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Otago shag

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270: 86: 42: 61: 382: 284:, with two plumages. Roughly one quarter of the individuals (a smaller percentage than in Foveaux shag) are pied, with dark and white feathers, and the rest, known as bronze shags, are dark all over. Both morphs breed together. These large, chunky birds are about 70 cm long and weigh about 2–3 kg, slightly larger than the closely related 291:
Otago shags vary in their facial ornamentation in the breeding season. Roughly half of breeding individuals have dark orange papillae on their face (as in Foveaux shag), while the other half have small bright orange caruncles above the base of the bill instead.
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Until 2016, Otago shags and the closely related shags living around Stewart Island and Foveaux Strait were considered to be a single species, called the Stewart Island shag. Mitochondrial DNA suggests Otago shags are actually more closely related to
378:. Restricted to a small area, and having little or no genetic variation, they require conservation efforts tailored to these extinction risk factors; this could include reintroduction to part of their former range. 362:, but when humans arrived the population was devastated, reduced by 99 percent within 100 years with a corresponding loss of genetic diversity. It became restricted to the rocky offshore islets off the 551:
Rawlence, Nicolas J.; Scofield, R. Paul; Spencer, Hamish G.; Lalas, Chris; Easton, Luke J.; Tennyson, Alan J.D.; Adams, Mark; Pasquet, Eric; Fraser, Cody; Waters, Jonathan M.; Kennedy, Martyn (2016).
485:
Rawlence, Nicolas J.; Till, Charlotte E.; Scofield, R. Paul; Tennyson, Alan J. D.; Collins, Catherine J.; Lalas, Chris; Loh, Graeme; Matisoo-Smith, Elizabeth; Waters, Jonathan M. (2014).
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on islands and sea cliffs. Colonies are large enough to be strikingly visible, and are used year after year. One notable colony is on the northern shore of
1651: 397:, at the mouth of the Otago Harbour. They feed in coastal waters less than 30 m deep and are rarely if ever seen inland or far out to sea. 735: 1805: 1777: 1612: 344:
is a distinct genus, which would contain amongst other species the Otago, Foveaux, and Chatham shags. Others place it in the genus
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Archaeological evidence shows that Otago shags were formerly found along the entire east coast of the South Island up to
1815: 366:, and has scarcely recovered since that time. There are less than 2500 Otago shags remaining, but they can be seen at 1820: 1656: 1551: 296:
colour in the breeding season also varies, from bright orange (as in Foveaux shag) to dark orange to purple.
624: 487:"Strong Phylogeographic Structure in a Sedentary Seabird, the Stewart Island Shag (Leucocarbo chalconotus)" 1591: 1586: 17: 389:
Otago shags breed colonially from May to September, making raised cup nests out of organic material and
438: 85: 325:. Foveaux and Otago shags probably diverged when populations were split up by lower sea levels in the 1676: 1565: 1205: 650: 1340: 1235: 755: 677: 848: 1728: 1573: 1513: 1215: 281: 1643: 1723: 1421: 1302: 1276: 1210: 858: 853: 588:
Kennedy, Martyn; Spencer, Hamish G. (2014). "Classification of the cormorants of the world".
