781:
4360:
4355:
3445:
3459:
713:
4342:
873:
118:
3452:
136:
2399:
4366:
1511:
1199:
595:
4349:
2387:
2366:
1060:
1109:
and display an incorrect, more rectangular profile. The skull was in fact triangular with a pointed snout, equipped with a beak. Its teeth were more tightly packed in the jaw compared to other
Morrison euornithopods. Museum curator John Foster describes them as having "thick median ridges on their
1037:, consisting of an anterior cervical centrum; an anterior dorsal centrum; a dorsosacral centrum; four sacral centra; a caudosacral centrum; two anterior caudal centra; three medial caudal centra; and a distal fragment from the left humerus. The Fuentecillas specimen is seen to be closer to
815:
in 1909 based on the holotype and only known specimen USNM 4753, a fragmentary postcranium, by the "narrowness or depressed nature of the ilia". Carpenter and Wilson (2008) referred this species to
2012:
Sachs, S.; Hornung, J.J. (2006). "Juvenile ornithopod (Dinosauria: Rhabdodontidae) remains from the Upper
Cretaceous (Lower Campanian, Gosau Group) of Muthmannsdorf (Lower Austria)".
1096:. The largest fragments from later strata indicate adult individuals around 6.79 metres (22.3 ft) long. The Quarry 13 individuals are smaller though. It is estimated that
1826:
Brill, K.; K. Carpenter (2007). "A description of a new ornithopod from the Lytle Member of the
Purgatoire Formation (Lower Cretaceous) and a reassessment of the skull of
1002:
by
Lydekker in 1889. However, Naish & Martill (2008), McDonald and colleagues (2010), and McDonald (2011) found that Seeley's original generic distinction was valid.
934:
dryosaurid, but a more recent analysis contends that the diagnostic features of
Dryosauridae do not overlap with its material, instead considering it an iguanodontian.
1713:
Bakker, R.T. (1998). "Dinosaur mid-life crisis: the
Jurassic-Cretaceous transition in Wyoming and Colorado". In Lucas, S.G.; Kirkland, J.I.; Estep, J.W. (eds.).
1525:), scientists believe they may have been able to achieve running speeds of 25 km per hour (15 mph). A tiny 9 inch fossilized embryo, referred to
1866:(Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb".
2231:"The largest ornithopod (Dinosauria: Ornithischia) from the Upper Jurassic of Europe sheds light on the evolutionary history of basal ankylopollexians"
780:
1570:"Tracing Late Jurassic ornithopod diversity in the eastern Iberian Peninsula: Camptosaurus-like postcranial remains from Alpuente (Valencia, Spain)"
930:
consists of the holotype and only known specimen NHMUK R167, a poorly preserved left femur lacking the distal end. It was earlier believed to be a
4851:
2452:
4925:
4910:
4940:
2155:
Norman D.B. and
Barrett, P.M. 2002. Ornithischian dinosaurs from the Lower Cretaceous (Berriasian) of England. Palaeontology 68:161-189
2230:
2131:, the earliest dryosaurid dinosaur (Ornithischia: Euornithopoda) from the Middle Jurassic of England". In Carpenter, Kenneth (ed.).
4838:
1120:
teeth frequently exhibit extensive wear, which indicates that individuals in the genus had a diet of relatively tough vegetation.
888:
species in North
America, numerous species from Europe were also referred to the genus in the late 19th and early 20th centuries:
2080:
Fonseca, André O.; Reid, Iain J.; Venner, Alexander; Duncan, Ruairidh J.; Garcia, Mauricio S.; Müller, Rodrigo T. (2024-12-31).
1070:
is a relatively heavily built form, with robust hindlimbs and broad feet, still having four toes. Due to the separate status of
4920:
4915:
4359:
1154:, of which group it would then be a basal member. This would mean that the genus is closely related to the ancestor of later
3444:
2431:
2174:
2140:
1839:
1694:
Galton, P.M.; Powell, H.P. (1980). "The ornithischian dinosaur
Camptosaurus prestwichii from the Upper Jurassic of England".
685:. Throughout the 1880 and 1890s, he continued to receive specimens from Quarry 13 and in 1894 named two additional species:
1543:
4354:
1110:
lateral sides and denticles along their edges," these features were similar to, but "more fully developed" than those in
855:
ending in a small "foot" among other features. An analysis by Andrew McDonald and colleagues in 2010 suggested that like
3458:
627:, or "flexible back", from Greek κάμπτω, "to bend" and νῶτον, "back", in reference to the presumed flexibility of the
517:
4930:
2303:
McDonald, A.T.; Kirkland, J.I.; DeBlieux, D.D.; Madsen, S.K.; Cavin, J.; Milner, A.R.C. & Panzarin, L. (2010).
829:. However, McDonald and colleagues (2010), and McDonald (2011) found that the horizontal postacetabular process of
4341:
4720:
2192:"Draconyx loureiroi, a new camptosauridae (Dinosauria, Ornithopoda) from the Late Jurassic of Lourinhã, Portugal"
794:
4743:
4730:
1030:
1906:
McDonald, A.T.; Kirkland, J.I.; DeBlieux, D.D.; Madsen, S.K.; Cavin, J.; Milner, A.R.C.; Panzarin, L. (2010).
1879:
4935:
4735:
2370:
2305:"New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs"
1908:"New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs"
1132:
to a family of its own: the
Camptosauridae. Alternatively some authors considered it an early member of the
912:(Nopcsa, 1900) consists of fragmentary material, a dentary and articular from Upper Cretaceous rocks of the
4365:
1100:
typically reached 5–6 metres (16–20 ft) long and weighed 500–1,000 kilograms (1,100–2,200 lb).
135:
2377:
844:
2424:
808:
3451:
2531:
4945:
2191:
4882:
4816:
4778:
2890:
2742:
610:
4856:
4877:
4869:
4700:
3256:
2558:
618:
4905:
4825:
4120:
2417:
2403:
2316:
2203:
2093:
2021:
1919:
1741:
1636:
1581:
985:
598:
Historical skeletal restoration by O.C. Marsh, with skull based on remains now referred to
315:
4348:
1717:. Vol. 14. New Mexico Museum of Natural History and Science Bulletin. pp. 67–77.
8:
773:
606:
404:
2320:
2207:
2097:
2025:
1975:
1923:
1745:
1640:
1585:
1103:
Earlier reconstructions, such as those by Marsh and Gilmore, were based on the skull of
4472:
4083:
2668:
2339:
2304:
1942:
1907:
1883:
1757:
1652:
1605:
1569:
1174:
1077:
833:
is more likely a product of distortion. Therefore, McDonald put it into its own genus,
812:
712:
694:
664:
390:
130:
4518:
2302:
2215:
1529:, was retrieved from Morrison Formation strata at Dinosaur National Monument in Utah.
4864:
4479:
4218:
4139:
4099:
2983:
2869:
2815:
2786:
2344:
2170:
2136:
2109:
2081:
1947:
1835:
1761:
1656:
1609:
1597:
1538:
1093:
877:
2061:, "Camptosaurus", "Iguanodon" – and referred specimens from Romania and elsewhere".
1887:
1799:
1088:, from Quarry 13, have been recovered from very deep layers, probably dating to the
4620:
4613:
4423:
4416:
3945:
3856:
3694:
3595:
3581:
3295:
3263:
2898:
2807:
2721:
2656:
2391:
2334:
2324:
2242:
2211:
2101:
2029:
1994:
1937:
1927:
1875:
1795:
1784:, with a revision of the species of the genus, and descriptions of two new species"
1749:
1644:
1589:
1428:
804:
640:
628:
544:
473:
2105:
2053:(2009). "Notes on Neocomian (Late Cretaceous) ornithopod dinosaurs from England -
2033:
4705:
4500:
4451:
4314:
4245:
4148:
3968:
3687:
3673:
3652:
3270:
3143:
2910:
2678:
2516:
2329:
2246:
2127:
Ruiz-Omeñaca, José Ignacio; Pereda Suberbiola, Xabier; Galton, Peter M. (2007). "
1932:
1779:
872:
755:
was a valid species. They also considered a skull, YPM 1887, in 1886 referred to
4715:
4710:
4648:
4585:
4458:
4307:
4270:
4252:
4197:
3829:
3645:
3638:
3567:
3489:
3476:
3404:
3392:
3361:
3336:
3200:
3122:
3083:
3069:
2946:
2857:
2779:
2757:
2750:
2491:
1593:
1340:
1218:
1180:
1151:
1133:
260:
212:
117:
1998:
1648:
1568:
Cubells-Alarcó, M.; Verdú, F. J.; Suñer, M.; de Santisteban, C. (2022-01-08).
