:Ecological energetics 3633:Niche differentiation 3496:Habitat fragmentation 3264:Ecological extinction 3211:Small population size 2963:Feed conversion ratio 2943:Ecological succession 2875:San Francisco Estuary 2789:Ecological efficiency 2731:Microbial cooperation 1848:10.1051/apido:2000109 930:10.2317/JKES-704.23.1 622:Foraging Arena Theory 590: 548: 526:economic anthropology 371: 271:Presence of predators 225: 156: 45: 3814:Evolutionary ecology 3779:Ecological footprint 3774:Ecological economics 3698:Ecological threshold 3693:Ecological indicator 3563:Source–sink dynamics 3516:Land change modeling 3511:Insular biogeography 3363:Species distribution 3102:Modelling ecosystems 2761:Microbial metabolism 2600:Intraguild predation 2389:Biogeochemical cycle 2355:Modelling ecosystems 1328:www.encyclopedia.com 568:Melipona scutellaris 398:New Caledonian crows 210:Studies showed that 206:Trigona fulviventris 111:, who developed the 55:Denali National Park 3864:Theoretical ecology 3839:Natural environment 3703:Ecosystem diversity 3673:Ecological collapse 3663:Bateman's principle 3618:Limiting similarity 3531:Landscape limnology 3353:Species homogeneity 3191:Population modeling 3186:Population dynamics 3003:Trophic state index 2217:2017JEMBE.487...86T 2182:2012AqFF...13...41A 2072:2009ASAJ..125..125B 2025:American Naturalist 1914:2009PLoSO...4.5116G 1695:American Naturalist 1655:1964Natur.201.1264G 1601:2010Sci...329.1523R 1546:2011PLoSO...622243P 1515:1990AmSci..78..152O 1429:2007JCEco..33..435T 1282:2022PLSCB..18E9934L 1118:2018FishR.204..172R 964:2007NW.....94..247H 952:Naturwissenschaften 709:Behavioural Ecology 574:Chimpanzees in the 364:Tool use by animals 201:time-place learning 3875:Outline of ecology 3824:Industrial ecology 3819:Functional ecology 3683:Ecological deficit 3628:Niche construction 3591:Ecosystem engineer 3368:Species–area curve 3289:Introduced species 3104:: Other components 3036:Deimatic behaviour 2938:Ecological network 2870:North Pacific Gyre 2855:hydrothermal vents 2794:Ecological pyramid 2741:Microbial food web 2552:Primary production 2497:Foundation species 2312:2013-05-31 at the 2299:2009-02-25 at the 2170:Fish and Fisheries 1503:American Scientist 1206:10.1079/pns2003243 1106:Fisheries Research 831:10.1002/evan.20180 651:(edible by humans) 597: 576:TaĂŻ Forest in CĂ´te 563:red harvester ants 559: 404:out of trees, and 382: 236: 228:European honey bee 171: 78:behavioral ecology 63: 3881: 3880: 3764:Balance of nature 3521:Landscape ecology 3406:Community ecology 3348:Species diversity 3284:Indicator species 3279:Gradient analysis 3156:Logistic function 3064: 3063: 3021:Animal coloration 2998:Trophic mutualism 2736:Microbial ecology 2527:Photoheterotrophs 2512:Myco-heterotrophy 2424:Ecosystem ecology 2409:Carrying capacity 2374:Abiotic component 2257:978-0-08-045405-4 2154:978-0-306-46334-1 2080:10.1121/1.2967480 1819:978-1-4822-4210-2 1663:10.1038/2011264a0 1649:(4926): 1264–66. 1595:(5998): 1523–26. 1362:978-0-520-06497-3 870:978-0-393-97659-5 751:978-3-642-75120-2 718:978-0-19-920629-2 551:northern cardinal 384:Some examples of 336:Solitary foraging 327:Types of foraging 281:foraging behavior 242:Apoica flavissima 134:John Goss-Custard 3906: 3899:Eating behaviors 3581:Ecological niche 3553:selection theory 3373:Umbrella species 3358:Species richness 3294:Invasive species 3274:Flagship species 3181:Population cycle 3176:Overexploitation 3141:Ecological yield 3091: 3084: 3077: 3068: 3067: 2973:Mesotrophic soil 2913:Climax community 2845:Marine food webs 2784:Biomagnification 2585:Chemoorganotroph 2439:Keystone species 2399:Biotic component 2344: 2337: 2330: 2321: 2320: 2262: 2261: 2235: 2229: 2228: 2200: 2194: 2193: 2165: 2159: 2158: 2133: 2127: 2126: 2109:(5): 1325–1339. 