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shag (Aves, Pelecaniformes, Phalacrocoracidae) from southern South Island of New Zealand"
415: 246: 50: 1741: 502: 1488: 1240: 868: 521: 486: 359: 206: 80: 65: 1710: 1599: 1463: 1307: 1261: 1185: 985: 942: 863: 707: 605: 526: 269: 234: 1604: 1416: 1266: 1220: 1138: 1042: 962: 895: 793: 597: 568: 516: 506: 433: 197: 1768: 1684: 1536: 1271: 1230: 1101: 1005: 771: 511: 363: 330: 601: 1483: 1478: 1473: 1281: 1225: 1190: 1106: 947: 337:. Other taxonomists have kept the Otago shag and the Foveaux shag conspecific. 157: 1799: 1638: 1457: 1447: 1442: 1370: 1176: 424: 367: 346: 70: 1437: 1427: 1381: 1200: 952: 609: 530: 394: 318: 306: 285: 242: 1689: 1545: 1468: 1452: 1331: 1010: 967: 957: 839: 789: 375: 326: 293: 41: 1782: 1736: 1617: 1401: 1252: 775: 137: 788: 573: 552: 1410: 1195: 1134: 314: 254: 147: 97: 1507: 1762: 1530: 117: 1702: 1560: 1361: 1162: 1096: 1073: 919: 891: 825: 1037: 1087: 1038: 996: 371: 107: 1293: 1151: 1062: 1046: 933: 908: 814: 797: 390: 381: 334: 258: 770: 550: 484: 407: 127: 1133: 890: 553:"Genetic and morphological evidence for two species of 413: 273:
An 1844 painting by Hullmandel of the bronze phase of
1317: 439:10.2305/IUCN.UK.2018-2.RLTS.T22696853A133556225.en 317:and genetic differences supported separating off 1797: 701: 461:"Otago shag | Matapo | New Zealand Birds Online" 353: 587: 756: 704:The Field Guide to the Birds of New Zealand 675: 625:"Meet our 'newest' endangered bird species" 763: 749: 333:were subsequently colonised by shags from 59: 40: 702:Heather, Barrie; Robertson, Hugh (2015). 695: 572: 561:Zoological Journal of the Linnean Society 520: 510: 437: 380: 340:A recent taxonomic revision argues that 268: 14: 1798: 676:Chamberlain, Rhys (18 February 2016). 622: 1512: 1511: 1132: 1036: 889: 787: 744: 706:. New Zealand: Penguin. p. 290. 590:Molecular Phylogenetics and Evolution 1677:142ade09-25a7-4fcf-9abd-0f0bb6704dd7 1460:(imperial cormorant, blue-eyed shag) 546: 544: 542: 540: 651:"New Otago shag species discovered" 425:IUCN Red List of Threatened Species 24: 623:Morton, Jamie (17 February 2016). 25: 1842: 724: 537: 299: 1806:IUCN Red List vulnerable species 730:Otago shag discussed on RadioNZ 84: 414:BirdLife International (2018). 1826:Taxa named by John Edward Gray 669: 643: 616: 581: 478: 453: 264: 13: 1: 400: 354:Distribution and conservation 249:and in its dark phase as the 1831:Endemic birds of New Zealand 859:Christmas Island frigatebird 512:10.1371/journal.pone.0090769 7: 1160: 1071: 917: 823: 602:10.1016/j.ympev.2014.06.020 10: 1847: 374:, and as far south as the 257:now found only in coastal 1816:Birds of the South Island 1752: 1520: 1399: 1379: 1359: 1329: 1291: 1250: 1174: 1159: 1150: 1146: 1128: 1085: 1070: 1061: 1057: 1032: 994: 976: 931: 916: 907: 903: 885: 837: 822: 813: 809: 783: 432:: e.T22696853A133556225. 