327:
85:
31:
59:
838:
376:
It has subsequently been discovered that the browsing herds of giant tortoises filled an essential role in the island's ecosystem and the regeneration of its forests. In recognition of this fact, measures have been undertaken to introduce replacements, in the form of similar species of tortoises from
364:
At the time of the arrival of human settlers, dense giant tortoise herds of many thousands were reported on
Rodrigues. Like many island species, they were reported to have been friendly, curious, and unafraid of humans. However, in the ensuing years, massive harvesting and exporting for food, as well
372:
A surviving giant tortoise was reported on
Rodrigues in 1795, at the bottom of a ravine. As late as 1802, there is mention of survivors reportedly being killed in the large fires used to clear the island's vegetation for agriculture.
338:
The domed
Rodrigues giant tortoise was one of the smallest of the giant tortoises of the Indian Ocean, reaching a length of just over 40 cm (16 in) and an estimated weight of around 12 kg (26 lb).
4033:
887:
369:, rapidly exterminated the giant tortoises. Hundreds of thousands were loaded into ships' holds for food, or to be transported to Mauritius, where they were burnt for fat and oil.
377:
other parts of the world, to assist in the rebuilding of
Rodrigues' devastated environment. The replacement species for the domed Rodrigues tortoise was chosen to be the
1170:
1043:
1182:
1176:
1730:
1724:
1682:
1742:
1688:
1382:
1364:
1037:
1718:
1388:
1370:
1799:
1754:
1736:
1700:
1376:
1025:
1760:
1394:
1358:
1712:
4128:
1748:
550:
1706:
1694:
1676:
4123:
1161:
2750:
2661:
686:
2669:
2423:
507:
Pedrono, Miguel; Griffiths, Owen L.; Clausen, Alison; Smith, Lora L.; Griffiths, Christine J.; Wilmé, Lucienne; Burney, David A. (2013).
634:
4103:
660:
1083:
4093:
1272:
1266:
739:
343:
4074:
4003:
Turtles of the World 2017 Update: Annotated
Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status
1812:
turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status
665:
1843:
4118:
356:), which colonised Rodrigues by sea many millions of years ago, and then differentiated into the two Rodrigues species.
4001:
1811:
894:
691:
625:
1125:
3876:
2959:
1508:
1323:
1248:
1120:
879:
1493:
1483:
1145:
3853:
4108:
4012:
2535:
1919:
1633:
1498:
1150:
554:
84:
4048:
2268:
1488:
1094:
732:
4067:
1503:
911:
334:
and a lizard on a palm trunk. The giant tortoises are most likely the domed
Rodrigues giant tortoise.
1901:
1260:
4113:
2914:
1836:
1414:
1135:
1104:
1073:
961:
3770:
1783:
1333:
704:
456:
433:
346:, which browsed the taller vegetation. Both species were descended from an ancestral species on
3742:
2542:
2500:
2140:
1140:
1078:
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1007:
979:
973:
967:
942:
872:
587:
4133:
4098:
2486:
1988:
1793:
1778:
1587:
1575:
1544:
1460:
1290:
1223:
1130:
1110:
903:
725:
699:
203:
1058:
1053:
4060:
3728:
2394:
2018:
1295:
1031:
1019:
520:
299:
678:
330:
1707 illustration showing a settlement in
Rodrigues, with giant tortoises, rats, crabs, a
8:
3713:
2072:
1829:
1788:
1581:
1465:
1228:
1002:
509:"Using a surviving lineage of Madagascar's vanished megafauna for ecological restoration"
352:
331:
48:
524:
3120:
3085:
2952:
2691:
1605:
1569:
1549:
1455:
1349:
1068:
489:
342:
A low grazer of grasses, it shared
Rodrigues Island with its much larger relative, the
315:
228:
79:
3663:
2646:
2598:
2589:
2472:
2049:
1610:
1437:
1432:
1200:
1089:
935:
930:
508:
481:
378:
318:. It appears to have become extinct around 1800, as a result of human exploitation.
493:
410:
3455:
2843:
2818:
2779:
2727:
2458:
2003:
1656:
1526:
1218:
1098:
854:
573:
Lost Land of the Dodo, An
Ecological History of Mauritius, Réunion & Rordrigues
528:
471:
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366:
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3551:
3280:
3248:
2885:
2794:
2570:
2156:
1770:
1638:
1328:
1318:
1313:
1114:
532:
4044:
3798:
3788:
3685:
3535:
3375:
3077:
3022:
2764:
2699:
2239:
2230:
2199:
2107:
1996:
1910:
1625:
1405:
1338:
997:
326:
30:
4087:
3978:
3861:
3761:
3750:
3721:
3635:
3519:
3503:
3463:
3397:
3307:
3272:
3216:
3200:
3184:
3158:
3146:
3130:
3108:
3100:
2907:
2618:
2451:
2312:
2259:
2207:
1615:
1518:
864:
797:
673:
610:
485:
419:
156:
69:
3913:
3838:
3814:
3331:
3208:
3192:
3168:
3069:
3014:
2938:
2854:
2828:
2742:
2653:
2632:
2556:
2521:
2331:
2297:
2164:
2080:
2026:
1648:
1560:
1239:
953:
803:
536:
307:
303:
179:
3891:
3846:
3692:
3677:
3599:
3591:
3583:
3559:
3439:
3423:
3407:
3323:
3315:
3264:
3240:
3232:
3224:
3092:
3044:
2967:
2892:
2757:
2713:
2639:
2625:
2577:
2434:
2352:
2290:
2283:
2276:
2191:
2133:
2125:
2042:
2034:
1981:
1971:
1597:
1447:
989:
922:
647:
619:
476:
3970:
3883:
3869:
3830:
3607:
3527:
3495:
3479:
3447:
3385:
3358:
3339:
3256:
3061:
2981:
2974:
2922:
2899:
2809:
2676:
2408:
2215:
2148:
2065:
2057:
1960:
1951:
1282:
1210:
846:
791:
785:
639:
386:
146:
63:
652:
3962:
3822:
3806:
3735:
3699:
3649:
3642:
3567:
3543:
3487:
3431:
3138:
2930:
2878:
2836:
2549:
2507:
2415:
2401:
2387:
2380:
2373:
2366:
2338:
2183:
2011:
347:
96:
581:
408:
3777:
3706:
3626:
3615:
3511:
3176:
3053:
2945:
2787:
2772:
2734:
2683:
2609:
2514:
2493:
2479:
2465:
2321:
2248:
2117:
2096:
2088:
1870:
1667:
1536:
1305:
1192:
823:
767:
748:
604:
311:
166:
116:
717:
3921:
3670:
3575:
3471:
3415:
2995:
2988:
2870:
2720:
2706:
2563:
2528:
2443:
2345:
1885:
1876:
1475:
1424:
779:
773:
126:
4040:
3656:
3299:
1891:
1864:
1852:
761:
136:
106:
404:
402:
4032:
2863:
1930:
1821:
506:
2359:
399:
454:
820:
4085:
567:
565:
4068:
4000:Phylogenetic arrangement of turtles based on
2238:
1837:
1810:Phylogenetic arrangement of turtles based on
733:
562:
409:World Conservation Monitoring Centre (1996).
