403:
356:
94:
42:
69:
736:
443:
was concerned about the vanishing species and, in his enquiries, was told about the 1844 expedition by one of its members, Mr. William Kerr. Kerr informed the
Governor that Mr. Corby, one of the other 1844 explorers, "captured a female land tortoise in one of the caves on Round Island and brought it
450:
Round Island itself, already badly damaged by rabbits, had goats introduced to it soon afterwards. This—or some other factor—led to the total extinction of the tortoises in their last refuge. At the unknown point when the last of Corby's hatchlings died or was killed, the species would have become
410:
This species was previously numerous throughout
Mauritius—both on the main island and on all of the surrounding islets. As Mauritius was the first of the Mascarene Islands to be settled, it was also the first to face the extermination of its native biodiversity—including the giant tortoises. The
435:
The Lloyd expedition in 1844 found several very large specimens of giant tortoise surviving on Round Island, although the island was by then already overrun with enormous numbers of introduced rabbits.
447:
The
Governor was unable to locate any of the progeny and, although hypothetically an 1845 hatchling could easily have lived into the 21st century, it is not known what happened to the hatchlings.
785:
377:
grazed on grass, as well as fallen leaves and fruit on forest floors. Although similarly sized, the two species differed substantially in their body shape and bone structure.
1068:
941:
1080:
1074:
414:
With the arrival of the Dutch, vast numbers of both giant tortoise species were slaughtered—either for food (for humans or pigs) or to be burned for fat and oil.
1628:
1622:
1580:
1640:
1586:
1280:
1262:
935:
1616:
1286:
1268:
1697:
1652:
1634:
1598:
1274:
923:
1658:
1292:
1256:
1610:
1646:
1604:
1592:
1574:
373:
to
Mauritius, this saddle-backed species seems to have specialized in browsing higher bushes and low-hanging branches of trees. Its lower, flatter
4133:
1059:
4039:
2648:
2559:
2567:
2321:
417:
In addition, they introduced invasive alien species such as rats, cats and pigs, which ate the giant tortoises' eggs and hatchlings.
3987:
4123:
4103:
4013:
981:
4093:
1170:
637:
589:
390:, and to have accidentally drifted to the surrounding islands of Reunion and Rodrigues in order to do so. Its species name of
4113:
420:
The species was likely extinct on the main island of
Mauritius by about 1700, and on most of the surrounding islets by 1735.
4018:
3901:
Turtles of the World 2017 Update: Annotated
Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status
1710:
turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status
1741:
4118:
17:
3899:
1709:
4044:
792:
1023:
4128:
3774:
2857:
1406:
1221:
1152:
1146:
1018:
777:
411:
giant tortoise species, like many island species, were reportedly friendly, curious and not afraid of humans.
3965:
1391:
1381:
1043:
3751:
3910:
2433:
1817:
1531:
1396:
1048:
93:
2166:
1386:
992:
630:
231:
1401:
809:
1799:
1158:
444:
to
Mauritius, where it produced a numerous progeny, which were distributed among his acquaintance."
4108:
2812:
1734:
1312:
1033:
1002:
971:
859:
3668:
1681:
1231:
4070:
492:
429:
3927:
3640:
2440:
2398:
2038:
1038:
976:
911:
905:
877:
871:
865:
840:
770:
402:
4098:
4065:
2384:
1886:
1691:
1676:
1485:
1473:
1442:
1358:
1188:
1121:
1028:
1008:
801:
623:
590:"Non-native megaherbivores: The case for novel function to manage plant invasions on islands"
212:
4057:
956:
951:
380:
This species in particular seems to have been the ancestor of all the other four species of
3974:
3626:
2292:
1916:
1193:
929:
917:
328:
4031:
8:
3611:
1970:
1727:
1686:
1479:
1363:
1126:
900:
597:
577:
Unequal struggle—how humans displaced the tortoise's dominant place in island ecosystems.
355:
58:
3018:
2983:
2850:
2589:
1503:
1467:
1447:
1353:
1247:
966:
545:
240:
88:
4052:
3561:
2544:
2496:
2487:
2370:
1947:
1508:
1335:
1330:
1098:
987:
833:
828:
537:
387:
549:
469:
3353:
2741:
2716:
2677:
2625:
2356:
1901:
1554:
1424:
1116:
996:
752:
527:
487:
359:
3801:
3449:
3178:
3146:
2783:
2692:
2468:
2054:
1668:
1536:
1226:
1216:
1211:
1012:
564:
Lost Land of the Dodo: The
Ecological History of Mauritius, Réunion and Rodrigues
406:
Engraving from 1601 showing the two
Mauritian species of giant tortoise, far left
432:(just north of Mauritius) until much later, according to the 1846 Lloyd report.
428:
At least one of the two
Mauritian giant tortoise species might have survived on
3696:
3686:
3583:
3433:
3273:
2975:
2920:
2662:
2597:
2137:
2128:
2097:
2005:
1894:
1808:
1523:
1303:
1236:
895:
363:
4087:
4026:
3950:
3876:
3759:
3659:
3648:
3619:
3533:
3417:
3401:
3361:
3295:
3205:
3170:
3114:
3098:
3082:
3056:
3044:
3028:
3006:
2998:
2805:
2516:
2349:
2210:
2157:
2105:
1513:
1416:
762:
695:
541:
478:
257:
165:
78:
41:
3811:
3736:
3712:
3229:
3106:
3090:
3066:
2967:
2912:
2836:
2752:
2726:
2640:
2551:
2530:
2454:
2419:
2229:
2195:
2062:
1978:
1924:
1546:
1458:
1137:
851:
701:
440:
382:
374:
336:
332:
188:
4000:
3959:
3789:
3744:
3590:
3575:
3497:
3489:
3481:
3457:
3337:
3321:
3305:
3221:
3213:
3162:
3138:
3130:
3122:
2990:
2942:
2865:
2790:
2655:
2611:
2537:
2523:
2475:
2332:
2250:
2188:
2181:
2174:
2089:
2031:
2023:
1940:
1932:
1879:
1869:
1495:
1345:
887:
820:
532:
515:
3992:
3868:
3781:
3767:
3728:
3505:
3425:
3393:
3377:
3345:
3283:
3256:
3237:
3154:
2959:
2879:
2872:
2820:
2797:
2707:
2574:
2306:
2113:
2046:
1963:
1955:
1858:
1849:
1180:
1108:
744:
689:
683:
155:
73:
4005:
579:
In: Gerlach J, ed. Western Indian Ocean Tortoises: biodiversity. 2014.