913:
767:, but the specimen was recently shown by Brill and Carpenter not to belong to
4899:
4801:
4655:
4599:
4578:
4549:
4465:
4402:
4300:
4293:
4279:
4189:
4169:
4162:
4059:
4009:
4002:
3986:
3961:
3934:
3927:
3920:
3891:
3842:
3804:
3769:
3750:
3722:
3666:
3329:
3322:
3277:
3207:
3076:
2935:
2920:
2882:
2793:
2686:
2649:
2642:
2113:
1753:
1601:
1364:
1307:
1245:
1140:
1105:
1007:
940:
860:
789:
705:
in his 1909 redescription of the Marsh specimens. In the Morrison Formation,
614:
556:
548:
541:
534:
75:
37:
4563:
4539:
4527:
4493:
4486:
4437:
4430:
4236:
4180:
4155:
4127:
4016:
3818:
3783:
3729:
3659:
3631:
3517:
3510:
3350:
3315:
3242:
3235:
3098:
3053:
2955:
2829:
2822:
2771:
2735:
2693:
2543:
2482:
2348:
2050:
1951:
1522:
1163:
1155:
977:
956:
931:
922:
920:. It is almost certainly a rhabdodontid and is no longer considered valid (
835:
825:, on the basis of similarities between its ilium and the holotype ilium of
728:
672:
648:
432:
323:
196:
1834:. Bloomington and Indianapolis: Indiana University Press. pp. 49–67.
963:
by Norman and Barrett in 2002. It has since been transferred to the genus
4810:
4752:
4641:
4606:
4444:
4286:
4259:
4211:
4204:
4037:
4023:
3995:
3975:
3954:
3898:
3877:
3870:
3863:
3797:
3790:
3743:
3708:
3680:
3616:
3609:
3378:
3228:
3214:
3186:
3164:
3106:
2969:
2800:
2764:
2728:
2624:
2611:
2570:
2440:
2135:. Bloomington and Indianapolis: Indiana University Press. pp. 3–16.
1159:
988:
was driven into the side of a hill. It was described by Hulke in 1880 as
981:
623:
417:
228:
50:
20:
2082:"A comprehensive phylogenetic analysis on early ornithischian evolution"
1047:
from the Jurassic of Portugal, hence its tentative placement within the
4843:
4694:
4669:
4662:
4634:
4627:
4507:
4393:
4092:
4072:
4030:
3905:
3757:
3624:
3574:
3539:
3532:
3503:
3343:
3221:
3193:
3062:
3035:
3010:
2850:
2843:
2836:
2714:
1388:
1290:
1203:
1168:
1112:
681:
656:
379:
244:
95:
60:
2278:
Jurassic West: The Dinosaurs of the Morrison Formation and Their World
1814:
Jurassic West: The Dinosaurs of the Morrison Formation and Their World
1567:
1510:
1198:
4676:
4592:
4571:
4409:
4106:
4044:
3553:
3525:
3385:
3249:
3179:
3150:
2976:
2962:
2705:
1188:
1089:
950:
817:
594:
147:
100:
44:
4772:
2229:
Sánchez-Fenollosa, Sergio; Verdú, Francisco; Cobos, Alberto (2023).
1729:
1624:
938:
is probably a valid dryosaurid that has been moved to the new genus
4795:
4556:
4225:
4113:
3913:
3884:
3811:
3736:
3701:
3602:
3588:
3546:
3370:
3171:
3136:
3042:
3028:
2990:
2500:
2473:
2464:
2126:
1412:
1043:
994:
965:
632:
537:
425:
372:
180:
167:
90:
85:
70:
65:
55:
639:
by him in 1885 because the original name was already in use for a
4830:
3849:
3284:
1259:
1072:
917:
865:
852:
660:
449:
357:
344:
105:
80:
1029:. A single specimen was found in the Fuentecillas member of the
1021:
In 2022, the first confirmed European remains belonging to aff.
747:
to be different growth stages or different gender of the larger
3715:
3308:
2458:
2365:
1059:
859:
is actually more closely related to more advanced iguanodonts (
843:, was named by Carpenter and Wilson in 2008 for specimens from
552:
157:
2409:
1905:
763:
as well. Gilmore had used this skull to describe the skull of
651:
of the genus) for material he received from his collectors at
3560:
3090:
1148:
1034:
580:
573:
566:
559:
530:
503:
2928:
2228:
787:
skeleton mounted in outdated quadrupedal posture, and with
506:
494:
720:
skeleton at the AMNH, now thought to be a growth stage of
1880:
10.2992/0097-4463(2008)76[227:ansoco]2.0.co;2
851:
in the lower jaw, shorter neck vertebrae, and straighter
482:
2190:
Mateus, Octávio; Telles Antunes, Miguel (January 2001).
1693:
1517:(blue) compared in size to a human and other ornithopods
635:
was YPM 1877, a partial skeleton. The genus was renamed
2079:
1861:
1010:, more closely related to advanced iguanodonts than to
771:. In 2007, they put it into its own genus and species,
2268:
2266:
2264:
2262:
2260:
2258:
2256:
2011:
2375:
2133:
Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs
1832:
Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs
1825:
518:
509:
500:
479:
476:
2189:
1862:
Carpenter, K.; Wilson, Y. (2008). "A new species of
491:
488:
368:
353:
2253:
759:by Marsh, later confirmed by Gilmore, to belong to
497:
485:
1973:
1857:
1855:
1853:
1851:
1730:"The typical Ornithopoda of the American Jurassic"
1715:Lower and Middle Cretaceous Terrestrial Ecosystems
1076:it has become problematic which material from the
1901:
1899:
1897:
1670:Marsh, O.C. (1885). "Names of extinct reptiles".
4897:
1788:Proceedings of the United States National Museum
709:fossils are present in stratigraphic zones 2–6.
671:based on the holotype YPM 1879, a foot found by
1848:
679:. The foot was later shown to have belonged to
2515:
1894:
1806:
1689:
1687:
1685:
1563:
1561:
1559:
3487:
2425:
2164:
1819:
1777:
1521:Based on studies of other iguanodonts (clade
2222:
2005:
1712:
1084:. The specimens with certainty belonging to
2296:
1969:
1967:
1965:
1963:
1961:
1773:
1771:
1682:
1556:
2622:
2432:
2418:
998:by Seeley in 1888, but was soon sunk into
731:and H.P. Powell in their redescription of
116:
2338:
2328:
2235:Zoological Journal of the Linnean Society
2120:
1941:
1931:
2045:
2043:
1958:
1816:. Indiana University Press. pp. 327-329.
1768:
1509:
1197:
1058:
1025:.sp were described by Sánchez-Fenollosa
984:in England. The fossil was found when a
871:
779:
711:
593:
2280:. Indiana University Press. p. 219-221.
976:was recovered from Chawley Brick Pits,
589:
4898:
2291:The Princeton Field Guide to Dinosaurs
2049:
863:). It has been moved to the new genus
4777:
4776:
4381:
3474:
2609:
2451:
2413:
2040:
1976:"The taxonomy of species assigned to
1727:
1706:
1669:
1622:
1191:analyses of McDonald and colleagues,
4382:
1629:American Journal of Science and Arts
1544:Paleobiota of the Morrison Formation
555:. The name means 'flexible lizard' (
4926:Taxa named by Othniel Charles Marsh
4911:Dinosaurs of the Morrison Formation
2293:, Princeton University Press p. 284
2086:Journal of Systematic Palaeontology
1780:"Osteology of the Jurassic reptile
992:, and then placed in its own genus
13:
3475:
14:
4957:
4941:Late Jurassic dinosaurs of Europe
2359:
1625:"Notice of new Jurassic reptiles"
1123:
4364:
4358:
4353:
4347:
4340:
3457:
3450:
3443:
2397:
2385:
2364:
2167:Fossil Reptiles of Great Britain
1830:". In Carpenter, Kenneth (ed.).
621:described and named the find as
472:
134:
125:Reconstructed skeleton in Japan
48:
4721:List of informal dinosaur names
2610:
2439:
2283:
2183:
2158:
2149:
2073:
1812:Foster, J. (2007). "Appendix."