275:Phalacrocorax chalconotus 215:Phalacrocorax chalconotus 212: 205: 186: 179: 81:Scientific classification 79: 57: 48: 39: 34: 1341:Double-crested cormorant 1236:White-breasted cormorant 678:"Otago shag new species" 1821:Birds described in 1845 1284:(long-tailed cormorant) 849:Magnificent frigatebird 321:as a distinct species, 1742:Leucocarbo-chalconotus 1552:Leucocarbo chalconotus 1522:Leucocarbo chalconotus 1413:(Magellanic cormorant) 1243:(Temminck's cormorant) 1216:Little black cormorant 778:(Phalacrocoraciformes) 629:The New Zealand Herald 418:Leucocarbo chalconotus 386: 277: 245:formerly known as the 238: 230:Leucocarbo chalconotus 190:Leucocarbo chalconotus 1724:Paleobiology Database 1422:New Zealand king shag 1277:Little pied cormorant 1211:Black-faced cormorant 854:Ascension frigatebird 655:39 Dunedin Television 385:Otago Shag - Otago NZ 384: 272: 241:), together with the 1754:Graculus chalconotus 1391:Red-legged cormorant 1351:Flightless cormorant 1321:Spectacled cormorant 1141:: Phalacrocoracidae) 1045:: Anhingidae · 796:: Fregatidae · 465:nzbirdsonline.org.nz 1667:stewart-island-shag 1346:Neotropic cormorant 1313:Red-faced cormorant 1112:Australasian darter 1016:Australasian gannet 892:Gannets and boobies 732:Critter of the Week 503:2014PLoSO...990769R 247:Stewart Island shag 172:L. chalconotus 51:Conservation status 1489:South Georgia shag 1424:(rough-faced shag) 1303:Brandt's cormorant 1241:Japanese cormorant 869:Lesser frigatebird 657:. 18 February 2016 387: 370:, as far north as 311:Leucocarbo onslowi 278: 253:, is a species of 1793: 1792: 1711:Open Tree of Life 1514:Taxon identifiers 1505: 1504: 1501: 1500: 1497: 1496: 1464:Heard Island shag 1308:Pelagic cormorant 1262:Crowned cormorant 1186:Socotra cormorant 1170: 1169: 1124: 1123: 1120: 1119: 1081: 1080: 1028: 1027: 1024: 1023: 943:Blue-footed booby 927: 926: 881: 880: 877: 876: 864:Great frigatebird 833: 832: 713:978-0-143-57092-9 682:Otago Daily Times 574:10.1111/zoj.12376 221: 220: 148:Phalacrocoracidae 74: 16:(Redirected from 1838: 1786: 1785: 1773: 1772: 1771: 1745: 1744: 1732: 1731: 1719: 1718: 1706: 1705: 1693: 1692: 1680: 1679: 1670: 1669: 1660: 1659: 1647: 1646: 1634: 1633: 1621: 1620: 1608: 1607: 1595: 1594: 1582: 1581: 1569: 1568: 1566:483FA17D5EC47483 1556: 1555: 1554: 1541: 1540: 1539: 1509: 1508: 1417:Guanay cormorant 1319: 1267:Little cormorant 1221:Indian cormorant 1157: 1156: 1148: 1147: 1130: 1129: 1068: 1067: 1059: 1058: 1034: 1033: 963:Red-footed booby 914: 913: 905: 904: 887: 886: 820: 819: 811: 810: 785: 784: 765: 758: 751: 742: 741: 718: 717: 699: 693: 692: 690: 688: 673: 667: 666: 664: 662: 647: 641: 640: 638: 636: 620: 614: 613: 585: 579: 578: 576: 548: 535: 534: 524: 514: 482: 476: 475: 473: 471: 457: 451: 450: 448: 446: 441: 411: 192: 89: 88: 68: 63: 62: 44: 32: 31: 21: 1846: 1845: 1841: 1840: 1839: 1837: 1836: 1835: 1796: 1795: 1794: 1789: 1781: 1776: 1767: 1766: 1761: 1748: 1740: 1735: 1727: 1722: 1714: 1709: 1701: 1696: 1688: 1685:Observation.org 1683: 1675: 1673: 1665: 1663: 1655: 1650: 1642: 1637: 1629: 1624: 1616: 1611: 1603: 1598: 1590: 1585: 1577: 1572: 1564: 1559: 1550: 1549: 1544: 1535: 1534: 1529: 1516: 1506: 1493: 1395: 1375: 1355: 1325: 1287: 1272:Pygmy cormorant 1246: 1231:Great cormorant 1166: 1142: 1116: 1102:Oriental darter 1077: 1053: 1020: 1006:Northern gannet 990: 972: 923: 899: 873: 829: 805: 779: 769: 727: 722: 721: 714: 700: 696: 686: 684: 674: 670: 660: 658: 649: 648: 644: 634: 632: 621: 617: 586: 582: 549: 538: 483: 479: 469: 467: 459: 458: 454: 444: 442: 412: 408: 403: 364:Otago Peninsula 356: 331:Chatham Islands 302: 280:The species is 267: 261:, New Zealand. 