3860:
3720:
3704:
3633:
3597:
3547:
3540:
3465:
3441:
3385:
3329:
3036:
2828:
2776:
2734:
2447:
2405:
2313:
2299:
2285:
2278:
2271:
2264:
2236:
2081:
1909:
347:. The last records of this tortoise date to the early 18th century.
344:
105:
3921:
467:
3944:
3675:
3604:
3524:
3513:
3409:
3074:
2951:
2843:
2685:
2670:
2632:
2581:
2507:
2412:
2391:
2377:
2363:
2219:
2146:
2015:
1994:
1986:
1768:
1565:
1434:
1203:
1090:
721:
665:
646:
370:
340:
175:
125:
615:
3819:
3568:
3473:
3369:
3313:
2893:
2886:
2768:
2618:
2604:
2461:
2426:
2341:
2243:
1783:
1774:
1373:
1322:
677:
671:
135:
3979:
3554:
3197:
1789:
1762:
1750:
659:
145:
115:
394:
is due to its supposed propensity for falling into the ocean.
2761:
1828:
1719:
2257:
461:
369:
One of two different giant tortoise species which were
513:
718:
4085:
3898:Phylogenetic arrangement of turtles based on
2136:
1735:
1708:Phylogenetic arrangement of turtles based on
631:
468:World Conservation Monitoring Centre (1996).
493:10.2305/IUCN.UK.1996.RLTS.T6062A12385198.en
2941:
1868:
1807:
1742:
1728:
638:
624:
67:
40:
531:
507:
491:
401:
354:
35:Saddle-backed Mauritius giant tortoise
14:
4134:Species endangered by invasive species
4086:
1171:Saddle-backed Rodrigues giant tortoise
1165:Saddle-backed Mauritius giant tortoise
516:"Checklist of Chelonians of the World"
423:
319:saddle-backed Mauritius giant tortoise
3926:
3925:
1816:
1723:
619:
566:. Bloomsbury Publishing. 2010. p.211.
645:
479:IUCN Red List of Threatened Species
24:
25:
4145:
734:
92:
3844:
3264:
514:Fritz Uwe; Peter Havaš (2007).
4104:Reptile extinctions since 1500
1749:
1153:Domed Rodrigues giant tortoise
1147:Domed Mauritius giant tortoise
1019:Santiago Island giant tortoise
582:
569:
556:
350:
13:
1:
4094:IUCN Red List extinct species
972:Chatham Island giant tortoise
454:
397:
4124:Taxa named by Albert Günther
4114:Extinct animals of Mauritius
1039:Volcán Darwin giant tortoise
1034:Volcán Alcedo giant tortoise
1024:Pinzón Island giant tortoise
7:
3911:List of Testudines families
1532:Flat-backed spider tortoise
1407:Speke's hinge-back tortoise
1397:Lobatse hinge-back tortoise
1222:Goode's thornscrub tortoise
1049:Western Santa Cruz tortoise
1029:Sierra Negra giant tortoise
977:Eastern Santa Cruz tortoise
27:Extinct species of tortoise
10:
4150:
4119:Reptiles described in 1873
1392:Home's hinge-back tortoise
1387:Forest hinge-back tortoise
1382:Bell's hinge-back tortoise
1044:Volcán Wolf giant tortoise
993:Hood Island giant tortoise
982:Fernandina Island tortoise
732:
439:In 1870, the Governor Sir
3934:
3894:
3851:
3837:
3799:
3685:
3658:
3523:
3293:
3271:
3255:
3188:
3054:
3016:
2934:
2903:
2751:
2715:
2706:
2506:
2485:
2331:
2218:
2209:
2156:
2127:
2072:
2004:
1857:
1848:
1826:
1825:
1798:
1757:
1706:
1667:
1563:
1545:
1522:
1494:
1456:
1433:
1415:
1402:Natal hinge-back tortoise
1372:
1344:
1321:
1301:
1245:
1202:
1179:
1135:
1107:
1089:
1057:
967:Cerro Azul giant tortoise
950:
886:
850:
819:
810:Aldabrachelys grandidieri
761:
743:
717:
710:
654:
594:aobpla.oxfordjournals.org
246:
239:
218:
211:
89:Scientific classification
87:
65:
56:
48:
39:
34:
1313:Hesperotestudo percrassa
1069:Cheirogaster bacharidisi
1003:Santa Fe Island tortoise
988:Floreana Island tortoise
942:Chelonoidis sombrerensis
860:African spurred tortoise
1232:Sonoran Desert tortoise
1159:Réunion giant tortoise
1081:Cheirogaster schafferi
1075:Cheirogaster gymnesica
912:Chelonoidis alburyorum
906:Yellow-footed tortoise
878:Centrochelys vulcanica
872:Centrochelys burchardi
866:Centrochelys atlantica
841:Astrochelys rogerbouri
771:Aldabra giant tortoise
407:
386:giant tortoise of the
366:
4129:Reptiles of Mauritius
4066:Paleobiology Database
1692:Spur-thighed tortoise
1629:Stylemys nebrascensis
1623:Stylemys karakolensis
1581:Stylemys calaverensis
1486:Megalochelys sondaari
1474:Megalochelys cautleyi
1443:Asian forest tortoise
1189:Burmese star tortoise
1009:Pinta Island tortoise
802:Aldabrachelys abrupta
405:
358:
1682:Kleinmann's tortoise
1641:Stylemys oregonensis
1587:Stylemys canetotiana
1281:Hadrianus schucherti
1263:Hadrianus majusculus
1194:Indian star tortoise
936:Chelonoidis sellovii
930:Chelonoidis monensis
918:Chelonoidis cubensis
533:10.3897/vz.57.e30895
486:: e.T6062A12385198.