1800:10.5479/si.00963801.36-1666.197
1505:
972:The remaining European species
795:Natural History Museum of Milan
4731:Timeline of hadrosaur research
2165:Benton MJ, Spencer PS (1995).
1721:
1663:
1616:
1054:
1031:Villar del Arzobispo Formation
697:named two additional species,
1:
4921:Fossil taxa described in 1885
4916:Fossil taxa described in 1879
2216:10.1016/S0753-3969(01)88003-4
2106:10.1080/14772019.2024.2346577
2034:10.1016/j.geobios.2005.01.003
1868:Annals of the Carnegie Museum
1549:
613:found the remains of a small
2330:10.1371/journal.pone.0014075
1933:10.1371/journal.pone.0014075
1166:than contemporaries such as
735:(see following), considered
581:
567:
364:Carpenter & Wilson, 2008
7:
1974:Andrew T. McDonald (2011).
1734:American Journal of Science
1672:American Journal of Science
1532:
884:While Marsh was describing
617:. That same year Professor
10:
4962:
2247:10.1093/zoolinnean/zlad076
1980:(Dinosauria: Ornithopoda)"
1594:10.1007/s41513-021-00182-z
1574:Journal of Iberian Geology
1147:by definition part of the
845:Dinosaur National Monument
574:
560:
18:
16:Extinct genus of dinosaurs
4785:
4686:
4537:
4517:
4392:
4388:
4377:
4338:
4269:
4235:
4179:
4138:
4082:
4067:
4058:
3985:
3944:
3837:
3828:
3778:
3767:
3498:
3483:
3470:
3441:
3403:
3360:
3303:
3294:
3131:
3120:
3052:
3023:
3019:
3008:
2945:
2909:
2877:
2868:
2704:
2667:
2637:
2633:
2618:
2605:
2569:
2551:
2542:
2524:
2511:
2447:
1999:10.11646/zootaxa.2783.1.4
1649:10.2475/ajs.s3-18.108.501
1426:
1409:
1402:
1385:
1378:
1361:
1354:
1337:
1330:
1304:
1287:
1280:
1273:
1256:
1249:
1229:
1222:
1014:, similar to the case of
959:in 1874 and was moved to
841:Camptosaurus aphanoecetes
839:. An additional species,
809:Hot Springs, South Dakota
693:, based in part on size.
396:
389:
350:
342:
329:
322:
131:Scientific classification
129:
124:
115:
30:
2196:Annales de Paléontologie
1754:10.2475/ajs.s3-48.283.85
1006:has been recovered as a
974:Camptosaurus prestwichii
533:of plant-eating, beaked
19:Not to be confused with
4931:Paleontology in Wyoming
1195:was placed as follows:
1162:dinosaurs and was more
1128:Marsh in 1885 assigned
803:was recovered from the
643:. In 1879, Marsh named
2169:. Chapman & Hall.
2063:Revue de Paléobiologie
1778:Gilmore, C.W. (1909).
1518:
1211:
1064:
881:
811:. It was described by
801:Camptosaurus depressus
797:
724:
611:Albany County, Wyoming
602:
4878:Paleobiology Database
4701:Basturs Poble bonebed
2373:at Wikimedia Commons
1513:
1206:dinosaurs, including
1201:
1187:In the 2010 and 2011
1062:
990:Iguanodon prestwichii
948:was originally named
876:Cast of a skull from
875:
783:
751:, and therefore only
715:
619:Othniel Charles Marsh
605:On September 4, 1879
597:
586:) meaning 'lizard').
572:) meaning 'bent' and
4936:Ornithischian genera
4723:(contains hadrosaur
4121:Probrachylophosaurus
2272:Foster, J. (2007). "
2129:Callovosaurus leedsi
1728:Marsh, O.C. (1894).
1623:Marsh, O.C. (1879).
716:Outdated mount of a
590:History of discovery
2321:2010PLoSO...514075M
2274:Camptosaurus dispar
2208:2001AnPal..87...61M
2098:2024JSPal..2246577F
2026:2006Geobi..39..415S
1924:2010PLoSO...514075M
1746:1894AmJS...48...85M
1641:1879AmJS...18..501M
1586:2022JIbG...48...65S
1515:Camptosaurus dispar
1202:Restoration of six
1086:Camptosaurus dispar
1012:Camptosaurus dispar
774:Theiophytalia kerri
607:William Harlow Reed
4473:Ornithopodichnites
4084:Brachylophosaurini
2669:Hypsilophodontidae
2289:Paul, G.S., 2010,
1519:
1212:
1078:Morrison Formation
1065:
882:
847:. It differs from
827:Planicoxa venenica
798:
725:
695:Charles W. Gilmore
665:Morrison Formation
603:
551:and possibly also
332:Camptonotus dispar
4893:
4892:
4865:Open Tree of Life
4779:Taxon identifiers
4770:
4769:
4766:
4765:
4762:
4761:
4480:Shiraminesauropus
4373:
4372:
4336:
4335:
4332:
4331:
4328:
4327:
4324:
4323:
4219:Naashoibitosaurus
4140:Austrokritosauria
4100:Brachylophosaurus
4054:
4053:
3466:
3465:
3439:
3438:
3435:
3434:
3431:
3430:
3427:
3426:
3423:
3422:
3116:
3115:
3004:
3003:
3000:
2999:
2984:Transylvanosaurus
2870:Rhabdodontomorpha
2816:Notohypsilophodon
2787:Macrogryphosaurus
2601:
2600:
2597:
2596:
2593:
2592:
2589:
2588:
2369:Media related to
2176:978-0-412-62040-9
2142:978-0-253-34817-3
1841:978-0-253-34817-3
1794:(1666): 197–332.
1539:List of dinosaurs
1502:
1501:
1493:
1492:
1484:
1483:
1475:
1474:
1466:
1465:
1457:
1456:
1448:
1447:
1439:
1438:
1319:
1318:
878:Bone Cabin Quarry
807:near the town of
463:
462:
457:
445:
436:
421:
408:
383:
365:
318:
296:
279:
43:156.3–146.8
4953:
4886:
4885:
4873:
4872:
4860:
4859:
4847:
4846:
4834:
4833:
4821:
4820:
4819:
4806:
4805:
4804:
4774:
4773:
4747:mummy S.M. R4036
4621:Microhadrosaurus
4614:Mandschurosaurus
4424:Camptosaurichnus
4417:Bonaparteichnium
4390:
4389:
4383:Related articles
4379:
4378:
4368:
4362:
4357:
4352:
4351:
4344:
4065:
4064:
3946:Parasaurolophini
3857:Angulomastacator
3835:
3834:
3776:
3775:
3695:Shuangmiaosaurus
3596:Huehuecanauhtlus
3582:Gongpoquansaurus
3496:
3495:
3485:
3484:
3472:
3471:
3461:
3455:
3454:
3447:
3301:
3300:
3296:Hadrosauriformes
3264:Ratchasimasaurus
3129:
3128:
3021:
3020:
3017:
3016:
2899:Muttaburrasaurus
2875:
2874:
2808:Muttaburrasaurus
2722:Atlascopcosaurus
2657:Thescelosauridae
2635:
2634:
2631:
2630:
2620:
2619:
2607:
2606:
2549:
2548:
2522:
2521:
2513:
2512:
2506:
2505:
2449:
2448:
2434:
2427:
2420:
2411:
2410:
2402:
2401:
2400:
2390:
2389:
2388:
2381:
2368:
2353:
2352:
2342:
2332:
2300:
2294:
2287:
2281:
2270:
2251:
2250:
2241:(4): 1013–1033.
2226:
2220:
2219:
2187:
2181:
2180:
2162:
2156:
2153:
2147:
2146:
2124:
2118:
2117:
2077:
2071:
2070:
2051:Galton, Peter M.
2047:
2038:
2037:
2009:
2003:
2002:
1984:
1971:
1956:
1955:
1945:
1935:
1903:
1892:
1891:
1859:
1846:
1845:
1823:
1817:
1810:
1804:
1803:
1775:
1766:
1765:
1725:
1719:
1718:
1710:
1704:
1703:
1691:
1680:
1679:
1667:
1661:
1660:
1635:(108): 501–505.
