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Ceratophyllum

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64: 44: 257:, which are narrow and often much-branched. The forked leaves are brittle and stiff to the touch in some species, softer in others. Roots are completely absent and are missing even in the embryonic stage, but sometimes they develop modified leaves with a rootlike appearance, which anchor the plant to the bottom. Also 549:
into its appropriately recognized species, subspecies, and varieties is not settled. More than 30 species have been described and published. A narrow interpretation of this work rejects over 24 of these taxa as variants, accepting only 6 species. This narrow interpretation lumps to the point of
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grows completely submerged, usually, though not always, floating on the surface, and does not tolerate drought. The plant stems can reach 1–3 m in length. At intervals along nodes of the stem they produce rings of bright green
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failing to give these potential species the taxonomic importance of even being named on a subspecific or varietal level. The genus as narrowly defined in this manner contains the following six species:
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Yang, Yongzhi; Sun, Pengchuan; Lv, Leke; Wang, Donglei; Ru, Dafu; Li, Ying; Ma, Tao; Zhang, Lei; Shen, Xingxing; Meng, Fanbo; Jiao, Beibei; Shan, Lanxing; Liu, Man; Wang, Qingfeng; Qin, Zhiji (2020).
269:) in the autumn, which sink to the bottom and give the impression that the plant has been killed by the frost; but come spring, these will grow back into the long stems, slowly filling up the pond. 989: 1009: 598: 292:, but recent research has shown that it is not closely related to Nymphaeaceae or any other extant plant family. Some early molecular phylogenies suggested it was the 620: 555: 222:
including four accepted living species in 2016, commonly found in ponds, marshes, and quiet streams in tropical and in temperate regions. It is the only
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Frontiers | Refined Interpretation of the Pistillate Flower in Ceratophyllum Sheds Fresh Light on Gynoecium Evolution in Angiosperms
1198: 1263: 1211: 975: 1268: 963: 616:(soft/tropical hornwort) – Europe, Middle-East, Central Asia, northern and central Africa, Florida, and Dominican Republic 1107: 922:"An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III" 850: 830: 735:"An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG IV" 1413: 1392: 1294: 1237: 17: 1387: 265:
are small and inconspicuous, with the male and female flowers on the same plant. In ponds it forms thick buds (
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is considered distinctive enough to warrant its own family, Ceratophyllaceae. It was considered a relative of
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to all other angiosperms, but more recent research suggests that it is the sister group to the
1330: 1255: 1325: 1177: 1317: 1120: 8: 1151: 1400: 1172: 897: 865:"Prickly waterlily and rigid hornwort genomes shed light on early angiosperm evolution" 864: 718: 158: 58: 1312: 1094: 938: 921: 902: 884: 846: 826: 1405: 943: 933: 892: 876: 746: 694: 376: 289: 243: 227: 121: 111: 1099: 1073: 1086: 1013: 266: 231: 219: 101: 1281: 1379: 699: 88: 880: 647: 1435: 888: 413: 309: 138: 1369: 1047: 906: 332: 293: 281: 223: 1224: 1056: 683:"The number of known plants species in the world and its annual increase" 354: 319: 285: 43: 1203: 948: 398: 751: 734: 242:, although hornwort is also used for unrelated plants of the division 1229: 1146: 1133: 1018: 308:
system placed the family in its own order, the Ceratophyllales. The