3966:Cylindraspis inepta
3936:Cylindraspis inepta
1687:Marginated tortoise
1617:Stylemys frizaciana
1480:Megalochelys margae
1364:Travancore tortoise
1287:Hadrianus utahensis
1269:Hadrianus octonaria
957:Chelonoidis complex
901:Red-footed tortoise
562:A.Cheke, J.P.Hume:
472:Cylindraspis inepta
424:Round Island refuge
324:Cylindraspis inepta
305:Cylindraspis inepta
297:Geochelone sauzieri
225:Cylindraspis inepta
59:Conservation status
51:Cylindraspis inepta
18:Cylindraspis inepta
3019:Nanhsiungchelyidae
2984:Macrocephalochelys
1698:Testudo brevitesta
1677:Hermann's tortoise
1653:Stylemys uintensis
1635:Stylemys neglectus
1599:Stylemys conspecta
1504:Geometric tortoise
1468:Megalochelys atlas
1448:Impressed tortoise
1359:Forsten's tortoise
1354:Elongated tortoise
1275:Hadrianus robustus
952:Galápagos tortoise
924:Chelonoidis lutzae
575:Cheke AS, Bour R:
520:Vertebrate Zoology
408:
367:
4081:
4080:
4053:Open Tree of Life
3928:Taxon identifiers
3919:
3918:
3890:
3889:
3886:
3885:
3833:
3832:
3829:
3828:
3251:
3250:
3247:
3246:
2930:
2929:
2702:
2701:
2205:
2204:
2123:
2122:
1844:
1843:
1840:
1839:
1836:
1835:
1717:
1716:
1712:. Key: †=extinct.
1659:Stylemys undabuna
1509:Serrated tortoise
1293:Hadrianus vialovi
1257:Hadrianus corsoni
1127:Speckled padloper
1099:Angulate tortoise
834:Radiated tortoise
829:Angonoka tortoise
730:
729:
451:totally extinct.
388:Mascarene Islands
315:
314:
309:
301:
293:
289:Geochelone inepta
285:
277:
269:
254:
82:
16:(Redirected from
4141:
4074:
4073:
4061:
4060:
4048:
4047:
4035:
4034:
4022:
4021:
4009:
4008:
3996:
3995:
3983:
3982:
3970:
3969:
3968:
3955:
3954:
3953:
3923:
3922:
3849:
3848:
3842:
3841:
3354:Chupacabrachelys
3269:
3268:
3262:
3261:
2939:
2938:
2717:Carettochelyidae
2713:
2712:
2216:
2215:
2134:
2133:
1866:
1865:
1855:
1854:
1823:
1822:
1814:
1813:
1805:
1804:
1744:
1737:
1730:
1721:
1720:
1611:Stylemys emiliae
1555:Leopard tortoise
1425:Pancake tortoise
1336:Greater padloper
753:Russian tortoise
738:
737:
715:
714:
640:
633:
626:
617:
616:
610:
609:
607:
605:
596:. Archived from
586:
580:
573:
567:
560:
554:
553:
535:
511:
505:
504:
502:
500:
495:
465:
360:Life restoration
307:
299:
291:
283:
281:Testudo sauzieri
275:
273:Testudo boutonii
267:
252:
250:Testudo neraudii
227:
223:
187:
97:
96:
76:
71:
70:
44:
32:
31:
21:
4149:
4148:
4144:
4143:
4142:
4140:
4139:
4138:
4109:Extinct turtles
4084:
4083:
4082:
4077:
4069:
4064:
4056:
4051:
4043:
4038:
4030:
4025:
4017:
4012:
4004:
3999:
3991:
3986:
3978:
3973:
3964:
3963:
3958:
3949:
3948:
3943:
3930:
3920:
3915:
3882:
3825:
3802:Sahonachelyidae
3795:
3681:
3654:
3519:
3450:Phosphatochelys
3289:
3243:
3184:
3179:Terlinguachelys
3147:Pneumatoarthrus
3050:
3012:
2926:
2899:
2747:
2698:
2502:
2481:
2469:Siebenrockiella
2327:
2201:
2152:
2119:
2068:
2055:Mesodermochelys
2000:
1861:
1832:
1794:
1753:
1748:
1718:
1713:
1702:
1663:
1647:Stylemys pygmea
1559:
1541:
1537:Spider tortoise
1518:
1490:
1452:
1429:
1411:
1368:
1340:
1331:Common padloper
1317:
1297:
1241:
1227:Gopher tortoise
1217:Desert tortoise
1212:Bolson tortoise
1198:
1175:
1131:
1103:
1085:
1053:
1013:Lonesome George
946:
882:
846:
815:
793:A. g. hololissa
757:
739:
735:
726:
719:Species of the
706:
650:
644:
614:
613:
603:
601:
588:
587:
583:
574:
570:
561:
557:
512:
508:
498:
496:
466:
462:
457:
426:
400:
353:
300:Pritchard, 1967
292:Pritchard, 1967
235:
229:
221:
220:
207:
185:
91:
83:
72:
68:
61:
28:
23:
22:
15:
12:
11:
5:
4147:
4137:
4136:
4131:
4126:
4121:
4116:
4111:
4106:
4101:
4096:
4079:
4078:
4076:
4075:
4062:
4049:
4036:
4023:
4010:
3997:
3984:
3971:
3956:
3940:
3938:
3932:
3931:
3917:
3916:
3914:
3913:
3906:
3905:
3904:. † = extinct.