1620:
1614:
1613:
1565:
1429:Hadrosauriformes
1405:
1404:
1381:
1380:
1357:
1356:
1333:
1332:
1283:
1282:
1276:
1275:
1252:
1251:
1225:
1224:
1215:
1214:
805:Lakota Formation
629:sacral vertebrae
584:
577:
576:
570:
563:
562:
525:
521:
516:
515:
512:
511:
508:
505:
502:
499:
496:
493:
490:
487:
484:
481:
478:
455:
443:
431:
415:
403:
378:
370:
363:
355:
313:
306:
294:
289:
277:
272:
259:
243:
227:
211:
195:
139:
138:
120:
110:
47:
36:Temporal range:
28:
27:
4961:
4960:
4956:
4955:
4954:
4952:
4951:
4950:
4896:
4895:
4894:
4889:
4881:
4876:
4868:
4863:
4855:
4850:
4842:
4837:
4829:
4824:
4815:
4814:
4809:
4800:
4799:
4794:
4781:
4771:
4758:
4739:mummy AMNH 5060
4706:Dakota (fossil)
4682:
4533:
4513:
4501:Tetrapodosaurus
4452:Hadrosaurichnus
4384:
4369:
4346:
4320:
4315:Shantungosaurus
4265:
4246:Augustynolophus
4231:
4175:
4149:Bonapartesaurus
4134:
4078:
4050:
3981:
3969:Parasaurolophus
3940:
3824:
3772:
3763:
3688:Qianjiangsaurus
3674:Probactrosaurus
3653:Penelopognathus
3492:
3479:
3462:
3449:
3419:
3399:
3356:
3290:
3271:Riabininohadros
3144:Bayannurosaurus
3125:
3112:
3048:
3013:
2996:
2941:
2911:Tenontosauridae
2905:
2864:
2700:
2679:Gideonmantellia
2663:
2627:
2614:
2585:
2565:
2538:
2532:Avemetatarsalia
2517:Avemetatarsalia
2507:
2454:
2453:
2443:
2438:
2408:
2398:
2396:
2386:
2384:
2376:
2362:
2357:
2356:
2301:
2297:
2288:
2284:
2271:
2254:
2227:
2223:
2188:
2184:
2177:
2163:
2159:
2154:
2150:
2143:
2125:
2121:
2078:
2074:
2048:
2041:
2010:
2006:
1982:
1972:
1959:
1904:
1895:
1860:
1849:
1842:
1824:
1820:
1811:
1807:
1776:
1769:
1726:
1722:
1711:
1707:
1692:
1683:
1668:
1664:
1621:
1617:
1566:
1557:
1552:
1535:
1508:
1503:
1494:
1485:
1476:
1467:
1458:
1449:
1440:
1320:
1210:at the far left
1126:
1057:
857:C. aphanoecetes
813:Charles Gilmore
592:
523:
519:
475:
471:
454:
442:
430:
414:
402:
377:
362:
358:C. aphanoecetes
338:
335:
312:
304:
293:
287:
276:
274:Camptosauroidea
270:
257:
241:
225:
209:
193:
133:
111:
109:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
42:
41:
34:
24:
17:
12:
11:
5:
4959:
4949:
4948:
4946:Jurassic Spain
4943:
4938:
4933:
4928:
4923:
4918:
4913:
4908:
4891:
4890:
4888:
4887:
4874:
4861:
4848:
4835:
4822:
4807:
4791:
4789:
4783:
4782:
4768:
4767:
4764:
4763:
4760:
4759:
4757:
4756:
4749:
4741:
4733:
4728:
4718:
4716:Hypsilophodont
4713:
4711:Hadrosaur diet
4708:
4703:
4698:
4690:
4688:
4687:Other articles
4684:
4683:
4681:
4680:
4673:
4666:
4659:
4652:
4649:Stephanosaurus
4645:
4638:
4631:
4624:
4617:
4610:
4603:
4596:
4589:
4586:Fulgurotherium
4582:
4575:
4568:
4560:
4553:
4545:
4543:
4535:
4534:
4532:
4531:
4523:
4521:
4515:
4514:
4512:
4511:
4504:
4497:
4490:
4483:
4476:
4469:
4462:
4459:Iguanodontipus
4455:
4448:
4441:
4434:
4427:
4420:
4413:
4406:
4398:
4396:
4386:
4385:
4375:
4374:
4371:
4370:
4339:
4337:
4334:
4333:
4330:
4329:
4326:
4325:
4322:
4321:
4319:
4318:
4311:
4308:Laiyangosaurus
4304:
4297:
4290:
4283:
4275:
4273:
4271:Edmontosaurini
4267:
4266:
4264:
4263:
4256:
4253:Prosaurolophus
4249:
4241:
4239:
4233:
4232:
4230:
4229:
4222:
4215:
4208:
4201:
4198:Coahuilasaurus
4194:
4185:
4183:
4177:
4176:
4174:
4173:
4166:
4159:
4152:
4144:
4142:
4136:
4135:
4133:
4132:
4124:
4117:
4110:
4103:
4096:
4088:
4086:
4080:
4079:
4077:
4076:
4068:
4062:
4056:
4055:
4052:
4051:
4049:
4048:
4041:
4034:
4027:
4020:
4013:
4006:
3999:
3991:
3989:
3983:
3982:
3980:
3979:
3972:
3965:
3958:
3950:
3948:
3942:
3941:
3939:
3938:
3931:
3924:
3917:
3910:
3902:
3895:
3888:
3881:
3874:
3867:
3860:
3853:
3846:
3838:
3832:
3830:Lambeosaurinae
3826:
3825:
3823:
3822:
3815:
3808:
3801:
3794:
3787:
3779:
3773:
3768:
3765:
3764:
3762:
3761:
3754:
3747:
3740:
3733:
3726:
3719:
3712:
3705:
3698:
3691:
3684:
3677:
3670:
3663:
3656:
3649:
3646:Nanyangosaurus
3642:
3639:Nanningosaurus
3635:
3628:
3621:
3613:
3606:
3599:
3592:
3585:
3578:
3571:
3568:Gilmoreosaurus
3564:
3557:
3550:
3543:
3536:
3529:
3522:
3514:
3507:
3499:
3493:
3490:Hadrosauroidea
3488:
3481:
3480:
3477:Hadrosauroidea
3468:
3467:
3464:
3463:
3442:
3440:
3437:
3436:
3433:
3432:
3429:
3428:
3425:
3424:
3421:
3420:
3418:
3417:
3416:
3415:
3409:
3407:
3405:Hadrosauroidea
3401:
3400:
3398:
3397:
3393:Mantellisaurus
3389:
3382:
3375:
3366:
3364:
3362:Iguanodontidae
3358:
3357:
3355:
3354:
3347:
3340:
3337:Mantellisaurus
3333:
3326:
3319:
3312:
3304:
3298:
3292:
3291:
3289:
3288:
3281:
3274:
3267:
3260:
3253:
3246:
3239:
3232:
3225:
3218:
3211:
3204:
3201:Iguanacolossus
3197:
3190:
3183:
3176:
3168:
3161:
3154:
3147:
3140:
3132:
3126:
3123:Ankylopollexia
3121:
3118:
3117:
3114:
3113:
3111:
3110:
3103:
3095:
3087:
3084:Eousdryosaurus
3080:
3073:
3070:Dysalotosaurus
3066:
3058:
3056:
3050:
3049:
3047:
3046:
3039:
3032:
3024:
3014:
3009:
3006:
3005:
3002:
3001:
2998:
2997:
2995:
2994:
2987:
2980:
2973:
2966:
2959:
2951:
2949:
2947:Rhabdodontidae
2943:
2942:
2940:
2939:
2932:
2925:
2916:
2914:
2907:
2906:
2904:
2903:
2895:
2887:
2878:
2872:
2866:
2865:
2863:
2862:
2858:Weewarrasaurus
2854:
2847:
2840:
2833:
2826:
2819:
2812:
2804:
2797:
2790:
2783:
2780:Leaellynasaura
2776:
2768:
2761:
2758:Gasparinisaura
2754:
2751:Galleonosaurus
2747:
2739:
2732:
2725:
2718:
2710:
2708:
2702:
2701:
2699:
2698:
2690:
2683:
2674:
2672:
2665:
2664:
2662:
2661:
2653:
2646:
2638:
2628:
2623:
2616:
2615:
2603:
2602:
2599:
2598:
2595:
2594:
2591:
2590:
2587:
2586:
2584:
2583:
2582:
2581:
2575:
2573:
2567:
2566:
2564:
2563:
2562:
2561:
2552:
2546:
2540:
2539:
2537:
2536:
2535:
2534:
2525:
2519:
2509:
2508:
2504:
2503:
2494:
2492:Neornithischia
2485:
2476:
2467:
2461:
2445:
2444:
2437:
2436:
2429:
2422:
2414:
2407:
2406:
2394:
2361:
2360:External links
2358:
2355:
2354:
2315:(11): e14075.