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(1989). 750: 707: 698: 637: 926:Botanical Journal of the Linnean Society 739:Botanical Journal of the Linnean Society 14: 1434: 728: 726: 594:(prickly hornwort) – Near-cosmopolitan 572:(rigid/common hornwort) – cosmopolitan 1023: 1022: 956: 815: 803: 801: 786: 784: 545:The subgeneric division of the genus 1393:5979fbfa-624f-4b54-9d4e-f9478b853dce 1305:1dcef8b1-6aee-477e-88bf-ba7dd20c3a3a 968: 823:Flora of Britain and Northern Europe 920:Angiosperm Phylogeny Group (2009). 835: 766:Competition Science Vision nov 2003 733:Angiosperm Phylogeny Group (2016). 723: 312:accepts the phylogeny shown below: 24: 982: 798: 781: 25: 1463: 1000: 1357:urn:lsid:ipni.org:names:330840-2 939:10.1111/j.1095-8339.2009.00996.x 583:(spiny hornwort) – North America 62: 856: 843:New RHS Dictionary of Gardening 770: 759: 13: 1: 630: 7: 599:Ceratophyllum platyacanthum 272: 10: 1468: 962:Angiosperm Phylogeny Web: 700:10.11646/phytotaxa.261.3.1 661:Royal Botanic Gardens, Kew 656:Plants of the World Online 540: 234:. They are usually called 1031: 881:10.1038/s41477-020-0594-6 470: 455: 448: 433: 426: 411: 396: 389: 374: 367: 352: 345: 330: 323: 164: 157: 150: 145: 59:Scientific classification 57: 50: 41: 34: 841:Huxley, A., ed. (1992). 807:Flora of North America: 621:Ceratophyllum tanaiticum 610:Ceratophyllum submersum 588:Ceratophyllum muricatum 577:Ceratophyllum echinatum 52:Ceratophyllum submersum 566:Ceratophyllum demersum 556:Ceratophyllum australe 1326:Paleobiology Database 304:genome in 2020. The 226:genus in the family 1012:2012-02-26 at the 168:Hydroceratophyllon 1452:Freshwater plants 1447:Angiosperm genera 1429: 1428: 1313:Open Tree of Life 1025:Taxon identifiers 752:10.1111/boj.12385 626: 615: 605:– Europe and Asia 604: 593: 582: 571: 561: 537: 536: 528: 527: 519: 518: 510: 509: 501: 500: 492: 491: 483: 482: 261:are missing. The 204: 203: 198: 190: 182: 174: 141: 16:(Redirected from 1459: 1422: 1421: 1409: 1408: 1396: 1395: 1383: 1382: 1373: 1372: 1360: 1359: 1347: 1346: 1334: 1333: 1321: 1320: 1308: 1307: 1298: 1297: 1285: 1284: 1282:NHMSYS0000457093 1272: 1271: 1259: 1258: 1246: 1245: 1233: 1232: 1220: 1219: 1207: 1206: 1194: 1193: 1181: 1180: 1168: 1167: 1155: 1154: 1142: 1141: 1129: 1128: 1116: 1115: 1103: 1102: 1090: 1089: 1077: 1076: 1067: 1066: 1065: 1052: 1051: 1050: 1020: 1019: 994: 988:Flora Europaea: 986: 980: 972: 966: 960: 954: 953: 951: 941: 917: 911: 910: 900: 860: 854: 839: 833: 819: 813: 805: 796: 790:Flora of China: 788: 779: 774: 768: 763: 757: 756: 754: 730: 721: 716: 705: 704: 702: 678: 672: 671: 669: 667: 644: 624: 613: 602: 591: 580: 569: 559: 451: 450: 429: 428: 392: 391: 377:Austrobaileyales 370: 369: 348: 347: 326: 325: 316: 315: 290:Cronquist system 284:and included in 244:Anthocerotophyta 228:Ceratophyllaceae 220:flowering plants 196: 188: 180: 170: 137: 122:Ceratophyllaceae 67: 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4: 3: 2: 1464: 1453: 1450: 1448: 1445: 1443: 1442:Ceratophyllum 1440: 1439: 1437: 1420: 1415: 1411: 1407: 1402: 1398: 1394: 1389: 1385: 1381: 1375: 1371: 1366: 1362: 1358: 1353: 1349: 1345: 1340: 1336: 1332: 1327: 1323: 1319: 1314: 1310: 1306: 1300: 1296: 1291: 1287: 1283: 1278: 1274: 1270: 1265: 1261: 1257: 1252: 1248: 1244: 1239: 1235: 1231: 1226: 1222: 1218: 1213: 1209: 1205: 1200: 1196: 1192: 1187: 1183: 1179: 1178:Ceratophyllum 1174: 1170: 1166: 1161: 1157: 1153: 1148: 1144: 1140: 1135: 1131: 1127: 1122: 1118: 1114: 1113:Ceratophyllum 1109: 1105: 1101: 1096: 1092: 1088: 1083: 1079: 1075: 1069: 1064: 1063:Ceratophyllum 1058: 1054: 1049: 1043: 1039: 1038: 1036: 1034: 1033:Ceratophyllum 1030: 1026: 1021: 1015: 1011: 1008: 1005: 1004: 993: 992: 991:Ceratophyllum 985: 979: 978: 977:Ceratophyllum 971: 965: 959: 950: 945: 940: 935: 931: 927: 923: 916: 908: 