3895:
3892:
3891:
3888:
3887:
3884:
3883:
3881:
3880:
3872:
3864:
3855:
3853:
3846:
3839:
3835:
3834:
3831:
3830:
3827:
3826:
3824:
3823:
3815:
3806:
3804:
3797:
3796:
3794:
3793:
3785:
3778:
3771:
3763:
3755:
3748:
3740:
3732:
3724:
3716:
3708:
3700:
3697:Albertwoodemys
3691:
3689:
3687:Podocnemididae
3683:
3682:
3680:
3679:
3672:
3664:
3662:
3656:
3655:
3653:
3652:
3644:
3637:
3630:
3623:
3615:
3608:
3601:
3594:
3587:
3584:Lomalatachelys
3579:
3572:
3565:
3558:
3551:
3544:
3537:
3529:
3527:
3521:
3520:
3518:
3517:
3509:
3501:
3493:
3485:
3477:
3469:
3461:
3453:
3445:
3437:
3434:Labrostochelys
3429:
3421:
3413:
3405:
3397:
3389:
3381:
3373:
3365:
3357:
3349:
3341:
3333:
3325:
3317:
3309:
3300:
3298:
3291:
3290:
3288:
3287:
3278:
3276:
3274:Araripemydidae
3266:
3259:
3253:
3252:
3249:
3248:
3245:
3244:
3242:
3241:
3233:
3225:
3217:
3209:
3201:
3192:
3190:
3186:
3185:
3183:
3182:
3174:
3166:
3158:
3150:
3142:
3134:
3126:
3118:
3110:
3102:
3094:
3086:
3078:
3070:
3061:
3059:
3052:
3051:
3049:
3048:
3040:
3032:
3023:
3021:
3014:
3013:
3011:
3010:
3002:
2994:
2987:
2979:
2976:Emarginachelys
2971:
2963:
2955:
2947:
2945:
2936:
2932:
2931:
2928:
2927:
2925:
2924:
2921:Sinaspideretes
2916:
2907:
2905:
2901:
2900:
2898:
2897:
2890:
2883:
2876:
2869:
2861:
2854:
2847:
2840:
2832:
2824:
2816:
2809:
2801:
2794:
2787:
2780:
2772:
2765:
2757:
2755:
2749:
2748:
2746:
2745:
2738:
2730:
2721:
2719:
2710:
2704:
2703:
2700:
2699:
2697:
2696:
2689:
2681:
2674:
2666:
2659:
2652:
2644:
2636:
2629:
2622:
2615:
2608:
2601:
2598:Hesperotestudo
2593:
2585:
2578:
2571:
2563:
2555:
2548:
2541:
2534:
2527:
2520:
2512:
2510:
2504:
2503:
2501:
2500:
2492:
2490:
2488:Platysternidae
2483:
2482:
2480:
2479:
2472:
2465:
2458:
2451:
2444:
2437:
2430:
2423:
2416:
2409:
2402:
2395:
2388:
2381:
2374:
2367:
2360:
2353:
2345:
2337:
2335:
2329:
2328:
2326:
2325:
2317:
2310:
2303:
2296:
2289:
2282:
2275:
2268:
2261:
2254:
2247:
2240:
2233:
2224:
2222:
2213:
2207:
2206:
2203:
2202:
2200:
2199:
2192:
2185:
2178:
2170:
2162:
2160:
2154:
2153:
2151:
2150:
2142:
2140:
2138:Dermatemydidae
2131:
2129:Kinosternoidea
2125:
2124:
2121:
2120:
2118:
2117:
2109:
2101:
2098:Procolpochelys
2093:
2085:
2076:
2074:
2070:
2069:
2067:
2066:
2058:
2050:
2042:
2035:
2027:
2019:
2010:
2008:
2006:Dermochelyidae
2002:
2001:
1999:
1998:
1990:
1982:
1974:
1967:
1959:
1951:
1944:
1936:
1928:
1920:
1913:
1905:
1898:
1895:Carolinochelys
1890:
1883:
1874:
1872:
1863:
1852:
1846:
1845:
1842:
1841:
1838:
1837:
1834:
1833:
1820:
1811:
1802:
1796:
1795:
1793:
1792:
1786:
1777:
1771:
1765:
1758:
1755:
1754:
1747:
1746:
1739:
1732:
1724:
1715:
1714:
1707:
1704:
1703:
1701:
1700:
1694:
1689:
1684:
1679:
1673:
1671:
1665:
1664:
1662:
1661:
1655:
1649:
1643:
1637:
1631:
1625:
1619:
1613:
1607:
1605:Stylemys copei
1601:
1595:
1593:Stylemys capax
1589:
1583:
1577:
1575:Stylemys botti
1570:
1568:
1561:
1560:
1558:
1557:
1551:
1549:
1543:
1542:
1540:
1539:
1534:
1528:
1526:
1520:
1519:
1517:
1516:
1511:
1506:
1500:
1498:
1492:
1491:
1489:
1488:
1482:
1476:
1470:
1463:
1461:
1454:
1453:
1451:
1450:
1445:
1439:
1437:
1431:
1430:
1428:
1427:
1421:
1419:
1413:
1412:
1410:
1409:
1404:
1399:
1394:
1389:
1384:
1378:
1376:
1370:
1369:
1367:
1366:
1361:
1356:
1350:
1348:
1342:
1341:
1339:
1338:
1333:
1327:
1325:
1319:
1318:
1316:
1315:
1308:
1306:
1304:Hesperotestudo
1299:
1298:
1296:
1295:
1289:
1283:
1277:
1271:
1265:
1259:
1252:
1250:
1243:
1242:
1240:
1239:
1237:Texas tortoise
1234:
1229:
1224:
1219:
1214:
1208:
1206:
1200:
1199:
1197:
1196:
1191:
1185:
1183:
1177:
1176:
1174:
1173:
1167:
1161:
1155:
1149:
1142:
1140:
1133:
1132:
1130:
1129:
1124:
1122:Karoo padloper
1119:
1113:
1111:
1105:
1104:
1102:
1101:
1095:
1093:
1087:
1086:
1084:
1083:
1077:
1071:
1064:
1062:
1055:
1054:
1052:
1051:
1046:
1041:
1036:
1031:
1026:
1021:
1016:
1005:
1000:
990:
984:
979:
974:
969:
963:
961:
960:
959:
948:
947:
945:
944:
938:
932:
926:
920:
914:
908:
903:
898:
896:Chaco tortoise
892:
890:
884:
883:
881:
880:
874:
868:
862:
856:
854:
848:
847:
845:
844:
836:
831:
825:
823:
817:
816:
814:
813:
805:
797:
789:
786:A. g. daudinii
781:
774:
767:
765:
759:
758:
756:
755:
749:
747:
741:
740:
733:
731:
728:
727:
712:
708:
707:
705:
704:
698:
692:
686:
680:
674:
668:
662:
655:
652:
651:
643:
642:
635:
628:
620:
612:
611:
600:on 5 June 2016
581:
568:
555:
526:(2): 277–278.