2103:Animal Behaviour 2098: 2092: 2091: 2055: 2049: 2048: 2020: 2009: 2008: 2002: 1994: 1966: 1960: 1959: 1953: 1945: 1935: 1925: 1893: 1887: 1886: 1863:Animal Behaviour 1858: 1852: 1851: 1830: 1824: 1823: 1805: 1799: 1798: 1790: 1769: 1768: 1733: 1727: 1726: 1689: 1683: 1682: 1635: 1629: 1628: 1584: 1578: 1577: 1567: 1557: 1525: 1519: 1518: 1500: 1491: 1482: 1481: 1479: 1478: 1463: 1457: 1456: 1411: 1405: 1404: 1387:(8): 1097–1107. 1376: 1370: 1369: 1354: 1344: 1338: 1337: 1335: 1334: 1320: 1314: 1313: 1303: 1293: 1261: 1255: 1254: 1233: 1227: 1226: 1208: 1184: 1175: 1174: 1172: 1144: 1138: 1137: 1101: 1095: 1094: 1084: 1052: 1046: 1045: 1035: 1003: 994: 993: 983: 943: 934: 933: 913: 902: 901: 886:Animal Behaviour 881: 875: 874: 856: 843: 842: 814: 808: 807: 797: 780:(1636): 803–08. 765: 756: 755: 729: 723: 722: 704: 514:hunter-gatherers 426:Robert MacArthur 320:Blepharida rhois 3914: 3913: 3909: 3908: 3907: 3905: 3904: 3903: 3884: 3883: 3882: 3877: 3868: 3854:Systems ecology 3742: 3713:Extinction debt 3678:Ecological debt 3668:Bioluminescence 3649: 3642: 3611:marine habitats 3586:Ecological trap 3567: 3447: 3440: 3383: 3377: 3333:Rapoport's rule 3328:Priority effect 3269:Endemic species 3237: 3196:Population size 3112: 3105: 3095: 3065: 3060: 3013: 3007: 2993:Trophic cascade 2903:Bioaccumulation 2886: 2813: 2770: 2692: 2659: 2556: 2468: 2429:Ecosystem model 2362: 2348: 2314:Wayback Machine 2301:Wayback Machine 2270: 2265: 2258: 2236: 2232: 2201: 2197: 2166: 2162: 2155: 2145:10.1007/b110584 2135: 2134: 2130: 2099: 2095: 2056: 2052: 2021: 2012: 1996: 1995: 1977:(11): 2183–96. 1967: 1963: 1947: 1946: 1894: 1890: 1859: 1855: 1831: 1827: 1820: 1806: 1802: 1791: 1772: 1744:(916): 611–17, 1734: 1730: 1702:(916): 603–09, 1690: 1686: 1636: 1632: 1585: 1581: 1526: 1522: 1498: 1492: 1485: 1476: 1474: 1465: 1464: 1460: 1412: 1408: 1377: 1373: 1363: 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W.W. Norton. 857: 846: 815: 811: 766: 759: 752: 730: 726: 719: 705: 692: 688: 655:Chesson's index 645: 633: 624: 585: 534: 518:sociobiological 448: 446:Versions of OFT 434:decision theory 418: 366: 360: 351: 349:Search Behavior 338: 329: 314: 306:self-medicating 293: 273: 220: 212:T. fulviventris 179: 167:Laikipia, Kenya 151: 76:is a branch of 74:Foraging theory 40: 21:Hunter-gatherer 17: 12: 11: 5: 3912: 3902: 3901: 3896: 3879: 3878: 3873: 3870: 3869: 3867: 3866: 3861: 3856: 3851: 3846: 3841: 3836: 3834:Microecosystem 3831: 3826: 3821: 3816: 3811: 3806: 3801: 3796: 3791: 3786: 3781: 3776: 3771: 3766: 3761: 3756: 3750: 3748: 3744: 3743: 3741: 3740: 3735: 3733:Thorson's rule 3730: 3725: 3720: 3715: 3710: 3705: 3700: 3695: 3690: 3685: 3680: 3675: 3670: 3665: 3660: 3658:Assembly rules 3654: 3652: 3644: 3643: 3641: 3640: 3635: 3630: 3625: 3620: 3615: 3614: 3613: 3603: 3598: 3593: 3588: 3583: 3577: 3575: 3569: 3568: 3566: 3565: 3560: 3555: 3543: 3541:Patch dynamics 3538: 3536:Metapopulation 3533: 3528: 3523: 3518: 3513: 3508: 3503: 3498: 3493: 3488: 3483: 3478: 3473: 3468: 3463: 3458: 