201: 194: 188: 175: 83: 75: 64: 60: 53: 28: 27:Species of bird 23: 22: 15: 12: 11: 5: 1844: 1834: 1833: 1828: 1823: 1818: 1813: 1808: 1791: 1790: 1788: 1787: 1774: 1758: 1756: 1750: 1749: 1747: 1746: 1733: 1720: 1707: 1694: 1681: 1671: 1661: 1648: 1635: 1622: 1609: 1596: 1583: 1570: 1557: 1542: 1526: 1524: 1518: 1517: 1503: 1502: 1499: 1498: 1495: 1494: 1492: 1491: 1486: 1484:Macquarie shag 1481: 1479:Kerguelen shag 1476: 1474:Antarctic shag 1471: 1466: 1461: 1455: 1450: 1445: 1440: 1435: 1430: 1425: 1419: 1414: 1407: 1405: 1397: 1396: 1394: 1393: 1387: 1385: 1377: 1376: 1374: 1373: 1367: 1365: 1357: 1356: 1354: 1353: 1348: 1343: 1337: 1335: 1327: 1326: 1324: 1323: 1315: 1310: 1305: 1299: 1297: 1289: 1288: 1286: 1285: 1282:Reed cormorant 1279: 1274: 1269: 1264: 1258: 1256: 1248: 1247: 1245: 1244: 1238: 1233: 1228: 1226:Cape cormorant 1223: 1218: 1213: 1208: 1206:Pied cormorant 1203: 1198: 1193: 1191:Bank cormorant 1188: 1182: 1180: 1172: 1171: 1168: 1167: 1154: 1144: 1143: 1126: 1125: 1122: 1121: 1118: 1117: 1115: 1114: 1109: 1107:African darter 1104: 1099: 1093: 1091: 1083: 1082: 1079: 1078: 1065: 1055: 1054: 1030: 1029: 1026: 1025: 1022: 1021: 1019: 1018: 1013: 1008: 1002: 1000: 992: 991: 989: 988: 986:Abbott's booby 982: 980: 974: 973: 971: 970: 965: 960: 955: 950: 948:Peruvian booby 945: 939: 937: 929: 928: 925: 924: 911: 901: 900: 883: 882: 879: 878: 875: 874: 872: 871: 866: 861: 856: 851: 845: 843: 835: 834: 831: 830: 817: 807: 806: 781: 780: 768: 767: 760: 753: 745: 739: 738: 726: 725:External links 723: 720: 719: 712: 694: 668: 642: 615: 580: 567:(3): 676–694. 536: 477: 452: 405: 404: 402: 399: 355: 352: 301: 300:Classification 298: 266: 263: 219: 218: 210: 209: 203: 202: 195: 184: 183: 177: 176: 169: 167: 163: 162: 155: 151: 150: 145: 141: 140: 135: 131: 130: 125: 121: 120: 115: 111: 110: 105: 101: 100: 95: 91: 90: 77: 76: 58: 55: 54: 49: 46: 45: 37: 36: 26: 9: 6: 4: 3: 2: 1843: 1832: 1829: 1827: 1824: 1822: 1819: 1817: 1814: 1812: 1809: 1807: 1804: 1803: 1801: 1784: 1779: 1775: 1770: 1764: 1760: 1759: 1757: 1755: 1751: 1743: 1738: 1734: 1730: 1725: 1721: 1717: 1712: 1708: 1704: 1699: 1695: 1691: 1686: 1682: 1678: 1672: 1668: 1662: 1658: 1653: 1649: 1645: 1640: 1636: 1632: 1627: 1623: 1619: 1614: 1610: 1606: 1601: 1597: 1593: 1588: 1584: 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320: 319:Foveaux shags 316: 312: 308: 307:Chatham shags 297: 295: 289: 287: 283: 276: 271: 262: 260: 256: 252: 248: 244: 240: 236: 232: 231: 226: 217: 216: 211: 208: 204: 199: 193: 191: 185: 182: 181:Binomial name 178: 174: 173: 168: 165: 164: 161: 160: 156: 153: 152: 149: 146: 143: 142: 139: 136: 133: 132: 129: 126: 123: 122: 119: 116: 113: 112: 109: 106: 103: 102: 99: 96: 93: 92: 87: 82: 78: 72: 67: 56: 52: 47: 43: 38: 33: 30: 19: 1753: 1521: 1438:Chatham shag 1432: 1428:Foveaux shag 1400: 1382:Poikilocarbo 1380: 1360: 1330: 1292: 1251: 1201:Spotted