434:10.2305/IUCN.UK.1996.RLTS.T6063A12388776.en
4075:
4061:
3043:
1970:
1909:
1844:
1830:
740:
726:
57:
29:
475:
448:
432:
359:
4129:Species made extinct by human activities
325:
389:, which is similar in size and habits.
4124:Species endangered by invasive species
4086:
1273:Saddle-backed Rodrigues giant tortoise
1267:Saddle-backed Mauritius giant tortoise
553:. The Sixth Extinction. Archived from
457:"Checklist of Chelonians of the World"
344:saddle-backed Rodrigues giant tortoise
1918:
1825:
721:
586:
585:
4027:
747:
420:IUCN Red List of Threatened Species
13:
14:
4145:
575:. T & AD Poyser, London.2008.
4031:
836:
83:
3946:
3366:
455:Fritz Uwe; Peter Havaš (2007).
262:Geochelone (Geochelone) rotunda
24:Domed Rodrigues giant tortoise
4104:Reptile extinctions since 1500
1851:
1255:Domed Rodrigues giant tortoise
1249:Domed Mauritius giant tortoise
1121:Santiago Island giant tortoise
543:
500:
321:
292:domed Rodrigues giant tortoise
1:
4094:IUCN Red List extinct species
1074:Chatham Island giant tortoise
392:
4047:. You can help Knowledge by
1141:Volcán Darwin giant tortoise
1136:Volcán Alcedo giant tortoise
1126:Pinzón Island giant tortoise
533:10.1016/j.biocon.2012.11.027
7:
4013:List of Testudines families
1634:Flat-backed spider tortoise
1509:Speke's hinge-back tortoise
1499:Lobatse hinge-back tortoise
1324:Goode's thornscrub tortoise
1151:Western Santa Cruz tortoise
1131:Sierra Negra giant tortoise
1079:Eastern Santa Cruz tortoise
16:Extinct species of tortoise
10:
4150:
4119:Reptiles described in 1835
4026:
1494:Home's hinge-back tortoise
1489:Forest hinge-back tortoise
1484:Bell's hinge-back tortoise
1146:Volcán Wolf giant tortoise
1095:Hood Island giant tortoise
1084:Fernandina Island tortoise
834:
257:Duméril & Bibron, 1835
222:Dumeril & Bibron, 1835
3996:
3953:
3939:
3901:
3787:
3760:
3625:
3395:
3373:
3357:
3290:
3156:
3118:
3036:
3005:
2853:
2817:
2808:
2608:
2587:
2433:
2320:
2311:
2258:
2229:
2174:
2106:
1959:
1950:
1928:
1927:
1900:
1859:
1808:
1769:
1665:
1647:
1624:
1596:
1558:
1535:
1517:
1504:Natal hinge-back tortoise
1474:
1446:
1423:
1403:
1347:
1304:
1281:
1237:
1209:
1191:
1159:
1069:Cerro Azul giant tortoise
1052:
988:
952:
921:
912:Aldabrachelys grandidieri
863:
845:
819:
812:
756:
594:
559:Retrieved on 15 May 2015.
234:
227:
209:
202:
80:Scientific classification
78:
55:
46:
37:
28:
23:
1415:Hesperotestudo percrassa
1171:Cheirogaster bacharidisi
1105:Santa Fe Island tortoise
1090:Floreana Island tortoise
1044:Chelonoidis sombrerensis
962:African spurred tortoise
551:"Cylindraspis peltastes"
1334:Sonoran Desert tortoise
513:Biological Conservation
365:as the introduction of
1261:Réunion giant tortoise
1183:Cheirogaster schafferi
1177:Cheirogaster gymnesica
1014:Chelonoidis alburyorum
1008:Yellow-footed tortoise
980:Centrochelys vulcanica
974:Centrochelys burchardi
968:Centrochelys atlantica
943:Astrochelys rogerbouri
873:Aldabra giant tortoise
626:Cylindraspis peltastes
596:Cylindraspis peltastes
413:Cylindraspis peltastes
360:Ecology and extinction
335:
296:Cylindraspis peltastes
278:Cylindraspis peltastes
216:Cylindraspis peltastes
4039:This article about a
1794:Spur-thighed tortoise
1731:Stylemys nebrascensis
1725:Stylemys karakolensis
1683:Stylemys calaverensis
1588:Megalochelys sondaari
1576:Megalochelys cautleyi
1545:Asian forest tortoise
1291:Burmese star tortoise
1111:Pinta Island tortoise
904:Aldabrachelys abrupta
700:Paleobiology Database
571:Cheke, A., Hume, J.:
329:
1784:Kleinmann's tortoise
1743:Stylemys oregonensis
1689:Stylemys canetotiana
1383:Hadrianus schucherti
1365:Hadrianus majusculus
1296:Indian star tortoise
1038:Chelonoidis sellovii
1032:Chelonoidis monensis
1020:Chelonoidis cubensis
477:10.3897/vz.57.e30895
427:: e.T6063A12388776.