2295:
2282:
2252:
2221:
2182:
2175:
2157:
2148:
2141:
2119:
2072:
2039:
2020:(3): 415–425.
2004:
1957:
1918:(11): e14075.
1893:
1874:(4): 227–263.
1847:
1840:
1818:
1805:
1767:
1740:(283): 85–90.
1720:
1705:
1681:
1662:
1615:
1554:
1553:
1551:
1548:
1547:
1546:
1541:
1534:
1531:
1507:
1504:
1500:
1499:
1496:
1495:
1491:
1490:
1487:
1486:
1482:
1481:
1478:
1477:
1473:
1472:
1469:
1468:
1464:
1463:
1460:
1459:
1455:
1454:
1451:
1450:
1446:
1445:
1442:
1441:
1437:
1436:
1433:
1432:
1425:
1422:
1421:
1418:
1417:
1408:
1403:
1401:
1398:
1397:
1394:
1393:
1384:
1379:
1377:
1374:
1373:
1370:
1369:
1360:
1355:
1353:
1350:
1349:
1346:
1345:
1341:Iguanacolossus
1336:
1331:
1329:
1326:
1325:
1322:
1321:
1317:
1316:
1313:
1312:
1303:
1300:
1299:
1296:
1295:
1286:
1281:
1279:
1274:
1272:
1269:
1268:
1265:
1264:
1255:
1250:
1248:
1242:
1241:
1238:
1237:
1228:
1223:
1221:
1219:Ankylopollexia
1213:
1181:Othnielosaurus
1152:Ankylopollexia
1134:Iguanodontidae
1125:
1124:Classification
1122:
1056:
1053:
1008:styracosternan
727:Then in 1980,
591:
588:
461:
460:
459:
458:
456:McDonald, 2011
446:
437:
422:
409:
394:
393:
387:
386:
385:
384:
373:C. prestwichii
366:
348:
347:
340:
339:
336:
327:
326:
320:
319:
302:
298:
297:
291:Camptosauridae
285:
281:
280:
268:
264:
263:
261:Ankylopollexia
255:
248:
247:
239:
232:
231:
223:
216:
215:
213:Neornithischia
207:
200:
199:
191:
184:
183:
178:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
127:
126:
122:
121:
113:
112:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
35:
15:
9:
6:
4:
3:
2:
4958:
4947:
4944:
4942:
4939:
4937:
4934:
4932:
4929:
4927:
4924:
4922:
4919:
4917:
4914:
4912:
4909:
4907:
4904:
4903:
4901:
4884:
4879:
4875:
4871:
4866:
4862:
4858:
4853:
4849:
4845:
4840:
4836:
4832:
4827:
4823:
4818:
4812:
4808:
4803:
4797:
4793:
4792:
4790:
4788:
4784:
4780:
4775:
4755:
4754:
4750:
4748:
4746:
4745:Edmontosaurus
4742:
4740:
4738:
4737:Edmontosaurus
4734:
4732:
4729:
4726:
4722:
4719:
4717:
4714:
4712:
4709:
4707:
4704:
4702:
4699:
4697:
4696:
4692:
4691:
4689:
4685:
4679:
4678:
4674:
4672:
4671:
4667:
4665:
4664:
4660:
4658:
4657:
4656:Syngonosaurus
4653:
4651:
4650:
4646:
4644:
4643:
4639:
4637:
4636:
4632:
4630:
4629:
4625:
4623:
4622:
4618:
4616:
4615:
4611:
4609:
4608:
4604:
4602:
4601:
4600:Koutalisaurus
4597:
4595:
4594:
4590:
4588:
4587:
4583:
4581:
4580:
4579:Eucercosaurus
4576:
4574:
4573:
4569:
4566:
4565:
4561:
4559:
4558:
4554:
4552:
4551:
4550:Arstanosaurus
4547:
4546:
4544:
4542:
4541:
4536:
4530:
4529:
4525:
4524:
4522:
4520:
4516:
4510:
4509:
4505:
4503:
4502:
4498:
4496:
4495:
4491:
4489:
4488:
4484:
4482:
4481:
4477:
4475:
4474:
4470:
4468:
4467:
4466:Orcauichnites
4463:
4461:
4460:
4456:
4454:
4453:
4449:
4447:
4446:
4442:
4440:
4439:
4435:
4433:
4432:
4428:
4426:
4425:
4421:
4419:
4418:
4414:
4412:
4411:
4407:
4405:
4404:
4403:Amblydactylus
4400:
4399:
4397:
4395:
4391:
4387:
4380:
4376:
4367:
4363:
4361:
4356:
4350:
4343:
4317:
4316:
4312:
4310:
4309:
4305:
4303:
4302:
4301:Kundurosaurus
4298:
4296:
4295:
4294:Kerberosaurus
4291:
4289:
4288:
4284:
4282:
4281:
4280:Edmontosaurus
4277:
4276:
4274:
4272:
4268:
4262:
4261:
4257:
4255:
4254:
4250:
4248:
4247:
4243:
4242:
4240:
4238:
4234:
4228:
4227:
4223:
4221:
4220:
4216:
4214:
4213:
4209:
4207:
4206:
4202:
4200:
4199:
4195:
4192:
4191:
4190:Anasazisaurus
4187:
4186:
4184:
4182:
4178:
4172:
4171:
4170:Secernosaurus
4167:
4165:
4164:
4163:Kelumapusaura
4160:
4158:
4157:
4153:
4151:
4150:
4146:
4145:
4143:
4141:
4137:
4130:
4129:
4125:
4123:
4122:
4118:
4116:
4115:
4111:
4109:
4108:
4104:
4102:
4101:
4097:
4095:
4094:
4090:
4089:
4087:
4085:
4081:
4075:
4074:
4070:
4069:
4066:
4063:
4061:
4060:Saurolophinae
4057:
4047:
4046:
4042:
4040:
4039:
4035:
4033:
4032:
4028:
4026:
4025:
4021:
4019:
4018:
4014:
4012:
4011:
4010:Hypacrosaurus
4007:
4005:
4004:
4003:Corythosaurus
4000:
3998:
3997:
3993:
3992:
3990:
3988:
3987:Lambeosaurini
3984:
3978:
3977:
3973:
3971:
3970:
3966:
3964:
3963:
3962:Charonosaurus
3959:
3957:
3956:
3952:
3951:
3949:
3947:
3943:
3937:
3936:
3935:Tsintaosaurus
3932:
3930:
3929:
3928:Pararhabdodon
3925:
3923:
3922:
3921:Nipponosaurus
3918:
3916:
3915:
3911:
3908:
3907:
3903:
3901:
3900:
3896:
3894:
3893:
3892:Jaxartosaurus
3889:
3887:
3886:
3882:
3880:
3879:
3875:
3873:
3872:
3868:
3866:
3865:
3861:
3859:
3858:
3854:
3852:
3851:
3847:
3845:
3844:
3843:Adynomosaurus
3840:
3839:
3836:
3833:
3831:
3827:
3821:
3820:
3816:
3814:
3813:
3809:
3807:
3806:
3805:Lapampasaurus
3802:
3800:
3799:
3795:
3793:
3792:
3788:
3786:
3785:
3781:
3780:
3777:
3774:
3771:
3770:Hadrosauridae
3766:
3760:
3759:
3755:
3753:
3752:
3751:Zhanghenglong
3748:
3746:
3745:
3741:
3739:
3738:
3734:
3732:
3731:
3727:
3725:
3724:
3723:Telmatosaurus
3720:
3718:
3717:
3713:
3711:
3710:
3706:
3704:
3703:
3699:
3697:
3696:
3692:
3690:
3689:
3685:
3683:
3682:
3678:
3676:
3675:
3671:
3669:
3668:
3667:Portellsaurus
3664:
3662:
3661:
3657:
3655:
3654:
3650:
3648:
3647:
3643:
3641:
3640:
3636:
3634:
3633:
3629:
3627:
3626:
3622:
3619:
3618:
3614:
3612:
3611:
3607:
3605:
3604:
3600:
3598:
3597:
3593:
3591:
3590:
3586:
3584:
3583:
3579:
3577:
3576:
3572:
3570:
3569:
3565:
3563:
3562:
3558:
3556:
3555:
3551:
3549:
3548:
3544:
3542:
3541:
3537:
3535:
3534:
3530:
3528:
3527:
3523:
3520:
3519:
3515:
3513:
3512:
3508:
3506:
3505:
3501:
3500:
3497:
3494:
3491:
3486:
3482:
3478:
3473:
3469:
3460:
3456:
3453:
3446:
3413:
3412:
3411:
3410:
3408:
3406:
3402:
3395:
3394:
3390:
3388:
3387:
3383:
3381:
3380:
3376:
3373:
3372:
3368:
3367:
3365:
3363:
3359:
3353:
3352:
3348:
3346:
3345:
3341:
3339:
3338:
3334:
3332:
3331:
3330:Jinzhousaurus
3327:
3325:
3324:
3323:Hypselospinus
3320:
3318:
3317:
3313:
3311:
3310:
3306:
3305:
3302:
3299:
3297:
3293:
3287:
3286:
3282:
3280:
3279:
3278:Theiophytalia
3275:
3273:
3272:
3268:
3266:
3265:
3261:
3259:
3258:
3254:
3252:
3251:
3247:
3245:
3244:
3240:
3238:
3237:
3233:
3231:
3230:
3226:
3224:
3223:
3219:
3217:
3216:
3212:
3210:
3209:
3208:Lanzhousaurus
3205:
3203:
3202:
3198:
3196:
3195:
3191:
3189:
3188:
3184:
3182:
3181:
3177:
3174:
3173:
3169:
3167:
3166:
3162:
3160:
3159:
3155:
3153:
3152:
3148:
3146:
3145:
3141:
3139:
3138:
3134:
3133:
3130:
3127:
3124:
3119:
3109:
3108:
3104:
3101:
3100:
3096:
3093:
3092:
3088:
3086:
3085:
3081:
3079:
3078:
3077:Elrhazosaurus
3074:
3072:
3071:
3067:
3065:
3064:
3060:
3059:
3057:
3055:
3051:
3045:
3044:
3040:
3038:
3037:
3033:
3031:
3030:
3026:
3025:
3022:
3018:
3015:
3012:
3007:
2993:
2992:
2988:
2986:
2985:
2981:
2979:
2978:
2974:
2972:
2971:
2967:
2965:
2964:
2960:
2958:
2957:
2953:
2952:
2950:
2948:
2944:
2938:
2937:
2936:Tenontosaurus
2933:
2931:
2930:
2926:
2923:
2922:
2921:Convolosaurus
2918:
2917:
2915:
2912:
2908:
2901:
2900:
2896:
2893:
2892:
2888:
2885:
2884:
2883:Ampelognathus
2880:
2879:
2876:
2873:
2871:
2867:
2860:
2859:
2855:
2853:
2852:
2848:
2846:
2845:
2841:
2839:
2838:
2834:
2832:
2831:
2827:
2825:
2824:
2820:
2818:
2817:
2813:
2810:
2809:
2805:
2803:
2802:
2798:
2796:
2795:
2794:Mahuidacursor
2791:
2789:
2788:
2784:
2782:
2781:
2777:
2774:
2773:
2769:
2767:
2766:
2762:
2760:
2759:
2755:
2753:
2752:
2748:
2745:
2744:
2740:
2738:
2737:
2733:
2731:
2730:
2726:
2724:
2723:
2719:
2717:
2716:
2712:
2711:
2709:
2707:
2703:
2696:
2695:
2691:
2689:
2688:
2687:Hypsilophodon
2684:
2681:
2680:
2676:
2675:
2673:
2670:
2666:
2659:
2658:
2654:
2652:
2651:
2650:Callovosaurus
2647:
2645:
2644:
2643:Burianosaurus
2640:
2639:
2636:
2632:
2629:
2626:
2621:
2617:
2613:
2608:
2604:
2579:
2578:
2577:
2576:
2574:
2572:
2568:
2560:
2556:
2555:
2554:
2553:
2550:
2547:
2545:
2541:
2533:
2529:
2528:
2527:
2526:
2523:
2520:
2518:
2514:
2510:
2502:
2498:
2495:
2493:
2489:
2486:
2484:
2480:
2477:
2475:
2471:
2468:
2466:
2462:
2460:
2456:
2455:
2450:
2446:
2442:
2435:
2430:
2428:
2423:
2421:
2416:
2415:
2412:
2405:
2404:United States
2395:
2393:
2383:
2382:
2379:
2374:
2372:
2367:
2350:
2346:
2341:
2336:
2331:
2326:
2322:
2318:
2314:
2310:
2306:
2299:
2292:
2286:
2279:
2275:
2269:
2267:
2265:
2263:
2261:
2259:
2257:
2248:
2244:
2240:
2236:
2232:
2225:
2217:
2213:
2209:
2205:
2201:
2197:
2193:
2186:
2178:
2172:
2168:
2161:
2152:
2144:
2138:
2134:
2130:
2123:
2115:
2111:
2107:
2103:
2099:
2095:
2091:
2087:
2083:
2076:
2069:(1): 211–273.
2068:
2064:
2060:
2056:
2055:Hypsilophodon
2052:
2046:
2044:
2035:
2031:
2027:
2023:
2019:
2015:
2008:
2000:
1996:
1992:
1988:
1981:
1979:
1970:
1968:
1966:
1964:
1962:
1953:
1949:
1944:
1939:
1934:
1929:
1925:
1921:
1917:
1913:
1909:
1902:
1900:
1898:
1889:
1885:
1881:
1877:
1873:
1869:
1865:
1858:
1856:
1854:
1852:
1843:
1837:
1833:
1829:
1822:
1815:
1809:
1801:
1797:
1793:
1789:
1785:
1783:
1774:
1772:
1763:
1759:
1755:
1751:
1747:
1743:
1739:
1735:
1731:
1724:
1716:
1709:
1701:
1697:
1696:Palaeontology
1690:
1688:
1686:
1677:
1673:
1666:
1658:
1654:
1650:
1646:
1642:
1638:
1634:
1630:
1626:
1619:
1611:
1607:
1603:
1599:
1595:
1591:
1587:
1583:
1579:
1575:
1571:
1564:
1562:
1560:
1555:
1545:
1542:
1540:
1537:
1536:
1530:
1528:
1524:
1516:
1512:
1498:
1497:
1489:
1488:
1480:
1479:
1471:
1470:
1462:
1461:
1453:
1452:
1444:
1443:
1435:
1434:
1431:
1430:
1424:
1423:
1420:
1419:
1416:
1415:
1414:
1407:
1406:
1400:
1399:
1396:
1395:
1392:
1391:
1390:
1383:
1382:
1376:
1375:
1372:
1371:
1368:
1367:
1366:
1365:Lanzhousaurus
1359:
1358:
1352:
1351:
1348:
1347:
1344:
1343:
1342:
1335:
1334:
1328:
1327:
1324:
1323:
1315:
1314:
1311:
1310:
1309:
1308:Theiophytalia
1302:
1301:
1298:
1297:
1294:
1293:
1292:
1285:
1284:
1278:
1277:
1271:
1270:
1267:
1266:
1263:
1262:
1261:
1254:
1253:
1247:
1246:Styracosterna
1244:
1243:
1240:
1239:
1236:
1235:
1234:
1227:
1226:
1220:
1217:
1216:
1209:
1205:
1200:
1196:
1194:
1190:
1185:
1183:
1182:
1177:
1176:
1171:
1170:
1165:
1161:
1157:
1153:
1150:
1146:
1142:
1141:phylogenetics
1137:
1135:
1131:
1121:
1119:
1115:
1114:
1108:
1107:
1106:Theiophytalia
1101:
1099:
1095:
1091:
1087:
1083:
1079:
1075:
1074:
1069:
1061:
1052:
1050:
1046:
1045:
1040:
1036:
1032:
1028:
1024:
1019:
1017:
1013:
1009:
1005:
1001:
997:
996:
991:
987:
983:
979:
975:
970:
968:
967:
962:
958:
954:
952:
947:
943:
942:
941:Callovosaurus
937:
933:
929:
925:
924:
919:
915:
911:
907:
903:
902:C. prestwichi
899:
895:
891:
887:
880:West, Wyoming
879:
874:
870:
868:
867:
862:
861:Styracosterna
858:
854:
850:
846:
842:
838:
837:
832:
828:
824:
820:
819:
814:
810:
806:
802:
796:
792:
791:
790:Theiophytalia
786:
782:
778:
776:
775:
770:
766:
762:
758:
754:
750:
746:
742:
738:
734:
733:C. prestwichi
730:
723:
719:
714:
710:
708:
704:
700:
696:
692:
688:
684:
683:
678:
674:
670:
666:
662:
658:
654:
650:
646:
642:
638:
634:
630:
626:
625:
620:
616:
612:
608:
601:
600:Theiophytalia
596:
587:
585:
583:
571:
569:
558:
554:
550:
549:North America
546:
543:
542:Late Jurassic
539:
536:
535:ornithischian
532:
528:
527:
514:
469:
468:
452:
451:
447:
441:
438:
434:
428:
427:
423:
419:
416:Marsh, 1879 (
413:
410:
406:
401:
398:
397:
395:
392:
388:
381:
375:
374:
367:
360:
359:
352:
351:
349:
346:
341:
334:
333:
328:
325:
321:
317:
311:
310:
303:
300:
299:
292:
286:
283:
282:
275:
269:
267:Superfamily:
266:
265:
262:
256:
253:
250:
249:
246:
240:
237:
234:
233:
230:
224:
221:
218:
217:
214:
208:
205:
202:
201:
198:
192:
189:
186:
185:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