shag 1175: 1161: 1086: 1072: 1049: 995: 977: 953:Masked booby 932: 918: 838: 824: 801: 790:Frigatebirds 736:4 March 2016 731: 703: 697: 685:. Retrieved 681: 671: 659:. Retrieved 654: 645: 633:. Retrieved 628: 618: 593: 589: 583: 564: 560: 554: 494: 490: 480: 468:. Retrieved 464: 455: 443:. Retrieved 429: 423: 417: 409: 395:Taiaroa Head 388: 357: 345: 341: 339: 322: 315:osteological 310: 303: 290: 286:Foveaux shag 279: 274: 250: 243:Foveaux shag 229: 228: 224: 222: 214: 213: 189: 187: 171: 170: 158: 29: 1546:Wikispecies 1469:Crozet shag 1453:Bounty shag 1332:Nannopterum 1011:Cape gannet 968:Brown booby 958:Nazca booby 596:: 249–257. 470:26 November 445:18 November 360:Marlborough 327:Pleistocene 323:L. stewarti 294:Gular pouch 265:Description 251:bronze shag 35:Otago shag 18:Bronze Shag 1811:Leucocarbo 1800:Categories 1737:Xeno-canto 1433:Otago shag 1402:Leucocarbo 1253:Microcarbo 1135:Cormorants 898:: Sulidae) 776:Suliformes 555:Leucocarbo 401:References 342:Leucocarbo 329:, and the 225:Otago shag 159:Leucocarbo 138:Suliformes 66:Vulnerable 1769:Q50877964 1537:Q27074618 1411:Rock shag 1196:Pitt shag 282:dimorphic 166:Species: 104:Kingdom: 98:Eukaryota 1763:Wikidata 1644:22696853 1579:22696853 1574:BirdLife 1531:Wikidata 978:Papasula 610:24994028 531:24614677 491:PLOS ONE 207:Synonyms 144:Family: 118:Chordata 114:Phylum: 108:Animalia 94:Domain: 71:IUCN 3.1 1783:8815392 1703:1610891 1618:5789268 1561:Avibase 1362:Gulosus 1163:Species 1097:Anhinga 1088:Anhinga 1074:Species 1050:Anhinga 1039:Darters 920:Species 840:Fregata 826:Species 802:Fregata 687:2 March 661:2 March 635:2 March 522:3948693 499:Bibcode 376:Catlins 313:), and 200:, 1845) 154:Genus: 134:Order: 124:Class: 69: ( 1729:399998 1716:480890 1674:NZOR: 1664:NZBO: 1657:146625 1631:824007 1592:198413 1139:family 1043:family 896:family 794:family 710:  631:. NZME 608:  529:  519:  372:Oamaru 239:Matapo 1690:73435 1605:6Q5KH 1294:Urile 1152:Genus 1063:Genus 1047:genus 997:Morus 909:Genus 815:Genus 798:genus 772:Order 391:guano 335:Otago 259:Otago 235:Māori 1778:GBIF 1698:OBIS 1652:NCBI 1639:IUCN 1626:ITIS 1613:GBIF 1587:BOLD 934:Sula 708:ISBN 689:2016 663:2016 637:2016 606:PMID 527:PMID 472:2023 447:2021 430:2018 255:shag 223:The 198:Gray 128:Aves 1600:CoL 598:doi 569:doi 565:177 517:PMC 507:doi 434:doi 233:) ( 1802:: 1780:: 1765:: 1739:: 1726:: 1713:: 1700:: 1687:: 1654:: 1641:: 1628:: 1615:: 1602:: 1589:: 1576:: 1563:: 1548:: 1533:: 800:: 774:: 734:, 680:. 653:. 627:. 604:. 594:79 592:. 563:. 559:. 539:^ 525:. 515:. 505:. 493:. 489:. 463:. 428:. 422:. 350:. 288:. 237:: 1318:† 1137:( 1052:) 1041:( 894:( 804:) 792:( 764:e 757:t 750:v 716:. 691:. 665:. 639:. 612:. 600:: 577:. 571:: 533:. 509:: 501:: 495:9 474:. 449:. 436:: 420:" 416:" 309:( 227:( 196:( 73:) 20:)

Index

Bronze Shag

Conservation status
Vulnerable
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Aves
Suliformes
Phalacrocoracidae
Leucocarbo
Binomial name
Gray
Synonyms
Māori
Foveaux shag
Stewart Island shag
shag
Otago

dimorphic
Foveaux shag
Gular pouch
Chatham shags
osteological
Foveaux shags
Pleistocene

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