270:Geochelone peltastes
66: (around 1800)
1789:Marginated tortoise
1719:Stylemys frizaciana
1582:Megalochelys margae
1466:Travancore tortoise
1389:Hadrianus utahensis
1371:Hadrianus octonaria
1059:Chelonoidis complex
1003:Red-footed tortoise
525:2013BCons.159..501P
383:Astrochelys radiata
353:Cylindraspis inepta
332:Rodrigues solitaire
49:Conservation status
4109:Fauna of Rodrigues
3121:Nanhsiungchelyidae
3086:Macrocephalochelys
1800:Testudo brevitesta
1779:Hermann's tortoise
1755:Stylemys uintensis
1737:Stylemys neglectus
1701:Stylemys conspecta
1606:Geometric tortoise
1570:Megalochelys atlas
1550:Impressed tortoise
1461:Forsten's tortoise
1456:Elongated tortoise
1377:Hadrianus robustus
1054:Galápagos tortoise
1026:Chelonoidis lutzae
464:Vertebrate Zoology
336:
38:Illustration from
4056:
4055:
4021:
4020:
3992:
3991:
3988:
3987:
3935:
3934:
3931:
3930:
3353:
3352:
3349:
3348:
3032:
3031:
2804:
2803:
2307:
2306:
2225:
2224:
1946:
1945:
1942:
1941:
1938:
1937:
1819:
1818:
1814:. Key: †=extinct.
1761:Stylemys undabuna
1611:Serrated tortoise
1395:Hadrianus vialovi
1359:Hadrianus corsoni
1229:Speckled padloper
1201:Angulate tortoise
936:Radiated tortoise
931:Angonoka tortoise
832:
831:
715:
714:
588:Taxon identifiers
379:radiated tortoise
288:
287:
282:
274:
266:
258:
254:Testudo peltastes
250:
242:
195:C. peltastes
73:
42:1770 by Jossigny
4141:
4077:
4070:
4063:
4035:
4028:
3951:
3950:
3944:
3943:
3456:Chupacabrachelys
3371:
3370:
3364:
3363:
3041:
3040:
2819:Carettochelyidae
2815:
2814:
2318:
2317:
2236:
2235:
1968:
1967:
1957:
1956:
1925:
1924:
1916:
1915:
1907:
1906:
1846:
1839:
1832:
1823:
1822:
1713:Stylemys emiliae
1657:Leopard tortoise
1527:Pancake tortoise
1438:Greater padloper
855:Russian tortoise
840:
839:
817:
816:
742:
735:
728:
719:
718:
708:
707:
695:
694:
682:
681:
669:
668:
656:
655:
643:
642:
630:
629:
628:
615:
614:
613:
583:
582:
576:
569:
560:
558:
557:on 13 July 2013.
547:
541:
540:
504:
498:
497:
479:
461:
452:
446:
445:
443:
441:
436:
406:
367:invasive species
350:(an ancestor of
280:
272:
264:
256:
248:
246:Chersine rotunda
240:
218:
214:
178:
88:
87:
67:
61:
60:
33:
21:
20:
4149:
4148:
4144:
4143:
4142:
4140:
4139:
4138:
4114:Extinct turtles
4084:
4083:
4082:
4081:
4024:
4022:
4017:
3984:
3927:
3904:Sahonachelyidae
3897:
3783:
3756:
3621:
3552:Phosphatochelys
3391:
3345:
3286:
3281:Terlinguachelys
3249:Pneumatoarthrus
3152:
3114:
3028:
3001:
2849:
2800:
2604:
2583:
2571:Siebenrockiella
2429:
2303:
2254:
2221:
2170:
2157:Mesodermochelys
2102:
1963:
1934:
1896:
1855:
1850:
1820:
1815:
1804:
1765:
1749:Stylemys pygmea
1661:
1643:
1639:Spider tortoise
1620:
1592:
1554:
1531:
1513:
1470:
1442:
1433:Common padloper
1419:
1399:
1343:
1329:Gopher tortoise
1319:Desert tortoise
1314:Bolson tortoise
1300:
1277:
1233:
1205:
1187:
1155:
1115:Lonesome George
1048:
984:
948:
917:
895:A. g. hololissa
859:
841:
837:
828:
821:Species of the
808:
752:
746:
716:
711:
703:
698:
690:
685:
677:
672:
664:
659:
651:
646:
638:
633:
624:
623:
618:
609:
608:
603:
590:
580:
579:
570:
563:
549:
548:
544:
505:
501:
459:
453:
449:
439:
437:
407:
400:
395:
362:
324:
273:Pritchard, 1967
265:Fitzinger, 1835
241:Latreille, 1801
238:Testudo rotunda
223:
220:
212:
211:
198:
176:
82:
74:
62:
58:
51:
17:
12:
11:
5:
4147:
4137:
4136:
4131:
4126:
4121:
4116:
4111:
4106:
4101:
4096:
4080:
4079:
4072:
4065:
4057:
4054:
4053:
4036:
4019:
4018:
4016:
4015:
4008:
4007:
4006:. † = extinct.