3452: 3450: 3442: 3441: 3439: 3438: 3433: 3431:Storage effect 3428: 3423: 3418: 3413: 3408: 3403: 3398: 3393: 3387: 3385: 3379: 3378: 3376: 3375: 3370: 3365: 3360: 3355: 3350: 3345: 3340: 3335: 3330: 3325: 3320: 3315: 3313:Neutral theory 3310: 3305: 3300: 3298:Native species 3291: 3286: 3281: 3276: 3271: 3266: 3261: 3256: 3251: 3245: 3243: 3239: 3238: 3236: 3235: 3230: 3229: 3228: 3223: 3213: 3208: 3203: 3198: 3193: 3188: 3183: 3178: 3173: 3171:Overpopulation 3168: 3163: 3158: 3153: 3148: 3143: 3138: 3133: 3128: 3123: 3117: 3115: 3107: 3106: 3094: 3093: 3086: 3079: 3071: 3062: 3061: 3059: 3058: 3053: 3048: 3043: 3038: 3033: 3028: 3023: 3017: 3015: 3009: 3008: 3006: 3005: 3000: 2995: 2990: 2985: 2980: 2978:Nutrient cycle 2975: 2970: 2968:Feeding frenzy 2965: 2960: 2955: 2950: 2948:Energy quality 2945: 2940: 2935: 2930: 2925: 2920: 2915: 2910: 2908:Cascade effect 2905: 2900: 2894: 2892: 2888: 2887: 2885: 2884: 2883: 2882: 2877: 2872: 2867: 2862: 2857: 2852: 2842: 2837: 2832: 2827: 2821: 2819: 2815: 2814: 2812: 2811: 2806: 2801: 2796: 2791: 2786: 2780: 2778: 2772: 2771: 2769: 2768: 2763: 2758: 2753: 2751:Microbial loop 2748: 2743: 2738: 2733: 2728: 2723: 2718: 2716:Lithoautotroph 2713: 2708: 2702: 2700: 2698:Microorganisms 2694: 2693: 2691: 2690: 2685: 2680: 2675: 2669: 2667: 2661: 2660: 2658: 2657: 2655:Prey switching 2652: 2647: 2642: 2637: 2632: 2627: 2622: 2617: 2612: 2607: 2602: 2597: 2592: 2587: 2582: 2577: 2572: 2566: 2564: 2558: 2557: 2555: 2554: 2549: 2544: 2539: 2534: 2532:Photosynthesis 2529: 2524: 2519: 2514: 2509: 2504: 2499: 2494: 2489: 2487:Chemosynthesis 2484: 2478: 2476: 2470: 2469: 2467: 2466: 2461: 2456: 2451: 2446: 2441: 2436: 2431: 2426: 2421: 2416: 2411: 2406: 2401: 2396: 2391: 2386: 2381: 2379:Abiotic stress 2376: 2370: 2368: 2364: 2363: 2347: 2346: 2339: 2332: 2324: 2318: 2317: 2303: 2291: 2286: 2281: 2276: 2269: 2268:External links 2266: 2264: 2263: 2256: 2230: 2195: 2160: 2153: 2128: 2093: 2066:(1): 125–137. 2050: 2037:10.1086/285079 2010: 1961: 1888: 1853: 1825: 1818: 1800: 1770: 1750:10.1086/282455 1728: 1708:10.1086/282454 1684: 1630: 1579: 1520: 1509:(2): 152–160. 1483: 1458: 1406: 1371: 1361: 1339: 1315: 1256: 1228: 1199:(2): 361–370. 1176: 1155:(1): 201–219. 1139: 1096: 1067:(1): 263–270. 1047: 1018:(1): 157–163. 995: 935: 903: 892:(3): 549–560. 876: 869: 844: 825:(4): 189–201. 809: 757: 750: 724: 717: 689: 687: 684: 683: 682: 677: 672: 667: 665:Avian foraging 662: 657: 652: 644: 641: 632: 629: 623: 620: 584: 581: 533: 532:Group foraging 530: 471: 470: 464: 458: 447: 444: 417: 414: 359: 356: 350: 347: 337: 334: 328: 325: 313: 310: 292: 289: 272: 269: 219: 216: 178: 175: 165:) foraging in 150: 147: 146: 145: 131: 116: 15: 9: 6: 4: 3: 2: 3911: 3900: 3897: 3895: 3892: 3891: 3889: 3876: 3871: 3865: 3862: 3860: 3859:Urban ecology 3857: 3855: 3852: 3850: 3847: 3845: 3842: 3840: 3837: 3835: 3832: 3830: 3827: 3825: 3822: 3820: 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Index

Hunter-gatherer
Euell Gibbons
Samuel Thayer
Ethnobotany
Forage

Grizzly bear
Denali National Park
Alaska
fitness
Foraging theory
behavioral ecology
Eric Charnov
marginal value theorem
John Krebs
tits
chickadees
John Goss-Custard
redshank
common pied oystercatcher

olive baboons
Laikipia, Kenya
Learning
primates
time-place learning
Trigona fulviventris

European honey bee
nectar

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