137:
132:
128:
123:
119:
114:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
46:
39:
38:Late Jurassic
33:
29:
26:
22:
4817:Camptosaurus
4787:Camptosaurus
4786:
4751:
4744:
4736:
4724:
4693:
4675:
4668:
4661:
4654:
4647:
4640:
4633:
4626:
4619:
4612:
4605:
4598:
4591:
4584:
4577:
4570:
4564:Claorhynchus
4562:
4555:
4548:
4540:Nomina dubia
4538:
4528:Guegoolithus
4526:
4506:
4499:
4494:Staurichnium
4492:
4487:Sousaichnium
4485:
4478:
4471:
4464:
4457:
4450:
4443:
4438:Dakotasaurus
4436:
4431:Caririchnium
4429:
4422:
4415:
4408:
4401:
4345:
4313:
4306:
4299:
4292:
4285:
4278:
4258:
4251:
4244:
4237:Saurolophini
4224:
4217:
4210:
4203:
4196:
4188:
4181:Kritosaurini
4168:
4161:
4156:Huallasaurus
4154:
4147:
4128:Wulagasaurus
4126:
4119:
4112:
4105:
4098:
4091:
4071:
4043:
4036:
4029:
4022:
4017:Lambeosaurus
4015:
4008:
4001:
3994:
3974:
3967:
3960:
3953:
3933:
3926:
3919:
3912:
3904:
3897:
3890:
3883:
3876:
3869:
3862:
3855:
3848:
3841:
3819:Yamatosaurus
3817:
3810:
3803:
3796:
3789:
3784:Aquilarhinus
3782:
3756:
3749:
3742:
3735:
3730:Tethyshadros
3728:
3721:
3714:
3707:
3700:
3693:
3686:
3679:
3672:
3665:
3660:Plesiohadros
3658:
3651:
3644:
3637:
3632:Lophorhothon
3630:
3623:
3615:
3608:
3601:
3594:
3587:
3580:
3573:
3566:
3559:
3552:
3545:
3538:
3531:
3524:
3518:Batyrosaurus
3516:
3511:Bactrosaurus
3509:
3502:
3448:
3391:
3384:
3377:
3369:
3351:Ouranosaurus
3349:
3342:
3335:
3328:
3321:
3316:Brighstoneus
3314:
3307:
3283:
3276:
3269:
3262:
3255:
3248:
3243:Osmakasaurus
3241:
3236:Oblitosaurus
3234:
3227:
3220:
3213:
3206:
3199:
3192:
3185:
3178:
3170:
3163:
3158:Camptosaurus
3157:
3156:
3149:
3142:
3135:
3105:
3099:Kangnasaurus
3097:
3089:
3082:
3075:
3068:
3061:
3054:Dryosauridae
3041:
3034:
3027:
2989:
2982:
2975:
2968:
2961:
2956:Matheronodon
2954:
2934:
2927:
2919:
2897:
2889:
2881:
2856:
2849:
2842:
2835:
2830:Sektensaurus
2828:
2823:Qantassaurus
2821:
2814:
2806:
2799:
2792:
2785:
2778:
2772:Kangnasaurus
2770:
2763:
2756:
2749:
2741:
2736:Diluvicursor
2734:
2727:
2720:
2713:
2694:Vectidromeus
2692:
2685:
2677:
2655:
2648:
2641:
2559:Ornithischia
2544:Ornithischia
2496:
2487:
2483:Ornithischia
2478:
2469:
2371:Camptosaurus
2363:
2312:
2308:
2298:
2290:
2285:
2277:
2273:
2238:
2234:
2224:
2202:(1): 61–73.
2199:
2195:
2185:
2166:
2160:
2151:
2132:
2128:
2122:
2089:
2085:
2075:
2066:
2062:
2058:
2054:
2017:
2013:
2007:
1990:
1986:
1978:Camptosaurus
1977:
1915:
1911:
1871:
1867:
1864:Camptosaurus
1863:
1831:
1828:Camptosaurus
1827:
1821:
1813:
1808:
1791:
1787:
1782:Camptosaurus
1781:
1737:
1733:
1723:
1714:
1708:
1699:
1695:
1675:
1671:
1665:
1632:
1628:
1618:
1580:(1): 65–78.
1577:
1573:
1527:Camptosaurus
1526:
1523:Iguanodontia
1520:
1514:
1506:Paleobiology
1427:
1411:
1410:
1387:
1386:
1363:
1362:
1339:
1338:
1306:
1305:
1289:
1288:
1258:
1257:
1233:Camptosaurus
1232:
1231:
1230:
1208:Camptosaurus
1207:
1193:Camptosaurus
1192:
1186:
1179:
1173:
1167:
1156:iguanodontid
1145:Camptosaurus
1144:
1138:
1130:Camptosaurus
1129:
1127:
1118:Camptosaurus
1117:
1111:
1104:
1102:
1098:Camptosaurus
1097:
1085:
1082:Camptosaurus
1081:
1071:
1068:Camptosaurus
1067:
1066:
1049:Camptosaurus
1048:
1042:
1039:Camptosaurus
1038:
1026:
1023:Camptosaurus
1022:
1020:
1015:
1011:
1003:
1000:Camptosaurus
999:
993:
989:
978:Cumnor Hurst
973:
971:
964:
961:Camptosaurus
960:
957:Richard Owen
949:
945:
939:
935:
928:C. valdensis
927:
923:nomen dubium
921:
909:
906:C. valdensis
905:
901:
897:
893:
889:
886:Camptosaurus
885:
883:
864:
856:
848:
840:
836:Osmakasaurus
834:
831:C. depressus
830:
826:
822:
816:
800:
799:
788:
785:Camptosaurus
784:
772:
769:Camptosaurus
768:
765:Camptosaurus
764:
760:
756:
752:
748:
744:
740:
736:
732:
729:Peter Galton
726:
721:
717:
707:Camptosaurus
706:
703:C. depressus
702:
698:
690:
686:
680:
676:
673:Arthur Lakes
668:
652:
649:type species
644:
637:Camptosaurus
636:
622:
615:euornithopod
604:
599:
579:
565:
467:Camptosaurus
466:
465:
464:
448:
440:Symphyrophus
439:
424:
411:
400:Brachyrophus
399:
371:
356:
331:
330:
324:Type species
309:Camptosaurus
308:
307:
290:
273:
251:
235:
219:
203:
197:Ornithischia
187:
174:
32:Camptosaurus
31:
25:
4906:Iguanodonts
4811:Wikispecies
4753:Willinakaqe
4725:nomina nuda
4642:Siluosaurus
4607:Loncosaurus
4445:Dinehichnus
4287:Kamuysaurus
4260:Saurolophus
4212:Kritosaurus
4205:Gryposaurus
4038:Sahaliyania
4024:Magnapaulia
3996:Amurosaurus
3976:Tlatolophus
3955:Adelolophus
3899:Kazaklambia
3878:Blasisaurus
3871:Arenysaurus
3864:Aralosaurus
3798:Hadrosaurus
3791:Eotrachodon
3744:Yunganglong
3709:Sirindhorna
3681:Protohadros
3617:Koshisaurus
3610:Jintasaurus
3379:Comptonatus
3229:Napaisaurus
3215:Lurdusaurus
3187:Fukuisaurus
3165:Cedrorestes
3107:Valdosaurus
2970:Pareisactus
2801:Morrosaurus
2765:Isasicursor
2729:Chakisaurus
2625:Ornithopoda
2612:Ornithopoda
2571:Ornithopoda
2441:Ornithopoda
2059:Valdosaurus
1160:hadrosaurid
1080:belongs to
1063:Restoration
1055:Description
982:Oxfordshire
914:Haţeg Basin
823:P. depressa
624:Camptonotus
547:of western
418:preoccupied
412:Camptonotus
337:Marsh, 1879
295:Marsh, 1885
278:Marsh, 1885
229:Ornithopoda
21:Camposaurus
4900:Categories
4695:Arkharavia
4670:Tichosteus
4663:Thespesius
4635:Pteropelyx
4628:Orthomerus
4508:Wintonopus
4093:Acristavus
4073:Barsboldia
4031:Olorotitan
3906:Latirhinus
3758:Zuoyunlong
3625:Levnesovia
3575:Gobihadros
3540:Datonglong
3533:Claosaurus
3504:Altirhinus
3414:see below↓
3344:Morelladon
3222:Magnamanus
3194:Hippodraco
3063:Dryosaurus
3036:Hesperonyx
3011:Dryomorpha
2851:Trinisaura
2844:Tietasaura
2837:Talenkauen
2715:Anabisetia
2580:see below↓
2474:Dinosauria
1702:: 411–443.