3997:
3994:
3993:
3990:
3989:
3986:
3985:
3983:
3982:
3974:
3966:
3957:
3955:
3948:
3941:
3937:
3936:
3933:
3932:
3929:
3928:
3926:
3925:
3917:
3908:
3906:
3899:
3898:
3896:
3895:
3887:
3880:
3873:
3865:
3857:
3850:
3842:
3834:
3826:
3818:
3810:
3802:
3799:Albertwoodemys
3793:
3791:
3789:Podocnemididae
3785:
3784:
3782:
3781:
3774:
3766:
3764:
3758:
3757:
3755:
3754:
3746:
3739:
3732:
3725:
3717:
3710:
3703:
3696:
3689:
3686:Lomalatachelys
3681:
3674:
3667:
3660:
3653:
3646:
3639:
3631:
3629:
3623:
3622:
3620:
3619:
3611:
3603:
3595:
3587:
3579:
3571:
3563:
3555:
3547:
3539:
3536:Labrostochelys
3531:
3523:
3515:
3507:
3499:
3491:
3483:
3475:
3467:
3459:
3451:
3443:
3435:
3427:
3419:
3411:
3402:
3400:
3393:
3392:
3390:
3389:
3380:
3378:
3376:Araripemydidae
3368:
3361:
3355:
3354:
3351:
3350:
3347:
3346:
3344:
3343:
3335:
3327:
3319:
3311:
3303:
3294:
3292:
3288:
3287:
3285:
3284:
3276:
3268:
3260:
3252:
3244:
3236:
3228:
3220:
3212:
3204:
3196:
3188:
3180:
3172:
3163:
3161:
3154:
3153:
3151:
3150:
3142:
3134:
3125:
3123:
3116:
3115:
3113:
3112:
3104:
3096:
3089:
3081:
3078:Emarginachelys
3073:
3065:
3057:
3049:
3047:
3038:
3034:
3033:
3030:
3029:
3027:
3026:
3023:Sinaspideretes
3018:
3009:
3007:
3003:
3002:
3000:
2999:
2992:
2985:
2978:
2971:
2963:
2956:
2949:
2942:
2934:
2926:
2918:
2911:
2903:
2896:
2889:
2882:
2874:
2867:
2859:
2857:
2851:
2850:
2848:
2847:
2840:
2832:
2823:
2821:
2812:
2806:
2805:
2802:
2801:
2799:
2798:
2791:
2783:
2776:
2768:
2761:
2754:
2746:
2738:
2731:
2724:
2717:
2710:
2703:
2700:Hesperotestudo
2695:
2687:
2680:
2673:
2665:
2657:
2650:
2643:
2636:
2629:
2622:
2614:
2612:
2606:
2605:
2603:
2602:
2594:
2592:
2590:Platysternidae
2585:
2584:
2582:
2581:
2574:
2567:
2560:
2553:
2546:
2539:
2532:
2525:
2518:
2511:
2504:
2497:
2490:
2483:
2476:
2469:
2462:
2455:
2447:
2439:
2437:
2431:
2430:
2428:
2427:
2419:
2412:
2405:
2398:
2391:
2384:
2377:
2370:
2363:
2356:
2349:
2342:
2335:
2326:
2324:
2315:
2309:
2308:
2305:
2304:
2302:
2301:
2294:
2287:
2280:
2272:
2264:
2262:
2256:
2255:
2253:
2252:
2244:
2242:
2240:Dermatemydidae
2233:
2231:Kinosternoidea
2227:
2226:
2223:
2222:
2220:
2219:
2211:
2203:
2200:Procolpochelys
2195:
2187:
2178:
2176:
2172:
2171:
2169:
2168:
2160:
2152:
2144:
2137:
2129:
2121:
2112:
2110:
2108:Dermochelyidae
2104:
2103:
2101:
2100:
2092:
2084:
2076:
2069:
2061:
2053:
2046:
2038:
2030:
2022:
2015:
2007:
2000:
1997:Carolinochelys
1992:
1985:
1976:
1974:
1965:
1954:
1948:
1947:
1944:
1943:
1940:
1939:
1936:
1935:
1922:
1913:
1904:
1898:
1897:
1895:
1894:
1888:
1879:
1873:
1867:
1860:
1857:
1856:
1849:
1848:
1841:
1834:
1826:
1817:
1816:
1809:
1806:
1805:
1803:
1802:
1796:
1791:
1786:
1781:
1775:
1773:
1767:
1766:
1764:
1763:
1757:
1751:
1745:
1739:
1733:
1727:
1721:
1715:
1709:
1707:Stylemys copei
1703:
1697:
1695:Stylemys capax
1691:
1685:
1679:
1677:Stylemys botti
1672:
1670:
1663:
1662:
1660:
1659:
1653:
1651:
1645:
1644:
1642:
1641:
1636:
1630:
1628:
1622:
1621:
1619:
1618:
1613:
1608:
1602:
1600:
1594:
1593:
1591:
1590:
1584:
1578:
1572:
1565:
1563:
1556:
1555:
1553:
1552:
1547:
1541:
1539:
1533:
1532:
1530:
1529:
1523:
1521:
1515:
1514:
1512:
1511:
1506:
1501:
1496:
1491:
1486:
1480:
1478:
1472:
1471:
1469:
1468:
1463:
1458:
1452:
1450:
1444:
1443:
1441:
1440:
1435:
1429:
1427:
1421:
1420:
1418:
1417:
1410:
1408:
1406:Hesperotestudo
1401:
1400:
1398:
1397:
1391:
1385:
1379:
1373:
1367:
1361:
1354:
1352:
1345:
1344:
1342:
1341:
1339:Texas tortoise
1336:
1331:
1326:
1321:
1316:
1310:
1308:
1302:
1301:
1299:
1298:
1293:
1287:
1285:
1279:
1278:
1276:
1275:
1269:
1263:
1257:
1251:
1244:
1242:
1235:
1234:
1232:
1231:
1226:
1224:Karoo padloper
1221:
1215:
1213:
1207:
1206:
1204:
1203:
1197:
1195:
1189:
1188:
1186:
1185:
1179:
1173:
1166:
1164:
1157:
1156:
1154:
1153:
1148:
1143:
1138:
1133:
1128:
1123:
1118:
1107:
1102:
1092:
1086:
1081:
1076:
1071:
1065:
1063:
1062:
1061:
1050:
1049:
1047:
1046:
1040:
1034:
1028:
1022:
1016:
1010:
1005:
1000:
998:Chaco tortoise
994:
992:
986:
985:
983:
982:
976:
970:
964:
958:
956:
950:
949:
947:
946:
938:
933:
927:
925:
919:
918:
916:
915:
907:
899:
891:
888:A. g. daudinii
883:
876:
869:
867:
861:
860:
858:
857:
851:
849:
843:
842:
835:
833:
830:
829:
814:
810:
809:
807:
806:
800:
794:
788:
782:
776:
770:
764:
757:
754:
753:
745:
744:
737:
730:
722:
713:
712:
710:
709:
696:
683:
670:
657:
644:
631:
616:
600:
598:
592:
591:
578:
577:
561:
542:
499:
447:
397:
396:
394:
391:
361:
358:
323:
320:
306:in the family
304:giant tortoise
286:
285:
284:
283:
275:
267:
259:
251:
243:
232:
231:
225:
224:
221:
207:
206:
200:
199:
191:
189:
185:
184:
174:
170:
169:
164:
160:
159:
154:
150:
149:
144:
140:
139:
134:
130:
129:
124:
120:
119:
114:
110:
109:
104:
100:
99:
94:
90:
89:
76:
75:
56:
53:
52:
47:
44:
43:
35:
34:
26:
25:
15:
9:
6:
4:
3:
2:
4146:
4135:
4132:
4130:
4127:
4125:
4122:
4120:
4117:
4115:
4112:
4110:
4107:
4105:
4102:
4100:
4097:
4095:
4092:
4091:
4089:
4078:
4073:
4071:
4066:
4064:
4059:
4058:
4052:
4050:
4046:
4042:
4037:
4034:
4030:
4029:
4025:
4014:
4010:
4009:
4005:
4004:
3999:
3998:
3995:
3981:
3980:
3979:Tacuarembemys
3975:
3973:
3972:
3967:
3965:
3964:
3959:
3958:
3956:
3952:
3949:
3945:
3942:
3938:
3924:
3923:
3918:
3916:
3915:
3914:Sahonachelys
3910:
3909:
3907:
3905:
3900:
3894:
3893:
3888:
3886:
3885:
3881:
3879:
3878:
3877:Peltocephalus
3874:
3872:
3871:
3866:
3864:
3863:
3862:Lapparentemys
3858:
3856:
3855:
3851:
3849:
3848:
3843:
3841:
3840:
3835:
3833:
3832:
3827:
3825:
3824:
3819:
3817:
3816:
3811:
3809:
3808:
3803:
3801:
3800:
3795:
3794:
3792:
3790:
3786:
3780:
3779:
3775:
3773:
3772:
3768:
3767:
3765:
3763:
3762:Pelomedusidae
3759:
3753:
3752:
3751:Yaminuechelys
3747:
3745:
3744:
3740:
3738:
3737:
3733:
3731:
3730:
3726:
3724:
3723:
3722:Prochelidella
3718:
3716:
3715:
3711:
3709:
3708:
3704:
3702:
3701:
3697:
3695:
3694:
3690:
3688:
3687:
3682:
3680:
3679:
3675:
3673:
3672:
3668:
3666:
3665:
3661:
3659:
3658:
3654:
3652:
3651:
3647:
3645:
3644:
3640:
3638:
3637:
3636:Acanthochelys
3633:
3632:
3630:
3628:
3624:
3618:
3617:
3612:
3610:
3609:
3604:
3602:
3601:
3596:
3594:
3593:
3588:
3586:
3585:
3580:
3578:
3577:
3572:
3570:
3569:
3564:
3562:
3561:
3556:
3554:
3553:
3548:
3546:
3545:
3540:
3538:
3537:
3532:
3530:
3529:
3524:
3522:
3521:
3520:Kinkonychelys
3516:
3514:
3513:
3508:
3506:
3505:
3504:Itapecuruemys
3500:
3498:
3497:
3492:
3490:
3489:
3484:
3482:
3481:
3476:
3474:
3473:
3468:
3466:
3465:
3464:Eotaphrosphys
3460:
3458:
3457:
3452:
3450:
3449:
3444:
3442:
3441:
3436:
3434:
3433:
3428:
3426:
3425:
3420:
3418:
3417:
3412:
3410:
3409:
3404:
3403:
3401:
3399:
3398:Bothremydidae
3394:
3388:
3387:
3382:
3381:
3379:
3377:
3372:
3369:
3365:
3362:
3360:
3356:
3342:
3341:
3336:
3334:
3333:
3328:
3326:
3325:
3320:
3318:
3317:
3312:
3310:
3309:
3308:Argillochelys
3304:
3302:
3301:
3296:
3295:
3293:
3289:
3283:
3282:
3277:
3275:
3274:
3273:Santanachelys
3269:
3267:
3266:
3261:
3259:
3258:
3253:
3251:
3250:
3245:
3243:
3242:
3237:
3235:
3234:
3229:
3227:
3226:
3221:
3219:
3218:
3217:Desmatochelys
3213:
3211:
3210:
3205:
3203:
3202:
3201:Calcarichelys
3197:
3195:
3194:
3189:
3187:
3186:
3185:Atlantochelys
3181:
3179:
3178:
3173:
3171:
3170:
3165:
3164:
3162:
3160:
3159:Protostegidae
3155:
3149:
3148:
3147:Jiangxichelys
3143:
3141:
3140:
3135:
3133:
3132:
3131:Anomalochelys
3127:
3126:
3124:
3122:
3117:
3111:
3110:
3109:Protochelydra
3105:
3103:
3102:
3101:Planiplastron
3097:
3095:
3094:
3090:
3088:
3087:
3082:
3080:
3079:
3074:
3072:
3071:
3066:
3064:
3063:
3058:
3056:
3055:
3051:
3050:
3048:
3046:
3042:
3039:
3035:
3025:
3024:
3019:
3017:
3016:
3011:
3010:
3008:
3004:
2998:
2997:
2993:
2991:
2990:
2986:
2984:
2983:
2979:
2977:
2976:
2972:
2970:
2969:
2964:
2962:
2961:
2957:
2955:
2954:
2950:
2948:
2947:
2943:
2941:
2940:
2935:
2933:
2932:
2927:
2925:
2924:
2919:
2917:
2916:
2912:
2910:
2909:
2908:Drazinderetes
2904:
2902:
2901:
2897:
2895:
2894:
2890:
2888:
2887:
2883:
2881:
2880:
2875:
2873:
2872:
2868:
2866:
2865:
2861:
2860:
2858:
2856:
2852:
2846:
2845:
2844:Carettochelys
2841:
2839:
2838:
2833:
2831:
2830:
2825:
2824:
2822:
2820:
2816:
2813:
2811:
2807:
2797:
2796:
2792:
2790:
2789:
2784:
2782:
2781:
2777:
2775:
2774:
2769:
2767:
2766:
2762:
2760:
2759:
2755:
2753:
2752:
2747:
2745:
2744:
2739:
2737:
2736:
2732:
2730:
2729:
2728:Malacochersus
2725:
2723:
2722:
2718:
2716:
2715:
2711:
2709:
2708:
2704:
2702:
2701:
2696:
2694:
2693:
2688:
2686:
2685:
2681:
2679:
2678:
2674:
2672:
2671:
2666:
2664:
2663:
2658:
2656:
2655:
2651:
2649:
2648:
2644:
2642:
2641:
2637:
2635:
2634:
2630:
2628:
2627:
2623:
2621:
2620:
2619:Aldabrachelys
2616:
2615:
2613:
2611:
2607:
2601:
2600:
2596:
2595:
2593:
2591:
2586:
2580:
2579:
2575:
2573:
2572:
2568:
2566:
2565:
2561:
2559:
2558:
2554:
2552:
2551:
2547:
2545:
2544:
2540:
2538:
2537:
2533:
2531:
2530:
2526:
2524:
2523:
2519:
2517:
2516:
2512:
2510:
2509:
2505:
2503:
2502:
2501:Leucocephalon
2498:
2496:
2495:
2491:
2489:
2488:
2484:
2482:
2481:
2477:
2475:
2474:
2470:
2468:
2467:
2463:
2461:
2460:
2456:
2454:
2453:
2452:Banhxeochelys
2448:
2446:
2445:
2441:
2440:
2438:
2436:
2432:
2426:
2425:
2420:
2418:
2417:
2413:
2411:
2410:
2406:
2404:
2403:
2399:
2397:
2396:
2392:
2390:
2389:
2385:
2383:
2382:
2378:
2376:
2375:
2371:
2369:
2368:
2364:
2362:
2361:
2357:
2355:
2354:
2350:
2348:
2347:
2343:
2341:
2340:
2336:
2334:
2333:
2328:
2327:
2325:
2323:
2319:
2316:
2314:
2313:Testudinoidea
2310:
2300:
2299:
2295:
2293:
2292:
2288:
2286:
2285:
2281:
2279:
2278:
2273:
2271:
2270:
2266:
2265:
2263:
2261:
2260:Kinosternidae
2257:
2251:
2250:
2246:
2245:
2243:
2241:
2237:
2234:
2232:
2228:
2218:
2217:
2212:
2210:
2209:
2208:Protosphargis
2204:
2202:
2201:
2196:
2194:
2193:
2188:
2186:
2185:
2180:
2179:
2177:
2173:
2167:
2166:
2161:
2159:
2158:
2153:
2151:
2150:
2145:
2143:
2142:
2138:
2136:
2135:
2130:
2128:
2127:
2122:
2120:
2119:
2114:
2113:
2111:
2109:
2105:
2099:
2098:
2093:
2091:
2090:
2085:
2083:
2082:
2077:
2075:
2074:
2070:
2068:
2067:
2062:
2060:
2059:
2054:
2052:
2051:
2047:
2045:
2044:
2039:
2037:
2036:
2031:
2029:
2028:
2023:
2021:
2020:
2016:
2014:
2013:
2008:
2006:
2005:
2001:
1999:
1998:
1993:
1991:
1990:
1986:
1984:
1983:
1978:
1977:
1975:
1973:
1969:
1966:
1964:(Sea turtles)
1962:
1958:
1955:
1953:
1949:
1933:
1932:
1926:
1923:
1921:
1917:
1914:
1912:
1908:
1905:
1903:
1899:
1893:
1889:
1887:
1883:
1880:
1878:
1874:
1872:
1868:
1866:
1862:
1861:
1858:
1854:
1847:
1842:
1840:
1835:
1833:
1828:
1827:
1824:
1813:
1807:
1801:
1797:
1795:
1792:
1790:
1787:
1785:
1782:
1780:
1777:
1776:
1774:
1772:
1768:
1762:
1758:
1756:
1752:
1750:
1746:
1744:
1740:
1738:
1734:
1732:
1728:
1726:
1722:
1720:
1716:
1714:
1710:
1708:
1704:
1702:
1698:
1696:
1692:
1690:
1686:
1684:
1680:
1678:
1674:
1673:
1671:
1669:
1664:
1658:
1655:
1654:
1652:
1650:
1646:
1640:
1637:
1635:
1632:
1631:
1629:
1627:
1623:
1617:
1616:Tent tortoise
1614:
1612:
1609:
1607:
1604:
1603:
1601:
1599:
1595:
1589:
1585:
1583:
1579:
1577:
1573:
1571:
1567:
1566:
1564:
1562:
1557:
1551:
1548:
1546:
1543:
1542:
1540:
1538:
1534:
1528:
1525:
1524:
1522:
1520:
1519:Malacochersus
1516:
1510:
1507:
1505:
1502:
1500:
1497:
1495:
1492:
1490:
1487:
1485:
1482:
1481:
1479:
1477:
1473:
1467:
1464:
1462:
1459:
1457:
1454:
1453:
1451:
1449:
1445:
1439:
1436:
1434:
1431:
1430:
1428:
1426:
1422:
1416:
1412:
1411:
1409:
1407:
1402:
1396:
1392:
1390:
1386:
1384:
1380:
1378:
1374:
1372:
1368:
1366:
1362:
1360:
1356:
1355:
1353:
1351:
1346:
1340:
1337:
1335:
1332:
1330:
1327:
1325:
1322:
1320:
1317:
1315:
1312:
1311:
1309:
1307:
1303:
1297:
1294:
1292:
1289:
1288:
1286:
1284:
1280:
1274:
1270:
1268:
1264:
1262:
1258:
1256:
1252:
1250:
1246:
1245:
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881:
880:A. g. arnoldi
877:
874:
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870:
868:
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865:Aldabrachelys
862:
856:
853:
852:
850:
848:
844:
826:
825:
818:
815:
811:
805:
801:
799:
798:Testudinoidea
796:Superfamily:
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793:
789:
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783:
781:
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279:
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255:
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204:Binomial name
201:
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182:
175:
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157:Testudinoidea
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153:Superfamily:
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138:
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86:
81:
77:
71:
65:
54:
50:
45:
41:
36:
32:
27:
22:
19:
4134:Turtle stubs
4099:Cylindraspis
4049:expanding it
4038:
4023:
4002:
3977:
3969:
3961:
3920:
3912:
3890:
3882:
3875:
3868:
3860:
3854:Erymnochelys
3852:
3845:
3839:Cerrejonemys
3837:
3829:
3821:
3815:Brontochelys
3813:
3805:
3797:
3776:
3769:
3749:
3741:
3734:
3729:Pseudemydura
3727:
3720:
3712:
3705:
3698:
3691:
3684:
3676:
3669:
3662:
3655:
3648:
3641:
3634:
3614:
3606:
3598:
3590:
3582:
3574:
3566:
3558:
3550:
3542:
3534:
3526:
3518:
3510:
3502:
3494:
3486:
3478:
3470:
3462:
3454:
3446:
3438:
3430:
3422:
3414:
3406:
3384:
3338:
3332:Prionochelys
3330:
3322:
3314:
3306:
3298:
3279:
3271:
3263:
3255:
3247:
3239:
3231:
3223:
3215:
3209:Cratochelone
3207:
3199:
3193:Bouliachelys
3191:
3183:
3175:
3169:Alienochelys
3167:
3145:
3137:
3129:
3107:
3099:
3091:
3084:
3076:
3070:Chelydropsis
3068:
3060:
3052:
3021:
3015:Basilochelys
3013:
2994:
2987:
2980:
2973:
2966:
2958:
2951:
2944:
2939:Khunnuchelys
2937:
2929:
2921:
2913:
2906:
2898:
2891:
2884:
2877:
2869:
2862:
2855:Trionychidae
2842:
2835:
2829:Allaeochelys
2827:
2793:
2786:
2780:Stigmochelys
2778:
2771:
2763:
2756:
2749:
2743:Megalochelys
2741:
2733:
2726:
2719:
2712:
2705:
2698:
2690:
2682:
2675:
2668:
2660:
2654:Cylindraspis
2652:
2645:
2638:
2633:Centrochelys
2631:
2624:
2617:
2610:Testudinidae
2599:Platysternon
2597:
2578:Vijayachelys
2576:
2569:
2562:
2557:Rhinoclemmys
2555:
2548:
2541:
2534:
2527:
2522:Melanochelys
2520:
2513:
2506:
2499:
2492:
2485:
2478:
2471:
2464:
2457:
2450:
2442:
2422:
2414:
2407:
2400:
2393:
2386:
2379:
2372:
2365:
2358:
2351:
2344:
2337:
2332:Acherontemys
2330:
2298:Sternotherus
2296:
2289:
2282:
2275:
2267:
2247:
2214:
2206:
2198:
2190:
2182:
2165:Psephophorus
2163:
2155:
2147:
2139:
2132:
2124:
2116:
2095:
2087:
2081:Pacifichelys
2079:
2071:
2064:
2056:
2050:Lepidochelys
2048:
2041:
2033:
2027:Gigantatypus
2025:
2019:Eretmochelys
2017:
2010:
2002:
1995:
1987:
1980:
1961:Chelonioidea
1929:
1881:
1649:Stigmochelys
1561:Megalochelys
1254:
1240:Cylindraspis
1162:Cheirogaster
954:Centrochelys
941:
910:
902:
893:
886:
878:
875:(subspecies:
822:
804:Testudinidae
595:
572:
555:the original
545:
537:ResearchGate
535:– via
516:
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467:
463:
450:
438:. Retrieved
424:
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375:
371:
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308:Testudinidae
295:
291:
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277:
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253:
249:Merrem, 1820
245:
237:
215:
210:
194:
193:
181:Cylindraspis
180:
167:Testudinidae
39:
18:
3892:Stupendemys
3847:Cordichelys
3693:Mesoclemmys
3678:Hydromedusa
3600:Taphrosphys
3592:Sankuchemys
3584:Rhothonemys
3560:Polysternon
3440:Cearachelys
3424:Azabbaremys
3408:Araiochelys
3324:Ctenochelys
3316:Bashuchelys
3265:Rhinochelys
3241:Ocepechelon
3233:Notochelone
3225:Iserosaurus
3093:Macrochelys
3045:Chelydridae
2968:Palaeoamyda
2893:Cyclanorbis
2758:Psammobates
2751:Oligopherus
2714:Indotestudo
2662:Cymatholcus
2640:Chelonoidis
2626:Astrochelys
2435:Geoemydidae
2353:Deirochelys
2291:Staurotypus
2284:Kinosternon
2277:Hoplochelys
2192:Peritresius
2141:Dermochelys
2134:Cosmochelys
2126:Corsochelys
2043:Itilochelys
2035:Glarichelys
1982:Allopleuron
1972:Cheloniidae
1911:Superfamily
1598:Psammobates
1448:Indotestudo
990:Chelonoidis
923:Astrochelys
648:iNaturalist
620:Wikispecies
519:: 501–506.