506:
459:
458:
456:
453:
425:
422:
399:
396:
375:sister species
364:Julian P. Hume
352:
349:
335:in the family
333:giant tortoise
313:
312:
311:
310:
302:
294:
286:
278:
270:
265:Testudo inepta
262:
244:
243:
237:
236:
230:
216:
215:
209:
208:
204:C. inepta
200:
198:
194:
193:
183:
179:
178:
173:
169:
168:
163:
159:
158:
153:
149:
148:
143:
139:
138:
133:
129:
128:
123:
119:
118:
113:
109:
108:
103:
99:
98:
85:
84:
66:
63:
62:
57:
54:
53:
46:
45:
37:
36:
26:
9:
6:
4:
3:
2:
4146:
4135:
4132:
4130:
4127:
4125:
4122:
4120:
4117:
4115:
4112:
4110:
4107:
4105:
4102:
4100:
4097:
4095:
4092:
4091:
4089:
4072:
4067:
4063:
4059:
4054:
4050:
4046:
4041:
4037:
4033:
4028:
4024:
4020:
4015:
4011:
4007:
4002:
3998:
3994:
3989:
3985:
3981:
3976:
3972:
3967:
3961:
3957:
3952:
3946:
3942:
3941:
3939:
3937:
3933:
3929:
3924:
3912:
3908:
3907:
3903:
3902:
3897:
3896:
3893:
3879:
3878:
3877:Tacuarembemys
3873:
3871:
3870:
3865:
3863:
3862:
3857:
3856:
3854:
3850:
3847:
3843:
3840:
3836:
3822:
3821:
3816:
3814:
3813:
3812:Sahonachelys
3808:
3807:
3805:
3803:
3798:
3792:
3791:
3786:
3784:
3783:
3779:
3777:
3776:
3775:Peltocephalus
3772:
3770:
3769:
3764:
3762:
3761:
3760:Lapparentemys
3756:
3754:
3753:
3749:
3747:
3746:
3741:
3739:
3738:
3733:
3731:
3730:
3725:
3723:
3722:
3717:
3715:
3714:
3709:
3707:
3706:
3701:
3699:
3698:
3693:
3692:
3690:
3688:
3684:
3678:
3677:
3673:
3671:
3670:
3666:
3665:
3663:
3661:
3660:Pelomedusidae
3657:
3651:
3650:
3649:Yaminuechelys
3645:
3643:
3642:
3638:
3636:
3635:
3631:
3629:
3628:
3624:
3622:
3621:
3620:Prochelidella
3616:
3614:
3613:
3609:
3607:
3606:
3602:
3600:
3599:
3595:
3593:
3592:
3588:
3586:
3585:
3580:
3578:
3577:
3573:
3571:
3570:
3566:
3564:
3563:
3559:
3557:
3556:
3552:
3550:
3549:
3545:
3543:
3542:
3538:
3536:
3535:
3534:Acanthochelys
3531:
3530:
3528:
3526:
3522:
3516:
3515:
3510:
3508:
3507:
3502:
3500:
3499:
3494:
3492:
3491:
3486:
3484:
3483:
3478:
3476:
3475:
3470:
3468:
3467:
3462:
3460:
3459:
3454:
3452:
3451:
3446:
3444:
3443:
3438:
3436:
3435:
3430:
3428:
3427:
3422:
3420:
3419:
3418:Kinkonychelys
3414:
3412:
3411:
3406:
3404:
3403:
3402:Itapecuruemys
3398:
3396:
3395:
3390:
3388:
3387:
3382:
3380:
3379:
3374:
3372:
3371:
3366:
3364:
3363:
3362:Eotaphrosphys
3358:
3356:
3355:
3350:
3348:
3347:
3342:
3340:
3339:
3334:
3332:
3331:
3326:
3324:
3323:
3318:
3316:
3315:
3310:
3308:
3307:
3302:
3301:
3299:
3297:
3296:Bothremydidae
3292:
3286:
3285:
3280:
3279:
3277:
3275:
3270:
3267:
3263:
3260:
3258:
3254:
3240:
3239:
3234:
3232:
3231:
3226:
3224:
3223:
3218:
3216:
3215:
3210:
3208:
3207:
3206:Argillochelys
3202:
3200:
3199:
3194:
3193:
3191:
3187:
3181:
3180:
3175:
3173:
3172:
3171:Santanachelys
3167:
3165:
3164:
3159:
3157:
3156:
3151:
3149:
3148:
3143:
3141:
3140:
3135:
3133:
3132:
3127:
3125:
3124:
3119:
3117:
3116:
3115:Desmatochelys
3111:
3109:
3108:
3103:
3101:
3100:
3099:Calcarichelys
3095:
3093:
3092:
3087:
3085:
3084:
3083:Atlantochelys
3079:
3077:
3076:
3071:
3069:
3068:
3063:
3062:
3060:
3058:
3057:Protostegidae
3053:
3047:
3046:
3045:Jiangxichelys
3041:
3039:
3038:
3033:
3031:
3030:
3029:Anomalochelys
3025:
3024:
3022:
3020:
3015:
3009:
3008:
3007:Protochelydra
3003:
3001:
3000:
2999:Planiplastron
2995:
2993:
2992:
2988:
2986:
2985:
2980:
2978:
2977:
2972:
2970:
2969:
2964:
2962:
2961:
2956:
2954:
2953:
2949:
2948:
2946:
2944:
2940:
2937:
2933:
2923:
2922:
2917:
2915:
2914:
2909:
2908:
2906:
2902:
2896:
2895:
2891:
2889:
2888:
2884:
2882:
2881:
2877:
2875:
2874:
2870:
2868:
2867:
2862:
2860:
2859:
2855:
2853:
2852:
2848:
2846:
2845:
2841:
2839:
2838:
2833:
2831:
2830:
2825:
2823:
2822:
2817:
2815:
2814:
2810:
2808:
2807:
2806:Drazinderetes
2802:
2800:
2799:
2795:
2793:
2792:
2788:
2786:
2785:
2781:
2779:
2778:
2773:
2771:
2770:
2766:
2764:
2763:
2759:
2758:
2756:
2754:
2750:
2744:
2743:
2742:Carettochelys
2739:
2737:
2736:
2731:
2729:
2728:
2723:
2722:
2720:
2718:
2714:
2711:
2709:
2705:
2695:
2694:
2690:
2688:
2687:
2682:
2680:
2679:
2675:
2673:
2672:
2667:
2665:
2664:
2660:
2658:
2657:
2653:
2651:
2650:
2645:
2643:
2642:
2637:
2635:
2634:
2630:
2628:
2627:
2626:Malacochersus
2623:
2621:
2620:
2616:
2614:
2613:
2609:
2607:
2606:
2602:
2600:
2599:
2594:
2592:
2591:
2586:
2584:
2583:
2579:
2577:
2576:
2572:
2570:
2569:
2564:
2562:
2561:
2556:
2554:
2553:
2549:
2547:
2546:
2542:
2540:
2539:
2535:
2533:
2532:
2528:
2526:
2525:
2521:
2519:
2518:
2517:Aldabrachelys
2514:
2513:
2511:
2509:
2505:
2499:
2498:
2494:
2493:
2491:
2489:
2484:
2478:
2477:
2473:
2471:
2470:
2466:
2464:
2463:
2459:
2457:
2456:
2452:
2450:
2449:
2445:
2443:
2442:
2438:
2436:
2435:
2431:
2429:
2428:
2424:
2422:
2421:
2417:
2415:
2414:
2410:
2408:
2407:
2403:
2401:
2400:
2399:Leucocephalon
2396:
2394:
2393:
2389:
2387:
2386:
2382:
2380:
2379:
2375:
2373:
2372:
2368:
2366:
2365:
2361:
2359:
2358:
2354:
2352:
2351:
2350:Banhxeochelys
2346:
2344:
2343:
2339:
2338:
2336:
2334:
2330:
2324:
2323:
2318:
2316:
2315:
2311:
2309:
2308:
2304:
2302:
2301:
2297:
2295:
2294:
2290:
2288:
2287:
2283:
2281:
2280:
2276:
2274:
2273:
2269:
2267:
2266:
2262:
2260:
2259:
2255:
2253:
2252:
2248:
2246:
2245:
2241:
2239:
2238:
2234:
2232:
2231:
2226:
2225:
2223:
2221:
2217:
2214:
2212:
2211:Testudinoidea
2208:
2198:
2197:
2193:
2191:
2190:
2186:
2184:
2183:
2179:
2177:
2176:
2171:
2169:
2168:
2164:
2163:
2161:
2159:
2158:Kinosternidae
2155:
2149:
2148:
2144:
2143:
2141:
2139:
2135:
2132:
2130:
2126:
2116:
2115:
2110:
2108:
2107:
2106:Protosphargis
2102:
2100:
2099:
2094:
2092:
2091:
2086:
2084:
2083:
2078:
2077:
2075:
2071:
2065:
2064:
2059:
2057:
2056:
2051:
2049:
2048:
2043:
2041:
2040:
2036:
2034:
2033:
2028:
2026:
2025:
2020:
2018:
2017:
2012:
2011:
2009:
2007:
2003:
1997:
1996:
1991:
1989:
1988:
1983:
1981:
1980:
1975:
1973:
1972:
1968:
1966:
1965:
1960:
1958:
1957:
1952:
1950:
1949:
1945:
1943:
1942:
1937:
1935:
1934:
1929:
1927:
1926:
1921:
1919:
1918:
1914:
1912:
1911:
1906:
1904:
1903:
1899:
1897:
1896:
1891:
1889:
1888:
1884:
1882:
1881:
1876:
1875:
1873:
1871:
1867:
1864:
1862:(Sea turtles)
1860:
1856:
1853:
1851:
1847:
1831:
1830:
1824:
1821:
1819:
1815:
1812:
1810:
1806:
1803:
1801:
1797:
1791:
1787:
1785:
1781:
1778:
1776:
1772:
1770:
1766:
1764:
1760:
1759:
1756:
1752:
1745:
1740:
1738:
1733:
1731:
1726:
1725:
1722:
1711:
1705:
1699:
1695:
1693:
1690:
1688:
1685:
1683:
1680:
1678:
1675:
1674:
1672:
1670:
1666:
1660:
1656:
1654:
1650:
1648:
1644:
1642:
1638:
1636:
1632:
1630:
1626:
1624:
1620:
1618:
1614:
1612:
1608:
1606:
1602:
1600:
1596:
1594:
1590:
1588:
1584:
1582:
1578:
1576:
1572:
1571:
1569:
1567:
1562:
1556:
1553:
1552:
1550:
1548:
1544:
1538:
1535:
1533:
1530:
1529:
1527:
1525:
1521:
1515:
1514:Tent tortoise
1512:
1510:
1507:
1505:
1502:
1501:
1499:
1497:
1493:
1487:
1483:
1481:
1477:
1475:
1471:
1469:
1465:
1464:
1462:
1460:
1455:
1449:
1446:
1444:
1441:
1440:
1438:
1436:
1432:
1426:
1423:
1422:
1420:
1418:
1417:Malacochersus
1414:
1408:
1405:
1403:
1400:
1398:
1395:
1393:
1390:
1388:
1385:
1383:
1380:
1379:
1377:
1375:
1371:
1365:
1362:
1360:
1357:
1355:
1352:
1351:
1349:
1347:
1343:
1337:
1334:
1332:
1329:
1328:
1326:
1324:
1320:
1314:
1310:
1309:
1307:
1305:
1300:
1294:
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1128:
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1120:
1118:
1117:Nama padloper
1115:
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975:
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904:
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873:
869:
867:
863:
861:
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853:
849:
843:
842:
837:
835:
832:
830:
827:
826:
824:
822:
818:
812:
811:
806:
804:
803:
798:
795:
794:
790:
788:
787:
782:
780:
779:
778:A. g. arnoldi
775:
772:
769:
768:
766:
764:
763:Aldabrachelys
760:
754:
751:
750:
748:
746:
742:
724:
723:
716:
713:
709:
703:
699:
697:
696:Testudinoidea
694:Superfamily:
693:
691:
687:
685:
681:
679:
675:
673:
669:
667:
663:
661:
657:
656:
653:
648:
641:
636:
634:
629:
627:
622:
621:
618:
599:
595:
591:
585:
578:
572:
565:
559:
551:
547:
543:
539:
534:
529:
525:
521:
517:
510:
494:
489:
485:
481:
480:
475:
473:
464:
460:
452:
448:
445:
442:
437:
433:
431:
421:
418:
415:
412:
404:
395:
393:
389:
385:
384:
378:
376:
372:
365:
361:
357:
348:
346:
342:
338:
334:
330:
326:
325:
320:
306:
303:
298:
295:
290:
287:
282:
279:
276:Günther, 1875
274:
271:
268:Günther, 1873
266:
263:
261:
259:
258:nomen oblitum
251:
248:
247:
245:
242:
238:
233:
228:
226:
217:
214:
213:Binomial name
210:
206:
205:
199:
196:
195:
192:
191:
184:
181:
180:
177:
174:
171:
170:
167:
166:Testudinoidea
164:
162:Superfamily:
161:
160:
157:
154:
151:
150:
147:
144:
141:
140:
137:
134:
131:
130:
127:
124:
121:
120:
117:
114:
111:
110:
107:
104:
101:
100:
95:
90:
86:
80:
75:
64:
60:
55:
52:
47:
43:
38:
33:
30:
19:
4099:Cylindraspis
3935:
3900:
3875:
3867:
3859:
3818:
3810:
3788:
3780:
3773:
3766:
3758:
3752:Erymnochelys
3750:
3743:
3737:Cerrejonemys
3735:
3727:
3719:
3713:Brontochelys
3711:
3703:
3695:
3674:
3667:
3647:
3639:
3632:
3627:Pseudemydura
3625:
3618:
3610:
3603:
3596:
3589:
3582:
3574:
3567:
3560:
3553:
3546:
3539:
3532:
3512:
3504:
3496:
3488:
3480:
3472:
3464:
3456:
3448:
3440:
3432:
3424:
3416:
3408:
3400:
3392:
3384:
3376:
3368:
3360:
3352:
3344:
3336:
3328:
3320:
3312:
3304:
3282:
3236:
3230:Prionochelys
3228:
3220:
3212:
3204:
3196:
3177:
3169:
3161:
3153:
3145:
3137:
3129:
3121:
3113:
3107:Cratochelone
3105:
3097:
3091:Bouliachelys
3089:
3081:
3073:
3067:Alienochelys
3065:
3043:
3035:
3027:
3005:
2997:
2989:
2982:
2974:
2968:Chelydropsis
2966:
2958:
2950:
2919:
2913:Basilochelys
2911:
2892:
2885:
2878:
2871:
2864:
2856:
2849:
2842:
2837:Khunnuchelys
2835:
2827:
2819:
2811:
2804:
2796:
2789:
2782:
2775:
2767:
2760:
2753:Trionychidae
2740:
2733:
2727:Allaeochelys
2725:
2691:
2684:
2678:Stigmochelys
2676:
2669:
2661:
2654:
2647:
2641:Megalochelys
2639:
2631:
2624:
2617:
2610:
2603:
2596:
2588:
2580:
2573:
2566:
2558:
2552:Cylindraspis
2550:
2543:
2536:
2531:Centrochelys
2529:
2522:
2515:
2508:Testudinidae
2497:Platysternon
2495:
2476:Vijayachelys
2474:
2467:
2460:
2455:Rhinoclemmys
2453:
2446:
2439:
2432:
2425:
2420:Melanochelys
2418:
2411:
2404:
2397:
2390:
2383:
2376:
2369:
2362:
2355:
2348:
2340:
2320:
2312:
2305:
2298:
2291:
2284:
2277:
2270:
2263:
2256:
2249:
2242:
2235:
2230:Acherontemys
2228:
2196:Sternotherus
2194:
2187:
2180:
2173:
2165:
2145:
2112:
2104:
2096:
2088:
2080:
2063:Psephophorus
2061:
2053:
2045:
2037:
2030:
2022:
2014:
1993:
1985:
1979:Pacifichelys
1977:
1969:
1962:
1954:
1948:Lepidochelys
1946:
1939:
1931:
1925:Gigantatypus
1923:
1917:Eretmochelys
1915:
1908:
1900:
1893:
1885:
1878:
1859:Chelonioidea
1827:
1779:
1547:Stigmochelys
1459:Megalochelys
1164:
1138:Cylindraspis
1060:Cheirogaster
852:Centrochelys
839:
808:
800:
791:
784:
776:
773:(subspecies:
720:
702:Testudinidae
602:. Retrieved
598:the original
593:
584:
576:
571:
563:
558:
523:
519:
509:
497:. Retrieved
483:
477:
471:
463:
449:
446:
441:Henry Barkly
438:
434:
430:Round Island
427:
419:
416:
413:
409:
391:
383:Cylindraspis
381:
379:
368:
337:Testudinidae
323:
322:
318:
316:
304:
296:
288:
280:
272:
264:
255:
249:
224:
219:
203:
202:
190:Cylindraspis
189:
176:Testudinidae
50:
29:
4001:iNaturalist
3960:Wikispecies
3790:Stupendemys
3745:Cordichelys
3591:Mesoclemmys
3576:Hydromedusa
3498:Taphrosphys
3490:Sankuchemys
3482:Rhothonemys
3458:Polysternon
3338:Cearachelys
3322:Azabbaremys
3306:Araiochelys
3222:Ctenochelys
3214:Bashuchelys
3163:Rhinochelys
3139:Ocepechelon
3131:Notochelone
3123:Iserosaurus
2991:Macrochelys
2943:Chelydridae
2866:Palaeoamyda
2791:Cyclanorbis
2656:Psammobates
2649:Oligopherus
2612:Indotestudo
2560:Cymatholcus
2538:Chelonoidis
2524:Astrochelys
2333:Geoemydidae
2251:Deirochelys
2189:Staurotypus
2182:Kinosternon
2175:Hoplochelys
2090:Peritresius
2039:Dermochelys
2032:Cosmochelys
2024:Corsochelys
1941:Itilochelys
1933:Glarichelys
1880:Allopleuron
1870:Cheloniidae
1809:Superfamily
1496:Psammobates
1346:Indotestudo
888:Chelonoidis
821:Astrochelys
499:14 November
351:Description
331:species of
284:Gadow, 1894
4088:Categories
3869:Caririemys
3782:Podocnemis
3768:Latentemys
3729:Carbonemys
3669:Pelomedusa
3506:Ummulisani
3426:Kurmademys
3394:Inaechelys
3378:Galianemys
3346:Chedighaii
3284:Araripemys
3257:Pleurodira
3238:Toxochelys
3155:Protostega
2960:Chelydrops
2880:Pelodiscus
2873:Pelochelys
2821:Gilmoremys
2798:Cycloderma
2708:Trionychia