904: 899: 894: 890: 886: 882: 878: 874: 870: 869:Nature Plants 866: 859: 852: 851:0-333-47494-5 848: 844: 838: 832: 831:0-340-40170-2 828: 824: 818: 812: 811: 810:Ceratophyllum 804: 802: 795: 794: 793:Ceratophyllum 787: 785: 778: 773: 767: 762: 753: 748: 744: 740: 736: 729: 727: 720: 715: 713: 711: 701: 696: 692: 688: 684: 677: 662: 658: 657: 653: 651: 650:Ceratophyllum 643: 641: 636: 623: 622: 618: 612: 611: 607: 601: 600: 596: 590: 589: 585: 579: 578: 574: 568: 567: 563: 558: 557: 553: 552: 551: 548: 547:Ceratophyllum 533: 532: 524: 523: 515: 514: 506: 505: 497: 496: 488: 487: 479: 478: 475: 474: 468: 467: 464: 463: 460: 459: 453: 452: 446: 445: 442: 441: 438: 437: 431: 430: 424: 423: 420: 419: 416: 415: 414:Chloranthales 409: 408: 405: 404: 401: 400: 394: 393: 387: 386: 383: 382: 379: 378: 372: 371: 365: 364: 361: 360: 357: 356: 350: 349: 343: 342: 339: 338: 335: 334: 328: 327: 321: 318: 317: 313: 311: 310:APG IV system 307: 303: 299: 295: 291: 287: 283: 279: 278:Ceratophyllum 270: 268: 264: 260: 256: 251: 250:Ceratophyllum 247: 245: 241: 237: 233: 229: 225: 221: 217: 214: 210: 209: 208:Ceratophyllum 195: 192: 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Macmillan 842: 837: 822: 817: 809: 792: 772: 761: 742: 738: 690: 686: 676: 664:. Retrieved 654: 649: 619: 608: 597: 586: 575: 564: 554: 546: 544: 471: 457: 456: 434: 412: 397: 375: 353: 333:Amborellales 331: 301: 294:sister group 282:Nymphaeaceae 277: 276: 249: 248: 239: 235: 213:cosmopolitan 207: 206: 205: 193: 185: 177: 167: 151: 132: 131: 95: 82: 51: 35: 29: 1225:iNaturalist 1057:Wikispecies 949:10654/18083 745:(1): 1–20. 355:Nymphaeales 320:angiosperms 302:C. demersum 286:Nymphaeales 102:Angiosperms 1436:Categories 631:References 399:magnoliids 1147:FloraBase 889:2055-0278 687:Phytotaxa 240:hornworts 236:coontails 197:Mavrodiev 194:Fassettia 152:See text 72:Kingdom: 1388:VicFlora 1377:VASCAN: 1370:40004488 1365:Tropicos 1243:330840-2 1108:eFloraSA 1042:Wikidata 1010:Archived 907:32094642 625:Sapjegin 473:eudicots 436:monocots 298:eudicots 273:Taxonomy 159:Synonyms 146:Species 118:Family: 1256:1341195 1204:2882396 898:8075997 666:25 July 560:Griseb. 541:Species 306:APG III 288:in the 267:turions 263:flowers 259:stomata 128:Genus: 108:Order: 76:Plantae 1419:394610 1339:PLANTS 1302:NZOR: 1191:106192 1165:106192 1074:189275 1071:APDB: 1048:Q21795 905:  895:  887:  849:  829:  581:A.Gray 255:leaves 224:extant 181:Moench 1414:WoRMS 1344:CERAT 1318:10945 1269:18402 1251:IRMNG 1230:60999 1173:FoAO2 1152:21431 1139:1CEYG 1126:61981 1087:60104 603:Cham. 592:Cham. 216:genus 211:is a 189:Röhl. 96:Clade 83:Clade 1352:POWO 1331:5268 1295:4427 1290:NCBI 1264:ITIS 1238:IPNI 1217:2302 1212:GRIN 1199:GBIF 1134:EPPO 1100:3KV7 1082:APNI 903:PMID 885:ISSN 847:ISBN 827:ISBN 668:2023 172:Ség. 1401:WFO 1380:973 1277:NBN 1186:FoC 1160:FNA 1121:EoL 1095:CoL 944:hdl 934:doi 930:161 893:PMC 877:doi 747:doi 743:181 695:doi 691:261 652:L." 238:or 218:of 1438:: 1416:: 1403:: 1390:: 1367:: 1354:: 1341:: 1328:: 1315:: 1292:: 1279:: 1266:: 1253:: 1240:: 1227:: 1214:: 1201:: 1188:: 1175:: 1162:: 1149:: 1136:: 1123:: 1110:: 1097:: 1084:: 1059:: 1044:: 942:. 928:. 924:. 901:. 891:. 883:. 871:. 867:. 825:. 800:^ 783:^ 741:. 737:. 725:^ 709:^ 689:. 685:. 659:. 639:^ 614:L. 570:L. 246:. 139:L. 98:: 85:: 952:. 946:: 936:: 909:. 879:: 873:6 853:. 755:. 749:: 703:. 697:: 670:. 648:" 20:)

Index

Ceratophyllanae

Scientific classification
Edit this classification
Plantae
Tracheophytes
Angiosperms
Ceratophyllales
Ceratophyllaceae
Ceratophyllum
L.
Synonyms
Ség.
cosmopolitan
genus
flowering plants
extant
Ceratophyllaceae
Ceratophyllales
Anthocerotophyta
leaves
stomata
flowers
turions
Nymphaeaceae
Nymphaeales
Cronquist system
sister group
eudicots
APG III

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