440:15 November
322:Description
302:species of
4088:Categories
3971:Caririemys
3884:Podocnemis
3870:Latentemys
3831:Carbonemys
3771:Pelomedusa
3608:Ummulisani
3528:Kurmademys
3496:Inaechelys
3480:Galianemys
3448:Chedighaii
3386:Araripemys
3359:Pleurodira
3340:Toxochelys
3257:Protostega
3062:Chelydrops
2982:Pelodiscus
2975:Pelochelys
2923:Gilmoremys
2900:Cycloderma
2810:Trionychia
2677:Geochelone
2670:Floridemys
2536:Notochelys
2424:Wilburemys
2395:Malaclemys
2249:Dermatemys
2216:Puppigerus
2149:Eosphargis
2066:Miocaretta
2058:Mexichelys
1952:Cryptodira
1892:Testudines
1853:Testudines
1283:Geochelone
1211:Chersobius
847:Agrionemys
792:Cryptodira
790:Suborder:
786:Testudines
778:Subclass:
470:(2): 278.
393:References
387:Madagascar
281:Bour, 1980
147:Cryptodira
143:Suborder:
137:Testudines
4011:See also
3963:Caribemys
3823:Caninemys
3807:Bauruemys
3736:Rheodytes
3700:Myuchelys
3643:Chelodina
3568:Puentemys
3544:Nigeremys
3488:Ilatardia
3432:Bothremys
3139:Basilemys
2953:Nilssonia
2931:Hutchemys
2879:Axestemys
2837:Anosteira
2692:Hadrianus
2550:Pangshura
2508:Malayemys
2473:Geoclemys
2416:Trachemys
2409:Terrapene
2402:Pseudemys
2388:Graptemys
2381:Glyptemys
2374:Emydoidea
2367:Actinemys
2339:Chrysemys
2184:Euclastes
2012:Eochelone
1863:Kingdom:
1350:Hadrianus
760:Kingdom:
486:1864-5755
348:Mauritius
316:Rodrigues
310:. It was
188:Species:
103:Kingdom:
97:Eukaryota
3778:Pelusios
3743:Rhinemys
3714:Platemys
3707:Phrynops
3627:Chelidae
3616:Zolhafah
3512:Jainemys
3177:Archelon
3054:Chelydra
2946:Lissemys
2788:Stylemys
2773:Solitudo
2735:Manouria
2684:Gopherus
2647:Chersina
2515:Mauremys
2494:Heosemys
2487:Hardella
2480:Geoemyda
2466:Cyclemys
2322:Emydidae
2269:Claudius
2118:Arabemys
2097:Tasbacka
2089:Syllomus
2004:Chelonia
1902:Suborder
1886:Diapsida
1877:Reptilia
1871:Chordata
1869:Phylum:
1865:Animalia
1668:Stylemys
1537:Manouria
1306:Gopherus
1193:Chersina
824:tortoise
802:Family:
780:Anapsida
774:Reptilia
768:Chordata
766:Phylum:
762:Animalia
749:Tortoise
605:Wikidata
494:87809001
298:) is an
229:Synonyms
163:Family:
127:Reptilia
117:Chordata
113:Phylum:
107:Animalia
93:Domain:
70:IUCN 2.3
3922:Sokatra
3671:Emydura
3576:Rosasia
3472:Foxemys
3416:Arenila
2996:Trionyx
2989:Rafetus
2915:Dogania
2871:Apalone
2795:Testudo
2721:Kinixys
2707:Homopus
2564:Sacalia
2543:Orlitia
2529:Morenia
2444:Batagur
2346:Clemmys
2073:Natator
1989:Caretta
1890:Order:
1875:Class:
1771:Testudo
1476:Kinixys
1425:Homopus
784:Order:
772:Class:
640:5789422
611:Q899794
521:Bibcode
312:endemic
300:extinct
213:†
192:†
173:Genus:
133:Order:
123:Class:
68: (
64:Extinct
4041:turtle
3954:
3947:
3940:
3664:Elusor
3657:Elseya
3650:Chelus
3367:
3300:Adocus
3291:
3037:
3006:
2886:Chitra
2588:
2175:
1920:Family
827:family
813:Genera
751:family
705:377727
692:180176
666:949459
653:121114
492:
484:
4043:is a
2960:Palea
2864:Amyda
2765:Pyxis
2459:Cuora
1931:Genus
1882:Clade
1626:Pyxis
1099:Diego
490:S2CID
460:(PDF)
385:) of
40:circa
4045:stub
2360:Emys
687:NCBI
679:6063
674:IUCN
661:ITIS
635:GBIF
482:ISSN
442:2021
425:1996
290:The
529:doi
517:159
472:doi
429:doi
314:to
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