2575:Geochelone
2568:Floridemys
2434:Notochelys
2322:Wilburemys
2293:Malaclemys
2147:Dermatemys
2114:Puppigerus
2047:Eosphargis
1964:Miocaretta
1956:Mexichelys
1850:Cryptodira
1790:Testudines
1751:Testudines
1181:Geochelone
1109:Chersobius
745:Agrionemys
690:Cryptodira
688:Suborder:
684:Testudines
676:Subclass:
604:3 February
455:References
398:Extinction
339:. It was
308:Bour, 1981
253:Gray, 1831
156:Cryptodira
152:Suborder:
146:Testudines
3909:See also
3861:Caribemys
3721:Caninemys
3705:Bauruemys
3634:Rheodytes
3598:Myuchelys
3541:Chelodina
3466:Puentemys
3442:Nigeremys
3386:Ilatardia
3330:Bothremys
3037:Basilemys
2851:Nilssonia
2829:Hutchemys
2777:Axestemys
2735:Anosteira
2590:Hadrianus
2448:Pangshura
2406:Malayemys
2371:Geoclemys
2314:Trachemys
2307:Terrapene
2300:Pseudemys
2286:Graptemys
2279:Glyptemys
2272:Emydoidea
2265:Actinemys
2237:Chrysemys
2082:Euclastes
1910:Eochelone
1761:Kingdom:
1248:Hadrianus
658:Kingdom:
542:1864-5755
345:Mauritius
197:Species:
112:Kingdom:
106:Eukaryota
49:Skull of
3945:Wikidata
3676:Pelusios
3641:Rhinemys
3612:Platemys
3605:Phrynops
3525:Chelidae
3514:Zolhafah
3410:Jainemys
3075:Archelon
2952:Chelydra
2844:Lissemys
2686:Stylemys
2671:Solitudo
2633:Manouria
2582:Gopherus
2545:Chersina
2413:Mauremys
2392:Heosemys
2385:Hardella
2378:Geoemyda
2364:Cyclemys
2220:Emydidae
2167:Claudius
2016:Arabemys
1995:Tasbacka
1987:Syllomus
1902:Chelonia
1800:Suborder
1784:Diapsida
1775:Reptilia
1769:Chordata
1767:Phylum:
1763:Animalia
1566:Stylemys
1435:Manouria
1204:Gopherus
1091:Chersina
722:tortoise
700:Family:
678:Anapsida
672:Reptilia
666:Chordata
664:Phylum:
660:Animalia
647:Tortoise
550:87809001
392:"inepta"
327:) is an
241:Synonyms
172:Family:
136:Reptilia
126:Chordata
122:Phylum:
116:Animalia
102:Domain:
79:IUCN 2.3
3993:5789418
3980:7221770
3951:Q305519
3820:Sokatra
3569:Emydura
3474:Rosasia
3370:Foxemys
3314:Arenila
2894:Trionyx
2887:Rafetus
2813:Dogania
2769:Apalone
2693:Testudo
2619:Kinixys
2605:Homopus
2462:Sacalia
2441:Orlitia
2427:Morenia
2342:Batagur
2244:Clemmys
1971:Natator
1887:Caretta
1788:Order:
1773:Class:
1669:Testudo
1374:Kinixys
1323:Homopus
682:Order:
670:Class:
371:endemic
341:endemic
329:extinct
232:Günther
222:†
201:†
182:Genus:
142:Order:
132:Class:
77: (
74:Extinct
4071:377726
4058:302061
4045:180174
4019:949458
4006:121113
3852:
3845:
3838:
3562:Elusor
3555:Elseya
3548:Chelus
3265:
3198:Adocus
3189:
2935:
2904:
2784:Chitra
2486:
2073:
1818:Family
725:family
711:Genera
649:family
548:
540:
234:, 1873
2858:Palea
2762:Amyda
2663:Pyxis
2357:Cuora
1829:Genus
1780:Clade
1524:Pyxis
997:Diego
546:S2CID
4040:NCBI
4032:6062
4027:IUCN
4014:ITIS
3988:GBIF
2258:Emys
606:2022
538:ISSN
501:2021
484:1996
317:The
3975:EoL
528:doi
488:doi
362:by
343:to
4090::
4068::
4055::
4042::
4029::
4016::
4003::
3990::
3977::
3962::
3947::
1782::
1011:†(
592:.
544:.
536:.
524:57
522:.
518:.
482:.
476:.
3874:†
3866:†
3858:†
3817:†
3809:†
3800:†
3787:†
3765:†
3757:†
3742:†
3734:†
3726:†
3718:†
3710:†
3702:†
3694:†
3646:†
3617:†
3581:†
3511:†
3503:†
3495:†
3487:†
3479:†
3471:†
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3455:†
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3439:†
3431:†
3423:†
3415:†
3407:†
3399:†
3391:†
3383:†
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3367:†
3359:†
3351:†
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3335:†
3327:†
3319:†
3311:†
3303:†
3294:†
3281:†
3272:†
3235:†
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3112:†
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3088:†
3080:†
3072:†
3064:†
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3042:†
3034:†
3026:†
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2996:†
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2957:†
2918:†
2910:†
2863:†
2834:†
2826:†
2818:†
2803:†
2774:†
2732:†
2724:†
2683:†
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2029:†
2021:†
2013:†
1992:†
1984:†
1976:†
1961:†
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1938:†
1930:†
1922:†
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1743:e
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