1550:References
1389:Kukufeldia
1291:Hippodraco
1204:ornithopod
1169:Dryosaurus
1113:Dryosaurus
682:Allosaurus
657:Como Bluff
444:Cope, 1878
245:Dryomorpha
181:Dinosauria
4677:Trachodon
4593:Glishades
4572:Diclonius
4410:Anomoepus
4394:Ichnotaxa
4107:Maiasaura
4045:Velafrons
3554:Equijubus
3526:Choyrodon
3386:Iguanodon
3250:Planicoxa
3180:Dakotadon
3151:Calvarius
2977:Rhabdodon
2963:Mochlodon
2706:Elasmaria
2457:Kingdom:
2392:Dinosaurs
2114:1477-2019
1993:: 52–68.
1762:130777820
1657:131001110
1610:245804125
1602:1698-6180
1189:cladistic
1143:has made
1094:Oxfordian
1090:Callovian
951:Iguanodon
946:C. hoggii
936:C. leedsi
910:C. inkeyi
898:C. leedsi
894:C. hoggii
890:C. inkeyi
849:C. dispar
818:Planicoxa
761:C. dispar
757:C. amplus
753:C. dispar
749:C. dispar
745:C. browni
741:C. medius
722:C. dispar
699:C. browni
687:C. medius
677:Quarry 1A
669:C. amplus
653:Quarry 13
645:C. dispar
538:dinosaurs
154:Kingdom:
148:Eukaryota
4831:52571997
4796:Wikidata
4557:Cionodon
4226:Rhinorex
4114:Ornatops
3914:Minqaria
3885:Canardia
3812:Malefica
3737:Xuwulong
3702:Siamodon
3603:Jeyawati
3589:Gonkoken
3547:Eolambia
3371:Barilium
3172:Cumnoria
3137:Barilium
3043:Owenodon
3029:Draconyx
2991:Zalmoxes
2891:Fostoria
2743:Fostoria
2501:Cerapoda
2465:Chordata
2463:Phylum:
2459:Animalia
2349:21124919
2309:PLOS ONE
1952:21124919
1912:PLOS ONE
1888:86197297
1533:See also
1413:Barilium
1044:Draconyx
1041:than to
1004:Cumnoria
995:Cumnoria
966:Owenodon
737:C. nanus
718:C. nanus
691:C. nanus
633:holotype
426:Cumnoria
391:Synonyms
284:Family:
168:Chordata
164:Phylum:
158:Animalia
144:Domain:
4870:4946706
4857:1421899
4844:3238881
4802:Q133203
3850:Ajnabia
3285:Uteodon
2378:Portals
2340:2989904
2317:Bibcode
2204:Bibcode
2094:Bibcode
2022:Bibcode
2014:Geobios
1987:Zootaxa
1943:2989904
1920:Bibcode
1742:Bibcode
1637:Bibcode
1582:Bibcode
1260:Uteodon
1175:Drinker
1164:derived
1139:Modern
1073:Uteodon
1051:genus.
1016:Uteodon
986:tramway
932:dubious
918:Romania
866:Uteodon
853:ischium
793:skull,
663:in the
661:Wyoming
641:cricket
568:kamptos
561:καμπτος
540:of the
529:) is a
450:Uteodon
382:, 1880
345:species
314:Marsh,
301:Genus:
4519:Ootaxa
3716:Tanius
3309:Bolong
2347:
2337:
2173:
2139:
2112:
1950:
1940:
1886:
1838:
1760:
1678:: 169.
1655:
1608:
1600:
1178:, and
1027:et al.
953:hoggii
904:, and
631:. The
582:sauros
575:σαυρος
553:Europe
545:period
435:, 1888
433:Seeley
407:, 1878
343:Other
4883:38748
4852:IRMNG
3561:Fylax
3091:Iyuku
2497:Clade
2488:Clade
2479:Clade
2470:Clade
2092:(1).
1983:(PDF)
1884:S2CID
1758:S2CID
1736:. 3.
1653:S2CID
1606:S2CID
1149:clade
1035:Spain
821:, as
655:near
557:Greek
531:genus
380:Hulke
252:Clade
236:Clade
220:Clade
204:Clade
188:Clade
175:Clade
4839:GBIF
3257:Proa
2929:Iani
2557:see
2530:see
2345:PMID
2171:ISBN
2137:ISBN
2110:ISSN
1991:2783
1948:PMID
1836:ISBN
1598:ISSN
1158:and
743:and
701:and
689:and
667:and
522:-tə-
520:KAMP
405:Cope
316:1885
51:PreꞒ
4826:EoL
2335:PMC
2325:doi
2276:."
2243:doi
2239:199
2212:doi
2102:doi
2030:doi
1995:doi
1938:PMC
1928:doi
1876:doi
1796:doi
1750:doi
1645:doi
1590:doi
1033:in
980:in
955:by
926:).
916:in
675:at
609:in
526:-əs
524:SOR
504:ɔːr
4902::
4880::
4867::
4854::
4841::
4828::
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4798::
2499::
2490::
2481::
2472::
2343:.
2333:.
2323:.
2311:.
2307:.
2255:^
2237:.
2233:.
2210:.
2200:87
2198:.
2194:.
2108:.
2100:.
2090:22
2088:.
2084:.
2067:28
2065:.
2057:,
2042:^
2028:.
2018:39
2016:.
1989:.
1985:.
1960:^
1946:.
1936:.
1926:.
1914:.
1910:.
1896:^
1882:.
1872:76
1870:.
1850:^
1792:36
1790:.
1786:.
1770:^
1756:.
1748:.
1738:48
1732:.
1700:23
1698:.
1684:^
1676:29
1674:.
1651:.
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1631:.
1627:.
1604:.
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1576:.
1572:.
1558:^
1184:.
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1136:.
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969:.
944:.
908:.
900:,
896:,
892:,
869:.
777:.
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659:,
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361:?
254::
238::
222::
206::
190::
177::
101:Pg
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4131:?
3909:?
3620:?
3521:?
3396:?
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2327::
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2249:.
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2206::
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2001:.
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1922::
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1659:.
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492:t
489:p
486:m
483:æ
480:k
477:ˌ
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470:(
453:?
429:?
420:)
369:†
354:†
305:†
288:†
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242:†
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194:†
106:N
96:K
91:J
86:T
81:P
76:C
71:D
66:S
61:O
56:Ꞓ
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